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<document id="903E22C48E7E35C0E99202000BF5E135" ID-CLB-Dataset="2636" ID-DOI="10.11646/zootaxa.5091.1.1" ID-GBIF-Dataset="51b42715-ddb1-4b4a-aad4-8be0f600a287" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840434" ID-ZooBank="ABC8AF70-F691-4D07-8F20-70934642C8BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1641986550580" checkinUser="plazi" docAuthor="Bergh, Eugene W. &amp; Compton, John S." docDate="2022" docId="197787BAFFF1930D7FC99D28FAF6FE24" docLanguage="en" docName="zootaxa.5091.1.1.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orbulina universa d'Orbigny 1839" docType="treatment" docVersion="6" lastPageNumber="36" masterDocId="E54EFFC2FFD3932E7F5E9B0CFFC4FF88" masterDocTitle="Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf" masterLastPageNumber="55" masterPageNumber="1" pageNumber="35" updateTime="1699301522325" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="B636E770C9B9776793DC676889CD9D73">Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf</mods:title>
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<mods:namePart id="E443CA727C821BBC16309AAF439C1A60">Bergh, Eugene W.</mods:namePart>
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<mods:affiliation id="A5A445BB49C3B0E57EBF668CEC0CB9E9">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa. &amp; Department of Environmental and Geographical Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa; &amp; Invertebrate palaeontology and Geology, Iziko South African Museum, P. O. Box 61, Cape Town 8000, South Africa. john. compton @ uct. ac. za; https: // orcid. org / 0000 - 0002 - 0765 - 4141</mods:affiliation>
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<paragraph id="916136ACFFF1930C7FC99D28FD8DF9B7" blockId="34.[151,585,1572,1635]" box="[151,585,1572,1599]" pageId="34" pageNumber="35">
<heading id="CA2981C0FFF1930C7FC99D28FD8DF9B7" bold="true" box="[151,585,1572,1599]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<taxonomicName id="56DE4D2FFFF1930C7FC99D28FD8DF9B7" ID-CoL="74TBL" authority="d'Orbigny, 1839" authorityName="d'Orbigny" authorityYear="1839" box="[151,585,1572,1599]" class="Globothalamea" family="Globigerinidae" genus="Orbulina" kingdom="Chromista" order="Rotaliida" pageId="34" pageNumber="35" phylum="Foraminifera" rank="species" species="universa">
<emphasis id="A3AAEABEFFF1930C7FC99D28FD8DF9B7" bold="true" box="[151,585,1572,1599]" pageId="34" pageNumber="35">
<emphasis id="A3AAEABEFFF1930C7FC99D28FEB1F9B7" bold="true" box="[151,373,1572,1599]" italics="true" pageId="34" pageNumber="35">Orbulina universa</emphasis>
<bibRefCitation id="F54F4B5DFFF1930C7E239D29FD8DF9B7" author="d'Orbigny, A. D." box="[381,585,1573,1599]" pageId="34" pageNumber="35" pagination="1 - 224" refId="ref30554" refString="d'Orbigny, A. D. (1839) Foraminiferes. In: de la Sagra, R. (Ed.), Histoire physique, politique et naturelle de I'ile de Cuba. A. Bertrand, Paris, pp. 1 - 224." type="book chapter" year="1839">dOrbigny, 1839</bibRefCitation>
</emphasis>
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</paragraph>
<paragraph id="916136ACFFF1930C7FC99D45FED1F9EB" blockId="34.[151,585,1572,1635]" box="[151,277,1609,1635]" pageId="34" pageNumber="35">
<heading id="CA2981C0FFF1930C7FC99D45FED1F9EB" bold="true" box="[151,277,1609,1635]" fontSize="11" level="1" pageId="34" pageNumber="35" reason="1">
<emphasis id="A3AAEABEFFF1930C7FC99D45FED1F9EB" bold="true" box="[151,277,1609,1635]" pageId="34" pageNumber="35">Pl. 8, fig. 7</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFF1930C7FC99D8BFD5CF935" pageId="34" pageNumber="35" type="reference_group">
<paragraph id="916136ACFFF1930C7FC99D8BFD5CF935" blockId="34.[151,1437,1670,1725]" pageId="34" pageNumber="35">
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<taxonomicName id="56DE4D2FFFF1930C7FC99D8BFDC0F917" ID-CoL="74TBL" authority="d'Orbigny, 1839" authorityName="d'Orbigny" authorityYear="1839" box="[151,516,1671,1695]" class="Globothalamea" family="Globigerinidae" genus="Orbulina" kingdom="Chromista" order="Rotaliida" pageId="34" pageNumber="35" phylum="Foraminifera" rank="species" species="universa">
<emphasis id="A3AAEABEFFF1930C7FC99D8BFE96F916" box="[151,338,1671,1695]" italics="true" pageId="34" pageNumber="35">Orbulina universa</emphasis>
<bibRefCitation id="F54F4B5DFFF1930C7E049D8BFDC0F917" author="d'Orbigny, A. D." box="[346,516,1671,1695]" pageId="34" pageNumber="35" pagination="1 - 224" refId="ref30554" refString="d'Orbigny, A. D. (1839) Foraminiferes. In: de la Sagra, R. (Ed.), Histoire physique, politique et naturelle de I'ile de Cuba. A. Bertrand, Paris, pp. 1 - 224." type="book chapter" year="1839">dOrbigny, 1839</bibRefCitation>
</taxonomicName>
, pl. 28, fig. 2;
<treatmentCitation id="107F10BDFFF1930C7DC39D8BFCB4F916" author="LeRoy, L. W." box="[669,880,1670,1695]" page="501" pageId="34" pageNumber="35" year="1948">
<bibRefCitation id="F54F4B5DFFF1930C7DC39D8BFCE7F917" author="LeRoy, L. W." box="[669,803,1671,1695]" pageId="34" pageNumber="35" pagination="500 - 508" refId="ref34447" refString="LeRoy, L. W. (1948) The foraminifer Orbulina universa d'Orbigny, a suggested Middle Tertiary time indicator. Journal of Paleontology, 22, 500 - 508." type="journal article" year="1948">LeRoy, 1948</bibRefCitation>
, p. 501
</treatmentCitation>
, fig.1a;
<treatmentCitation id="107F10BDFFF1930C7C9B9D8BFAE1F917" author="Kennett, J. P. &amp; Srinivasan, M. S." box="[965,1317,1671,1695]" page="86" pageId="34" pageNumber="35" year="1983">
<bibRefCitation id="F54F4B5DFFF1930C7C9B9D8BFB21F917" author="Kennett, J. P. &amp; Srinivasan, M. S." box="[965,1253,1671,1695]" pageId="34" pageNumber="35" refId="ref33851" refString="Kennett, J. P. &amp; Srinivasan, M. S. (1983) Neogene Planktonic Foraminifera. A Phylogenetic Atlas. Hutchinson Ross, Stroudsburg, 265 pp." type="book" year="1983">Kennett &amp; Srinivasan, 1983</bibRefCitation>
, p. 86
</treatmentCitation>
, pl. 20, fig. 46;
<treatmentCitation id="107F10BDFFF1930C7FA59DA9FDCFF935" author="Kender, S. &amp; Kaminski, M. A. &amp; Jones, R. W." box="[251,523,1701,1725]" page="522" pageId="34" pageNumber="35" year="2008">
<bibRefCitation id="F54F4B5DFFF1930C7FA59DA9FE04F935" author="Kender, S. &amp; Kaminski, M. A. &amp; Jones, R. W." box="[251,448,1701,1725]" pageId="34" pageNumber="35" pagination="477 - 568" refId="ref33808" refString="Kender, S., Kaminski, M. A. &amp; Jones, R. W. (2008) Early to middle Miocene foraminifera from the deep-sea Congo Fan, offshore Angola. Micropalaeontology, 54, 477 - 568." type="journal article" year="2008">
Kender
<emphasis id="A3AAEABEFFF1930C7E159DAAFEBCF935" box="[331,376,1701,1725]" italics="true" pageId="34" pageNumber="35">et al</emphasis>
., 2008
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, p. 522
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, pl. 28, fig. 2.
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</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFF1930C7FC99DE9FABDF8AB" pageId="34" pageNumber="35" type="description">
<paragraph id="916136ACFFF1930C7FC99DE9FABDF8AB" blockId="34.[151,1437,1765,2044]" pageId="34" pageNumber="35">
<emphasis id="A3AAEABEFFF1930C7FC99DE9FEE5F977" bold="true" box="[151,289,1765,1791]" pageId="34" pageNumber="35">Description</emphasis>
: The test surface is spinose and covered with many perforations of different sizes across the entire test. The test is spherical, which may enclose an earlier globigerinid stage. There is no single large primary aperture.
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFF1930D7F999C21FAF6FE24" lastPageId="35" lastPageNumber="36" pageId="34" pageNumber="35" type="discussion">
<paragraph id="916136ACFFF1930C7F999C21FE7CF83B" blockId="34.[151,1437,1765,2044]" pageId="34" pageNumber="35">
<emphasis id="A3AAEABEFFF1930C7F999C21FEF9F8CF" bold="true" box="[199,317,1837,1863]" pageId="34" pageNumber="35">Remarks:</emphasis>
The relative abundances of
<taxonomicName id="56DE4D2FFFF1930C7DD59C21FC98F8CF" authorityName="d'Orbigny" authorityYear="1839" box="[651,860,1837,1863]" class="Globothalamea" family="Globigerinidae" genus="Orbulina" kingdom="Chromista" order="Rotaliida" pageId="34" pageNumber="35" phylum="Foraminifera" rank="species" species="universa">
<emphasis id="A3AAEABEFFF1930C7DD59C21FC98F8CF" box="[651,860,1837,1863]" italics="true" pageId="34" pageNumber="35">Orbulina universa</emphasis>
</taxonomicName>
form a minor component (&lt;10%) of the planktic foraminiferal assemblage in core 2658. In the deeper cores (2670 and 2682), the relative abundances are mostly at &lt;1% in the middle Miocene component of the cores. The tests are relatively small to moderate in size, measuring up to
<quantity id="56269B49FFF1930C7F899C95FEE9F83B" box="[215,301,1945,1972]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="34" pageNumber="35" unit="mm" value="0.5">0.5 mm</quantity>
in diameter.
</paragraph>
<paragraph id="916136ACFFF1930C7F999CB1FC7AF873" blockId="34.[151,1437,1765,2044]" pageId="34" pageNumber="35">
<emphasis id="A3AAEABEFFF1930C7F999CB1FDD2F85F" bold="true" box="[199,534,1981,2007]" pageId="34" pageNumber="35">Global stratigraphic range:</emphasis>
<taxonomicName id="56DE4D2FFFF1930C7D7F9CB1FD36F85F" authorityName="d'Orbigny" authorityYear="1839" box="[545,754,1981,2007]" class="Globothalamea" family="Globigerinidae" genus="Orbulina" kingdom="Chromista" order="Rotaliida" pageId="34" pageNumber="35" phylum="Foraminifera" rank="species" species="universa">
<emphasis id="A3AAEABEFFF1930C7D7F9CB1FD36F85F" box="[545,754,1981,2007]" italics="true" pageId="34" pageNumber="35">Orbulina universa</emphasis>
</taxonomicName>
is an extant species with its first appearance datum in the middle Miocene (Langhian) (
<bibRefCitation id="F54F4B5DFFF1930C7EBE9CEDFCD9F873" author="Kennett, J. P. &amp; Srinivasan, M. S." box="[480,797,2017,2043]" pageId="34" pageNumber="35" refId="ref33851" refString="Kennett, J. P. &amp; Srinivasan, M. S. (1983) Neogene Planktonic Foraminifera. A Phylogenetic Atlas. Hutchinson Ross, Stroudsburg, 265 pp." type="book" year="1983">Kennett &amp; Srinivasan, 1983</bibRefCitation>
;
<bibRefCitation id="F54F4B5DFFF1930C7C779CEDFC75F873" author="Jones, R. W." box="[809,945,2017,2043]" pageId="34" pageNumber="35" refId="ref33343" refString="Jones, R. W. (1994) The Challenger Foraminifera. Oxford University Press, Oxford, 149 pp." type="book" year="1994">Jones, 1994</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFF0930D7F999B9AFBAEFF70" blockId="35.[151,1436,150,428]" pageId="35" pageNumber="36">
<emphasis id="A3AAEABEFFF0930D7F999B9AFDD2FF38" bold="true" box="[199,534,150,176]" pageId="35" pageNumber="36">Environmental preferences:</emphasis>
This species is cosmopolitan and occurs widespread (
<bibRefCitation id="F54F4B5DFFF0930D7B259B9AFA54FF39" author="BouDagher-Fadel, M. K." box="[1147,1424,150,177]" pageId="35" pageNumber="36" refId="ref28678" refString="BouDagher-Fadel, M. K. (2015) Biostratigraphic and geological significance of planktonic foraminifera. UCL Press, London, 306 pp. https: // doi. org / 10.2307 / j. ctt 1 g 69 xwk" type="book" year="2015">BouDagher-Fadel, 2015</bibRefCitation>
), but has been documented to be more adapted to subtropical conditions, preferring warmer waters (
<bibRefCitation id="F54F4B5DFFF0930D7B8D9BB6FF16FF70" author="Zachariasse, W. J. &amp; Jorissen, F. J. &amp; Perissoratis, C. &amp; Rohling, E. J. &amp; Tsapralis, V." pageId="35" pageNumber="36" pagination="391 - 394" refId="ref39285" refString="Zachariasse, W. J., Jorissen, F. J., Perissoratis, C., Rohling, E. J. &amp; Tsapralis, V. (1997) Late Quaternary foraminiferal changes and the nature of sapropel S 1 in Skopelos Basin. Proceedings of the 5 th Hellenic symposium on Oceanography and Fisheries, Kavalla, Greece, 1997, 391 - 394." type="journal article" year="1997">
Zachariasse
<emphasis id="A3AAEABEFFF0930D7A019BB7FA4AFF5C" box="[1375,1422,186,212]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
., 1997
</bibRefCitation>
;
<bibRefCitation id="F54F4B5DFFF0930D7F809BD2FE10FF70" author="Gallagher, S. J. &amp; Jonasson, K. &amp; Wallace, M. W. &amp; Holdgate, G. R. &amp; Taylor, D." box="[222,468,222,249]" pageId="35" pageNumber="36" pagination="53 - 80" refId="ref31944" refString="Gallagher, S. J., Smith. A. J., Jonasson, K., Wallace, M. W., Holdgate, G. R., Daniels. J. &amp; Taylor, D. (2001) The Miocene palaeoenvironmental and palaeoceanographic evolution of the Gippsland Basin, Southeast Australia: a record of Southern Ocean change. Palaeogeography, Palaeoclimatology, Palaeoecology, 172, 53 - 80. https: // doi. org / 10.1016 / S 0031 - 0182 (01) 00271 - 1" type="journal article" year="2001">
Gallagher
<emphasis id="A3AAEABEFFF0930D7E0D9BD3FE41FF70" box="[339,389,222,248]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
., 2001
</bibRefCitation>
;
<bibRefCitation id="F54F4B5DFFF0930D7EBE9BD3FDBEFF70" author="Kucera, M." box="[480,634,222,249]" pageId="35" pageNumber="36" pagination="213 - 262" refId="ref33999" refString="Kucera, M. (2007) Planktonic foraminifera as tracers of past oceanic conditions. In: Hillaire-Marcel, C. &amp; de Vernal, A. (Eds.), Proxies in Late Cenozoic Paleoceanography. Elsevier, Amsterdam, 213 - 262. https: // doi. org / 10.1016 / S 1572 - 5480 (07) 01011 - 1" type="book chapter" year="2007">Kucera, 2007</bibRefCitation>
;
<bibRefCitation id="F54F4B5DFFF0930D7DD89BD2FC91FF70" author="Drinia, H. A. &amp; Antonarakou, A. &amp; Kontakiotis, G." box="[646,853,222,249]" pageId="35" pageNumber="36" pagination="63 - 78" refId="ref31090" refString="Drinia, H. A., Antonarakou, A. &amp; Kontakiotis, G. (2008) On the occurrence of Early Pliocene marine deposits in the Ierapetra Basin, Eastern Crete, Greece. Bulletin of Geosciences, 83 (1), 63 - 78. https: // doi. org / 10.3140 / bull. geosci. 2008.01.063" type="journal article" year="2008">
Drinia
<emphasis id="A3AAEABEFFF0930D7D8A9BD3FCC1FF70" box="[724,773,222,248]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
., 2008
</bibRefCitation>
;
<bibRefCitation id="F54F4B5DFFF0930D7C3F9BD2FB99FF70" author="Hemleben, C. &amp; Spindler, M. &amp; Anderson, O. R." box="[865,1117,222,249]" pageId="35" pageNumber="36" refId="ref32764" refString="Hemleben, C., Spindler, M. &amp; Anderson, O. R. (1990) Modern Planktonic Foraminifera. Springer Science and Business Media, New York, 386 pp. https: // doi. org / 10.1007 / 978 - 1 - 4612 - 3544 - 6" type="book" year="1990">
Hemleben
<emphasis id="A3AAEABEFFF0930D7C859BD3FBC8FF70" box="[987,1036,222,248]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
., 1990
</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFF0930D7F999A0EFAF6FE24" blockId="35.[151,1436,150,428]" pageId="35" pageNumber="36">
<emphasis id="A3AAEABEFFF0930D7F999A0EFE02FE94" bold="true" box="[199,454,258,284]" pageId="35" pageNumber="36">Regional occurrence:</emphasis>
<taxonomicName id="56DE4D2FFFF0930D7E909A0EFD58FE94" authorityName="d'Orbigny" authorityYear="1839" box="[462,668,258,284]" class="Globothalamea" family="Globigerinidae" genus="Orbulina" kingdom="Chromista" order="Rotaliida" pageId="35" pageNumber="36" phylum="Foraminifera" rank="species" species="universa">
<emphasis id="A3AAEABEFFF0930D7E909A0EFD58FE94" box="[462,668,258,284]" italics="true" pageId="35" pageNumber="36">Orbulina universa</emphasis>
</taxonomicName>
is stratigraphically and spatially widespread along the southern and western margin of Africa. This species has been documented to occur in Miocene-aged strata of the
<collectingCountry id="E9C9763CFFF0930D7A579A2AFA90FEC8" box="[1289,1364,294,320]" name="Democratic Republic of the Congo" pageId="35" pageNumber="36">Congo</collectingCountry>
Basin (
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) to the northern Namibian outer continental shelf, south of the Kunene River mouth (this study) and the southwestern shelf of
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(
<bibRefCitation id="F54F4B5DFFF0930D7D0B9A62FC8EFE00" author="Compton, J. S. &amp; Wigley, R. &amp; McMillan, I. K." box="[597,842,366,392]" pageId="35" pageNumber="36" pagination="19 - 40" refId="ref29411" refString="Compton, J. S., Wigley, R. &amp; McMillan, I. K. (2004) Late Cenozoic phosphogenesis on the western shelf of South Africa in the vicinity of the Cape Canyon. Marine Geology, 206, 19 - 40. https: // doi. org / 10.1016 / j. margeo. 2004.02.004" type="journal article" year="2004">
Compton
<emphasis id="A3AAEABEFFF0930D7D989A63FCC1FE00" box="[710,773,366,392]" italics="true" pageId="35" pageNumber="36">et al.,</emphasis>
2004
</bibRefCitation>
).
<bibRefCitation id="F54F4B5DFFF0930D7C029A62FBF5FE00" author="Wefer, G. &amp; Berger, W. H. &amp; Richter, C. &amp; Adams, D. D. &amp; Anderon, L. D. &amp; Andreasen, D. J. &amp; Bruchert, V. &amp; Cambray, H. &amp; Christensen, B. A. &amp; Frost, G. M. &amp; Giraudeau, J. &amp; Gorgas, T. J. &amp; Hermelin, O. &amp; Lange, C. B. &amp; Laser, B. &amp; Lin, H-L. &amp; Maslin, M. &amp; Meyers, P. A. &amp; Motoyama, I. &amp; Murray, R. W. &amp; Pato, D. &amp; Perez, M. E. &amp; Pufahl, P. K. &amp; Spiess, V. &amp; Wigley, R. &amp; Yamazaki, T." box="[860,1073,366,393]" pageId="35" pageNumber="36" refId="ref38965" refString="Wefer, G., Berger, W. H., Richter, C., Adams, D. D., Anderon, L. D., Andreasen, D. J., Bruchert, V., Cambray, H., Christensen, B. A., Frost, G. M., Giraudeau, J., Gorgas, T. J., Hermelin, O., Lange, C. B., Laser, B., Lin, H-L., Maslin, M., Meyers, P. A., Motoyama, I., Murray, R. W., Pato, D., Perez, M. E., Pufahl, P. K., Spiess, V., Vidal. L., Wigley, R. &amp; Yamazaki, T. (1998) Proceedings of the Ocean Drilling Program, Initial Reports. Vol. 175. Ocean Drilling Program, College Station, Texas, 577 pp." type="book" year="1998">
Wefer
<emphasis id="A3AAEABEFFF0930D7CF99A63FC1CFE00" box="[935,984,366,392]" italics="true" pageId="35" pageNumber="36">et al</emphasis>
. (1998)
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reported the species to occur on the slope between central
<collectingCountry id="E9C9763CFFF0930D7EE49A9EFDDEFE24" box="[442,538,402,428]" name="Namibia" pageId="35" pageNumber="36">Namibia</collectingCountry>
and offshore of the Orange River during the Pliocene and Pleistocene.
</paragraph>
</subSubSection>
</treatment>
</document>