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<document id="6E3D1BA147DA4BEEA9704F38C947E7E1" ID-CLB-Dataset="2636" ID-DOI="10.11646/zootaxa.5091.1.1" ID-GBIF-Dataset="51b42715-ddb1-4b4a-aad4-8be0f600a287" ID-ISSN="1175-5326" ID-Zenodo-Dep="5840434" ID-ZooBank="ABC8AF70-F691-4D07-8F20-70934642C8BC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1641986550580" checkinUser="plazi" docAuthor="Bergh, Eugene W. &amp; Compton, John S." docDate="2022" docId="197787BAFFC3933E7FC99E71FB18F85B" docLanguage="en" docName="zootaxa.5091.1.1.pdf" docOrigin="Zootaxa 5091 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Saracenaria spinosa Eichenberg 1935" docType="treatment" docVersion="6" lastPageNumber="17" masterDocId="E54EFFC2FFD3932E7F5E9B0CFFC4FF88" masterDocTitle="Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf" masterLastPageNumber="55" masterPageNumber="1" pageNumber="17" updateTime="1699301522325" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A6C7EE8CDCBBA3A827B2C12097445CBD">Taxonomy of Middle Miocene foraminifera from the northern Namibian continental shelf</mods:title>
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<mods:namePart id="244486BAB6756F02986B2F05877F696F">Bergh, Eugene W.</mods:namePart>
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<mods:affiliation id="6B8795712E4867A0AF20750EBBD64233">Marine Research Institute and Department of Geological Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa. &amp; Department of Environmental and Geographical Sciences, University of Cape Town, Private Bag X 3, Rondebosch, 7701, South Africa; &amp; Invertebrate palaeontology and Geology, Iziko South African Museum, P. O. Box 61, Cape Town 8000, South Africa. john. compton @ uct. ac. za; https: // orcid. org / 0000 - 0002 - 0765 - 4141</mods:affiliation>
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<mods:namePart id="6609671E14CFCD53C9D0403E40A7A0D8">Compton, John S.</mods:namePart>
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<treatment id="197787BAFFC3933E7FC99E71FB18F85B" ID-DOI="http://doi.org/10.5281/zenodo.5840497" ID-GBIF-Taxon="192044121" ID-Zenodo-Dep="5840497" LSID="urn:lsid:plazi:treatment:197787BAFFC3933E7FC99E71FB18F85B" httpUri="http://treatment.plazi.org/id/197787BAFFC3933E7FC99E71FB18F85B" lastPageNumber="17" pageId="16" pageNumber="17">
<subSubSection id="D9C46527FFC3933E7FC99E71FE8CFA33" pageId="16" pageNumber="17" type="nomenclature">
<paragraph id="916136ACFFC3933E7FC99E71FDABFA1F" blockId="16.[151,623,1405,1467]" box="[151,623,1405,1431]" pageId="16" pageNumber="17">
<heading id="CA2981C0FFC3933E7FC99E71FDABFA1F" bold="true" box="[151,623,1405,1431]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<taxonomicName id="56DE4D2FFFC3933E7FC99E71FDABFA1F" ID-CoL="79KL2" authority="Eichenberg, 1935" authorityName="Eichenberg" authorityYear="1935" box="[151,623,1405,1431]" class="Nodosariata" family="Vaginulinidae" genus="Saracenaria" kingdom="Chromista" order="Vaginulinida" pageId="16" pageNumber="17" phylum="Foraminifera" rank="species" species="spinosa">
<emphasis id="A3AAEABEFFC3933E7FC99E71FDABFA1F" bold="true" box="[151,623,1405,1431]" pageId="16" pageNumber="17">
<emphasis id="A3AAEABEFFC3933E7FC99E71FE4AFA1F" bold="true" box="[151,398,1405,1431]" italics="true" pageId="16" pageNumber="17">Saracenaria spinosa</emphasis>
<bibRefCitation id="F54F4B5DFFC3933E7ECB9E71FDABFA1F" author="Eichenberg, W." box="[405,623,1405,1431]" pageId="16" pageNumber="17" pagination="1 - 40" refId="ref31338" refString="Eichenberg, W. (1935) Die Erforschung der Mikroorganismen insbesondere der Foraminiferen der norddeutschen Erdolfelder; Teil 1 - Die Foraminiferen der Unterkreide; Folge 4 - Foraminiferen aus dem Apt von Wenden am Mittellandkanal. Jahresbericht des Niedersachsischen Geologischen Vereins, 27, 1 - 40." type="journal article" year="1935">Eichenberg, 1935</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="916136ACFFC3933E7FC99EADFE8CFA33" blockId="16.[151,623,1405,1467]" box="[151,328,1441,1467]" pageId="16" pageNumber="17">
<heading id="CA2981C0FFC3933E7FC99EADFE8CFA33" bold="true" box="[151,328,1441,1467]" fontSize="11" level="1" pageId="16" pageNumber="17" reason="1">
<emphasis id="A3AAEABEFFC3933E7FC99EADFE8CFA33" bold="true" box="[151,328,1441,1467]" pageId="16" pageNumber="17">Pl. 3, figs. 3ac</emphasis>
</heading>
</paragraph>
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<subSubSection id="D9C46527FFC3933E7FC99EEAFBF5FA76" box="[151,1073,1510,1534]" pageId="16" pageNumber="17" type="reference_group">
<paragraph id="916136ACFFC3933E7FC99EEAFBF5FA76" blockId="16.[151,1073,1510,1534]" box="[151,1073,1510,1534]" pageId="16" pageNumber="17">
<treatmentCitationGroup id="B1CE1182FFC3933E7FC99EEAFBF5FA76" box="[151,1073,1510,1534]" pageId="16" pageNumber="17">
<taxonomicName id="56DE4D2FFFC3933E7FC99EEAFDE7FA76" ID-CoL="79KL2" authority="Eichenberg, 1935" authorityName="Eichenberg" authorityYear="1935" box="[151,547,1510,1534]" class="Nodosariata" family="Vaginulinidae" genus="Saracenaria" kingdom="Chromista" order="Vaginulinida" pageId="16" pageNumber="17" phylum="Foraminifera" rank="species" species="spinosa">
<emphasis id="A3AAEABEFFC3933E7FC99EEAFEACFA75" box="[151,360,1510,1533]" italics="true" pageId="16" pageNumber="17">Saracenaria spinosa</emphasis>
<bibRefCitation id="F54F4B5DFFC3933E7E319EEAFDE7FA76" author="Eichenberg, W." box="[367,547,1510,1534]" pageId="16" pageNumber="17" pagination="1 - 40" refId="ref31338" refString="Eichenberg, W. (1935) Die Erforschung der Mikroorganismen insbesondere der Foraminiferen der norddeutschen Erdolfelder; Teil 1 - Die Foraminiferen der Unterkreide; Folge 4 - Foraminiferen aus dem Apt von Wenden am Mittellandkanal. Jahresbericht des Niedersachsischen Geologischen Vereins, 27, 1 - 40." type="journal article" year="1935">Eichenberg, 1935</bibRefCitation>
</taxonomicName>
, pl. 4, fig.5;
<treatmentCitation id="107F10BDFFC3933E7DFB9EEAFC55FA76" author="Bolli, H. M. &amp; Beckmann, J. P. &amp; Sanders, J. B." box="[677,913,1510,1534]" page="28" pageId="16" pageNumber="17" year="1994">
<bibRefCitation id="F54F4B5DFFC3933E7DFB9EEAFC96FA76" author="Bolli, H. M. &amp; Beckmann, J. P. &amp; Sanders, J. B." box="[677,850,1510,1534]" pageId="16" pageNumber="17" refId="ref28525" refString="Bolli, H. M., Beckmann, J. P. &amp; Sanders, J. B. (1994) Benthic Foraminiferal Biostratigraphy of the South Caribbean Region. Cambridge University Press, Cambridge, 408 pp. https: // doi. org / 10.1017 / CBO 9780511564406" type="book" year="1994">
Bolli
<emphasis id="A3AAEABEFFC3933E7D809EEBFCCFFA76" box="[734,779,1510,1534]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
., 1994
</bibRefCitation>
, p. 28
</treatmentCitation>
, fig. 8.448.48.
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</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC3933E7FC99D20FB82F95E" pageId="16" pageNumber="17" type="description">
<paragraph id="916136ACFFC3933E7FC99D20FB82F95E" blockId="16.[151,1437,1580,2003]" pageId="16" pageNumber="17">
<emphasis id="A3AAEABEFFC3933E7FC99D20FEEFF9CE" bold="true" box="[151,299,1580,1606]" pageId="16" pageNumber="17">Description:</emphasis>
The test wall is calcareous, smooth and finely perforate. The test is planispiral, becoming rectilinear and is triangular in section, almost twice as long as wide. Ridges form along the test surface, ending in spines along margins and pointing away from the apertural end. Chambers enlarge toward the terminal end, with the terminal chamber being the largest. The initial end is slightly curved away from the apertural end. The apertural face is triangular and broad. The aperture is radiate at the dorsal angle of the final chamber.
</paragraph>
</subSubSection>
<subSubSection id="D9C46527FFC3933E7F999DECFB18F85B" pageId="16" pageNumber="17" type="discussion">
<paragraph id="916136ACFFC3933E7F999DECFDF9F896" blockId="16.[151,1437,1580,2003]" pageId="16" pageNumber="17">
<emphasis id="A3AAEABEFFC3933E7F999DECFEF9F972" bold="true" box="[199,317,1760,1786]" pageId="16" pageNumber="17">Remarks:</emphasis>
Specimens of this species are relatively large, measuring up to
<quantity id="56269B49FFC3933E7CA19DECFB91F973" box="[1023,1109,1760,1787]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="16" pageNumber="17" unit="mm" value="0.5">0.5 mm</quantity>
in width and
<quantity id="56269B49FFC3933E7BB29DECFAE9F973" box="[1260,1325,1760,1787]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="16" pageNumber="17" unit="mm" value="1.0">1 mm</quantity>
in length. The relative abundance is low (&lt;1%).
</paragraph>
<paragraph id="916136ACFFC3933E7F999C24FAE0F8CA" blockId="16.[151,1437,1580,2003]" box="[199,1316,1832,1859]" pageId="16" pageNumber="17">
<emphasis id="A3AAEABEFFC3933E7F999C24FEACF8CA" bold="true" box="[199,360,1832,1858]" pageId="16" pageNumber="17">Life strategy:</emphasis>
Species of the genus
<taxonomicName id="56DE4D2FFFC3933E7D059C24FD21F8CA" authorityName="Defrance" authorityYear="1824" box="[603,741,1832,1858]" class="Nodosariata" family="Vaginulinidae" genus="Saracenaria" kingdom="Chromista" order="Vaginulinida" pageId="16" pageNumber="17" phylum="Foraminifera" rank="genus">
<emphasis id="A3AAEABEFFC3933E7D059C24FD21F8CA" box="[603,741,1832,1858]" italics="true" pageId="16" pageNumber="17">Saracenaria</emphasis>
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are generally epifaunal (
<bibRefCitation id="F54F4B5DFFC3933E7CA29C24FADCF8CA" author="Barbieri, R. &amp; Panieri, G." box="[1020,1304,1832,1858]" pageId="16" pageNumber="17" pagination="257 - 275" refId="ref27706" refString="Barbieri, R. &amp; Panieri, G. (2004) How are benthic foraminiferal faunas influenced by cold seeps? Evidence from the Miocene of Italy. Palaeogeography, Palaeoclimatology, Palaeoecology, 204, 257 - 275. https: // doi. org / 10.1016 / S 0031 - 0182 (03) 00730 - 2" type="journal article" year="2004">Barbieri &amp; Panieri, 2004</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFC3933E7F999C40FF24F802" blockId="16.[151,1437,1580,2003]" pageId="16" pageNumber="17">
<emphasis id="A3AAEABEFFC3933E7F999C40FDDCF8EE" bold="true" box="[199,536,1868,1894]" pageId="16" pageNumber="17">Global stratigraphic range:</emphasis>
<taxonomicName id="56DE4D2FFFC3933E7D7D9C40FD5EF8EE" authorityName="Eichenberg" authorityYear="1935" box="[547,666,1868,1894]" class="Nodosariata" family="Vaginulinidae" genus="Saracenaria" kingdom="Chromista" order="Vaginulinida" pageId="16" pageNumber="17" phylum="Foraminifera" rank="species" species="spinosa">
<emphasis id="A3AAEABEFFC3933E7D7D9C40FD5EF8EE" box="[547,666,1868,1894]" italics="true" pageId="16" pageNumber="17">S. spinosa</emphasis>
</taxonomicName>
is recorded to occur from the Cretaceous to Recent (
<bibRefCitation id="F54F4B5DFFC3933E7A4A9C40FF17F802" author="Bolli, H. M. &amp; Beckmann, J. P. &amp; Sanders, J. B." pageId="16" pageNumber="17" refId="ref28525" refString="Bolli, H. M., Beckmann, J. P. &amp; Sanders, J. B. (1994) Benthic Foraminiferal Biostratigraphy of the South Caribbean Region. Cambridge University Press, Cambridge, 408 pp. https: // doi. org / 10.1017 / CBO 9780511564406" type="book" year="1994">
Bolli
<emphasis id="A3AAEABEFFC3933E7A079C41FA4AF8EE" box="[1369,1422,1868,1894]" italics="true" pageId="16" pageNumber="17">et al</emphasis>
., 1994
</bibRefCitation>
).
</paragraph>
<paragraph id="916136ACFFC3933E7F999C98FB18F85B" blockId="16.[151,1437,1580,2003]" pageId="16" pageNumber="17">
<emphasis id="A3AAEABEFFC3933E7F999C98FE07F826" bold="true" box="[199,451,1940,1966]" pageId="16" pageNumber="17">Regional occurrence:</emphasis>
This is the first documentation of this species in the region and occurs in middle Miocene sediments on the Namibian outer continental shelf, south of the
<collectingRegion id="531AF84EFFC3933E7C3D9CB5FBC4F85A" box="[867,1024,1976,2003]" isEnumeration="true" pageId="16" pageNumber="17">
<collectingRegion id="531AF84EFFC3933E7C3D9CB5FC7EF85B" box="[867,954,1977,2003]" country="Namibia" name="Kunene" pageId="16" pageNumber="17">Kunene</collectingRegion>
<collectingRegion id="531AF84EFFC3933E7C9C9CB4FBC4F85A" box="[962,1024,1976,2002]" country="Nigeria" name="Rivers" pageId="16" pageNumber="17">River</collectingRegion>
</collectingRegion>
mouth (this study).
</paragraph>
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