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<document id="9161E1FD40612F19008F75B0538904BF" ID-DOI="10.1590/S1679-62252008000200001" ID-ISSN="1982-0224" ID-Zenodo-Dep="10558366" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="admin" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="julia" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="julia" IM.treatments_approvedBy="admin" checkinTime="1698180736224" checkinUser="julia" docAuthor="Lucinda, Paulo Henrique Franco" docDate="2008" docId="0D2787D41D54FFA4DFAF51DCC19BFCEF" docLanguage="en" docName="NeotropIchthyol.6.2.113-158.pdf.imd" docOrigin="Neotropical Ichthyology 6 (2)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252008000200001&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Phalloceros titthos Lucinda 2008, new species" docType="treatment" docVersion="8" lastPageNumber="134" masterDocId="F11EFFAC1D40FFB1DC6D516EC41EFF81" masterDocTitle="Systematics and biogeography of the genus Phalloceros Eigenmann, 1907 (Cyprinodontiformes: Poeciliidae: Poeciliinae), with the description of twenty-one new species" masterLastPageNumber="158" masterPageNumber="113" pageNumber="133" updateTime="1706210653105" updateUser="ExternalLinkService">
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<mods:title id="C9447BC6D29932EC0B07D45A71E40D6B">Systematics and biogeography of the genus Phalloceros Eigenmann, 1907 (Cyprinodontiformes: Poeciliidae: Poeciliinae), with the description of twenty-one new species</mods:title>
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<mods:namePart id="8EDC80568EA311CFBB6135287012EA70">Lucinda, Paulo Henrique Franco</mods:namePart>
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<mods:title id="CA98BC14679B14E4C27DFBF1BF033308">Neotropical Ichthyology</mods:title>
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<mods:date id="01F85C8B34CB4441334EE3E9B6731BD5">2008</mods:date>
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<mods:number id="0CDB907A6CB9C07F0A62823C3F4F9BEF">2008-12-31</mods:number>
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<mods:classification id="8232DA41DAC0EB07C2AB7B0494C4C895">journal article</mods:classification>
<mods:identifier id="72A2379B0C1DE3CF0EE9480A83BED139" type="DOI">10.1590/S1679-62252008000200001</mods:identifier>
<mods:identifier id="77A2026161270BEC38E8B405A6436E98" type="ISSN">1982-0224</mods:identifier>
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<treatment id="0D2787D41D54FFA4DFAF51DCC19BFCEF" ID-DOI="http://doi.org/10.5281/zenodo.10064502" ID-Zenodo-Dep="10064502" LSID="urn:lsid:plazi:treatment:0D2787D41D54FFA4DFAF51DCC19BFCEF" httpUri="http://treatment.plazi.org/id/0D2787D41D54FFA4DFAF51DCC19BFCEF" lastPageId="21" lastPageNumber="134" pageId="20" pageNumber="133">
<subSubSection id="CD9465491D54FFA5DFAF51DCC13FFF48" box="[962,1313,178,201]" pageId="20" pageNumber="133" type="nomenclature">
<paragraph id="853136C21D54FFA5DFAF51DCC13FFF48" blockId="20.[962,1313,178,233]" box="[962,1313,178,201]" pageId="20" pageNumber="133">
<heading id="DE7981AE1D54FFA5DFAF51DCC13FFF48" box="[962,1313,178,201]" centered="true" fontSize="10" level="2" pageId="20" pageNumber="133" reason="2">
<emphasis id="B7FAEAD01D54FFA5DFAF51DCC13FFF48" bold="true" box="[962,1313,178,201]" pageId="20" pageNumber="133">
<taxonomicName id="428E4D411D54FFA5DFAF51DCC08FFF48" ID-CoL="777JX" authority="Lucinda, 2008" authorityName="Lucinda" authorityYear="2008" box="[962,1169,178,201]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="titthos" status="sp. nov.">
<emphasis id="B7FAEAD01D54FFA5DFAF51DCC08FFF48" bold="true" box="[962,1169,178,201]" italics="true" pageId="20" pageNumber="133">Phalloceros titthos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="ACC957AB1D54FFA5D8F351DCC13FFF48" box="[1182,1313,178,201]" pageId="20" pageNumber="133" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CD9465491D54FFA5D85B51BCC11BFEA8" pageId="20" pageNumber="133" type="description">
<paragraph id="853136C21D54FFA5D85B51BCC0B3FF68" blockId="20.[962,1313,178,233]" box="[1078,1197,210,233]" pageId="20" pageNumber="133">
<figureCitation id="1DB52A471D54FFA5D85B51BCC094FF68" box="[1078,1162,210,233]" captionStart="Fig" captionStartId="21.[129,167,700,723]" captionTargetBox="[340,1247,170,679]" captionTargetId="figure-497@21.[340,1247,170,679]" captionTargetPageId="21" captionText="Fig. 25. Phalloceros titthos. (A) Male, 17.1 mm SL, USNM 297945, rio da Praia, near Guaratuba; (B) holotype, female, 35.9 mm SL, MZUSP 79670, creek near beach, Guaratuba, Paraná, Brazil." figureDoi="http://doi.org/10.5281/zenodo.10558390" httpUri="https://zenodo.org/record/10558390/files/figure.png" pageId="20" pageNumber="133">Figs. 25</figureCitation>
,
<figureCitation id="1DB52A471D54FFA5D8FE51BCC0B3FF68" box="[1171,1197,210,233]" captionStart="Fig" captionStartId="21.[129,168,1904,1927]" captionTargetBox="[129,782,1506,1872]" captionTargetId="figure-476@21.[129,772,1529,1888]" captionText="Fig. 26. Upper and lower lip of Phalloceros titthos, MZUSP 43467, showing symphyseal papilla (SP). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10569764" httpUri="https://zenodo.org/record/10569764/files/figure.png" pageId="20" pageNumber="133">26</figureCitation>
</paragraph>
<paragraph id="853136C21D54FFA5DF5D507CC11BFEA8" blockId="20.[816,1285,274,297]" box="[816,1285,274,297]" pageId="20" pageNumber="133">
<taxonomicName id="428E4D411D54FFA5DF5D507CC7ACFEA8" ID-CoL="6MQG" ID-ENA="55107" authority="Lucinda, 2008" authorityName="Lucinda" authorityYear="2008" box="[816,946,274,297]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="genus" status="sp. nov.">
<emphasis id="B7FAEAD01D54FFA5DF5D507CC7ACFEA8" box="[816,946,274,297]" italics="true" pageId="20" pageNumber="133">Phalloceros</emphasis>
</taxonomicName>
<taxonomicNameLabel id="ACC957AB1D54FFA5DFD5507CC7EEFEA8" box="[952,1008,274,297]" pageId="20" pageNumber="133" rank="species">sp. n.</taxonomicNameLabel>
T
<bibRefCitation id="E11F4B331D54FFA5D867507CC11FFEA8" author="Lucinda, P. H. F. &amp; R. E. Reis" box="[1034,1281,274,297]" journalOrPublisher="Neotropical Ichthyology" pageId="20" pageNumber="133" pagination="1 - 60" part="3" refId="ref43822" refString="Lucinda, P. H. F. &amp; R. E. Reis. 2005. Systematics of the subfamily Poeciliinae Bonaparte (Cyprinodontiformes, Poeciliidae). Neotropical Ichthyology, 3: 1 - 60." title="Systematics of the subfamily Poeciliinae Bonaparte (Cyprinodontiformes, Poeciliidae)" type="journal article" year="2005">Lucinda &amp; Reis (2005)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CD9465491D54FFA5DF5D503FC0EFFD92" pageId="20" pageNumber="133" type="materials_examined">
<paragraph id="853136C21D54FFA5DF5D503FC7C5FE21" blockId="20.[816,1459,336,531]" pageId="20" pageNumber="133">
<materialsCitation id="35E63C9F1D54FFA5DF5D503FC7C5FE21" collectingDate="1977-09-23" collectionCode="MZUSP" country="Brazil" latitude="-25.883333" longitude="-48.616665" pageId="20" pageNumber="133" typeStatus="holotype">
<emphasis id="B7FAEAD01D54FFA5DF5D503FC78FFEE7" bold="true" box="[816,913,337,358]" pageId="20" pageNumber="133">
<typeStatus id="5A3588601D54FFA5DF5D503FC78FFEE7" box="[816,913,337,358]" pageId="20" pageNumber="133" type="holotype">Holotype</typeStatus>
</emphasis>
.
<collectionCode id="E39FAE071D54FFA5DFCF503FC7EAFEE7" box="[930,1012,337,358]" collectionName="MZUSP" pageId="20" pageNumber="133">MZUSP</collectionCode>
79670, creek near beach, approx.
<geoCoordinate id="E0BA50051D54FFA5D90E503FC1ADFEE7" box="[1379,1459,336,358]" degrees="25" direction="south" minutes="53" orientation="latitude" pageId="20" pageNumber="133" precision="925" value="-25.883333">
25
<superScript id="72FB9B8A1D54FFA5D910503EC19AFEDD" attach="right" box="[1405,1412,336,348]" fontSize="5" pageId="20" pageNumber="133">o</superScript>
53S
</geoCoordinate>
<geoCoordinate id="E0BA50051D54FFA5DF5D5000C797FE02" box="[816,905,365,387]" degrees="48" direction="west" minutes="37" orientation="longitude" pageId="20" pageNumber="133" precision="925" value="-48.616665">
48
<superScript id="72FB9B8A1D54FFA5DF245003C74EFEF8" attach="right" box="[841,848,365,377]" fontSize="5" pageId="20" pageNumber="133">o</superScript>
37W
</geoCoordinate>
, Guaratuba, Paraná,
<collectingCountry id="FD9976521D54FFA5D8095000C0BAFE02" box="[1124,1188,366,387]" name="Brazil" pageId="20" pageNumber="133" value="Brazil">Brazil</collectingCountry>
,
<date id="F13010021D54FFA5D8DC5000C129FE02" box="[1201,1335,366,387]" pageId="20" pageNumber="133" value="1977-09-23">23 Sep 1977</date>
, Expedition
<collectionCode id="E39FAE071D54FFA5DF5D50E5C79BFE21" box="[816,901,395,416]" collectionName="MZUSP" pageId="20" pageNumber="133">MZUSP</collectionCode>
/
<collectionCode id="E39FAE071D54FFA5DFE050E5C7C9FE21" box="[909,983,395,416]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="20" pageNumber="133">USNM</collectionCode>
.
</materialsCitation>
</paragraph>
<paragraph id="853136C21D54FFA5DF5D50C6C7ABFE58" blockId="20.[816,1459,336,531]" pageId="20" pageNumber="133">
<materialsCitation id="35E63C9F1D54FFA5DF5D50C6C7ABFE58" collectionCode="MZUSP" country="Brazil" pageId="20" pageNumber="133" stateProvince="Parana" typeStatus="paratype">
<emphasis id="B7FAEAD01D54FFA5DF5D50C6C789FE3C" bold="true" box="[816,919,424,445]" pageId="20" pageNumber="133">
<typeStatus id="5A3588601D54FFA5DF5D50C6C789FE3C" box="[816,919,424,445]" pageId="20" pageNumber="133" type="paratype">Paratypes</typeStatus>
</emphasis>
.
<collectingCountry id="FD9976521D54FFA5DFCF50C6C7E2FE3C" box="[930,1020,424,445]" name="Brazil" pageId="20" pageNumber="133" value="Brazil">BRAZIL</collectingCountry>
.
<collectingRegion id="474AF8201D54FFA5D86950C6C057FE3C" box="[1028,1097,424,445]" pageId="20" pageNumber="133">Paraná</collectingRegion>
.
<collectionCode id="E39FAE071D54FFA5D83D50C6C0BEFE3C" box="[1104,1184,424,445]" collectionName="MZUSP" pageId="20" pageNumber="133">MZUSP</collectionCode>
43467, 26/3*, collected with the holotype.
</materialsCitation>
</paragraph>
<paragraph id="853136C21D54FFA5DF5D508FC0EFFD92" blockId="20.[816,1459,336,531]" pageId="20" pageNumber="133">
<materialsCitation id="35E63C9F1D54FFA5DF5D508FC0EFFD92" collectingDate="1988-02-18" collectionCode="USNM" country="Brazil" pageId="20" pageNumber="133" stateProvince="Parana">
<emphasis id="B7FAEAD01D54FFA5DF5D508FC784FE77" bold="true" box="[816,922,481,502]" pageId="20" pageNumber="133">Non-types</emphasis>
.
<collectingCountry id="FD9976521D54FFA5DFCB508FC01FFE77" box="[934,1025,481,502]" name="Brazil" pageId="20" pageNumber="133" value="Brazil">BRAZIL</collectingCountry>
.
<collectingRegion id="474AF8201D54FFA5D867508FC04EFE77" box="[1034,1104,481,502]" pageId="20" pageNumber="133">Paraná</collectingRegion>
.
<collectionCode id="E39FAE071D54FFA5D835508FC080FE77" box="[1112,1182,481,502]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="20" pageNumber="133">USNM</collectionCode>
297945, 13/2*, rio da
<collectingRegion id="474AF8201D54FFA5D916508FC1AEFE77" box="[1403,1456,481,502]" pageId="20" pageNumber="133">Praia</collectingRegion>
, near Guaratuba,
<date id="F13010021D54FFA5DFB95090C04FFD92" box="[980,1105,510,531]" pageId="20" pageNumber="133" value="1988-02-18">18 Feb 1988</date>
, R.M.C. Castro.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="CD9465491D54FFA5DF5D5355C15BFD13" pageId="20" pageNumber="133" type="diagnosis">
<paragraph id="853136C21D54FFA5DF5D5355C15BFD13" blockId="20.[816,1459,571,658]" pageId="20" pageNumber="133">
<emphasis id="B7FAEAD01D54FFA5DF5D5355C781FDD3" bold="true" box="[816,927,571,594]" pageId="20" pageNumber="133">Diagnosis</emphasis>
.
<taxonomicName id="428E4D411D54FFA5DFC15355C067FDD3" authorityName="Lucinda" authorityYear="2008" box="[940,1145,571,594]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="titthos">
<emphasis id="B7FAEAD01D54FFA5DFC15355C067FDD3" box="[940,1145,571,594]" italics="true" pageId="20" pageNumber="133">Phalloceros titthos</emphasis>
</taxonomicName>
can be distinguished from its congeners by the possession of very large papilla at the mandibular symphysis of large adult females (
<figureCitation id="1DB52A471D54FFA5D8845315C127FD13" box="[1257,1337,635,658]" captionStart="Fig" captionStartId="21.[129,168,1904,1927]" captionTargetBox="[129,782,1506,1872]" captionTargetId="figure-476@21.[129,772,1529,1888]" captionText="Fig. 26. Upper and lower lip of Phalloceros titthos, MZUSP 43467, showing symphyseal papilla (SP). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10569764" httpUri="https://zenodo.org/record/10569764/files/figure.png" pageId="20" pageNumber="133">Fig. 26</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="CD9465491D54FFA4DF5D53D5C5C5FB2A" lastPageId="21" lastPageNumber="134" pageId="20" pageNumber="133" type="description">
<paragraph id="853136C21D54FFA5DF5D53D5C17DFBF3" blockId="20.[816,1459,699,1138]" pageId="20" pageNumber="133">
<emphasis id="B7FAEAD01D54FFA5DF5D53D5C7ABFD53" bold="true" box="[816,949,699,722]" pageId="20" pageNumber="133">Description</emphasis>
. Morphometric data in
<tableCitation id="C80C03791D54FFA5D8DD53D5C10BFD53" box="[1200,1301,699,722]" captionStart="Table 11" captionStartId="19.[129,192,1465,1488]" captionTargetBox="[136,1436,1550,1964]" captionTargetPageId="19" captionText="Table 11. Descriptive morphometrics of female specimens of Phalloceros ocellatus, P. pellos, P. spiloura, and P. titthos. Measurements 1-10 are percents of standard length and measurements 11-13 are percents of head length. H = holotype." httpUri="http://table.plazi.org/id/D1F1664A1D53FFA2DCEC54D7C19FFA71" pageId="20" pageNumber="133" tableUuid="D1F1664A1D53FFA2DCEC54D7C19FFA71">Tables 11</tableCitation>
and
<tableCitation id="C80C03791D54FFA5D92A53D5C17BFD53" box="[1351,1381,699,722]" captionStart="Table 12" captionStartId="20.[136,199,1464,1487]" captionTargetBox="[139,1445,1549,1962]" captionTargetPageId="20" captionText="Table 12. Descriptive morphometrics of male specimens of Phalloceros ocellatus, P. pellos, P. spiloura, and P. titthos. Measurements 1-10 are percents of standard length and measurements 11-13 are percents of head length. H = holotype." httpUri="http://table.plazi.org/id/D1F1664A1D54FFA5DCE554D6C197FA6E" pageId="20" pageNumber="133" tableUuid="D1F1664A1D54FFA5DCE554D6C197FA6E">12</tableCitation>
.
<collectingRegion id="474AF8201D54FFA5D90353D5C772FD73" pageId="20" pageNumber="133">Range of SL</collectingRegion>
: 21.0 to
<quantity id="42769B271D54FFA5DFD153B5C008FD73" box="[956,1046,731,754]" metricMagnitude="-2" metricUnit="m" metricValue="4.52" pageId="20" pageNumber="133" unit="mm" value="45.2">45.2 mm</quantity>
(females),
<quantity id="42769B271D54FFA5D8E853B5C136FD73" box="[1157,1320,731,754]" metricMagnitude="-2" metricUnit="m" metricValue="2.045" metricValueMax="2.58" metricValueMin="1.51" pageId="20" pageNumber="133" unit="mm" value="20.45" valueMax="25.8" valueMin="15.1">15.1 to 25.8 mm</quantity>
(males).
<collectingRegion id="474AF8201D54FFA5D9EF53B5C7B8FC93" pageId="20" pageNumber="133">Dorsal-fin rays</collectingRegion>
: 8* [21], 9 [2].
<collectingRegion id="474AF8201D54FFA5D8525395C14BFC93" box="[1087,1365,763,786]" pageId="20" pageNumber="133">Branched pectoral-fin rays</collectingRegion>
: 6 [4], 7* [14].
<collectingRegion id="474AF8201D54FFA5DF085275C7E4FCB3" box="[869,1018,795,818]" pageId="20" pageNumber="133">Pelvic-fin rays</collectingRegion>
: 5 [3] (males), 4 [1], 5* [19] (females).
<collectingRegion id="474AF8201D54FFA5D9145275C7E3FCD3" pageId="20" pageNumber="133">Anal-fin rays of females</collectingRegion>
: 11* [20].
<collectingRegion id="474AF8201D54FFA5D8005255C17CFCD3" box="[1133,1378,827,850]" pageId="20" pageNumber="133">Anal-fin rays of males</collectingRegion>
: 9* [3].
<collectingRegion id="474AF8201D54FFA5DF5D5235C05CFCF3" box="[816,1090,859,882]" pageId="20" pageNumber="133">Branched caudal-fin rays</collectingRegion>
: 10 [1], 12* [14], 13 [1].
<collectingRegion id="474AF8201D54FFA5D9205235C76BFC13" pageId="20" pageNumber="133">Predorsal scales</collectingRegion>
: 13* [8], 14 [11].
<collectingRegion id="474AF8201D54FFA5D85D5215C17AFC13" box="[1072,1380,891,914]" pageId="20" pageNumber="133">Longitudinal series of scales</collectingRegion>
: 27 [2], 28* [5], 29 [16].
<collectingRegion id="474AF8201D54FFA5DFB252F5C19DFC33" box="[991,1411,923,946]" pageId="20" pageNumber="133">Series of scales around caudal peduncle</collectingRegion>
: 16* [23].
<collectingRegion id="474AF8201D54FFA5DF0A52D5C060FC53" box="[871,1150,955,978]" pageId="20" pageNumber="133">Transverse series of scales</collectingRegion>
: 7* [23]. Serrae on R4p: 7 [1], 8 [1], 9 [3], 10 [1], 12 [1], 13 [2].
<collectingRegion id="474AF8201D54FFA5D8EC52B5C100FC73" box="[1153,1310,987,1010]" pageId="20" pageNumber="133">Epipleural ribs</collectingRegion>
: 12 [2], 13 [1].
<collectingRegion id="474AF8201D54FFA5DF5D5295C7B9FB93" box="[816,935,1019,1042]" pageId="20" pageNumber="133">Pleural ribs</collectingRegion>
: 13 [1], 14 [2].
<collectingRegion id="474AF8201D54FFA5D8565295C0BFFB93" box="[1083,1185,1019,1042]" pageId="20" pageNumber="133">Vertebrae</collectingRegion>
: 30 [1], 32 [2]. Symphyseal papillae present (
<figureCitation id="1DB52A471D54FFA5DF8B5575C02BFBB3" box="[998,1077,1051,1074]" captionStart="Fig" captionStartId="21.[129,168,1904,1927]" captionTargetBox="[129,782,1506,1872]" captionTargetId="figure-476@21.[129,772,1529,1888]" captionText="Fig. 26. Upper and lower lip of Phalloceros titthos, MZUSP 43467, showing symphyseal papilla (SP). Scale bar 1 mm." figureDoi="http://doi.org/10.5281/zenodo.10569764" httpUri="https://zenodo.org/record/10569764/files/figure.png" pageId="20" pageNumber="133">Fig. 26</figureCitation>
). Female urogenital papilla straight, located along midventral line and between anus and base of first anal-fin ray. Gonopodial appendix hook present.
</paragraph>
<paragraph id="853136C21D54FFA4DF5D55F5C5C5FB2A" blockId="20.[816,1459,1179,1426]" lastBlockId="21.[129,772,820,1195]" lastPageId="21" lastPageNumber="134" pageId="20" pageNumber="133">
<emphasis id="B7FAEAD01D54FFA5DF5D55F5C7FCFB33" bold="true" box="[816,994,1179,1202]" pageId="20" pageNumber="133">Color in alcohol</emphasis>
. Eye black with greenish brown pupil. Background coloration dark brown, darker in upper half. Overall appearance dark. Border of scales and subjacent skin replete with brown chromatophores, more concentrated at short distance from scale border, conferring reticulate pattern to whole flanks. Brown chromatophores scattered through whole body, more concentrated on dorsal portion, mainly on head, snout, opercle, and ventral surface of mandible. Margin of scales on prepelvic region clearly defined bordered by dark chromatophores. Lateral dark brown spot located over 17
<superScript id="72FB9B8A1D55FFA4DEFA523DC6BDFCE0" attach="left" box="[663,675,851,865]" fontSize="6" pageId="21" pageNumber="134">th</superScript>
and 18
<superScript id="72FB9B8A1D55FFA4DE9A523DC71DFCE0" attach="none" box="[759,771,851,865]" fontSize="6" pageId="21" pageNumber="134">th</superScript>
scale of longitudinal series (absent in
<specimenCount id="9388FD4B1D55FFA4DE62521AC6BFFC0A" box="[527,673,884,907]" count="1" pageId="21" pageNumber="134" type="generic">one specimen</specimenCount>
studied). Lateral spot covering approximately one or two scales on horizontal direction, and two or three scales on vertical. Inconspicuous vertical bars present in some individuals. Dorsal-fin membrane dusky, bearing dark band of chromatophores on distal border and dark band of chromatophores near dorsal-fin base. Pectoral-, pelvic- and caudal-fin rays dusky replete of brown chromatophores distributed along fin rays. First three anal-fin rays of females sometimes spotted with black. Dark brown line along R3.
</paragraph>
</subSubSection>
<caption id="D1F1664A1D54FFA5DCE554D6C197FA6E" ID-Table-UUID="D1F1664A1D54FFA5DCE554D6C197FA6E" httpUri="http://table.plazi.org/id/D1F1664A1D54FFA5DCE554D6C197FA6E" pageId="20" pageNumber="133" startId="20.[136,199,1464,1487]" targetBox="[139,1445,1549,1962]" targetIsTable="true" targetPageId="20" targetType="table">
<paragraph id="853136C21D54FFA5DCE554D6C197FA6E" blockId="20.[136,1459,1464,1519]" pageId="20" pageNumber="133">
<emphasis id="B7FAEAD01D54FFA5DCE554D6C4F5FA4E" bold="true" box="[136,235,1464,1487]" pageId="20" pageNumber="133">Table 12</emphasis>
. Descriptive morphometrics of male specimens of
<taxonomicName id="428E4D411D54FFA5DF4F54D6C010FA4E" authorityName="Lucinda" authorityYear="2008" box="[802,1038,1464,1487]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B7FAEAD01D54FFA5DF4F54D6C010FA4E" box="[802,1038,1464,1487]" italics="true" pageId="20" pageNumber="133">Phalloceros ocellatus</emphasis>
</taxonomicName>
,
<taxonomicName id="428E4D411D54FFA5D87354D6C060FA4E" authorityName="Lucinda" authorityYear="2008" box="[1054,1150,1464,1487]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="pellos">
<emphasis id="B7FAEAD01D54FFA5D87354D6C030FA4E" box="[1054,1070,1464,1487]" italics="true" pageId="20" pageNumber="133">P</emphasis>
.
<emphasis id="B7FAEAD01D54FFA5D85354D6C060FA4E" box="[1086,1150,1464,1487]" italics="true" pageId="20" pageNumber="133">pellos</emphasis>
</taxonomicName>
,
<taxonomicName id="428E4D411D54FFA5D8E354D6C116FA4E" authorityName="Lucinda" authorityYear="2008" box="[1166,1288,1464,1487]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="spiloura">
<emphasis id="B7FAEAD01D54FFA5D8E354D6C080FA4E" box="[1166,1182,1464,1487]" italics="true" pageId="20" pageNumber="133">P</emphasis>
.
<emphasis id="B7FAEAD01D54FFA5D8C354D6C116FA4E" box="[1198,1288,1464,1487]" italics="true" pageId="20" pageNumber="133">spiloura</emphasis>
</taxonomicName>
, and
<taxonomicName id="428E4D411D54FFA5D92554D6C1B2FA4E" authorityName="Lucinda" authorityYear="2008" box="[1352,1452,1464,1487]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="titthos">
<emphasis id="B7FAEAD01D54FFA5D92554D6C146FA4E" box="[1352,1368,1464,1487]" italics="true" pageId="20" pageNumber="133">P</emphasis>
.
<emphasis id="B7FAEAD01D54FFA5D90554D6C1B2FA4E" box="[1384,1452,1464,1487]" italics="true" pageId="20" pageNumber="133">titthos</emphasis>
</taxonomicName>
. Measurements 1-10 are percents of standard length and measurements 11-13 are percents of head length. H = holotype.
</paragraph>
</caption>
<paragraph id="853136C21D54FFA5DDA55763C179F82B" pageId="20" pageNumber="133">
<table id="F78EC4621D54004EDCE65763C1BBF82B" box="[139,1445,1549,1962]" gridcols="11" gridrows="17" pageId="20" pageNumber="133">
<tr id="3BBE34801D54004EDCE65763C1BBF9A1" box="[139,1445,1549,1568]" gridrow="0" pageId="20" pageNumber="133" rowspan-0="1">
<th id="786F5DFC1D54004EDDC45763C6BDF9A1" box="[425,675,1549,1568]" colspan="3" colspanRight="2" gridcol="1" gridrow="0" pageId="20" pageNumber="133">
<taxonomicName id="428E4D411D54FFA5DDA55763C69AF9A1" authorityName="Lucinda" authorityYear="2008" box="[456,644,1549,1568]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="ocellatus">
<emphasis id="B7FAEAD01D54FFA5DDA55763C69AF9A1" box="[456,644,1549,1568]" italics="true" pageId="20" pageNumber="133">Phalloceros ocellatus</emphasis>
</taxonomicName>
</th>
<th id="786F5DFC1D54004EDEA45763C7DDF9A1" box="[713,963,1549,1568]" colspan="3" colspanRight="2" gridcol="4" gridrow="0" pageId="20" pageNumber="133">
<taxonomicName id="428E4D411D54FFA5DE985763C789F9A1" authorityName="Lucinda" authorityYear="2008" box="[757,919,1549,1568]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="pellos">
<emphasis id="B7FAEAD01D54FFA5DE985763C789F9A1" box="[757,919,1549,1568]" italics="true" pageId="20" pageNumber="133">Phalloceros pellos</emphasis>
</taxonomicName>
</th>
<th id="786F5DFC1D54004EDF855763C0FDF9A1" box="[1000,1251,1549,1568]" colspan="3" colspanRight="2" gridcol="7" gridrow="0" pageId="20" pageNumber="133">
<taxonomicName id="428E4D411D54FFA5D8675763C0DEF9A1" authorityName="Lucinda" authorityYear="2008" box="[1034,1216,1549,1568]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="spiloura">
<emphasis id="B7FAEAD01D54FFA5D8675763C0DEF9A1" box="[1034,1216,1549,1568]" italics="true" pageId="20" pageNumber="133">Phalloceros spiloura</emphasis>
</taxonomicName>
</th>
<th id="786F5DFC1D54004ED96C5763C1BBF9A1" box="[1281,1445,1549,1568]" gridcol="10" gridrow="0" pageId="20" pageNumber="133">
<taxonomicName id="428E4D411D54FFA5D96C5763C1BBF9A1" authority="Character" authorityName="Character" box="[1281,1445,1549,1568]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="20" pageNumber="133" phylum="Chordata" rank="species" species="titthos">
<emphasis id="B7FAEAD01D54FFA5D96C5763C1BBF9A1" box="[1281,1445,1549,1568]" italics="true" pageId="20" pageNumber="133">Phalloceros titthos</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="3BBE34801D54004EDCE65748C1BBF9B8" box="[139,1445,1574,1593]" gridrow="1" pageId="20" pageNumber="133" rowspan-1="1" rowspan-3="1" rowspan-4="1" rowspan-6="1" rowspan-7="1" rowspan-9="1">
<th id="786F5DFC1D54004EDCE65748C594F9B8" box="[139,394,1574,1593]" gridcol="0" gridrow="1" pageId="20" pageNumber="133">Character</th>
<td id="786F5DFC1D54004EDD9E5748C64EF9B8" box="[499,592,1574,1593]" gridcol="2" gridrow="1" pageId="20" pageNumber="133">n = 11</td>
<td id="786F5DFC1D54004EDF7E5748C76EF9B8" box="[787,880,1574,1593]" gridcol="5" gridrow="1" pageId="20" pageNumber="133">n = 6</td>
<td id="786F5DFC1D54004ED85F5748C08EF9B8" box="[1074,1168,1574,1593]" gridcol="8" gridrow="1" pageId="20" pageNumber="133">n = 6</td>
<td id="786F5DFC1D54004ED96C5748C1BBF9B8" box="[1281,1445,1574,1593]" gridcol="10" gridrow="1" pageId="20" pageNumber="133">n = 1</td>
</tr>
<tr id="3BBE34801D54004EDCE65750C1BBF9D0" box="[139,1445,1598,1617]" gridrow="2" pageId="20" pageNumber="133" rowspan-0="1" rowspan-10="1">
<td id="786F5DFC1D54004EDDC45750C5CDF9D0" box="[425,467,1598,1617]" gridcol="1" gridrow="2" pageId="20" pageNumber="133">H</td>
<td id="786F5DFC1D54004EDD9E5750C64EF9D0" box="[499,592,1598,1617]" gridcol="2" gridrow="2" pageId="20" pageNumber="133">Range</td>
<td id="786F5DFC1D54004EDE035750C6BDF9D0" box="[622,675,1598,1617]" gridcol="3" gridrow="2" pageId="20" pageNumber="133">Mean</td>
<td id="786F5DFC1D54004EDEA45750C6EDF9D0" box="[713,755,1598,1617]" gridcol="4" gridrow="2" pageId="20" pageNumber="133">H</td>
<td id="786F5DFC1D54004EDF7E5750C76EF9D0" box="[787,880,1598,1617]" gridcol="5" gridrow="2" pageId="20" pageNumber="133">Range</td>
<td id="786F5DFC1D54004EDFE35750C7DDF9D0" box="[910,963,1598,1617]" gridcol="6" gridrow="2" pageId="20" pageNumber="133">Mean</td>
<td id="786F5DFC1D54004EDF855750C00DF9D0" box="[1000,1043,1598,1617]" gridcol="7" gridrow="2" pageId="20" pageNumber="133">H</td>
<td id="786F5DFC1D54004ED85F5750C08EF9D0" box="[1074,1168,1598,1617]" gridcol="8" gridrow="2" pageId="20" pageNumber="133">Range</td>
<td id="786F5DFC1D54004ED8C35750C0FDF9D0" box="[1198,1251,1598,1617]" gridcol="9" gridrow="2" pageId="20" pageNumber="133">Mean</td>
</tr>
<tr id="3BBE34801D54004EDCE65736C1BBF9EA" box="[139,1445,1624,1643]" gridrow="3" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE65736C594F9EA" box="[139,394,1624,1643]" gridcol="0" gridrow="3" pageId="20" pageNumber="133">Standard length (mm)</th>
<td id="786F5DFC1D54004EDDC45736C5CDF9EA" box="[425,467,1624,1643]" gridcol="1" gridrow="3" pageId="20" pageNumber="133">16.3</td>
<td id="786F5DFC1D54004EDD9E5736C64EF9EA" box="[499,592,1624,1643]" gridcol="2" gridrow="3" pageId="20" pageNumber="133">15.4-18.5</td>
<td id="786F5DFC1D54004EDE035736C6BDF9EA" box="[622,675,1624,1643]" gridcol="3" gridrow="3" pageId="20" pageNumber="133">16.7</td>
<td id="786F5DFC1D54004EDEA45736C6EDF9EA" box="[713,755,1624,1643]" gridcol="4" gridrow="3" pageId="20" pageNumber="133">26.4</td>
<td id="786F5DFC1D54004EDF7E5736C76EF9EA" box="[787,880,1624,1643]" gridcol="5" gridrow="3" pageId="20" pageNumber="133">21.5-26.4</td>
<td id="786F5DFC1D54004EDFE35736C7DDF9EA" box="[910,963,1624,1643]" gridcol="6" gridrow="3" pageId="20" pageNumber="133">24.3</td>
<td id="786F5DFC1D54004EDF855736C00DF9EA" box="[1000,1043,1624,1643]" gridcol="7" gridrow="3" pageId="20" pageNumber="133">22.0</td>
<td id="786F5DFC1D54004ED85F5736C08EF9EA" box="[1074,1168,1624,1643]" gridcol="8" gridrow="3" pageId="20" pageNumber="133">18.0-22.4</td>
<td id="786F5DFC1D54004ED8C35736C0FDF9EA" box="[1198,1251,1624,1643]" gridcol="9" gridrow="3" pageId="20" pageNumber="133">20.0</td>
<td id="786F5DFC1D54004ED96C5736C1BBF9EA" box="[1281,1445,1624,1643]" gridcol="10" gridrow="3" pageId="20" pageNumber="133">17.1</td>
</tr>
<tr id="3BBE34801D54004EDCE6571FC1BBF905" box="[139,1445,1649,1668]" gridrow="4" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE6571FC594F905" box="[139,394,1649,1668]" gridcol="0" gridrow="4" pageId="20" pageNumber="133">1-Head length</th>
<td id="786F5DFC1D54004EDDC4571FC5CDF905" box="[425,467,1649,1668]" gridcol="1" gridrow="4" pageId="20" pageNumber="133">23.5</td>
<td id="786F5DFC1D54004EDD9E571FC64EF905" box="[499,592,1649,1668]" gridcol="2" gridrow="4" pageId="20" pageNumber="133">22.2-24.8</td>
<td id="786F5DFC1D54004EDE03571FC6BDF905" box="[622,675,1649,1668]" gridcol="3" gridrow="4" pageId="20" pageNumber="133">23.6</td>
<td id="786F5DFC1D54004EDEA4571FC6EDF905" box="[713,755,1649,1668]" gridcol="4" gridrow="4" pageId="20" pageNumber="133">20.2</td>
<td id="786F5DFC1D54004EDF7E571FC76EF905" box="[787,880,1649,1668]" gridcol="5" gridrow="4" pageId="20" pageNumber="133">20.2-24.7</td>
<td id="786F5DFC1D54004EDFE3571FC7DDF905" box="[910,963,1649,1668]" gridcol="6" gridrow="4" pageId="20" pageNumber="133">21.4</td>
<td id="786F5DFC1D54004EDF85571FC00DF905" box="[1000,1043,1649,1668]" gridcol="7" gridrow="4" pageId="20" pageNumber="133">21.8</td>
<td id="786F5DFC1D54004ED85F571FC08EF905" box="[1074,1168,1649,1668]" gridcol="8" gridrow="4" pageId="20" pageNumber="133">21.5-24.2</td>
<td id="786F5DFC1D54004ED8C3571FC0FDF905" box="[1198,1251,1649,1668]" gridcol="9" gridrow="4" pageId="20" pageNumber="133">22.4</td>
<td id="786F5DFC1D54004ED96C571FC1BBF905" box="[1281,1445,1649,1668]" gridcol="10" gridrow="4" pageId="20" pageNumber="133">20.4</td>
</tr>
<tr id="3BBE34801D54004EDCE657E7C1BBF91D" box="[139,1445,1673,1692]" gridrow="5" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE657E7C594F91D" box="[139,394,1673,1692]" gridcol="0" gridrow="5" pageId="20" pageNumber="133">2-Snout-occipital distance</th>
<td id="786F5DFC1D54004EDDC457E7C5CDF91D" box="[425,467,1673,1692]" gridcol="1" gridrow="5" pageId="20" pageNumber="133">20.5</td>
<td id="786F5DFC1D54004EDD9E57E7C64EF91D" box="[499,592,1673,1692]" gridcol="2" gridrow="5" pageId="20" pageNumber="133">18.2-21.8</td>
<td id="786F5DFC1D54004EDE0357E7C6BDF91D" box="[622,675,1673,1692]" gridcol="3" gridrow="5" pageId="20" pageNumber="133">20.2</td>
<td id="786F5DFC1D54004EDEA457E7C6EDF91D" box="[713,755,1673,1692]" gridcol="4" gridrow="5" pageId="20" pageNumber="133">20.8</td>
<td id="786F5DFC1D54004EDF7E57E7C76EF91D" box="[787,880,1673,1692]" gridcol="5" gridrow="5" pageId="20" pageNumber="133">15.2-20.8</td>
<td id="786F5DFC1D54004EDFE357E7C7DDF91D" box="[910,963,1673,1692]" gridcol="6" gridrow="5" pageId="20" pageNumber="133">19.5</td>
<td id="786F5DFC1D54004EDF8557E7C00DF91D" box="[1000,1043,1673,1692]" gridcol="7" gridrow="5" pageId="20" pageNumber="133">21.0</td>
<td id="786F5DFC1D54004ED85F57E7C08EF91D" box="[1074,1168,1673,1692]" gridcol="8" gridrow="5" pageId="20" pageNumber="133">16.6-21.4</td>
<td id="786F5DFC1D54004ED8C357E7C0FDF91D" box="[1198,1251,1673,1692]" gridcol="9" gridrow="5" pageId="20" pageNumber="133">19.7</td>
<td id="786F5DFC1D54004ED96C57E7C1BBF91D" box="[1281,1445,1673,1692]" gridcol="10" gridrow="5" pageId="20" pageNumber="133">23.7</td>
</tr>
<tr id="3BBE34801D54004EDCE657CCC1BBF934" box="[139,1445,1698,1717]" gridrow="6" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE657CCC594F934" box="[139,394,1698,1717]" gridcol="0" gridrow="6" pageId="20" pageNumber="133">3-Predorsal distance</th>
<td id="786F5DFC1D54004EDDC457CCC5CDF934" box="[425,467,1698,1717]" gridcol="1" gridrow="6" pageId="20" pageNumber="133">58.9</td>
<td id="786F5DFC1D54004EDD9E57CCC64EF934" box="[499,592,1698,1717]" gridcol="2" gridrow="6" pageId="20" pageNumber="133">57.5-60.7</td>
<td id="786F5DFC1D54004EDE0357CCC6BDF934" box="[622,675,1698,1717]" gridcol="3" gridrow="6" pageId="20" pageNumber="133">59.5</td>
<td id="786F5DFC1D54004EDEA457CCC6EDF934" box="[713,755,1698,1717]" gridcol="4" gridrow="6" pageId="20" pageNumber="133">54.8</td>
<td id="786F5DFC1D54004EDF7E57CCC76EF934" box="[787,880,1698,1717]" gridcol="5" gridrow="6" pageId="20" pageNumber="133">54.8-58.8</td>
<td id="786F5DFC1D54004EDFE357CCC7DDF934" box="[910,963,1698,1717]" gridcol="6" gridrow="6" pageId="20" pageNumber="133">56.7</td>
<td id="786F5DFC1D54004EDF8557CCC00DF934" box="[1000,1043,1698,1717]" gridcol="7" gridrow="6" pageId="20" pageNumber="133">57.7</td>
<td id="786F5DFC1D54004ED85F57CCC08EF934" box="[1074,1168,1698,1717]" gridcol="8" gridrow="6" pageId="20" pageNumber="133">59.8-63.2</td>
<td id="786F5DFC1D54004ED8C357CCC0FDF934" box="[1198,1251,1698,1717]" gridcol="9" gridrow="6" pageId="20" pageNumber="133">61.2</td>
<td id="786F5DFC1D54004ED96C57CCC1BBF934" box="[1281,1445,1698,1717]" gridcol="10" gridrow="6" pageId="20" pageNumber="133">58.9</td>
</tr>
<tr id="3BBE34801D54004EDCE657D4C1BBF94C" box="[139,1445,1722,1741]" gridrow="7" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE657D4C594F94C" box="[139,394,1722,1741]" gridcol="0" gridrow="7" pageId="20" pageNumber="133">4-Dorsal-fin base length</th>
<td id="786F5DFC1D54004EDDC457D4C5CDF94C" box="[425,467,1722,1741]" gridcol="1" gridrow="7" pageId="20" pageNumber="133">7.7</td>
<td id="786F5DFC1D54004EDD9E57D4C64EF94C" box="[499,592,1722,1741]" gridcol="2" gridrow="7" pageId="20" pageNumber="133">7.7-13.1</td>
<td id="786F5DFC1D54004EDE0357D4C6BDF94C" box="[622,675,1722,1741]" gridcol="3" gridrow="7" pageId="20" pageNumber="133">10.9</td>
<td id="786F5DFC1D54004EDEA457D4C6EDF94C" box="[713,755,1722,1741]" gridcol="4" gridrow="7" pageId="20" pageNumber="133">11.5</td>
<td id="786F5DFC1D54004EDF7E57D4C76EF94C" box="[787,880,1722,1741]" gridcol="5" gridrow="7" pageId="20" pageNumber="133">8.4-11.5</td>
<td id="786F5DFC1D54004EDFE357D4C7DDF94C" box="[910,963,1722,1741]" gridcol="6" gridrow="7" pageId="20" pageNumber="133">10.7</td>
<td id="786F5DFC1D54004EDF8557D4C00DF94C" box="[1000,1043,1722,1741]" gridcol="7" gridrow="7" pageId="20" pageNumber="133">13.1</td>
<td id="786F5DFC1D54004ED85F57D4C08EF94C" box="[1074,1168,1722,1741]" gridcol="8" gridrow="7" pageId="20" pageNumber="133">10.4-12.4</td>
<td id="786F5DFC1D54004ED8C357D4C0FDF94C" box="[1198,1251,1722,1741]" gridcol="9" gridrow="7" pageId="20" pageNumber="133">11.1</td>
<td id="786F5DFC1D54004ED96C57D4C1BBF94C" box="[1281,1445,1722,1741]" gridcol="10" gridrow="7" pageId="20" pageNumber="133">11.5</td>
</tr>
<tr id="3BBE34801D54004EDCE657BDC1BBF967" box="[139,1445,1747,1766]" gridrow="8" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE657BDC594F967" box="[139,394,1747,1766]" gridcol="0" gridrow="8" pageId="20" pageNumber="133">5-Anal-fin base length</th>
<td id="786F5DFC1D54004EDDC457BDC5CDF967" box="[425,467,1747,1766]" gridcol="1" gridrow="8" pageId="20" pageNumber="133">8.1</td>
<td id="786F5DFC1D54004EDD9E57BDC64EF967" box="[499,592,1747,1766]" gridcol="2" gridrow="8" pageId="20" pageNumber="133">5.9-10.7</td>
<td id="786F5DFC1D54004EDE0357BDC6BDF967" box="[622,675,1747,1766]" gridcol="3" gridrow="8" pageId="20" pageNumber="133">8.6</td>
<td id="786F5DFC1D54004EDEA457BDC6EDF967" box="[713,755,1747,1766]" gridcol="4" gridrow="8" pageId="20" pageNumber="133">6.3</td>
<td id="786F5DFC1D54004EDF7E57BDC76EF967" box="[787,880,1747,1766]" gridcol="5" gridrow="8" pageId="20" pageNumber="133">6.3-8.9</td>
<td id="786F5DFC1D54004EDFE357BDC7DDF967" box="[910,963,1747,1766]" gridcol="6" gridrow="8" pageId="20" pageNumber="133">7.2</td>
<td id="786F5DFC1D54004EDF8557BDC00DF967" box="[1000,1043,1747,1766]" gridcol="7" gridrow="8" pageId="20" pageNumber="133">9.9</td>
<td id="786F5DFC1D54004ED85F57BDC08EF967" box="[1074,1168,1747,1766]" gridcol="8" gridrow="8" pageId="20" pageNumber="133">6.1-10.3</td>
<td id="786F5DFC1D54004ED8C357BDC0FDF967" box="[1198,1251,1747,1766]" gridcol="9" gridrow="8" pageId="20" pageNumber="133">7.8</td>
<td id="786F5DFC1D54004ED96C57BDC1BBF967" box="[1281,1445,1747,1766]" gridcol="10" gridrow="8" pageId="20" pageNumber="133">8.7</td>
</tr>
<tr id="3BBE34801D54004EDCE65785C1BBF97F" box="[139,1445,1771,1790]" gridrow="9" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE65785C594F97F" box="[139,394,1771,1790]" gridcol="0" gridrow="9" pageId="20" pageNumber="133">6-Body depth</th>
<td id="786F5DFC1D54004EDDC45785C5CDF97F" box="[425,467,1771,1790]" gridcol="1" gridrow="9" pageId="20" pageNumber="133">30.5</td>
<td id="786F5DFC1D54004EDD9E5785C64EF97F" box="[499,592,1771,1790]" gridcol="2" gridrow="9" pageId="20" pageNumber="133">30.5-34.0</td>
<td id="786F5DFC1D54004EDE035785C6BDF97F" box="[622,675,1771,1790]" gridcol="3" gridrow="9" pageId="20" pageNumber="133">31.9</td>
<td id="786F5DFC1D54004EDEA45785C6EDF97F" box="[713,755,1771,1790]" gridcol="4" gridrow="9" pageId="20" pageNumber="133">31.1</td>
<td id="786F5DFC1D54004EDF7E5785C76EF97F" box="[787,880,1771,1790]" gridcol="5" gridrow="9" pageId="20" pageNumber="133">31.1-34.5</td>
<td id="786F5DFC1D54004EDFE35785C7DDF97F" box="[910,963,1771,1790]" gridcol="6" gridrow="9" pageId="20" pageNumber="133">32.0</td>
<td id="786F5DFC1D54004EDF855785C00DF97F" box="[1000,1043,1771,1790]" gridcol="7" gridrow="9" pageId="20" pageNumber="133">35.0</td>
<td id="786F5DFC1D54004ED85F5785C08EF97F" box="[1074,1168,1771,1790]" gridcol="8" gridrow="9" pageId="20" pageNumber="133">29.9-35.9</td>
<td id="786F5DFC1D54004ED8C35785C0FDF97F" box="[1198,1251,1771,1790]" gridcol="9" gridrow="9" pageId="20" pageNumber="133">33.1</td>
<td id="786F5DFC1D54004ED96C5785C1BBF97F" box="[1281,1445,1771,1790]" gridcol="10" gridrow="9" pageId="20" pageNumber="133">33.9</td>
</tr>
<tr id="3BBE34801D54004EDCE6566AC1BBF896" box="[139,1445,1796,1815]" gridrow="10" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE6566AC594F896" box="[139,394,1796,1815]" gridcol="0" gridrow="10" pageId="20" pageNumber="133">7-Pre-pelvic length</th>
<td id="786F5DFC1D54004EDDC4566AC5CDF896" box="[425,467,1796,1815]" gridcol="1" gridrow="10" pageId="20" pageNumber="133">31.3</td>
<td id="786F5DFC1D54004EDD9E566AC64EF896" box="[499,592,1796,1815]" gridcol="2" gridrow="10" pageId="20" pageNumber="133">29.4-36.8</td>
<td id="786F5DFC1D54004EDE03566AC6BDF896" box="[622,675,1796,1815]" gridcol="3" gridrow="10" pageId="20" pageNumber="133">31.4</td>
<td id="786F5DFC1D54004EDEA4566AC6EDF896" box="[713,755,1796,1815]" gridcol="4" gridrow="10" pageId="20" pageNumber="133">27.9</td>
<td id="786F5DFC1D54004EDF7E566AC76EF896" box="[787,880,1796,1815]" gridcol="5" gridrow="10" pageId="20" pageNumber="133">27.9-32.1</td>
<td id="786F5DFC1D54004EDFE3566AC7DDF896" box="[910,963,1796,1815]" gridcol="6" gridrow="10" pageId="20" pageNumber="133">29.3</td>
<td id="786F5DFC1D54004EDF85566AC00DF896" box="[1000,1043,1796,1815]" gridcol="7" gridrow="10" pageId="20" pageNumber="133">31.4</td>
<td id="786F5DFC1D54004ED85F566AC08EF896" box="[1074,1168,1796,1815]" gridcol="8" gridrow="10" pageId="20" pageNumber="133">30.7-34.7</td>
<td id="786F5DFC1D54004ED8C3566AC0FDF896" box="[1198,1251,1796,1815]" gridcol="9" gridrow="10" pageId="20" pageNumber="133">32.1</td>
<td id="786F5DFC1D54004ED96C566AC1BBF896" box="[1281,1445,1796,1815]" gridcol="10" gridrow="10" pageId="20" pageNumber="133">28.4</td>
</tr>
<tr id="3BBE34801D54004EDCE65673C1BBF8B1" box="[139,1445,1821,1840]" gridrow="11" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE65673C594F8B1" box="[139,394,1821,1840]" gridcol="0" gridrow="11" pageId="20" pageNumber="133">8-Preanal length</th>
<td id="786F5DFC1D54004EDDC45673C5CDF8B1" box="[425,467,1821,1840]" gridcol="1" gridrow="11" pageId="20" pageNumber="133">42.4</td>
<td id="786F5DFC1D54004EDD9E5673C64EF8B1" box="[499,592,1821,1840]" gridcol="2" gridrow="11" pageId="20" pageNumber="133">39.5-45.7</td>
<td id="786F5DFC1D54004EDE035673C6BDF8B1" box="[622,675,1821,1840]" gridcol="3" gridrow="11" pageId="20" pageNumber="133">42.2</td>
<td id="786F5DFC1D54004EDEA45673C6EDF8B1" box="[713,755,1821,1840]" gridcol="4" gridrow="11" pageId="20" pageNumber="133">36.0</td>
<td id="786F5DFC1D54004EDF7E5673C76EF8B1" box="[787,880,1821,1840]" gridcol="5" gridrow="11" pageId="20" pageNumber="133">36.0-43.6</td>
<td id="786F5DFC1D54004EDFE35673C7DDF8B1" box="[910,963,1821,1840]" gridcol="6" gridrow="11" pageId="20" pageNumber="133">38.8</td>
<td id="786F5DFC1D54004EDF855673C00DF8B1" box="[1000,1043,1821,1840]" gridcol="7" gridrow="11" pageId="20" pageNumber="133">41.5</td>
<td id="786F5DFC1D54004ED85F5673C08EF8B1" box="[1074,1168,1821,1840]" gridcol="8" gridrow="11" pageId="20" pageNumber="133">39.6-44.3</td>
<td id="786F5DFC1D54004ED8C35673C0FDF8B1" box="[1198,1251,1821,1840]" gridcol="9" gridrow="11" pageId="20" pageNumber="133">41.4</td>
<td id="786F5DFC1D54004ED96C5673C1BBF8B1" box="[1281,1445,1821,1840]" gridcol="10" gridrow="11" pageId="20" pageNumber="133">38.3</td>
</tr>
<tr id="3BBE34801D54004EDCE6565BC1BBF8C9" box="[139,1445,1845,1864]" gridrow="12" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE6565BC594F8C9" box="[139,394,1845,1864]" gridcol="0" gridrow="12" pageId="20" pageNumber="133">9-Postanal length</th>
<td id="786F5DFC1D54004EDDC4565BC5CDF8C9" box="[425,467,1845,1864]" gridcol="1" gridrow="12" pageId="20" pageNumber="133">54.3</td>
<td id="786F5DFC1D54004EDD9E565BC64EF8C9" box="[499,592,1845,1864]" gridcol="2" gridrow="12" pageId="20" pageNumber="133">50.3-58.6</td>
<td id="786F5DFC1D54004EDE03565BC6BDF8C9" box="[622,675,1845,1864]" gridcol="3" gridrow="12" pageId="20" pageNumber="133">53.9</td>
<td id="786F5DFC1D54004EDEA4565BC6EDF8C9" box="[713,755,1845,1864]" gridcol="4" gridrow="12" pageId="20" pageNumber="133">61.7</td>
<td id="786F5DFC1D54004EDF7E565BC76EF8C9" box="[787,880,1845,1864]" gridcol="5" gridrow="12" pageId="20" pageNumber="133">53.9-61.7</td>
<td id="786F5DFC1D54004EDFE3565BC7DDF8C9" box="[910,963,1845,1864]" gridcol="6" gridrow="12" pageId="20" pageNumber="133">58.9</td>
<td id="786F5DFC1D54004EDF85565BC00DF8C9" box="[1000,1043,1845,1864]" gridcol="7" gridrow="12" pageId="20" pageNumber="133">55.4</td>
<td id="786F5DFC1D54004ED85F565BC08EF8C9" box="[1074,1168,1845,1864]" gridcol="8" gridrow="12" pageId="20" pageNumber="133">53.2-57.4</td>
<td id="786F5DFC1D54004ED8C3565BC0FDF8C9" box="[1198,1251,1845,1864]" gridcol="9" gridrow="12" pageId="20" pageNumber="133">55.2</td>
<td id="786F5DFC1D54004ED96C565BC1BBF8C9" box="[1281,1445,1845,1864]" gridcol="10" gridrow="12" pageId="20" pageNumber="133">56.9</td>
</tr>
<tr id="3BBE34801D54004EDCE65620C1BBF8E0" box="[139,1445,1870,1889]" gridrow="13" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE65620C594F8E0" box="[139,394,1870,1889]" gridcol="0" gridrow="13" pageId="20" pageNumber="133">10-Caudal peduncle depth</th>
<td id="786F5DFC1D54004EDDC45620C5CDF8E0" box="[425,467,1870,1889]" gridcol="1" gridrow="13" pageId="20" pageNumber="133">17.2</td>
<td id="786F5DFC1D54004EDD9E5620C64EF8E0" box="[499,592,1870,1889]" gridcol="2" gridrow="13" pageId="20" pageNumber="133">14.6-18.1</td>
<td id="786F5DFC1D54004EDE035620C6BDF8E0" box="[622,675,1870,1889]" gridcol="3" gridrow="13" pageId="20" pageNumber="133">16.4</td>
<td id="786F5DFC1D54004EDEA45620C6EDF8E0" box="[713,755,1870,1889]" gridcol="4" gridrow="13" pageId="20" pageNumber="133">15.3</td>
<td id="786F5DFC1D54004EDF7E5620C76EF8E0" box="[787,880,1870,1889]" gridcol="5" gridrow="13" pageId="20" pageNumber="133">14.3-16.6</td>
<td id="786F5DFC1D54004EDFE35620C7DDF8E0" box="[910,963,1870,1889]" gridcol="6" gridrow="13" pageId="20" pageNumber="133">15.5</td>
<td id="786F5DFC1D54004EDF855620C00DF8E0" box="[1000,1043,1870,1889]" gridcol="7" gridrow="13" pageId="20" pageNumber="133">17.0</td>
<td id="786F5DFC1D54004ED85F5620C08EF8E0" box="[1074,1168,1870,1889]" gridcol="8" gridrow="13" pageId="20" pageNumber="133">14.5-17.4</td>
<td id="786F5DFC1D54004ED8C35620C0FDF8E0" box="[1198,1251,1870,1889]" gridcol="9" gridrow="13" pageId="20" pageNumber="133">16.5</td>
<td id="786F5DFC1D54004ED96C5620C1BBF8E0" box="[1281,1445,1870,1889]" gridcol="10" gridrow="13" pageId="20" pageNumber="133">16.1</td>
</tr>
<tr id="3BBE34801D54004EDCE65608C1BBF8F8" box="[139,1445,1894,1913]" gridrow="14" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE65608C594F8F8" box="[139,394,1894,1913]" gridcol="0" gridrow="14" pageId="20" pageNumber="133">11-Snout length</th>
<td id="786F5DFC1D54004EDDC45608C5CDF8F8" box="[425,467,1894,1913]" gridcol="1" gridrow="14" pageId="20" pageNumber="133">28.3</td>
<td id="786F5DFC1D54004EDD9E5608C64EF8F8" box="[499,592,1894,1913]" gridcol="2" gridrow="14" pageId="20" pageNumber="133">20.4-30.7</td>
<td id="786F5DFC1D54004EDE035608C6BDF8F8" box="[622,675,1894,1913]" gridcol="3" gridrow="14" pageId="20" pageNumber="133">25.9</td>
<td id="786F5DFC1D54004EDEA45608C6EDF8F8" box="[713,755,1894,1913]" gridcol="4" gridrow="14" pageId="20" pageNumber="133">25.6</td>
<td id="786F5DFC1D54004EDF7E5608C76EF8F8" box="[787,880,1894,1913]" gridcol="5" gridrow="14" pageId="20" pageNumber="133">22.6-26.2</td>
<td id="786F5DFC1D54004EDFE35608C7DDF8F8" box="[910,963,1894,1913]" gridcol="6" gridrow="14" pageId="20" pageNumber="133">25.1</td>
<td id="786F5DFC1D54004EDF855608C00DF8F8" box="[1000,1043,1894,1913]" gridcol="7" gridrow="14" pageId="20" pageNumber="133">24.8</td>
<td id="786F5DFC1D54004ED85F5608C08EF8F8" box="[1074,1168,1894,1913]" gridcol="8" gridrow="14" pageId="20" pageNumber="133">22.0-25.9</td>
<td id="786F5DFC1D54004ED8C35608C0FDF8F8" box="[1198,1251,1894,1913]" gridcol="9" gridrow="14" pageId="20" pageNumber="133">24.1</td>
<td id="786F5DFC1D54004ED96C5608C1BBF8F8" box="[1281,1445,1894,1913]" gridcol="10" gridrow="14" pageId="20" pageNumber="133">19.5</td>
</tr>
<tr id="3BBE34801D54004EDCE65611C1BBF813" box="[139,1445,1919,1938]" gridrow="15" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE65611C594F813" box="[139,394,1919,1938]" gridcol="0" gridrow="15" pageId="20" pageNumber="133">12-Orbital diameter</th>
<td id="786F5DFC1D54004EDDC45611C5CDF813" box="[425,467,1919,1938]" gridcol="1" gridrow="15" pageId="20" pageNumber="133">41.9</td>
<td id="786F5DFC1D54004EDD9E5611C64EF813" box="[499,592,1919,1938]" gridcol="2" gridrow="15" pageId="20" pageNumber="133">39.6-48.1</td>
<td id="786F5DFC1D54004EDE035611C6BDF813" box="[622,675,1919,1938]" gridcol="3" gridrow="15" pageId="20" pageNumber="133">44.1</td>
<td id="786F5DFC1D54004EDEA45611C6EDF813" box="[713,755,1919,1938]" gridcol="4" gridrow="15" pageId="20" pageNumber="133">38.3</td>
<td id="786F5DFC1D54004EDF7E5611C76EF813" box="[787,880,1919,1938]" gridcol="5" gridrow="15" pageId="20" pageNumber="133">36.5-48.8</td>
<td id="786F5DFC1D54004EDFE35611C7DDF813" box="[910,963,1919,1938]" gridcol="6" gridrow="15" pageId="20" pageNumber="133">40.0</td>
<td id="786F5DFC1D54004EDF855611C00DF813" box="[1000,1043,1919,1938]" gridcol="7" gridrow="15" pageId="20" pageNumber="133">45.5</td>
<td id="786F5DFC1D54004ED85F5611C08EF813" box="[1074,1168,1919,1938]" gridcol="8" gridrow="15" pageId="20" pageNumber="133">38.3-46.3</td>
<td id="786F5DFC1D54004ED8C35611C0FDF813" box="[1198,1251,1919,1938]" gridcol="9" gridrow="15" pageId="20" pageNumber="133">42.1</td>
<td id="786F5DFC1D54004ED96C5611C1BBF813" box="[1281,1445,1919,1938]" gridcol="10" gridrow="15" pageId="20" pageNumber="133">45.3</td>
</tr>
<tr id="3BBE34801D54004EDCE656F9C1BBF82B" box="[139,1445,1943,1962]" gridrow="16" pageId="20" pageNumber="133">
<th id="786F5DFC1D54004EDCE656F9C594F82B" box="[139,394,1943,1962]" gridcol="0" gridrow="16" pageId="20" pageNumber="133">13-Postorbital length</th>
<td id="786F5DFC1D54004EDDC456F9C5CDF82B" box="[425,467,1943,1962]" gridcol="1" gridrow="16" pageId="20" pageNumber="133">34.3</td>
<td id="786F5DFC1D54004EDD9E56F9C64EF82B" box="[499,592,1943,1962]" gridcol="2" gridrow="16" pageId="20" pageNumber="133">29.3-38.2</td>
<td id="786F5DFC1D54004EDE0356F9C6BDF82B" box="[622,675,1943,1962]" gridcol="3" gridrow="16" pageId="20" pageNumber="133">34.3</td>
<td id="786F5DFC1D54004EDEA456F9C6EDF82B" box="[713,755,1943,1962]" gridcol="4" gridrow="16" pageId="20" pageNumber="133">42.2</td>
<td id="786F5DFC1D54004EDF7E56F9C76EF82B" box="[787,880,1943,1962]" gridcol="5" gridrow="16" pageId="20" pageNumber="133">30.2-44.0</td>
<td id="786F5DFC1D54004EDFE356F9C7DDF82B" box="[910,963,1943,1962]" gridcol="6" gridrow="16" pageId="20" pageNumber="133">40.1</td>
<td id="786F5DFC1D54004EDF8556F9C00DF82B" box="[1000,1043,1943,1962]" gridcol="7" gridrow="16" pageId="20" pageNumber="133">40.1</td>
<td id="786F5DFC1D54004ED85F56F9C08EF82B" box="[1074,1168,1943,1962]" gridcol="8" gridrow="16" pageId="20" pageNumber="133">33.6-42.4</td>
<td id="786F5DFC1D54004ED8C356F9C0FDF82B" box="[1198,1251,1943,1962]" gridcol="9" gridrow="16" pageId="20" pageNumber="133">38.5</td>
<td id="786F5DFC1D54004ED96C56F9C1BBF82B" box="[1281,1445,1943,1962]" gridcol="10" gridrow="16" pageId="20" pageNumber="133">40.4</td>
</tr>
</table>
</paragraph>
<caption id="D1F1664A1D55FFA4DCEC53D2C73FFD72" ID-DOI="http://doi.org/10.5281/zenodo.10558390" ID-Zenodo-Dep="10558390" httpUri="https://zenodo.org/record/10558390/files/figure.png" pageId="21" pageNumber="134" startId="21.[129,167,700,723]" targetBox="[340,1247,170,679]" targetPageId="21" targetType="figure">
<paragraph id="853136C21D55FFA4DCEC53D2C73FFD72" blockId="21.[129,1451,700,755]" pageId="21" pageNumber="134">
<emphasis id="B7FAEAD01D55FFA4DCEC53D2C4CFFD52" bold="true" box="[129,209,700,723]" pageId="21" pageNumber="134">Fig. 25.</emphasis>
<taxonomicName id="428E4D411D55FFA4DCBB53D2C584FD52" box="[214,410,700,723]" family="Poeciliidae" genus="Phalloceros" kingdom="Animalia" order="Cyprinodontiformes" pageId="21" pageNumber="134" phylum="Chordata" rank="species" species="titthos">
<emphasis id="B7FAEAD01D55FFA4DCBB53D2C584FD52" box="[214,410,700,723]" italics="true" pageId="21" pageNumber="134">Phalloceros titthos</emphasis>
</taxonomicName>
. (A) Male, 17.1 mm SL, USNM 297945, rio da Praia, near Guaratuba; (B) holotype, female, 35.9 mm SL, MZUSP 79670, creek near beach, Guaratuba, Paraná, Brazil.
</paragraph>
</caption>
<subSubSection id="CD9465491D55FFA4DCEC55BAC599FAEA" pageId="21" pageNumber="134" type="etymology">
<paragraph id="853136C21D55FFA4DCEC55BAC599FAEA" blockId="21.[129,771,1236,1387]" pageId="21" pageNumber="134">
<emphasis id="B7FAEAD01D55FFA4DCEC55BAC4E6FB6A" bold="true" box="[129,248,1236,1259]" pageId="21" pageNumber="134">Etymology</emphasis>
. From the Greek τιτθος [= titthós], n. m. nom. sg., a womans breast or animals teat. In allusion to the presence of small papillae at the mandibular symphysis of large adult females. Also alluding to the fact that this character is only present in adult females.
</paragraph>
</subSubSection>
<subSubSection id="CD9465491D55FFA4DCEC54FAC71AFA4A" pageId="21" pageNumber="134" type="discussion">
<paragraph id="853136C21D55FFA4DCEC54FAC71AFA4A" blockId="21.[129,772,1428,1483]" pageId="21" pageNumber="134">
<emphasis id="B7FAEAD01D55FFA4DCEC54FAC4F9FA2A" bold="true" box="[129,231,1428,1451]" pageId="21" pageNumber="134">Remarks</emphasis>
.The gonopodial tip of this species was not described and illustrated due to the lack of adult males available for study.
</paragraph>
</subSubSection>
<subSubSection id="CD9465491D55FFA4DF445259C19BFCEF" pageId="21" pageNumber="134" type="distribution">
<paragraph id="853136C21D55FFA4DF445259C19BFCEF" blockId="21.[809,1452,823,878]" pageId="21" pageNumber="134">
<emphasis id="B7FAEAD01D55FFA4DF445259C7ACFCCF" bold="true" box="[809,946,823,846]" pageId="21" pageNumber="134">Distribution</emphasis>
. Coastal drainages flowing to Baia de Guaratuba and Baia de Paranaguá in Paraná State of Brazil (
<figureCitation id="1DB52A471D55FFA4D9475239C167FCEF" box="[1322,1401,855,878]" captionStart="Fig" captionStartId="11.[129,168,1777,1800]" captionTargetBox="[130,773,1182,1752]" captionTargetId="figure-818@11.[130,773,1182,1752]" captionText="Fig. 12. Collection localities of Phalloceros ocellatus (circle, 1), P. tupinamba (triangle, 2), P. lucenorum (five-pointed star, 3), P. buckupi (eight-pointed star, 4), P. titthos (inverted triangle, 5), and P. caudimaculatus (square, 6). Some symbols represent more than one lot or locality. Numbers represent type localities. The inset is the range of the genus." figureDoi="http://doi.org/10.5281/zenodo.10569750" httpUri="https://zenodo.org/record/10569750/files/figure.png" pageId="21" pageNumber="134">Fig. 12</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>