132 lines
16 KiB
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132 lines
16 KiB
XML
<document id="9ACFC3D5D7F851915E89F59A3B8F2E97" ID-CLB-Dataset="35361" ID-DOI="http://dx.doi.org/10.3897/zookeys.468.8506" ID-GBIF-Dataset="df3f2d10-3b5e-480f-936c-368e01ffd7b1" ID-PMC="PMC4296520" ID-Pensoft-Pub="1313-2970-468-1" ID-PubMed="25610341" ID-ZooBank="AE5AB793FDC74DCD8A47AE96A141E2AD" ModsDocAuthor="" ModsDocDate="2014" ModsDocID="1313-2970-468-1" ModsDocOrigin="ZooKeys 468" ModsDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" checkinTime="1451244902156" checkinUser="pensoft" docAuthor="Shaverdo, Helena, Sagata, Katayo, Panjaitan, Rawati, Menufandu, Herlina & Balke, Michael" docDate="2014" docId="04BE798BDB8F8499DF2952C24ACF7968" docLanguage="en" docName="ZooKeys 468: 1-83" docOrigin="ZooKeys 468" docSource="http://dx.doi.org/10.3897/zookeys.468.8506" docTitle="Exocelina edeltraudae Shaverdo, Hendrich & Balke 2012" docType="treatment" docVersion="5" lastPageNumber="8" masterDocId="FFE9FFA5117AFFD17452FFB5FF94FFE3" masterDocTitle="Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)" masterLastPageNumber="83" masterPageNumber="1" pageNumber="6" updateTime="1701379654281" updateUser="ExternalLinkService">
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<mods:mods id="128B83259A0790EE7BE618FF6DBC22C5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="A6021BBDB78E90C1A25D49615E458553">
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<mods:title id="379F75EFA25EB322DFE402BD14C9B25B">Description of 23 new species of the Exocelinaekari-group from New Guinea, with a key to all representatives of the species group (Coleoptera, Dytiscidae, Copelatinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="F8D4AEF8CC3BAACF8AFAF8BD0B440D04" type="personal">
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<mods:roleTerm id="0F1860A960F07D5AF09FD357FB8F77B4">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="7B00D71C560CF10F770CEF961ECD2179">Shaverdo, Helena</mods:namePart>
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</mods:name>
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<mods:name id="8339FA5DF2184851009CB729A635CA09" type="personal">
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<mods:role id="C7564269835C89602791FDD3693DBEC0">
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<mods:roleTerm id="778CA49851329AA405C9638205C6F3EA">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="E41417F8F53F6A135562DEAA7E5F5AE4">Sagata, Katayo</mods:namePart>
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</mods:name>
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<mods:name id="299ADB152C1703C677414EFBD7A1273E" type="personal">
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<mods:role id="A8725BDE4F3EAE9AF649FBDDC4D46D46">
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<mods:roleTerm id="387631BC28DE66FC6658255D962A99BF">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="97CFD129D79E39288E543964C549BFBD">Panjaitan, Rawati</mods:namePart>
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</mods:name>
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<mods:name id="67137F4607670D8EA4E2D283AEEC9EDF" type="personal">
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<mods:role id="21679343DA1F73210799609C42DF9161">
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<mods:roleTerm id="E3A2429E7989D4BC3CB63F28B59246C1">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="81930C8771DF4FE43FBC07FC89004CF0">Menufandu, Herlina</mods:namePart>
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</mods:name>
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<mods:name id="5CEF57EEA2E21495117424B094C9EF48" type="personal">
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<mods:role id="AD88FF0CCB58CEE34439FC2C0F8FDFC7">
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<mods:roleTerm id="6EC85266714F1C96C5939CDDFE0F743A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="F2A094A32F8D7A1276746FA0D686FE6B">Balke, Michael</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="2E5F0FB23F0863293BB3527C9955EB71">text</mods:typeOfResource>
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<mods:title id="E285D15655C63650C3E68D9E3F4A5759">ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part id="1D26AEA5F70970DA8EFAE9517B3B1062">
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<mods:date id="3F5EAB80B0D2D2546AA1447B1D4B0983">2014</mods:date>
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<mods:detail id="F99D7B29015C5E323DB41CD7E18D9744" type="volume">
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<mods:number id="5096B1456EFBE3A2049D2F90FF0B8F5E">468</mods:number>
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</mods:detail>
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<mods:start id="B2547D1B352A3EB77966C7B5038E8EAF">1</mods:start>
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<mods:end id="6B0E9631CA1D7438BE816BC6D3901DE3">83</mods:end>
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<mods:location id="4658206FBFB8C412D7DCE882BABAE4CE">
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<mods:url id="E5DCA754E1BE281D7709FB7B745BFF05">http://dx.doi.org/10.3897/zookeys.468.8506</mods:url>
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</mods:location>
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<mods:classification id="37DC320B498C70ED0C9AF90C20A8D642">journal article</mods:classification>
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<mods:identifier id="FD614FCED0F23493648098C5AEF1260A" type="DOI">http://dx.doi.org/10.3897/zookeys.468.8506</mods:identifier>
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<mods:identifier id="FAF88BB64463544B4828031E585D93E8" type="Pensoft-Pub">1313-2970-468-1</mods:identifier>
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<mods:identifier id="6C7C9C7F4F4029AF93F9F39DCFEA1C78" type="ZooBank">AE5AB793FDC74DCD8A47AE96A141E2AD</mods:identifier>
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</mods:mods>
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<treatment id="04BE798BDB8F8499DF2952C24ACF7968" ID-GBIF-Taxon="152057389" LSID="urn:lsid:plazi:treatment:04BE798BDB8F8499DF2952C24ACF7968" httpUri="http://treatment.plazi.org/id/04BE798BDB8F8499DF2952C24ACF7968" lastPageId="7" lastPageNumber="8" pageId="5" pageNumber="6">
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<subSubSection id="A479A82EA910478AFECDADC831473CFA" pageId="5" pageNumber="6" type="multiple">
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<paragraph id="18B9CF83E70748071F9C4DEC6774F5F3" pageId="5" pageNumber="6">Taxon classification Animalia Coleoptera Dytiscidae</paragraph>
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</subSubSection>
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<subSubSection id="7ED6A0BDAA95DBDB1DAFC748916EF549" pageId="5" pageNumber="6" type="nomenclature">
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<paragraph id="98F40FD28093B944C89D2829D22143C8" pageId="5" pageNumber="6">
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<taxonomicName id="E91BB4F677A7A36BEB22CCC9CD5303F9" ID-CoL="8LVC9" authority="Shaverdo, Hendrich & Balke, 2012" authorityName="Shaverdo, Hendrich & Balke" authorityYear="2012" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae Shaverdo, Hendrich & Balke, 2012</taxonomicName>
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Figs 8, 37
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</paragraph>
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</subSubSection>
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<subSubSection id="33967DB8B366C31076D779EF1036B8C5" pageId="6" pageNumber="7" type="type locality">
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<paragraph id="561D19335EAF7CB1C916F38E150F7CC7" pageId="6" pageNumber="7">
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<pageBreakToken id="AF5865F6786803DB287C07BB8BC1E88F" pageId="6" pageNumber="7" start="start">Type</pageBreakToken>
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locality.
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</paragraph>
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<paragraph id="12A3527F405CC857B38D231EA5F74080" pageId="6" pageNumber="7">
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Papua New Guinea: Western Highlands Province, Kurumul, 6 km SW Kudjip,
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<geoCoordinate id="98564A0DCF6312AE5EF650F22E29E614" direction="south" orientation="latitude" precision="9" value="-5.8905">05°53.43'S</geoCoordinate>
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;
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<geoCoordinate id="4CBAD6D7570CA1F20AF75576F4CD4721" direction="east" orientation="longitude" precision="9" value="144.61">144°36.60'E</geoCoordinate>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="BCE711E5C38E591E142EE02F3769D8A1" pageId="6" pageNumber="7" type="type material">
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<paragraph id="98234A15CE636CAC5EE430557E6E8686" pageId="6" pageNumber="7">Type material.</paragraph>
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<paragraph id="5A9E7273BCF6048E682B017E5E0C28D2" pageId="6" pageNumber="7">Holotype: male "Papua New Guinea: Western Highlands, Kurumul, 6Km SW Kudjip, small stream, 1584m, 13.vi.1994, 05.53.426S 144.36.600E, John (PNG 78)" (ZSM). Paratypes: 11 males with the same label as the holotype, one of them additionally with a green label "DNA M.Balke 1341" (NHMW, ZSM). 2 males "Papua New Guinea: Western Highlands, Mt. Hagen town area, 1600m, 7.xii.1994 05.49.745S 144.22.357E Balke & Kinibel (PNG 131)" (ZSM).</paragraph>
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</subSubSection>
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<subSubSection id="374982682CBB83ACAF95861BC5756953" pageId="6" pageNumber="7" type="additional material">
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<paragraph id="27F4CE2F2A2FAD50E3FDE3D99678E1FC" pageId="6" pageNumber="7">Additional material.</paragraph>
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<paragraph id="4F9BA7CBAE73004EB6104A7B766448C7" pageId="6" pageNumber="7">
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2 males "Papua New Guinea: Western Highlands, Kurumul, 6Km SW Kudjip, small stream, 1584m, 13.vi.1994, 05.53.426S 144.36.600E, John (PNG 78)" (ZSM). 6 males "Papua New Guinea: Western Highlands, Gonzsidai-Sarup, 1700m, 4.iii.2007, 05.19.060S 144.28.671E, Kinibel (PNG 144)" (NHMW, ZSM). 26 females "Papua New Guinea: Western Highlands, Gonzsidai-Sarup, 1700m, 4.iii.2007, 05.19.060S 144.28.671E, Kinibel (PNG 144)" (ZSM), these females are most likely a mixture of two species:
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<taxonomicName id="F1A223FE0D9290679F0401AE7A8D4C2C" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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and a species from the
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<taxonomicName id="1FE5592EDD92E4C50DE6C432BC99CBF0" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
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-group. 142 females with the same label as the holotype (ZSM), these females are most likely a mixture of three species:
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<taxonomicName id="60058FDA2E02719D4BAB6CEEA2379B0E" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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and two species from the
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<taxonomicName id="2BBD6BED14E908553F76882852D943FA" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
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- and
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<taxonomicName id="2D23358963C1B4F189AF7F2CCFAF8FB1" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina rivulus" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="rivulus">Exocelina rivulus</taxonomicName>
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-groups. 30 females "Papua New Guinea: Western Highlands, Mt. Hagen town area, 1600m, 7.xii.1994 05.49.745S 144.22.357E Balke & Kinibel (PNG 131)" (ZSM), these females are most likely a mixture of two species:
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<taxonomicName id="F5F822E0DFF638305EA5C39D23884E81" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina edeltraudae" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="edeltraudae">Exocelina edeltraudae</taxonomicName>
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and a species from the
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<taxonomicName id="37ECBE2D03AD1FA046D2AB8C7EA28AC6" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina broschii" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="broschii">Exocelina broschii</taxonomicName>
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-group.
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</paragraph>
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</subSubSection>
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<subSubSection id="5D08CF8A81EB21CCA6EEB2ED6244FBF2" pageId="6" pageNumber="7" type="diagnosis">
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<paragraph id="0D36F58E91CB1A5608E51CD931F33164" pageId="6" pageNumber="7">Diagnosis.</paragraph>
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<paragraph id="67E462D7E7B470202A53BE8C1949F0B1" pageId="6" pageNumber="7">
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Beetle medium-sized, piceous, slightly submatt; pronotum with distinct lateral bead; male ventrite 6 slightly to distinctly concave apically; male antennomeres 3-5 distinctly enlarged, almost equal in size and shape, antennomeres 6-8 enlarged; male protarsomere 4 with medium-sized, slender, evidently curved anterolateral hook-like seta; median lobe with very strong submedian constriction and proximal part very broad in ventral view, apex of median lobe pointed and strongly curved downwards in lateral; paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, long, thin setae. The species is very similar to
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<taxonomicName id="D2B42C0E08F25DC2442549C012B29DA3" class="Insecta" family="Dytiscidae" genus="Exocelina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Exocelina pseudoedeltraudae" order="Coleoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="pseudoedeltraudae">Exocelina pseudoedeltraudae</taxonomicName>
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sp. n., from which differs with slightly shinier dorsal surface, due to weaker punctation and microreticulation, with smaller and less rounded male antennomeres 3-5 (for male antennomeres 3 and 4, ratio width/length: <0.92) and apex of median lobe broader in lateral view.
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</paragraph>
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</subSubSection>
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<subSubSection id="6BE831CD35FF80BB63BB9C66ED110226" lastPageId="7" lastPageNumber="8" pageId="6" pageNumber="7" type="redescription">
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<paragraph id="358EC30C6C936053E27E9D9CFF20238C" pageId="6" pageNumber="7">Redescription.</paragraph>
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<paragraph id="1AB5A19B62E34F49389BB3FE4D4A47B1" pageId="6" pageNumber="7">Size and shape: Beetle medium-sized (TL-H 3.5-3.85 mm, TL 3.9-4.3 mm, MW 1.9-2.05 mm), with oblong-oval habitus, broadest at elytral middle. Coloration: Dorsally piceous, with dark brown anterior margin of head and narrowly pronotal sides; head appendages and legs reddish to reddish brown, legs distally darker (Fig. 37). Teneral specimens paler.</paragraph>
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<paragraph id="E4D02D73693D321F453FA5804CC230A1" pageId="7" pageNumber="8">
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<pageBreakToken id="D8C4AA12D6441DAE0842E25C7CD51A91" pageId="7" pageNumber="8" start="start">Surface</pageBreakToken>
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sculpture: Head with dense, coarse punctation (spaces between punctures 1-3 times size of punctures). Pronotum with punctation finer, sparser, and more evenly distributed than on head. Elytra with punctation much finer, sparser than on pronotum. Pronotum and elytra with less strongly impressed microreticulation, dorsal surface slightly submatt. Head with microreticulation stronger. Metaventrite and metacoxa distinctly microreticulate, metacoxal figs with longitudinal strioles and transverse wrinkles. Abdominal ventrites with distinct microreticulation, long strioles, and fine sparse punctation, coarser and denser on two last abdominal ventrites.
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</paragraph>
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<paragraph id="05215D13621E5836C491F3735FF1FF60" pageId="7" pageNumber="8">Structures: Pronotum with distinct lateral bead. Base of prosternum and neck of prosternal process with distinct ridge, rounded and smooth anteriorly, with small anterolateral extensions. Blade of prosternal process lanceolate, relatively narrow, convex, with distinct lateral bead and few setae; neck and blade of prosternal process evenly jointed. Abdominal ventrite 6 broadly rounded, or slightly truncate, or concave apically.</paragraph>
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<paragraph id="C606BB93DE8C0282DAD17DFAFC1AF0A3" pageId="7" pageNumber="8">Male: Antennomeres 3-5 distinctly enlarged, almost equal in size, antennomeres 6-8 enlarged (Fig. 8A), antennomeres 3-7 rugose ventrally. Protarsomere 4 with medium-sized, slender, evidently curved anterolateral hook-like seta. Protarsomere 5 ventrally with anterior row of 16 and posterior row of 5 short setae (Fig. 8B). Abdominal ventrite 6 with 8-13 lateral striae on each side, slightly to distinctly concave apically. Median lobe with very strong submedian constriction and proximal part very broad in ventral view, apex of median lobe pointed and strongly curved downwards in lateral view (Fig. 8D, E). Paramere with distinct notch on dorsal side and subdistal part elongate, with numerous, dense, long, thin setae (Fig. 8F).</paragraph>
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<paragraph id="53C1037070FC9626D136D987A7D8554B" pageId="7" pageNumber="8">Holotype: TL-H 3.85 mm, TL 4.3 mm, MW 2.05 mm.</paragraph>
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<paragraph id="711ADD58B392A5C93D8E7568289417DA" pageId="7" pageNumber="8">Female: Antennae simple, abdominal ventrite 6 broadly rounded or slightly truncate apically, without striae.</paragraph>
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</subSubSection>
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<subSubSection id="8E5EE4B2621D8C8A867C8A53676C1234" pageId="7" pageNumber="8" type="distribution">
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<paragraph id="F61232517807F974B0EFD56A417943FA" pageId="7" pageNumber="8">Distribution.</paragraph>
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<paragraph id="D8624705E4368406AEC45990388EA07E" pageId="7" pageNumber="8">Papua New Guinea. The species is known only from Western Highlands Province (Fig. 53).</paragraph>
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</subSubSection>
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</treatment>
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</document> |