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<document id="533CDAF94371BFD3CB1BD36F62BAA27F" ID-CLB-Dataset="31425" ID-DOI="10.11646/zootaxa.4085.4.3" ID-GBIF-Dataset="67846749-8510-4ff2-85c1-d9a684cf72a8" ID-ISSN="1175-5326" ID-Zenodo-Dep="1052741" ID-ZooBank="39DEC898-B737-4ABC-A03C-BEF9DE2E4FC6" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1510873838092" checkinUser="plazi" docAuthor="Bezděk, Jan" docDate="2016" docId="03D9D645FFD5FF99FF33FDB6FEBEFAE4" docLanguage="en" docName="zootaxa.4085.4.3.pdf" docOrigin="Zootaxa 4085 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Coeligetoides trifurcatus Bezděk 2016" docType="treatment" docVersion="7" masterDocId="FFE0AE3DFFC5FF8AFFA4FFAFFFFFFFB1" masterDocTitle="Revision of the genus Coeligetes from Malaysia and Indonesia, and description of Coeligetoides gen. nov. (Coleoptera: Chrysomelidae: Galerucinae)" updateTime="1698499684459" updateUser="ExternalLinkService">
<mods:mods id="89AE6D54256C448DD9C583BF39A3A1DA" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C1D53734DE1FF9F44BCD8558D7B27527">
<mods:title id="81A0574AE391B7CE40343D39B2C3C392">Revision of the genus Coeligetes from Malaysia and Indonesia, and description of Coeligetoides gen. nov. (Coleoptera: Chrysomelidae: Galerucinae)</mods:title>
</mods:titleInfo>
<mods:name id="0C370870C322B5CD201D7C1C077C52F2" type="personal">
<mods:role id="A385CBD90019F5A581C162B8F7E97CCB">
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<mods:namePart id="FB8180A596CE5E9D477892B02C3EDCEF">Bezděk, Jan</mods:namePart>
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<mods:typeOfResource id="6DE231BA00375CDFF20F0D6C5B7F7376">text</mods:typeOfResource>
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<mods:title id="C7CACE3240A03685865D9A2F7D6F6F9B">Zootaxa</mods:title>
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<mods:part id="7466909A0D246DF072C6EAD623B10B8F">
<mods:date id="DA03DDD4CDEB94019BD6F8D6FF14F040">2016</mods:date>
<mods:detail id="C97A307B52B9EE5384B2093778776146" type="pubDate">
<mods:number id="AD0A9A4AABAB5CCB4BEFD24D2A80E130">2016-03-04</mods:number>
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<mods:number id="0936C35738079A1E0D98D7850E647A8E">4085</mods:number>
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<mods:number id="77B06B302E5577EB51379A518CB05796">4</mods:number>
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<mods:classification id="88FED46CF383E728B1A4FF43F42B72F9">journal volume</mods:classification>
<mods:identifier id="4BEDBD45AF15A7EE9D181DB11475899D" type="CLB-Dataset">31425</mods:identifier>
<mods:identifier id="7BF2DCC5F1FB46C92DB9BE975E40761E" type="DOI">10.11646/zootaxa.4085.4.3</mods:identifier>
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<treatment id="03D9D645FFD5FF99FF33FDB6FEBEFAE4" ID-DOI="http://doi.org/10.5281/zenodo.6086522" ID-GBIF-Taxon="137309166" ID-Zenodo-Dep="6086522" LSID="urn:lsid:plazi:treatment:03D9D645FFD5FF99FF33FDB6FEBEFAE4" httpUri="http://treatment.plazi.org/id/03D9D645FFD5FF99FF33FDB6FEBEFAE4" lastPageId="19" pageId="16">
<subSubSection id="C36A34D8FFD5FF9AFF33FDB6FED8FDE4" pageId="16" type="nomenclature">
<paragraph id="8BCF6753FFD5FF9AFF33FDB6FD71FD82" blockId="16.[151,654,537,597]" box="[151,654,537,563]" pageId="16">
<heading id="D087D03FFFD5FF9AFF33FDB6FD71FD82" bold="true" box="[151,654,537,563]" fontSize="11" level="1" pageId="16" reason="1">
<emphasis id="B904BB41FFD5FF9AFF33FDB6FD71FD82" bold="true" box="[151,654,537,563]" pageId="16">
<taxonomicName id="4C701CD0FFD5FF9AFF33FDB6FE40FD82" authorityName="Bezděk" authorityYear="2016" box="[151,447,537,563]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="trifurcatus">
<emphasis id="B904BB41FFD5FF9AFF33FDB6FE40FD82" bold="true" box="[151,447,537,563]" italics="true" pageId="16">Coeligetoides trifurcatus</emphasis>
</taxonomicName>
BezdĚK, SP. nov.
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCF6753FFD5FF9AFF33FD93FED8FDE4" blockId="16.[151,654,537,597]" box="[151,295,572,597]" pageId="16">
(
<figureCitation id="134B7BD6FFD5FF9AFF3BFD93FEE0FDE4" box="[159,287,572,597]" captionStart-0="FIGURES 43 44" captionStart-1="FIGURES 45 51" captionStartId-0="17.[151,264,1976,1999]" captionStartId-1="18.[151,264,1880,1903]" captionTargetBox-0="[175,1376,1111,1931]" captionTargetBox-1="[169,1416,212,1844]" captionTargetId-0="figure@17.[151,1436,1087,1955]" captionTargetId-1="figure@18.[151,1436,193,1860]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="FIGURES 43 44. Habitus of Coeligetoides gen. nov. 43 — C. trifurcatus sp. nov. (holotype, male, 8.4 mm); 44 — C. trifurcatus sp. nov. (paratype, female, 10.7 mm)." captionText-1="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri-0="https://zenodo.org/record/1052755/files/figure.png" httpUri-1="https://zenodo.org/record/1052757/files/figure.png" pageId="16">Figs 4351</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C36A34D8FFD5FF9BFF33FDD1FC80FED6" lastPageId="17" pageId="16" type="materials_examined">
<paragraph id="8BCF6753FFD5FF9AFF33FDD1FEE4FD0A" blockId="16.[151,1437,638,2037]" pageId="16">
<materialsCitation id="3B186D0EFFD5FF9AFF33FDD1FEE4FD0A" ID-GBIF-Occurrence="1701201102" country="Indonesia" latitude="-1.135" location="Sungai Wain Protection Forest" longLatPrecision="130" longitude="116.831665" pageId="16" specimenCount="1" stateProvince="East Kalimantan" typeStatus="holotype">
<emphasis id="B904BB41FFD5FF9AFF33FDD1FEC6FD26" bold="true" box="[151,313,638,663]" pageId="16">
<typeStatus id="54CBD9F1FFD5FF9AFF33FDD1FF2DFD26" box="[151,210,638,663]" pageId="16">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F36727C3FFD5FF9AFEE0FDD1FE4BFD26" box="[324,436,638,663]" name="Indonesia" pageId="16">Indonesia</collectingCountry>
,
<collectingRegion id="49B4A9B1FFD5FF9AFE67FDD0FD7CFD26" box="[451,643,638,663]" country="Indonesia" name="Kalimantan Timur" pageId="16">East Kalimantan</collectingRegion>
,
<locationDeviation id="9A7D7E71FFD5FF9AFD36FDD1FC54FD26" box="[658,939,638,663]" pageId="16">
<quantity id="4C88CAB6FFD5FF9AFD36FDD1FD22FD26" box="[658,733,638,663]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="16" unit="km" value="15.0">15 km</quantity>
N of Balikpapan
</locationDeviation>
,
<location id="8EAF3188FFD5FF9AFC1EFDD1FAE0FD27" LSID="urn:lsid:plazi:treatment:03D9D645FFD5FF99FF33FDB6FEBEFAE4:8EAF3188FFD5FF9AFC1EFDD1FAE0FD27" box="[954,1311,638,663]" country="Indonesia" latitude="-1.135" longLatPrecision="130" longitude="116.831665" name="Sungai Wain Protection Forest" pageId="16" stateProvince="East Kalimantan">Sungai Wain Protection Forest</location>
,
<geoCoordinate id="EE440194FFD5FF9AFA8AFDD1FA63FD26" box="[1326,1436,638,663]" direction="south" orientation="latitude" pageId="16" precision="92" value="-1.135">01°08.1S</geoCoordinate>
<geoCoordinate id="EE440194FFD5FF9AFF33FD0DFEE8FD0A" box="[151,279,674,699]" direction="east" orientation="longitude" pageId="16" precision="92" value="116.831665">116°49.9E</geoCoordinate>
.
</materialsCitation>
</paragraph>
<paragraph id="8BCF6753FFD5FF9BFF63FD69FC80FED6" blockId="16.[151,1437,638,2037]" lastBlockId="17.[151,1437,151,1043]" lastPageId="17" pageId="16">
<emphasis id="B904BB41FFD5FF9AFF63FD69FE88FD6E" bold="true" box="[199,375,710,735]" pageId="16">Type material.</emphasis>
Holotype: ♂ (NMPC), “INDONESIA, E Kalimantan / ca.
<quantity id="4C88CAB6FFD5FF9AFB86FD69FB93FD6E" box="[1058,1132,710,735]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="16" unit="km" value="15.0">15 km</quantity>
N of Balikpapan / Sungai Wain Protection Forest / 01°08.1
<emphasis id="B904BB41FFD5FF9AFDACFD45FDF2FCB5" bold="true" box="[520,525,746,772]" pageId="16">'</emphasis>
S, 116°49.9
<emphasis id="B904BB41FFD5FF9AFD34FD45FD6AFCB5" bold="true" box="[656,661,746,772]" pageId="16">'</emphasis>
E,
<quantity id="4C88CAB6FFD5FF9AFD11FD44FD10FCB2" box="[693,751,747,772]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="16" unit="m" value="35.0">35 m</quantity>
/ J. Hájek, J. Schneider &amp; / P. Votruba leg.
<date id="FFCE4193FFD5FF9AFB7CFD44FA8DFCB5" box="[1240,1394,747,772]" pageId="16" value="2011-12-08" valueMax="2011-12-11" valueMin="2011-12-08">811.xii.2011</date>
[w, p] // primary rainforest; individual / collecting in stream, puddles and / swamp, on vegetation and dead / wood; light trap at border of forest [w, p]”. Paratypes: 1 ♂ (NMPC), same label data as in holotype; 1 ♀ (NMPC), “MALAYSIA, Perak / Cameron Highlands / Batu (= Mile) 19 vill. env. / 04°22.2
<emphasis id="B904BB41FFD5FF9AFB84FCF8FBDAFCC0" bold="true" box="[1056,1061,855,881]" pageId="16">'</emphasis>
N, 101°20.0
<emphasis id="B904BB41FFD5FF9AFB0AFCF8FB4CFCC0" bold="true" box="[1198,1203,855,881]" pageId="16">'</emphasis>
E,
<quantity id="4C88CAB6FFD5FF9AFB76FCF7FAE6FCC1" box="[1234,1305,856,881]" metricMagnitude="2" metricUnit="m" metricValue="5.9" pageId="16" unit="m" value="590.0">590 m</quantity>
/ Jiří Hájek leg.
<date id="FFCE4193FFD5FF9AFF6EFCD2FEA4FC27" box="[202,347,893,918]" pageId="16" value="2009-05-05" valueMax="2009-05-15" valueMin="2009-05-05">5.15.v.2009</date>
[w, p]”; 1 ♀ (JBCB), “MALAYSIA W, 2013 / KELANTAN, Kg. Tunku / Mt. Noring Timur,
<quantity id="4C88CAB6FFD5FF9AFF33FC0DFF12FC08" box="[151,237,930,954]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="16" unit="m" value="1200.0">1200 m</quantity>
/
<quantity id="4C88CAB6FFD5FF9AFEA2FC0EFEA2FC0B" box="[262,349,929,954]" metricMagnitude="5" metricUnit="m" metricValue="1.5" pageId="16" unit="km" value="150.0">150 km</quantity>
S of JELI,
<date id="FFCE4193FFD5FF9AFE45FC0EFDE7FC0B" box="[481,536,929,954]" pageId="16">21.ii.</date>
/
<date id="FFCE4193FFD5FF9AFDE6FC0EFD7AFC0B" box="[578,645,929,954]" pageId="16">14.iii.</date>
, P. Čechovský leg. [w, p]”; 2 ♀ (JBCB), “MALAYSIA, prov. Johor / Tioman Isl. / jungle track
<quantity id="4C88CAB6FFD5FF9AFE60FC6AFDC8FC6C" box="[452,567,965,990]" metricMagnitude="2" metricUnit="m" metricValue="3.05" metricValueMax="6.0" metricValueMin="0.1" pageId="16" unit="m" value="305.0" valueMax="600.0" valueMin="10.0">10600 m</quantity>
a.s.l. / 15.
<date id="FFCE4193FFD5FF9AFD1EFC6AFCCDFC6F" box="[698,818,965,990]" pageId="16" value="2009-03-29">29.3.2009</date>
, V. Hula [w, p]”; 1 ♂ (JBCB), “MALAYSIA, Pahang distr., /
<quantity id="4C88CAB6FFD5FF9AFF49FC45FECAFBB2" box="[237,309,1002,1027]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="16" unit="km" value="30.0">30 km</quantity>
NE Raub, Lata Lembik / 3°56
<emphasis id="B904BB41FFD5FF9AFD37FC47FD67FBB3" bold="true" box="[659,664,1000,1026]" pageId="16">'</emphasis>
N 101°38
<emphasis id="B904BB41FFD5FF9AFCA2FC47FCF4FBB3" bold="true" box="[774,779,1000,1026]" pageId="16">'</emphasis>
E,
<quantity id="4C88CAB6FFD5FF9AFC8FFC45FC52FBB3" box="[811,941,1002,1026]" metricMagnitude="2" metricUnit="m" metricValue="3.0" metricValueMax="4.0" metricValueMin="2.0" pageId="16" unit="m" value="300.0" valueMax="400.0" valueMin="200.0">200400 m</quantity>
, /
<date id="FFCE4193FFD5FF9AFC6EFC45FB78FBB2" box="[970,1159,1002,1027]" pageId="16" value="2002-04-22" valueMax="2002-05-15" valueMin="2002-04-22">22.iv.15.v.2002</date>
, / E. Jendek &amp; O. Šauša leg. [w, p]”; 1 ♂ (NHMB), “MALAYSIA; Tioman;
<quantity id="4C88CAB6FFD5FF9AFD72FBA0FCE5FB96" box="[726,794,1039,1063]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="16" unit="m" value="400.0">400m</quantity>
; / Kampong Tekek—K. Juara /
<date id="FFCE4193FFD5FF9AFBDDFBA1FB17FB96" box="[1145,1256,1038,1063]" pageId="16" value="1998-03-09">9.iii.1998</date>
; 2,48N 104,11E / Dembický &amp; Pacholátko leg. [w, p]”; 3 ♀ (NHMB), “W SUMATRA prov.; Kerinci / Seblat N. P.;
<quantity id="4C88CAB6FFD5FF9AFA89FB9DFA92FBFA" box="[1325,1389,1074,1099]" metricMagnitude="4" metricUnit="m" metricValue="2.4" pageId="16" unit="km" value="24.0">24km</quantity>
NE Tapan; / MUARA SAKO→E env.; / 2°05
<emphasis id="B904BB41FFD5FF9AFD2AFBF9FD6CFBC1" bold="true" box="[654,659,1110,1136]" pageId="16">'</emphasis>
S 101°15
<emphasis id="B904BB41FFD5FF9AFCA5FBF9FCF9FBC1" bold="true" box="[769,774,1110,1136]" pageId="16">'</emphasis>
E;
<quantity id="4C88CAB6FFD5FF9AFC88FBF8FC58FBC1" box="[812,935,1111,1136]" metricMagnitude="2" metricUnit="m" metricValue="4.75" metricValueMax="5.5" metricValueMin="4.0" pageId="16" unit="m" value="475.0" valueMax="550.0" valueMin="400.0">400550m</quantity>
; / Dembický leg.;
<date id="FFCE4193FFD5FF9AFB2AFBF8FAD4FBC1" box="[1166,1323,1111,1136]" pageId="16" value="2003-03-04" valueMax="2003-03-18" valueMin="2003-03-04">4.18.iii.2003</date>
[w, p] // Collection / Naturhistorisches / Museum Basel [w, p]”; 2 ♀ (HNHM), “THAILAND, pr. Trang, / Kao Chong Nature / Wildlife Reserve / Center, rainforest, [w, p] // river Khao Chong, / swept &amp; beaten, /
<date id="FFCE4193FFD5FF9AFB09FB0FFAD1FB09" box="[1197,1326,1184,1208]" pageId="16" value="2003-11-22">22.XI.2003</date>
, No. 14, / A. Orosz &amp; Gy. Sziráki [w, p]”; 1 ♂ (BMNH), “59470 [w, h] // Doherty [w, p] // Borneo / Pengaron [w, p]”; 1 ♀ (ZMHB), “Dohrn / Sumatra / Liangagas [w, p] // 71514 [w, p] // Cotype [sic!] [o, p] //
<taxonomicName id="4C701CD0FFD5FF9AFBCDFB48FB12FAB1" authorityName="Jacoby" authorityYear="1893" box="[1129,1261,1255,1280]" class="Insecta" family="Chrysomelidae" genus="Coeligethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="genus">Coeligethes</taxonomicName>
/
<taxonomicName id="4C701CD0FFD5FF9AFAA0FB48FA9EFAB1" authorityName="Jacoby" authorityYear="1895" box="[1284,1377,1255,1280]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="unicolor">unicolor</taxonomicName>
/ Jac [w, h]”; 1 ♂ 1 ♀ (BMNH), “Dohrn / Sumatra / Liangagas [w, p] // Jacoby Coll. /
<date id="FFCE4193FFD5FF9AFB8EFAA3FB75FA95" box="[1066,1162,1292,1316]" pageId="16" value="1909" valueMax="1928">1909-28</date>
a. [w, p]”; 1 ♂ (MSNG), “Dohrn / Sumatra / Liangagas [w, p] //
<taxonomicName id="4C701CD0FFD5FF9AFDC1FA80FD16FAF9" authorityName="Jacoby" authorityYear="1893" box="[613,745,1327,1352]" class="Insecta" family="Chrysomelidae" genus="Coeligethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="genus">Coeligethes</taxonomicName>
/
<taxonomicName id="4C701CD0FFD5FF9AFCA1FA80FC45FAF9" authority="Jacoby" authorityName="Jacoby" authorityYear="1895" box="[773,954,1327,1352]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="unicolor">unicolor Jacoby</taxonomicName>
/ det. M. Jacoby, 1899 [w, p] // MUSEO GENOVA / dedit Dr. Heinrich / Dohrn (Stettin), 1900 [w, p]”; 1 ♂ (MSNG), “Soekaranda /
<date id="FFCE4193FFD5FF9AFB34FAFAFAE8FADD" box="[1168,1303,1364,1388]" pageId="16" value="1894-01">Januar 1894</date>
/ Dohrn [w, p] //
<taxonomicName id="4C701CD0FFD5FF9AFF6BFAD8FEACFA21" authorityName="Jacoby" authorityYear="1893" box="[207,339,1399,1424]" class="Insecta" family="Chrysomelidae" genus="Coeligethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="genus">Coeligethes</taxonomicName>
/
<taxonomicName id="4C701CD0FFD5FF9AFEC9FAD8FE35FA21" authorityName="Jacoby" authorityYear="1895" box="[365,458,1399,1424]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="unicolor">unicolor</taxonomicName>
/ Jac [w, h] //
<taxonomicName id="4C701CD0FFD5FF9AFDD0FAD8FD06FA21" authorityName="Jacoby" authorityYear="1893" box="[628,761,1399,1424]" class="Insecta" family="Chrysomelidae" genus="Coeligethes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="genus">Coeligethes</taxonomicName>
/
<taxonomicName id="4C701CD0FFD5FF9AFCB6FAD8FC39FA21" authority="Jacoby" authorityName="Jacoby" authorityYear="1895" box="[786,966,1399,1424]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="species" species="unicolor">unicolor Jacoby</taxonomicName>
/ det. M. Jacoby, 1899 [w, p] // MUSEO GENOVA / dedit Dr. Heinrich / Dohrn (Stettin), 1900 [w, p]”; 1 ♀ (BMNH), “Mt. Matang, / W. Sarawak. / G. E. Bryant. [p] /
<date id="FFCE4193FFD5FF9AFE88FA6FFE66FA69" box="[300,409,1472,1496]" pageId="16" value="2013-12-20">20.XII.13</date>
[w, h] // MSI [w, h] // G. Bryant Coll. / 1919-147 [w, p]”; 1 ♀ (BMNH), “PENINSULAR SIAM, / NAKON SRI TAMARAT / KHAO LUANG [p] / 15009000 [h] FT. [p] /
<date id="FFCE4193FFD5FF9AFBE9FA4BFB2FFA4C" box="[1101,1232,1508,1533]" pageId="16">March 20th</date>
[h] 1922. / H. M. PENDLEBURY [w, p] //
<taxonomicName id="4C701CD0FFD5FF9AFE12F9A8FDD3F991" authorityName="Bezděk" authorityYear="2016" box="[438,556,1543,1568]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="genus">Coeligetes</taxonomicName>
/ sp. [h] / Det. G. E. Bryant [w, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♀ (BMNH), “PENINSULAR SIAM, / NAKON SRI TAMARAT / KHAO LUANG [p] / 2000 [h] FT. [p] /
<date id="FFCE4193FFD5FF9AFF03F9E0FED3F9D9" box="[167,300,1615,1640]" pageId="16">March 25th</date>
[h] 1922. / H. M. PENDLEBURY [w, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♂ (BMNH), “PAHANG, F. M. S. / Lubok Tamang [p] / 3500 [h] ft. [p] / 10th [h] June, 1923 / H. M. Pendlebury. [w, p] // Ex F. M. S. / Museum. / B. M. 1955-354. [w, p]”; 1 ♂ 3 ♀ (UKM), “PERAK: Taiping / Bukit Larut /
<date id="FFCE4193FFD5FF9AFAF5F937FF30F965" pageId="16" value="1994-06-20">20 Jun 1994</date>
/ Ismail, Ruslan, Yusof [w, p]”; 1 ♀ (UKM), “PERAK: Bukit / Larut / 21 Juiai 1994 / Salleh, Ismail &amp; Ruslan [w, p]”; 1 ♂ (UKM), “PERAK: Taiping, / Bukit Larut /
<date id="FFCE4193FFD5FF9AFCABF970FC57F949" box="[783,936,1759,1784]" pageId="16" value="1991-06-13" valueMax="1991-06-15" valueMin="1991-06-13">1315 Jun 91</date>
/ Ismail &amp; Yusof [w, p] //
<taxonomicName id="4C701CD0FFD5FF9AFB71F94FFA63F949" authorityName="Latreille" authorityYear="1802" box="[1237,1436,1760,1784]" class="Insecta" family="Chrysomelidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="subFamily" subFamily="Galerucinae">GALERUCINAE</taxonomicName>
[p] /
<taxonomicName id="4C701CD0FFD5FF9AFF68F8ABFEDCF8AC" authorityName="Weise" authorityYear="1889" box="[204,291,1796,1821]" class="Insecta" family="Chrysomelidae" genus="Liroetis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="16" phylum="Arthropoda" rank="genus">Liroetis</taxonomicName>
[h] / det. Mohamedsaid 199 [p] 3 [w, h]”; 1 ♀ (UKM), “PERAK: Taiping, / Bukit Larut /
<date id="FFCE4193FFD5FF9AFAA3F8ABFA63F8AD" box="[1287,1436,1796,1821]" pageId="16" value="1991-06-13" valueMax="1991-06-15" valueMin="1991-06-13">1315 Jun 91</date>
/ Ismail &amp; Yusof [w, p]”; 1 ♀ (UKM), “PERAK: Bukit / Larut / 5 Ogos 1991 / Mohd Salleh [w, p]”; 1 ♀ (UKM), “PERAK: Bukit / Larut / 89 Mac 1990 / Ismail, Ruslan [w, p]”; 1 ♂ (UKM), “PERAK: Bukit / Larut [h] / 27 Mei 90 / Mohd Salleh [w, p]”; 1 ♀ (UKM), “TERENGGANU: / H. Lipur Sekayu / 24 Ogos 1994 / Ismail &amp; Zabidi [w, p]”; 1 ♀ (UKM), “SELANGOR: Hutan / Rekrasi Sg. Sendat /
<date id="FFCE4193FFD5FF9AFCF9F83BFC29F81D" box="[861,982,1940,1965]" pageId="16" value="1991-10-12">12 Okt. 91</date>
/ Ismail &amp; Ruslan [w, p]”; 1 ♀ (UKM), “PAHANG: Pulau Tioman / 2731 Ogos 91 / Yusof, J´din &amp; Mahbob [w, p]”; 1 ♀ (UKM), “PAHANG: Cameron / Highlands /
<date id="FFCE4193FFD5FF9AFE85F873FE4EF845" box="[289,433,2012,2037]" pageId="16" value="1990-04-04">4 April 1990</date>
/ Zaidi, Ismail, Ruslan [w, p]”; 1 ♂ (PRCS), “MALAYSIA, NW N Penang Island, / near Teluk Bahang vill., National Park / h~
<quantity id="4C88CAB6FFD4FF9BFD9BFF37FD7FFF00" box="[575,640,152,177]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="17" unit="m" value="160.0">160m</quantity>
,N 05°27
<emphasis id="B904BB41FFD4FF9BFD40FF38FD16FF00" bold="true" box="[740,745,151,177]" pageId="17">'</emphasis>
29'', E 100°11
<emphasis id="B904BB41FFD4FF9BFC21FF38FC75FF00" bold="true" box="[901,906,151,177]" pageId="17">'</emphasis>
36'' /
<date id="FFCE4193FFD4FF9BFC61FF36FBBFFF00" box="[965,1088,153,177]" pageId="17" value="2014-02-18">18. II.2014</date>
P. Romantsov leg. [w, p]”; 1 ♂ (PRCS), “MALAYSIA, N Borneo, Sabah, / Keningau dist., Trus Madi Mt., / h~
<quantity id="4C88CAB6FFD4FF9BFBB0FF11FB9BFF66" box="[1044,1124,190,215]" metricMagnitude="3" metricUnit="m" metricValue="1.25" pageId="17" unit="m" value="1250.0">1250m</quantity>
, N 05°26
<emphasis id="B904BB41FFD4FF9BFB6AFF13FB2CFF67" bold="true" box="[1230,1235,188,214]" pageId="17">'</emphasis>
35'', E 116°27
<emphasis id="B904BB41FFD4FF9BFAD4FF13FA8AFF67" bold="true" box="[1392,1397,188,214]" pageId="17">'</emphasis>
5'' /
<date id="FFCE4193FFD4FF9BFF33FF4CFEF4FF4A" box="[151,267,227,251]" pageId="17" value="2014-02-28">28.II.2014</date>
P. Romantsov leg. [w, p]”; 1 ♀ (BMNH), “BRUNEI:
<date id="FFCE4193FFD4FF9BFCC8FF4CFC44FF4A" box="[876,955,227,251]" pageId="17" value="1992-10">x-1992</date>
/ Temburong District / Ridge NE of Kuala / Belagong,
<quantity id="4C88CAB6FFD4FF9BFE84FEA8FE9CFEAE" box="[288,355,263,287]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="17" unit="m" value="300.0">300m</quantity>
. [w, p] // 125W MV Light Trap / J H Martin coll. / BMNH(E) 1992-172 [w, p]”. The specimens are provided with one additional printed red label: “HOLOTYPUS, [or PARATYPUS, respectively], /
<taxonomicName id="4C701CD0FFD4FF9BFF33FEE1FECEFED7" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[151,305,334,358]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B904BB41FFD4FF9BFF33FEE1FECEFED7" box="[151,305,334,358]" italics="true" pageId="17">Coeligetoides</emphasis>
</taxonomicName>
g. nov. /
<taxonomicName id="4C701CD0FFD4FF9BFE3DFEE0FDF1FED6" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[409,526,335,359]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="species" species="trifurcatus" status="sp. nov.">
<emphasis id="B904BB41FFD4FF9BFE3DFEE0FDF1FED6" box="[409,526,335,359]" italics="true" pageId="17">trifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD4FF9BFDB1FEE0FD91FED7" box="[533,622,335,358]" pageId="17" rank="species">sp. nov.</taxonomicNameLabel>
, / J. Bezděk det., 2015”.
</paragraph>
</subSubSection>
<subSubSection id="C36A34D8FFD4FF99FF63FEDDFE33FCEC" lastPageId="19" pageId="17" type="description">
<paragraph id="8BCF6753FFD4FF9BFF63FEDDFB2DFE3B" blockId="17.[151,1437,151,1043]" box="[199,1234,370,395]" pageId="17">
<emphasis id="B904BB41FFD4FF9BFF63FEDDFEAEFE3A" bold="true" box="[199,337,370,395]" pageId="17">Description</emphasis>
. Measurements. Males:
<quantity id="4C88CAB6FFD4FF9BFDC0FEDDFD14FE3B" box="[612,747,370,395]" metricMagnitude="-3" metricUnit="m" metricValue="8.6" metricValueMax="9.5" metricValueMin="7.7" pageId="17" unit="mm" value="8.6" valueMax="9.5" valueMin="7.7">7.79.5 mm</quantity>
(
<typeStatus id="54CBD9F1FFD4FF9BFD5DFEDDFCA2FE3A" box="[761,861,370,395]" pageId="17" type="holotype">holotype</typeStatus>
<quantity id="4C88CAB6FFD4FF9BFCC0FEDCFC41FE3A" box="[868,958,371,395]" metricMagnitude="-3" metricUnit="m" metricValue="8.4" pageId="17" unit="mm" value="8.4">8.4 mm</quantity>
), females: 8.0
<quantity id="4C88CAB6FFD4FF9BFBCCFEDCFB32FE3B" box="[1128,1229,371,395]" metricMagnitude="-2" metricUnit="m" metricValue="1.17" pageId="17" unit="mm" value="11.7">11.7 mm</quantity>
.
</paragraph>
<paragraph id="8BCF6753FFD4FF9BFF63FE39FF30FE62" blockId="17.[151,1437,151,1043]" pageId="17">Body almost completely yellowish brown, except black apices of mandibles and slightly infuscate tibiae and tarsi.</paragraph>
<paragraph id="8BCF6753FFD4FF9BFF63FE71FCEDFCA6" blockId="17.[151,1437,151,1043]" pageId="17">
Male (
<typeStatus id="54CBD9F1FFD4FF9BFEB0FE71FE85FE46" box="[276,378,478,503]" pageId="17" type="holotype">holotype</typeStatus>
,
<figureCitation id="134B7BD6FFD4FF9BFE2EFE71FE1DFE46" box="[394,482,478,503]" captionStart="FIGURES 43 44" captionStartId="17.[151,264,1976,1999]" captionTargetBox="[175,1376,1111,1931]" captionTargetId="figure@17.[151,1436,1087,1955]" captionTargetPageId="17" captionText="FIGURES 43 44. Habitus of Coeligetoides gen. nov. 43 — C. trifurcatus sp. nov. (holotype, male, 8.4 mm); 44 — C. trifurcatus sp. nov. (paratype, female, 10.7 mm)." httpUri="https://zenodo.org/record/1052755/files/figure.png" pageId="17">Fig. 43</figureCitation>
). Labrum transverse, with widely rounded anterior angles and shallowly concave anterior margin, surface with 8 setae in transverse row. Anterior part of head lustrous, flat, slightly uneven, posterior tip relatively wide, surface covered with several large punctures, and with several long pale setae along anterior margin and laterally along antennal insertions. Interantennal space wide, 1.70 times as wide as transverse diameter of antennal socket. Eyes large, interocular space 1.30 times as wide as transverse diameter of eye. Frontal tubercles subpentagonal, transverse, with anterior tips divergent, surface covered with several larger punctures, lustrous. Vertex distinctly impressed behind frontal tubercles, surface covered with distinct punctures and short pale setae, medially with narrow impressed line. Antennae filiform, 0.91 times as long as body, length ratio of antennomeres equals
<date id="FFCE4193FFD4FF9BFE2CFD51FDD3FCA6" box="[392,556,766,791]" pageId="17" value="2013-05-17" valueMax="2013-12-17" valueMin="2013-05-17">13-5-12-17-17</date>
-17-18-17-17-16-19.
</paragraph>
<paragraph id="8BCF6753FFD4FF9BFF63FC8CFD3BFBA2" blockId="17.[151,1437,151,1043]" pageId="17">Pronotum lustrous, transverse, 1.85 times as broad as long, widest in middle, covered with double fine punctation: very dense fine micropunctures and moderately dense larger punctures. Anterior margin slightly concave, lateral margins moderately rounded (in posterior half nearly parallel, in anterior half convergently rounded), posterior margin widely rounded. Anterior margin with indistinct border, posterior margin moderately bordered. Lateral margins with wider channel-like margins. All angles obtusangulate, anterior ones distinctly swollen, all angles with setigerous pore bearing long pale seta, additional four to five setae placed on lateral margins. Scutellum with very widely rounded tip.</paragraph>
<caption id="DF0F37DBFFD4FF9BFF33F817FDEAF85C" httpUri="https://zenodo.org/record/1052755/files/figure.png" pageId="17" targetBox="[175,1376,1111,1931]" targetPageId="17">
<paragraph id="8BCF6753FFD4FF9BFF33F817FDEAF85C" blockId="17.[151,1436,1976,2029]" pageId="17">
<emphasis id="B904BB41FFD4FF9BFF33F817FEACF87F" bold="true" box="[151,339,1976,1999]" pageId="17">FIGURES 4344.</emphasis>
Habitus of
<taxonomicName id="4C701CD0FFD4FF9BFE6DF816FDABF87E" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[457,596,1977,1999]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B904BB41FFD4FF9BFE6DF816FDABF87E" box="[457,596,1977,1999]" italics="true" pageId="17">Coeligetoides</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD4FF9BFDFEF816FD47F87F" box="[602,696,1977,1998]" pageId="17" rank="genus">
<emphasis id="B904BB41FFD4FF9BFDFEF816FD47F87F" bold="true" box="[602,696,1977,1998]" pageId="17">gen. nov.</emphasis>
</taxonomicNameLabel>
43—
<taxonomicName id="4C701CD0FFD4FF9BFD55F815FC87F87E" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[753,888,1977,1999]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="species" species="trifurcatus" status="sp. nov.">
<emphasis id="B904BB41FFD4FF9BFD55F815FC87F87E" box="[753,888,1977,1999]" italics="true" pageId="17">C. trifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD4FF9BFCDAF816FC31F87F" box="[894,974,1977,1998]" pageId="17" rank="species">
<emphasis id="B904BB41FFD4FF9BFCDAF816FC31F87F" bold="true" box="[894,974,1977,1998]" pageId="17">sp. nov.</emphasis>
</taxonomicNameLabel>
(holotype, male, 8.4 mm); 44—
<taxonomicName id="4C701CD0FFD4FF9BFAB1F815FA63F87E" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[1301,1436,1977,1999]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" phylum="Arthropoda" rank="species" species="trifurcatus" status="sp. nov.">
<emphasis id="B904BB41FFD4FF9BFAB1F815FA63F87E" box="[1301,1436,1977,1999]" italics="true" pageId="17">C. trifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD4FF9BFF33F877FF18F85C" box="[151,231,2008,2029]" pageId="17" rank="species">
<emphasis id="B904BB41FFD4FF9BFF33F877FF18F85C" bold="true" box="[151,231,2008,2029]" pageId="17">sp. nov.</emphasis>
</taxonomicNameLabel>
(paratype, female, 10.7 mm).
</paragraph>
</caption>
<caption id="DF0F37DBFFD7FF98FF33F8F7FF48F87A" httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="18" targetBox="[169,1416,212,1844]" targetPageId="18">
<paragraph id="8BCF6753FFD7FF98FF33F8F7FF48F87A" blockId="18.[151,1436,1880,1995]" pageId="18">
<emphasis id="B904BB41FFD7FF98FF33F8F7FEAAF8DF" bold="true" box="[151,341,1880,1903]" pageId="18">FIGURES 4551.</emphasis>
Details of
<taxonomicName id="4C701CD0FFD7FF98FE6DF8F6FD39F8DE" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[457,710,1881,1903]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" phylum="Arthropoda" rank="species" species="trifurcatus" status="sp. nov.">
<emphasis id="B904BB41FFD7FF98FE6DF8F6FD39F8DE" box="[457,710,1881,1903]" italics="true" pageId="18">Coeligetoides trifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD7FF98FD6AF8F6FCDFF8DF" box="[718,800,1881,1902]" pageId="18" rank="species">
<emphasis id="B904BB41FFD7FF98FD6AF8F6FCDFF8DF" bold="true" box="[718,800,1881,1902]" pageId="18">sp. nov.</emphasis>
</taxonomicNameLabel>
45—aedeagus (dorsal, lateral and ventral views); 46—claws; 47—last two abdominal ventrites of male; 48—female last abdominal ventrite; 49—female pygidium and tergite VIII; 50— spermatheca; 51—sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49.
</paragraph>
</caption>
<paragraph id="8BCF6753FFD6FF99FF63FF38FBA0FF46" blockId="19.[151,1437,151,1365]" pageId="19">Elytra long 1.85 times as long as wide and 0.75 times as long as body, almost glabrous, with several short setae visible on lateral and apical slopes, covered with dense confused punctures. Humeral calli well developed. Epipleura wide basally, gradually narrowing posteriorly and disappearing before apex.</paragraph>
<paragraph id="8BCF6753FFD6FF99FF63FEABFB2BFE1C" blockId="19.[151,1437,151,1365]" pageId="19">
Posterior margin of anterior coxal cavities open. Last ventrite short, with two short, wide, V-shaped incisions, median lobe wide with elongate oval impression, penultimate ventrite with small semicircular incision in middle of posterior margin (
<figureCitation id="134B7BD6FFD6FF99FEC6FEE3FE47FED5" box="[354,440,332,357]" captionStart="FIGURES 45 51" captionStartId="18.[151,264,1880,1903]" captionTargetBox="[169,1416,212,1844]" captionTargetId="figure@18.[151,1436,193,1860]" captionTargetPageId="18" captionText="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="19">Fig. 47</figureCitation>
). Protarsomeres I and II narrow, protarsomere I elongate subtriangular, twice as long as wide, length ratio of protarsomeres IIV equals 8-6-6-9. Metatarsomere I long and narrow, length ratio of metatarsomeres IIV equals
<date id="FFCE4193FFD6FF99FE7CFE3BFDB0FE1C" box="[472,591,404,429]" pageId="19" value="2014-06-07" valueMax="2014-10-07" valueMin="2014-06-07">14-7-6-10.</date>
Claws bifid, inner teeth shorter than outer ones (
<figureCitation id="134B7BD6FFD6FF99FBD7FE3BFB38FE1C" box="[1139,1223,404,429]" captionStart="FIGURES 45 51" captionStartId="18.[151,264,1880,1903]" captionTargetBox="[169,1416,212,1844]" captionTargetId="figure@18.[151,1436,193,1860]" captionTargetPageId="18" captionText="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="19">Fig. 46</figureCitation>
).
</paragraph>
<paragraph id="8BCF6753FFD6FF99FF63FE18FD45FDA9" blockId="19.[151,1437,151,1365]" pageId="19">
Aedeagus small, flat, apex with short sharp incision, in lateral view apical part bent downwards. Dorsal process with two branches: ventral, slightly wider, directed obliquely upwards, and dorsal, slightly narrower, widely bent, with apical third bifurcate (
<figureCitation id="134B7BD6FFD6FF99FDFEFE50FD51FDA9" box="[602,686,511,536]" captionStart="FIGURES 45 51" captionStartId="18.[151,264,1880,1903]" captionTargetBox="[169,1416,212,1844]" captionTargetId="figure@18.[151,1436,193,1860]" captionTargetPageId="18" captionText="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="19">Fig. 45</figureCitation>
).
</paragraph>
<paragraph id="8BCF6753FFD6FF99FF63FD8BFC51FCA4" blockId="19.[151,1437,151,1365]" pageId="19">
Female (
<figureCitation id="134B7BD6FFD6FF99FE8CFD8BFE83FD8D" box="[296,380,548,573]" captionStart="FIGURES 43 44" captionStartId="17.[151,264,1976,1999]" captionTargetBox="[175,1376,1111,1931]" captionTargetId="figure@17.[151,1436,1087,1955]" captionTargetPageId="17" captionText="FIGURES 43 44. Habitus of Coeligetoides gen. nov. 43 — C. trifurcatus sp. nov. (holotype, male, 8.4 mm); 44 — C. trifurcatus sp. nov. (paratype, female, 10.7 mm)." httpUri="https://zenodo.org/record/1052755/files/figure.png" pageId="19">Fig. 44</figureCitation>
). Interantennal space wider, 2.20 times as wide as transverse diameter of antennal socket. Eyes smaller, interocular space wider, 1.85 times as wide as transverse diameter of eye. Antennae shorter than in male, 0.65 times as long as body. Last ventrite with entire regularly rounded posterior margin (
<figureCitation id="134B7BD6FFD6FF99FB2FFDC3FB1FFD35" box="[1163,1248,620,645]" captionStart="FIGURES 45 51" captionStartId="18.[151,264,1880,1903]" captionTargetBox="[169,1416,212,1844]" captionTargetId="figure@18.[151,1436,193,1860]" captionTargetPageId="18" captionText="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="19">Fig. 48</figureCitation>
). Pygidium with posterior margin widely concave, tergite VIII with posterior margin widely shallowly concave (
<figureCitation id="134B7BD6FFD6FF99FB7EFD20FACEFD19" box="[1242,1329,655,680]" captionStart="FIGURES 45 51" captionStartId="18.[151,264,1880,1903]" captionTargetBox="[169,1416,212,1844]" captionTargetId="figure@18.[151,1436,193,1860]" captionTargetPageId="18" captionText="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="19">Fig. 49</figureCitation>
). Sternite VIII transversely rhomboid, with V-shaped incision in middle of posterior margin, with short pale setae cummulated along posterior margin except middle part, tignum twice longer than sternite VIII (
<figureCitation id="134B7BD6FFD6FF99FB7FFD78FACEFD41" box="[1243,1329,727,752]" captionStart="FIGURES 45 51" captionStartId="18.[151,264,1880,1903]" captionTargetBox="[169,1416,212,1844]" captionTargetId="figure@18.[151,1436,193,1860]" captionTargetPageId="18" captionText="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="19">Fig. 51</figureCitation>
). Vaginal sclerite missing. Spermatheca with poorly delimited nodulus (
<figureCitation id="134B7BD6FFD6FF99FCE9FD53FC5EFCA4" box="[845,929,764,789]" captionStart="FIGURES 45 51" captionStartId="18.[151,264,1880,1903]" captionTargetBox="[169,1416,212,1844]" captionTargetId="figure@18.[151,1436,193,1860]" captionTargetPageId="18" captionText="FIGURES 45 51. Details of Coeligetoides trifurcatus sp. nov. 45 — aedeagus (dorsal, lateral and ventral views); 46 — claws; 47 — last two abdominal ventrites of male; 48 — female last abdominal ventrite; 49 — female pygidium and tergite VIII; 50 — spermatheca; 51 — sternite VIII and tignum. Scale bars: 0.25 mm for Fig. 46, 0.5 mm for Figs 45, 50 and 51, 1 mm for Figs 47 49." httpUri="https://zenodo.org/record/1052757/files/figure.png" pageId="19">Fig. 50</figureCitation>
).
</paragraph>
<paragraph id="8BCF6753FFD6FF99FF63FCB0FE33FCEC" blockId="19.[151,1437,151,1365]" pageId="19">
<emphasis id="B904BB41FFD6FF99FF63FCB0FEB2FC89" bold="true" box="[199,333,799,824]" pageId="19">Variability.</emphasis>
Outer margin of tibiae usually darkened, tarsi rarely infuscate, rarely legs completely yellow or tibiae completely infuscate.
</paragraph>
</subSubSection>
<subSubSection id="C36A34D8FFD6FF99FF63FCC8FE14FB75" pageId="19" type="diagnosis">
<paragraph id="8BCF6753FFD6FF99FF63FCC8FE14FB75" blockId="19.[151,1437,151,1365]" pageId="19">
<emphasis id="B904BB41FFD6FF99FF63FCC8FE2DFC31" bold="true" box="[199,466,871,896]" pageId="19">Differential diagnosis.</emphasis>
<taxonomicName id="4C701CD0FFD6FF99FE7AFCC7FD06FC31" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[478,761,872,896]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="trifurcatus" status="sp. nov.">
<emphasis id="B904BB41FFD6FF99FE7AFCC7FD06FC31" box="[478,761,872,896]" italics="true" pageId="19">Coeligetoides trifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD6FF99FCA1FCC8FC9CFC31" box="[773,867,871,896]" pageId="19" rank="species">
<emphasis id="B904BB41FFD6FF99FCA1FCC8FC9CFC31" bold="true" box="[773,867,871,896]" pageId="19">sp. nov.</emphasis>
</taxonomicNameLabel>
is habitually similar to dorsally pale
<taxonomicName id="4C701CD0FFD6FF99FA82FCC7FA63FC31" authorityName="Bezděk" authorityYear="2016" box="[1318,1436,872,896]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FA82FCC7FA63FC31" box="[1318,1436,872,896]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
species (
<taxonomicName id="4C701CD0FFD6FF99FF5FFC23FE81FC15" authorityName="Jacoby" authorityYear="1895" box="[251,382,908,932]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="unicolor">
<emphasis id="B904BB41FFD6FF99FF5FFC23FE81FC15" box="[251,382,908,932]" italics="true" pageId="19">C. unicolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C701CD0FFD6FF99FE18FC23FDC4FC15" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[444,571,908,932]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="howardi" status="sp. nov.">
<emphasis id="B904BB41FFD6FF99FE18FC23FDC4FC15" box="[444,571,908,932]" italics="true" pageId="19">C. howardi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD6FF99FDE2FC23FD5BFC14" box="[582,676,908,933]" pageId="19" rank="species">
<emphasis id="B904BB41FFD6FF99FDE2FC23FD5BFC14" bold="true" box="[582,676,908,933]" pageId="19">sp. nov.</emphasis>
</taxonomicNameLabel>
). Most of specimens of
<taxonomicName id="4C701CD0FFD6FF99FC61FC23FB21FC15" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[965,1246,908,932]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="trifurcatus" status="sp. nov.">
<emphasis id="B904BB41FFD6FF99FC61FC23FB21FC15" box="[965,1246,908,932]" italics="true" pageId="19">Coeligetoides trifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD6FF99FB4DFC23FAB9FC14" box="[1257,1350,908,933]" pageId="19" rank="species">
<emphasis id="B904BB41FFD6FF99FB4DFC23FAB9FC14" bold="true" box="[1257,1350,908,933]" pageId="19">sp. nov.</emphasis>
</taxonomicNameLabel>
can be distinguished by the darkened tibiae which are always pale in both similar
<taxonomicName id="4C701CD0FFD6FF99FB9DFC1FFB50FC79" authorityName="Bezděk" authorityYear="2016" box="[1081,1199,944,968]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FB9DFC1FFB50FC79" box="[1081,1199,944,968]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
species. Additional differencies duplicate the generic characters:
<taxonomicName id="4C701CD0FFD6FF99FD06FC7BFC44FC5D" authority="Bezděk, 2016" authorityName="Bezděk" authorityYear="2016" box="[674,955,980,1004]" class="Insecta" family="Chrysomelidae" genus="Coeligetoides" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="species" species="trifurcatus" status="sp. nov.">
<emphasis id="B904BB41FFD6FF99FD06FC7BFC44FC5D" box="[674,955,980,1004]" italics="true" pageId="19">Coeligetoides trifurcatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A237063AFFD6FF99FC62FC7BFBDCFC5C" box="[966,1059,980,1005]" pageId="19" rank="species">
<emphasis id="B904BB41FFD6FF99FC62FC7BFBDCFC5C" bold="true" box="[966,1059,980,1005]" pageId="19">sp. nov.</emphasis>
</taxonomicNameLabel>
has posterior margin of anterior coxal cavities open (closed in
<taxonomicName id="4C701CD0FFD6FF99FE43FC57FDA2FBA1" authorityName="Bezděk" authorityYear="2016" box="[487,605,1016,1040]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FE43FC57FDA2FBA1" box="[487,605,1016,1040]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
), aedeagus is wide and flat with dorsal process thin and trifurcate, anterior part of head is flat with wide posterior tip (elevated and convex with posterior tip narrower in
<taxonomicName id="4C701CD0FFD6FF99FB69FBB3FABCFB85" authorityName="Bezděk" authorityYear="2016" box="[1229,1347,1052,1076]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FB69FBB3FABCFB85" box="[1229,1347,1052,1076]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
), claws bifid (claws appendiculate in
<taxonomicName id="4C701CD0FFD6FF99FE52FBEFFD93FBE9" authorityName="Bezděk" authorityYear="2016" box="[502,620,1088,1112]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FE52FBEFFD93FBE9" box="[502,620,1088,1112]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
), labrum with shallowly concave anterior margin (rounded or nearly straight in
<taxonomicName id="4C701CD0FFD6FF99FEBAFBCBFE6BFBCD" authorityName="Bezděk" authorityYear="2016" box="[286,404,1124,1148]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FEBAFBCBFE6BFBCD" box="[286,404,1124,1148]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
), female last ventrite with entire and regularly rounded posterior margin (incised in
<taxonomicName id="4C701CD0FFD6FF99FF33FB27FEF2FB11" authorityName="Bezděk" authorityYear="2016" box="[151,269,1160,1184]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FF33FB27FEF2FB11" box="[151,269,1160,1184]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
), vaginal sclerite absent (present in
<taxonomicName id="4C701CD0FFD6FF99FD15FB27FCD8FB11" authorityName="Bezděk" authorityYear="2016" box="[689,807,1160,1184]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FD15FB27FCD8FB11" box="[689,807,1160,1184]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
), spermatheca with poorly delimited nodulus (nodulus well developed in
<taxonomicName id="4C701CD0FFD6FF99FEC1FB03FE24FB75" authorityName="Bezděk" authorityYear="2016" box="[357,475,1196,1220]" class="Insecta" family="Chrysomelidae" genus="Coeligetes" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" phylum="Arthropoda" rank="genus">
<emphasis id="B904BB41FFD6FF99FEC1FB03FE24FB75" box="[357,475,1196,1220]" italics="true" pageId="19">Coeligetes</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C36A34D8FFD6FF99FF63FB60FEFDFABC" pageId="19" type="distribution">
<paragraph id="8BCF6753FFD6FF99FF63FB60FEFDFABC" blockId="19.[151,1437,151,1365]" pageId="19">
<emphasis id="B904BB41FFD6FF99FF63FB60FE9FFB59" bold="true" box="[199,352,1231,1256]" pageId="19">Distribution.</emphasis>
<collectingCountry id="F36727C3FFD6FF99FEC8FB60FE2CFB59" box="[364,467,1231,1256]" name="Malaysia" pageId="19">Malaysia</collectingCountry>
(Peninsular
<collectingCountry id="F36727C3FFD6FF99FDCEFB60FD2BFB59" box="[618,724,1231,1256]" name="Malaysia" pageId="19">Malaysia</collectingCountry>
,
<collectingRegion id="49B4A9B1FFD6FF99FD40FB60FCD4FB59" box="[740,811,1231,1256]" country="Malaysia" name="Sabah" pageId="19">Sabah</collectingRegion>
,
<collectingRegion id="49B4A9B1FFD6FF99FC9FFB60FC5FFB59" box="[827,928,1231,1256]" country="Malaysia" name="Sarawak" pageId="19">
<collectingCountry id="F36727C3FFD6FF99FC9FFB60FC5FFB59" box="[827,928,1231,1256]" name="Malaysia" pageId="19">Sarawak</collectingCountry>
</collectingRegion>
),
<collectingCountry id="F36727C3FFD6FF99FC13FB60FBF9FB59" box="[951,1030,1231,1256]" name="Brunei" pageId="19">Brunei</collectingCountry>
,
<collectingCountry id="F36727C3FFD6FF99FBB3FB60FB7BFB59" box="[1047,1156,1231,1256]" name="Indonesia" pageId="19">Indonesia</collectingCountry>
(
<collectingRegion id="49B4A9B1FFD6FF99FB33FB7FFB06FB59" box="[1175,1273,1232,1256]" country="Indonesia" name="Sumatera Utara" pageId="19">Sumatra</collectingRegion>
,
<collectingRegion id="49B4A9B1FFD6FF99FAADFB60FA6FFB59" box="[1289,1424,1231,1256]" country="Indonesia" name="Kalimantan Utara" pageId="19">Kalimantan</collectingRegion>
),
<collectingCountry id="F36727C3FFD6FF99FF33FB5BFF00FABC" box="[151,255,1268,1293]" name="Thailand" pageId="19">Thailand</collectingCountry>
.
</paragraph>
</subSubSection>
<subSubSection id="C36A34D8FFD6FF99FF63FAB8FEBEFAE4" pageId="19" type="etymology">
<paragraph id="8BCF6753FFD6FF99FF63FAB8FEBEFAE4" blockId="19.[151,1437,151,1365]" pageId="19">
<emphasis id="B904BB41FFD6FF99FF63FAB8FEB1FA81" bold="true" box="[199,334,1303,1328]" pageId="19">Etymology.</emphasis>
Trifurcatus (= trident) refers to the unusual shape of the dorsal process of the aedeagus, bearing three branches.
</paragraph>
</subSubSection>
</treatment>
</document>