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<document id="1730BF6903E4A263542F54D8AE3E9813" ID-DOI="10.5281/zenodo.204914" ID-GBIF-Dataset="035006f1-9010-4f0b-a5fb-83fed142d245" ID-ISSN="1175-5326" ID-Zenodo-Dep="204914" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460221844229" checkinUser="plazi" docAuthor="Ciccotto, Patrick J., Konings, Adrianus &amp; Stauffer, Jay R." docDate="2011" docId="03BB942A0D7E836B9DEFF9ABFD15FE31" docLanguage="en" docName="zt02738p025.pdf" docOrigin="Zootaxa 2738" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Metriaclima xanthos Ciccotto, Konings &amp; Stauffer, 2011, new species" docType="treatment" docVersion="10" lastPageNumber="14" masterDocId="FF82EC520D7483669D78FF83FFCBFFFE" masterDocTitle="Descriptions of five new species in the genus Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa" masterLastPageNumber="25" masterPageNumber="1" pageNumber="11" updateTime="1698237790047" updateUser="plazi">
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<mods:title id="40D777A3D0FD402AD0442EC885987C38">Descriptions of five new species in the genus Metriaclima (Teleostei: Cichlidae) from Lake Malaŵi, Africa</mods:title>
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<mods:namePart id="1614579B8747CC8E917828A5C402E5F1">Ciccotto, Patrick J.</mods:namePart>
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<mods:namePart id="3880131E575A173F2F6BF8C2B2C75E93">Konings, Adrianus</mods:namePart>
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<mods:namePart id="7623E16E3EA4127345984E195DF9E873">Stauffer, Jay R.</mods:namePart>
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<mods:date id="EA7DED42F4840D62A4F0214A040DE0A6">2011</mods:date>
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<treatment id="03BB942A0D7E836B9DEFF9ABFD15FE31" ID-DOI="http://doi.org/10.5281/zenodo.6192254" ID-GBIF-Taxon="119385413" ID-Zenodo-Dep="6192254" LSID="urn:lsid:plazi:treatment:03BB942A0D7E836B9DEFF9ABFD15FE31" httpUri="http://treatment.plazi.org/id/03BB942A0D7E836B9DEFF9ABFD15FE31" lastPageId="13" lastPageNumber="14" pageId="10" pageNumber="11">
<subSubSection id="C30876B70D7E836C9DEFF9ABFEA4F99D" pageId="10" pageNumber="11" type="nomenclature">
<paragraph id="8BAD253C0D7E836C9DEFF9ABFDFAF9BC" blockId="10.[151,561,1576,1635]" box="[151,561,1576,1602]" pageId="10" pageNumber="11">
<heading id="D0E592500D7E836C9DEFF9ABFDFAF9BC" bold="true" box="[151,561,1576,1602]" fontSize="11" level="1" pageId="10" pageNumber="11" reason="1">
<emphasis id="B966F92E0D7E836C9DEFF9ABFDFAF9BC" bold="true" box="[151,561,1576,1602]" pageId="10" pageNumber="11">
<taxonomicName id="4C125EBF0D7E836C9DEFF9ABFE59F9BC" ID-CoL="42HWY" box="[151,402,1576,1602]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="xanthos" status="sp. nov.">
<emphasis id="B966F92E0D7E836C9DEFF9ABFE59F9BC" bold="true" box="[151,402,1576,1602]" italics="true" pageId="10" pageNumber="11">Metriaclima xanthos</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A25544550D7E836C9CD9F9ABFDFAF9BC" box="[417,561,1576,1602]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
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<paragraph id="8BAD253C0D7E836C9DEFF9C8FEA4F99D" blockId="10.[151,561,1576,1635]" box="[151,367,1611,1635]" pageId="10" pageNumber="11">
<figureCitation id="132939B90D7E836C9DEFF9C8FF26F99D" box="[151,237,1611,1635]" captionStart="FIGURE 5. A" captionStartId="11.[151,250,1381,1404]" captionTargetBox="[276,1311,553,1360]" captionTargetId="figure@11.[276,1312,525,1360]" captionTargetPageId="11" captionText="FIGURE 5. A. Metriaclima xanthos, holotype, PSU 4494, 44.3 mm SL; Mozambique: Lake MalaŴi: Lumbaulo. B. M. xanthos, male in habitat at Lumbaulo, Mozambique; approx. 70 mm SL; not preserved. C. M. xanthos, female in habitat at Lumbaulo, Mozambique; approx. 50 mm SL; not preserved." httpUri="https://zenodo.org/record/204919/files/figure.png" pageId="10" pageNumber="11">Fig. 5A</figureCitation>
C;
<tableCitation id="C69010870D7E836C9C64F9C8FEA4F99D" box="[284,367,1611,1635]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1383,283,1959]" captionTargetPageId="12" captionText="TABLE 4. Morphometric and meristic values of Metriaclima xanthos from Lumbaulo. Mean and range include holotype." httpUri="http://table.plazi.org/id/DF6D75B40D78836A9DEFFF1BFAACFF50" pageId="10" pageNumber="11" tableUuid="DF6D75B40D78836A9DEFFF1BFAACFF50">Table 4</tableCitation>
</paragraph>
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<subSubSection id="C30876B70D7E836C9DEFF910FD1EF957" box="[151,725,1683,1705]" pageId="10" pageNumber="11" type="description">
<paragraph id="8BAD253C0D7E836C9DEFF910FD1EF957" blockId="10.[151,725,1683,1705]" box="[151,725,1683,1705]" pageId="10" pageNumber="11">
<treatmentCitationGroup id="AB0202120D7E836C9DEFF910FD1EF957" box="[151,725,1683,1705]" pageId="10" pageNumber="11">
<taxonomicName id="4C125EBF0D7E836C9DEFF910FEDFF957" box="[151,276,1683,1705]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D7E836C9DEFF910FEDFF957" box="[151,276,1683,1705]" italics="true" pageId="10" pageNumber="11">Metriaclima</emphasis>
</taxonomicName>
sp. &quot;
<taxonomicName id="4C125EBF0D7E836C9C3EF917FE42F957" box="[326,393,1684,1705]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="aurora">aurora</taxonomicName>
lumbaulo&quot;:
<treatmentCitation id="0AB3032D0D7E836C9F7EF910FD1AF957" author="Konings" box="[518,721,1683,1705]" page="155" pageId="10" pageNumber="11" year="2001">
<bibRefCitation id="EF8358CD0D7E836C9F7EF910FD54F957" author="Konings" box="[518,671,1683,1705]" pageId="10" pageNumber="25" refString="Konings, A. (2001) Mala w i cichlids in their natural habitat, Third edition. Cichlid Press, El Paso, Texas, 352 pp." type="book" year="2001">Konings, 2001</bibRefCitation>
: 155
</treatmentCitation>
.
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</paragraph>
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<subSubSection id="C30876B70D7E836C9DEFF955FC63F8C8" pageId="10" pageNumber="11" type="materials_examined">
<paragraph id="8BAD253C0D7E836C9DEFF955FD8DF8EF" blockId="10.[151,1436,1749,2026]" pageId="10" pageNumber="11">
<emphasis id="B966F92E0D7E836C9DEFF955FEC0F910" bold="true" box="[151,267,1750,1774]" pageId="10" pageNumber="11">
<typeStatus id="54A99B9E0D7E836C9DEFF955FECDF910" box="[151,262,1750,1774]" pageId="10" pageNumber="11" type="holotype">Holotype</typeStatus>
.
</emphasis>
<collectionCode id="ED03BDF90D7E836C9C6CF955FE83F910" box="[276,328,1750,1774]" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="10" pageNumber="11">PSU</collectionCode>
4494,
<quantity id="4CEA88D90D7E836C9CE3F955FDC9F910" box="[411,514,1750,1774]" metricMagnitude="-2" metricUnit="m" metricValue="4.43" pageId="10" pageNumber="11" unit="mm" value="44.3">44.3 mm</quantity>
SL;
<collectingCountry id="F30565AC0D7E836C9F46F955FD1DF910" box="[574,726,1750,1774]" name="Mozambique" pageId="10" pageNumber="11">Mozambique</collectingCountry>
: Lake
<collectingCountry id="F30565AC0D7E836C9E5DF956FCB4F910" box="[805,895,1749,1774]" name="Malawi" pageId="10" pageNumber="11">Malaŵi</collectingCountry>
: Lumbaulo,
<geoCoordinate id="EE2643FB0D7E836C9968F955FB66F910" box="[1040,1197,1750,1774]" direction="south" orientation="latitude" pageId="10" pageNumber="11" precision="1" value="-11.9004">11° 54.024' S</geoCoordinate>
,
<geoCoordinate id="EE2643FB0D7E836C99C4F955FA96F910" box="[1212,1373,1750,1774]" direction="east" orientation="longitude" pageId="10" pageNumber="11" precision="1" value="34.905117">34° 54.307' E</geoCoordinate>
; A.F. Konings &amp; J.R. Stauffer,
<date id="FFAC03FC0D7E836C9CCAF97AFD89F8EF" box="[434,578,1785,1809]" pageId="10" pageNumber="11" value="2002-02-19">19 Feb 2002</date>
.
</paragraph>
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<emphasis id="B966F92E0D7E836C9DBFF89DFE8DF8C8" bold="true" box="[199,326,1822,1846]" pageId="10" pageNumber="11">
<typeStatus id="54A99B9E0D7E836C9DBFF89DFE8AF8C8" box="[199,321,1822,1846]" pageId="10" pageNumber="11" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="ED03BDF90D7E836C9C34F89DFEB4F8C8" box="[332,383,1822,1846]" httpUri="http://grbio.org/cool/5mg9-yyc1" name="Portland State University, Vertebrate Biology Museum" pageId="10" pageNumber="11">PSU</collectionCode>
4495, 4;
<quantity id="4CEA88D90D7E836C9C92F89DFD47F8C8" box="[490,652,1822,1846]" metricMagnitude="-2" metricUnit="m" metricValue="4.695" metricValueMax="4.93" metricValueMin="4.46" pageId="10" pageNumber="11" unit="mm" value="46.95" valueMax="49.3" valueMin="44.6">44.649.3 mm</quantity>
SL; data as for
<typeStatus id="54A99B9E0D7E836C9E46F89DFC6FF8C8" box="[830,932,1822,1846]" pageId="10" pageNumber="11" type="holotype">holotype</typeStatus>
.
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7E836D9DBFF8C2FEA9FEE2" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11" type="diagnosis">
<paragraph id="8BAD253C0D7E836D9DBFF8C2FEA9FEE2" blockId="10.[151,1436,1749,2026]" lastBlockId="11.[151,1437,151,500]" lastPageId="11" lastPageNumber="12" pageId="10" pageNumber="11">
<emphasis id="B966F92E0D7E836C9DBFF8C2FE8AF8A4" bold="true" box="[199,321,1857,1882]" pageId="10" pageNumber="11">Diagnosis.</emphasis>
A moderately-sloped head and bicuspid teeth in the outer row of the jaws place this species in
<taxonomicName id="4C125EBF0D7E836C9DEFF8E5FEEAF880" box="[151,289,1894,1918]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D7E836C9DEFF8E5FEEAF880" box="[151,289,1894,1918]" italics="true" pageId="10" pageNumber="11">Metriaclima</emphasis>
</taxonomicName>
. The brown and blue/gray alternating lateral bars in conjunction with the yellow dorsal and dark anal fin of breeding males distinguish
<taxonomicName id="4C125EBF0D7E836C9F69F808FD46F85C" box="[529,653,1930,1954]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7E836C9F69F808FD46F85C" box="[529,653,1930,1954]" italics="true" pageId="10" pageNumber="11">M. xanthos</emphasis>
</taxonomicName>
from males of all other members of
<taxonomicName id="4C125EBF0D7E836C9954F809FB7DF85C" box="[1068,1206,1930,1954]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="genus">
<emphasis id="B966F92E0D7E836C9954F809FB7DF85C" box="[1068,1206,1930,1954]" italics="true" pageId="10" pageNumber="11">Metriaclima</emphasis>
</taxonomicName>
. Female
<taxonomicName id="4C125EBF0D7E836C9867F808FA57F85C" box="[1311,1436,1930,1954]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7E836C9867F808FA57F85C" box="[1311,1436,1930,1954]" italics="true" pageId="10" pageNumber="11">M. xanthos</emphasis>
</taxonomicName>
are yellow/brown, usually more yellow than females of any other member of the
<taxonomicName id="4C125EBF0D7E836C9956F82DFBB6F838" box="[1070,1149,1966,1990]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="23" phylum="Chordata" rank="species" species="aurora">Aurora</taxonomicName>
group, but not all female
<taxonomicName id="4C125EBF0D7E836C9DEFF850FEDFF814" box="[151,276,2002,2026]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7E836C9DEFF850FEDFF814" box="[151,276,2002,2026]" italics="true" pageId="10" pageNumber="11">M. xanthos</emphasis>
</taxonomicName>
can be distinguished according to the color pattern.
<taxonomicName id="4C125EBF0D7E836C9E10F851FB9AF814" box="[872,1105,2002,2026]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="10" pageNumber="11" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7E836C9E10F851FB9AF814" box="[872,1105,2002,2026]" italics="true" pageId="10" pageNumber="11">Metriaclima xanthos</emphasis>
</taxonomicName>
is further distinguished from
<taxonomicName id="4C125EBF0D7F836D9DEFFF1AFEC1FF4E" box="[151,266,153,176]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="aurora">
<emphasis id="B966F92E0D7F836D9DEFFF1AFEC1FF4E" box="[151,266,153,176]" italics="true" pageId="11" pageNumber="12">M. aurora</emphasis>
</taxonomicName>
males from the Mbweca and Tumbi Point populations that are similar but lack dark pigment in the anal fin, and from the neighboring
<taxonomicName id="4C125EBF0D7F836D9C9FFF3EFDA8FF2A" box="[487,611,188,212]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7F836D9C9FFF3EFDA8FF2A" box="[487,611,188,212]" italics="true" pageId="11" pageNumber="12">M. glaucos</emphasis>
</taxonomicName>
by the lower jaw length, which is longer in
<taxonomicName id="4C125EBF0D7F836D9937FF3EFB00FF2A" box="[1103,1227,188,212]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7F836D9937FF3EFB00FF2A" box="[1103,1227,188,212]" italics="true" pageId="11" pageNumber="12">M. xanthos</emphasis>
</taxonomicName>
(mean 35.4, range 34.636.7 % HL) than in
<taxonomicName id="4C125EBF0D7F836D9CBCFF62FDF1FF06" box="[452,570,225,248]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="aurora">
<emphasis id="B966F92E0D7F836D9CBCFF62FDF1FF06" box="[452,570,225,248]" italics="true" pageId="11" pageNumber="12">M. aurora</emphasis>
</taxonomicName>
(mean 31.2, range 28.433.5 % HL) and in
<taxonomicName id="4C125EBF0D7F836D993BFF62FB09FF06" box="[1091,1218,224,248]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7F836D993BFF62FB09FF06" box="[1091,1218,224,248]" italics="true" pageId="11" pageNumber="12">M. glaucos</emphasis>
</taxonomicName>
(mean 30.8, range 28.832.8 % HL).
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7F836D9DBFFEA4FA40F90A" pageId="11" pageNumber="12" type="description">
<paragraph id="8BAD253C0D7F836D9DBFFEA4FA85FE0A" blockId="11.[151,1437,151,500]" pageId="11" pageNumber="12">
<emphasis id="B966F92E0D7F836D9DBFFEA4FE92FEBE" bold="true" box="[199,345,295,320]" pageId="11" pageNumber="12">Description.</emphasis>
Morphometric and meristic data in
<tableCitation id="C69010870D7F836D9E78FEA4FC91FEC1" box="[768,858,295,319]" captionStart="TABLE 4" captionStartId="12.[151,235,152,173]" captionTargetBox="[151,1383,283,1959]" captionTargetPageId="12" captionText="TABLE 4. Morphometric and meristic values of Metriaclima xanthos from Lumbaulo. Mean and range include holotype." httpUri="http://table.plazi.org/id/DF6D75B40D78836A9DEFFF1BFAACFF50" pageId="11" pageNumber="12" tableUuid="DF6D75B40D78836A9DEFFF1BFAACFF50">Table 4</tableCitation>
. Dorsal snout profile slightly concave to straight; mouth cleft at slightly downward angle; jaws isognathous. Teeth on dentary in 2 or 3 rows, on premaxilla in 2 or 3 rows; outer row teeth typically bicuspid anteriorly and unicuspid posteriorly, inner rows tricuspid, innermost row unicuspid; lower pharyngeal jaw with numerous slender teeth with teeth in posterior row slightly larger. Portion of upper dental arcade normally visible in closed mouth. Tips of teeth in premaxilla and dentary in V-shaped line with anteriormost in upper and lower jaw furthest apart and not touching in closed mouth. Lateral scales ctenoid.
</paragraph>
<caption id="DF6D75B40D7F836D9DEFFAE6FD01FA47" httpUri="https://zenodo.org/record/204919/files/figure.png" pageId="11" pageNumber="12" targetBox="[276,1311,553,1360]" targetPageId="11">
<paragraph id="8BAD253C0D7F836D9DEFFAE6FD01FA47" blockId="11.[151,1436,1380,1465]" pageId="11" pageNumber="12">
<emphasis id="B966F92E0D7F836D9DEFFAE6FEFEFA85" bold="true" box="[151,309,1381,1404]" pageId="11" pageNumber="12">FIGURE 5. A.</emphasis>
<taxonomicName id="4C125EBF0D7F836D9C44FAE5FDDBFA82" box="[316,528,1382,1404]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7F836D9C44FAE5FDDBFA82" box="[316,528,1382,1404]" italics="true" pageId="11" pageNumber="12">Metriaclima xanthos</emphasis>
</taxonomicName>
, holotype, PSU 4494, 44.3 mm SL; Mozambique: Lake MalaŴi: Lumbaulo.
<emphasis id="B966F92E0D7F836D9854FAE5FA88FA85" bold="true" box="[1324,1347,1382,1403]" pageId="11" pageNumber="12">B.</emphasis>
<taxonomicName id="4C125EBF0D7F836D9834FAE4FF0AFA64" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7F836D9834FAE4FF0AFA64" italics="true" pageId="11" pageNumber="12">M. xanthos</emphasis>
</taxonomicName>
, male in habitat at Lumbaulo, Mozambique; approx. 70 mm SL; not preserved.
<emphasis id="B966F92E0D7F836D9E80FA07FBDBFA65" bold="true" box="[1016,1040,1412,1435]" pageId="11" pageNumber="12">C.</emphasis>
<taxonomicName id="4C125EBF0D7F836D9960FA06FB41FA64" box="[1048,1162,1412,1434]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7F836D9960FA06FB41FA64" box="[1048,1162,1412,1434]" italics="true" pageId="11" pageNumber="12">M. xanthos</emphasis>
</taxonomicName>
, female in habitat at Lumbaulo, Mozambique; approx. 50 mm SL; not preserved.
</paragraph>
</caption>
<paragraph id="8BAD253C0D7F836D9DBFFA63FC54F976" blockId="11.[151,1437,1504,2032]" pageId="11" pageNumber="12">Lateral body coloration of breeding male blue/gray with 6 or 7 brown bars below dorsal fin, yellow ventrally, yellow caudal peduncle with flecks of blue. Head gray with 2 blue/gray interorbital bars; ventral third of operculum yellow with gray opercular spot; gular region yellow. Dorsal fin yellow with blue lappets. Caudal-fin rays yellow with blue membranes. Anal fin dark gray/brown with light blue distal edge and 4 orange ocelli. Leading edge of pelvic fin white, first ray brown, remainder yellow. Pectoral fin clear.</paragraph>
<paragraph id="8BAD253C0D7F836D9DBFF917FA40F90A" blockId="11.[151,1437,1504,2032]" pageId="11" pageNumber="12">Female yellow/brown laterally with 5 faint brown bars, yellow breast, and white belly. Head orange/brown; gular region yellow; gray/black opercular spot. Dorsal fin yellow/brown with orange lappets. Caudal-fin rays orange with brown membrane. Anal fin orange with 14 small yellow ocelli. Pelvic fin orange. Pectoral fin clear.</paragraph>
</subSubSection>
<subSubSection id="C30876B70D7F836B9DBFF882FD5FFEC1" lastPageId="13" lastPageNumber="14" pageId="11" pageNumber="12" type="discussion">
<paragraph id="8BAD253C0D7F836A9DBFF882FC57FF21" blockId="11.[151,1437,1504,2032]" lastBlockId="12.[151,1383,152,264]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
<emphasis id="B966F92E0D7F836D9DBFF882FEF0F8E7" bold="true" box="[199,315,1793,1817]" pageId="11" pageNumber="12">Remarks.</emphasis>
We compared the minimum polygon clusters formed by plotting the second sheared principal component of the morphometric data against the first principal component of the meristic data for
<taxonomicName id="4C125EBF0D7F836D99CEF8A6FA5EF8C3" box="[1206,1429,1829,1853]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="aurora">
<emphasis id="B966F92E0D7F836D99CEF8A6FA5EF8C3" box="[1206,1429,1829,1853]" italics="true" pageId="11" pageNumber="12">Metriaclima aurora</emphasis>
</taxonomicName>
,
<taxonomicName id="4C125EBF0D7F836D9DEFF8CAFEDFF89E" box="[151,276,1864,1888]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D7F836D9DEFF8CAFEDFF89E" box="[151,276,1864,1888]" italics="true" pageId="11" pageNumber="12">M. glaucos</emphasis>
</taxonomicName>
, and
<taxonomicName id="4C125EBF0D7F836D9C2CF8CAFE19F89E" box="[340,466,1864,1888]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="11" pageNumber="12" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D7F836D9C2CF8CAFE19F89E" box="[340,466,1864,1888]" italics="true" pageId="11" pageNumber="12">M. xanthos</emphasis>
</taxonomicName>
(
<figureCitation id="132939B90D7F836D9C9AF8CBFDE3F89E" box="[482,552,1864,1888]" captionStart="FIGURE 6" captionStartId="13.[151,250,1629,1652]" captionTargetBox="[204,1383,534,1605]" captionTargetId="figure@13.[189,1398,521,1610]" captionTargetPageId="13" captionText="FIGURE 6. Plot of the first principal component of the meristic data (PC 1) and the sheared second principal component of the morphometric data (PC 2) of Metriaclima aurora (●, Likoma Island; x, Mara Point; , Mbweca; +, Tumbi Point; Ο, Nkolongwe), M. xanthos (˔), and M. glaucos (□)." httpUri="https://zenodo.org/record/204920/files/figure.png" pageId="11" pageNumber="12">Fig. 6</figureCitation>
). Size accounted for 90.9% of the observed variance and the second principal component accounted for 2.4%. Variables with the highest loadings on the sheared second principal components were horizontal eye diameter (0.59), vertical eye diameter (0.55), and snout length (-0.28). The first principal component of the meristic data accounted for 14.3% of the total variance. Variables with the highest loadings on the first principal component were tooth rows on lower jaw (0.34), tooth rows on upper jaw (0.32), and dorsal spines
<taxonomicName id="4C125EBF0D78836A9FB2FF4AFC57FF21" box="[714,924,201,223]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D78836A9FB2FF4AFC57FF21" box="[714,924,201,223]" italics="true" pageId="12" pageNumber="13">Metriaclima xanthos</emphasis>
</taxonomicName>
</paragraph>
<caption id="DF6D75B40D78836A9DEFFF1BFAACFF50" ID-Table-UUID="DF6D75B40D78836A9DEFFF1BFAACFF50" box="[151,1383,152,174]" httpUri="http://table.plazi.org/id/DF6D75B40D78836A9DEFFF1BFAACFF50" pageId="12" pageNumber="13" targetBox="[151,1383,283,1959]" targetIsTable="true" targetPageId="12">
<paragraph id="8BAD253C0D78836A9DEFFF1BFAACFF50" blockId="12.[151,1383,152,264]" box="[151,1383,152,174]" pageId="12" pageNumber="13">
<emphasis id="B966F92E0D78836A9DEFFF1BFECFFF53" bold="true" box="[151,260,152,173]" pageId="12" pageNumber="13">TABLE 4.</emphasis>
Morphometric and meristic values of
<taxonomicName id="4C125EBF0D78836A9FF0FF1BFC90FF50" box="[648,859,152,174]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="12" pageNumber="13" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D78836A9FF0FF1BFC90FF50" box="[648,859,152,174]" italics="true" pageId="12" pageNumber="13">Metriaclima xanthos</emphasis>
</taxonomicName>
from Lumbaulo. Mean and range include holotype.
</paragraph>
</caption>
<paragraph id="8BAD253C0D78836A9FB2FF71FD39FEF6" blockId="12.[151,1383,152,264]" box="[714,754,242,264]" pageId="12" pageNumber="13">n=5</paragraph>
<paragraph id="8BAD253C0D78836A9DE7FEC6FB64F859" pageId="12" pageNumber="13">
<table id="F912D79C0D787C999DEFFE98FAACF859" box="[151,1383,283,1959]" gridcols="4" gridrows="9" pageId="12" pageNumber="13">
<tr id="3522277E0D787C999DEFFE98FAACFE7A" box="[151,1383,283,388]" gridrow="0" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFFE98FDC2FE7A" box="[151,521,283,388]" gridcol="0" gridrow="0" pageId="12" pageNumber="13">Standard length, mm Head length, mm</th>
<th id="76F34E020D787C999FB2FE98FCE8FE7A" box="[714,803,283,388]" gridcol="1" gridrow="0" pageId="12" pageNumber="13">holotype 44.3 13.9</th>
<th id="76F34E020D787C999EBEFE98FBCEFE7A" box="[966,1029,283,388]" gridcol="2" gridrow="0" pageId="12" pageNumber="13">mean 45.7 14.7</th>
<th id="76F34E020D787C9999F1FE98FAACFE7A" box="[1161,1383,283,388]" gridcol="3" gridrow="0" pageId="12" pageNumber="13">range 44.349.3 13.915.6</th>
</tr>
<tr id="3522277E0D787C999DEFFE1BFAACFD82" box="[151,1383,408,636]" gridrow="1" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFFE1BFDC2FD82" box="[151,521,408,636]" gridcol="0" gridrow="1" pageId="12" pageNumber="13">Percentage of standard length Head length Snout to dorsal Snout to pelvic Greatest body depth Caudal peduncle length</th>
<td id="76F34E020D787C999FB2FE1BFCE8FD82" box="[714,803,408,636]" gridcol="1" gridrow="1" pageId="12" pageNumber="13">31.4 35.4 36.4 33.9 13.2</td>
<td id="76F34E020D787C999EBEFE1BFBCEFD82" box="[966,1029,408,636]" gridcol="2" gridrow="1" pageId="12" pageNumber="13">32.2 34.7 36.9 33.0 14.2</td>
<td id="76F34E020D787C9999F1FE1BFAACFD82" box="[1161,1383,408,636]" gridcol="3" gridrow="1" pageId="12" pageNumber="13">31.432.8 32.436.5 36.137.4 30.835.1 12.815.3</td>
</tr>
<tr id="3522277E0D787C999DEFFD13FAACFC8A" box="[151,1383,656,884]" gridrow="2" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFFD13FDC2FC8A" box="[151,521,656,884]" gridcol="0" gridrow="2" pageId="12" pageNumber="13">Least caudal peduncle depth Dorsal-fin base length Anterior dorsal to anterior anal Anterior dorsal to posterior anal Posterior dorsal to anterior anal Posterior dorsal to posterior anal</th>
<td id="76F34E020D787C999FB2FD13FCE8FC8A" box="[714,803,656,884]" gridcol="1" gridrow="2" pageId="12" pageNumber="13">11.4 62.8 52.6 65.6 30.3 15.9</td>
<td id="76F34E020D787C999EBEFD13FBCEFC8A" box="[966,1029,656,884]" gridcol="2" gridrow="2" pageId="12" pageNumber="13">11.9 61.6 51.3 64.1 29.8 15.4</td>
<td id="76F34E020D787C9999F1FD13FAACFC8A" box="[1161,1383,656,884]" gridcol="3" gridrow="2" pageId="12" pageNumber="13">11.412.4 58.663.9 49.653.1 62.265.6 28.331.1 14.816.7</td>
</tr>
<tr id="3522277E0D787C999DEFFC0BFAACFB92" box="[151,1383,904,1132]" gridrow="3" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFFC0BFDC2FB92" box="[151,521,904,1132]" gridcol="0" gridrow="3" pageId="12" pageNumber="13">Anterior dorsal to pelvic-fin origin Posterior dorsal to pelvic-fin origin Percentage of head length Horizontal eye diameter Vertical eye diameter Snout length</th>
<td id="76F34E020D787C999FB2FC0BFCE8FB92" box="[714,803,904,1132]" gridcol="1" gridrow="3" pageId="12" pageNumber="13">37.2 60.7 36.8 33.7 27.5</td>
<td id="76F34E020D787C999EBEFC0BFBCEFB92" box="[966,1029,904,1132]" gridcol="2" gridrow="3" pageId="12" pageNumber="13">35.9 57.8 37.1 35.9 26.2</td>
<td id="76F34E020D787C9999F1FC0BFAACFB92" box="[1161,1383,904,1132]" gridcol="3" gridrow="3" pageId="12" pageNumber="13">33.337.4 55.360.7 36.737.6 33.737.1 24.127.5</td>
</tr>
<tr id="3522277E0D787C999DEFFB03FAACFAEF" box="[151,1383,1152,1297]" gridrow="4" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFFB03FDC2FAEF" box="[151,521,1152,1297]" gridcol="0" gridrow="4" pageId="12" pageNumber="13">Postorbital head length Preorbital depth Lower-jaw length Cheek depth</th>
<td id="76F34E020D787C999FB2FB03FCE8FAEF" box="[714,803,1152,1297]" gridcol="1" gridrow="4" pageId="12" pageNumber="13">43.7 21.4 34.6 20.1</td>
<td id="76F34E020D787C999EBEFB03FBCEFAEF" box="[966,1029,1152,1297]" gridcol="2" gridrow="4" pageId="12" pageNumber="13">40.0 20.9 35.4 20.8</td>
<td id="76F34E020D787C9999F1FB03FAACFAEF" box="[1161,1383,1152,1297]" gridcol="3" gridrow="4" pageId="12" pageNumber="13">38.443.7 19.221.7 34.636.7 18.722.6</td>
</tr>
<tr id="3522277E0D787C999DEFFAA6FAACFA9A" box="[151,1383,1317,1380]" gridrow="5" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFFAA6FDC2FA9A" box="[151,521,1317,1380]" gridcol="0" gridrow="5" pageId="12" pageNumber="13">Head depth Meristics</th>
<td id="76F34E020D787C999FB2FAA6FCE8FA9A" box="[714,803,1317,1380]" gridcol="1" gridrow="5" pageId="12" pageNumber="13">92.2</td>
<td id="76F34E020D787C999EBEFAA6FBCEFA9A" box="[966,1029,1317,1380]" gridcol="2" gridrow="5" pageId="12" pageNumber="13">87.8 mode</td>
<td id="76F34E020D787C9999F1FAA6FAACFA9A" box="[1161,1383,1317,1380]" gridcol="3" gridrow="5" pageId="12" pageNumber="13">80.892.2 range</td>
</tr>
<tr id="3522277E0D787C999DEFFAFBFAACF9A2" box="[151,1383,1400,1628]" gridrow="6" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFFAFBFDC2F9A2" box="[151,521,1400,1628]" gridcol="0" gridrow="6" pageId="12" pageNumber="13">Dorsal-fin spines Dorsal-fin rays Anal-fin spines Anal-fin rays Pelvic-fin rays Pectoral-fin rays</th>
<td id="76F34E020D787C999FB2FAFBFCE8F9A2" box="[714,803,1400,1628]" gridcol="1" gridrow="6" pageId="12" pageNumber="13">17 9 3 9 5 13</td>
<td id="76F34E020D787C999EBEFAFBFBCEF9A2" box="[966,1029,1400,1628]" gridcol="2" gridrow="6" pageId="12" pageNumber="13">17 8, 9 3 8 5 14</td>
<td id="76F34E020D787C9999F1FAFBFAACF9A2" box="[1161,1383,1400,1628]" gridcol="3" gridrow="6" pageId="12" pageNumber="13">1718 810 33 89 55 1314</td>
</tr>
<tr id="3522277E0D787C999DEFF9F3FAACF8AA" box="[151,1383,1648,1876]" gridrow="7" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFF9F3FDC2F8AA" box="[151,521,1648,1876]" gridcol="0" gridrow="7" pageId="12" pageNumber="13">Lateral line scales Pored scales posterior to lateral line Cheek scales Gill rakers on first ceratobranchial Gill rakers on first epibranchial Teeth in outer row of left lower jaw</th>
<td id="76F34E020D787C999FB2F9F3FCE8F8AA" box="[714,803,1648,1876]" gridcol="1" gridrow="7" pageId="12" pageNumber="13">29 0 4 12 3 21</td>
<td id="76F34E020D787C999EBEF9F3FBCEF8AA" box="[966,1029,1648,1876]" gridcol="2" gridrow="7" pageId="12" pageNumber="13">29, 31 1, 2 4 11, 12 3 21</td>
<td id="76F34E020D787C9999F1F9F3FAACF8AA" box="[1161,1383,1648,1876]" gridcol="3" gridrow="7" pageId="12" pageNumber="13">2932 02 35 1113 23 1523</td>
</tr>
<tr id="3522277E0D787C999DEFF8EBFAACF859" box="[151,1383,1896,1959]" gridrow="8" pageId="12" pageNumber="13">
<th id="76F34E020D787C999DEFF8EBFDC2F859" box="[151,521,1896,1959]" gridcol="0" gridrow="8" pageId="12" pageNumber="13">Teeth rows on upper jaw Teeth rows on lower jaw</th>
<td id="76F34E020D787C999FB2F8EBFCE8F859" box="[714,803,1896,1959]" gridcol="1" gridrow="8" pageId="12" pageNumber="13">3 3</td>
<td id="76F34E020D787C999EBEF8EBFBCEF859" box="[966,1029,1896,1959]" gridcol="2" gridrow="8" pageId="12" pageNumber="13">3 2</td>
<td id="76F34E020D787C9999F1F8EBFAACF859" box="[1161,1383,1896,1959]" gridcol="3" gridrow="8" pageId="12" pageNumber="13">23 23</td>
</tr>
</table>
</paragraph>
<paragraph id="8BAD253C0D79836B9DEFFF14FD5FFEC1" blockId="13.[151,1436,151,463]" pageId="13" pageNumber="14">
(0.24). The minimum polygon clusters formed by
<taxonomicName id="4C125EBF0D79836B9FA9FF1AFC8FFF4E" box="[721,836,153,176]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="aurora">
<emphasis id="B966F92E0D79836B9FA9FF1AFC8FFF4E" box="[721,836,153,176]" italics="true" pageId="13" pageNumber="14">M. aurora</emphasis>
</taxonomicName>
and
<taxonomicName id="4C125EBF0D79836B9E04FF1AFC31FF4E" box="[892,1018,152,176]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D79836B9E04FF1AFC31FF4E" box="[892,1018,152,176]" italics="true" pageId="13" pageNumber="14">M. glaucos</emphasis>
</taxonomicName>
did not overlap.
<taxonomicName id="4C125EBF0D79836B99C4FF1BFA57FF4E" box="[1212,1436,152,176]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="aurora">
<emphasis id="B966F92E0D79836B99C4FF1BFA57FF4E" box="[1212,1436,152,176]" italics="true" pageId="13" pageNumber="14">Metriaclima aurora</emphasis>
</taxonomicName>
had a larger horizontal eye diameter (mean 37.0) and vertical eye diameter (mean 36.8) than
<taxonomicName id="4C125EBF0D79836B99E4FF3EFAD3FF2A" box="[1180,1304,188,212]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="glaucos">
<emphasis id="B966F92E0D79836B99E4FF3EFAD3FF2A" box="[1180,1304,188,212]" italics="true" pageId="13" pageNumber="14">M. glaucos</emphasis>
</taxonomicName>
(mean 33.7 for horizontal and vertical eye diameters).
<taxonomicName id="4C125EBF0D79836B9F0DFF63FC95FF06" box="[629,862,224,248]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D79836B9F0DFF63FC95FF06" box="[629,862,224,248]" italics="true" pageId="13" pageNumber="14">Metriaclima xanthos</emphasis>
</taxonomicName>
was intermediate between these two species, however a MANOVA in conjunction with a Hotelling-Lawley trace demonstrated that the three minimum polygon clusters were significantly (p&lt;0.05) different.
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D79836B9DBFFEC8FEA3FE79" pageId="13" pageNumber="14" type="distribution">
<paragraph id="8BAD253C0D79836B9DBFFEC8FEA3FE79" blockId="13.[151,1436,151,463]" pageId="13" pageNumber="14">
<emphasis id="B966F92E0D79836B9DBFFEC8FEABFE9A" bold="true" box="[199,352,331,356]" pageId="13" pageNumber="14">Distribution.</emphasis>
<taxonomicName id="4C125EBF0D79836B9C10FECFFD99FE9A" box="[360,594,332,356]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D79836B9C10FECFFD99FE9A" box="[360,594,332,356]" italics="true" pageId="13" pageNumber="14">Metriaclima xanthos</emphasis>
</taxonomicName>
was only collected at Lumbaulo,
<collectingCountry id="F30565AC0D79836B9EAAFECFFBA2FE9A" box="[978,1129,332,356]" name="Mozambique" pageId="13" pageNumber="14">Mozambique</collectingCountry>
, and is not known to occur elsewhere (
<figureCitation id="132939B90D79836B9C6EFEECFE9CFE79" box="[278,343,367,391]" captionStart="FIGURE 1" captionStartId="1.[151,250,1266,1289]" captionTargetBox="[154,1435,196,1245]" captionTargetId="figure@1.[151,1436,193,1245]" captionTargetPageId="1" captionText="FIGURE 1. Map of Lake MalaŴi showing the locations of species discussed throughout the text." httpUri="https://zenodo.org/record/204915/files/figure.png" pageId="13" pageNumber="14">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C30876B70D79836B9DBFFE17FD15FE31" pageId="13" pageNumber="14" type="etymology">
<paragraph id="8BAD253C0D79836B9DBFFE17FD15FE31" blockId="13.[151,1436,151,463]" pageId="13" pageNumber="14">
<emphasis id="B966F92E0D79836B9DBFFE17FE85FE52" bold="true" box="[199,334,404,428]" pageId="13" pageNumber="14">Etymology.</emphasis>
The specific epithet,
<taxonomicName id="4C125EBF0D79836B9F45FE17FD5FFE52" box="[573,660,404,428]" family="Cichlidae" genus="Metriaclima" higherTaxonomySource="CoL" kingdom="Animalia" order="Perciformes" pageId="13" pageNumber="14" phylum="Chordata" rank="species" species="xanthos">
<emphasis id="B966F92E0D79836B9F45FE17FD5FFE52" box="[573,660,404,428]" italics="true" pageId="13" pageNumber="14">xanthos</emphasis>
</taxonomicName>
, is Greek for yellow, in reference to the yellow belly and dorsal fin of breeding males. It is treated as a noun in apposition.
</paragraph>
</subSubSection>
</treatment>
</document>