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<mods:title id="4B5C1F573A03D40E393B77C3388A88E5">A revision of African Psammoecus (Coleoptera, Silvanidae) and descriptions of two new species from the collection of the Musée royal de lAfrique centrale</mods:title>
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<emphasis id="B966E017E10CFF80FF157CCF8588FC19" bold="true" box="[189,397,932,963]" pageId="27" pageNumber="28">
Key to
<taxonomicName id="4C124786E10CFF80FE8F7CCF8588FC19" ID-CoL="6YB5" ID-ENA="432911" authorityName="Latreille" authorityYear="1829" box="[295,397,932,963]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="genus">species</taxonomicName>
</emphasis>
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<paragraph id="8BAD3C05E10CFF80FF157CB78172FBC6" blockId="27.[189,1399,988,1090]" pageId="27" pageNumber="28">
<keyLead id="30E39F30E10CFF80FF157CB78172FBC6" pageId="27" pageNumber="28">1. Lateral margins of pronotum without teeth; only anterior and posterior groups of small teeth present ............................................................................................................................................................. 2</keyLead>
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<keyLead id="30E39F30E10CFF80FF6D7B438172FB98" box="[197,1399,1063,1090]" pageId="27" pageNumber="28"> Lateral margins of pronotum with various number of large or small teeth ......................................... 3</keyLead>
</paragraph>
</keyStep>
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<paragraph id="8BAD3C05E10CFF80FF157B078173FB71" blockId="27.[189,1399,1130,1271]" pageId="27" pageNumber="28">
<keyLead id="30E39F30E10CFF80FF157B078173FB71" pageId="27" pageNumber="28">
2. 3
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antennomere shorter than 2
<superScript id="7C67914DE10CFF80FDE87B018654FBA2" attach="none" box="[576,593,1130,1144]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
; elytral rows of punctures narrower than interstices.......................... ....................................................................................................................
<taxonomicName id="4C124786E10CFF80FB937BFA8173FB71" authority="(Grouvelle, 1899)" baseAuthorityName="Grouvelle" baseAuthorityYear="1899" box="[1083,1398,1169,1196]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="lateralis">
<emphasis id="B966E017E10CFF80FB937BFA8098FB76" bold="true" box="[1083,1181,1169,1196]" italics="true" pageId="27" pageNumber="28">lateralis</emphasis>
(
<bibRefCitation id="EF8341F4E10CFF80FB047BFA816BFB71" author="Grouvelle A." box="[1196,1390,1169,1195]" pageId="27" pageNumber="28" pagination="136 - 185" refId="ref15042" refString="Grouvelle A. 1899. Descriptions de Clavicornes d'Afrique et de la region Malgache, 2 e memoire. Annales de la Societe Entomologique de France 68: 136 - 185." type="journal article" year="1899">Grouvelle, 1899</bibRefCitation>
)
</taxonomicName>
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</paragraph>
<paragraph id="8BAD3C05E10CFF80FF6D7BD38172FB2D" blockId="27.[189,1399,1130,1271]" pageId="27" pageNumber="28">
<keyLead id="30E39F30E10CFF80FF6D7BD38172FB2D" pageId="27" pageNumber="28">
3
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antennomere longer than 2
<superScript id="7C67914DE10CFF80FD917BDE864FFB19" attach="none" box="[569,586,1205,1219]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
; elytral rows of punctures somewhat wider than interstices............... ..................................................................................................................
<taxonomicName id="4C124786E10CFF80FB8E7BB78172FB2D" authority="Schaufuss, 1872" authorityName="Schaufuss" authorityYear="1872" box="[1062,1399,1244,1271]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="longicornis">
<emphasis id="B966E017E10CFF80FB8E7BB780AAFB2D" bold="true" box="[1062,1199,1244,1271]" italics="true" pageId="27" pageNumber="28">longicornis</emphasis>
Schaufuss, 1872
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<keyLead id="30E39F30E10CFF80FF157A4A8172FABB" pageId="27" pageNumber="28">
3. Body length
<quantity id="4CEA91E0E10CFF80FEDA7A4A861EFAE1" box="[370,539,1313,1339]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" metricValueMax="4.2" metricValueMin="3.4" pageId="27" pageNumber="28" unit="mm" value="3.8" valueMax="4.2" valueMin="3.4">3.40-4.20 mm</quantity>
; lateral margins of pronotum with very small teeth; 1
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antennomere more than three times as long as 2
<superScript id="7C67914DE10CFF80FD807A2E863CFA89" attach="none" box="[552,569,1349,1363]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
.........................................................................
<taxonomicName id="4C124786E10CFF80FBFD7A2D8172FABB" authority="Grouvelle, 1908" authorityName="Grouvelle" authorityYear="1908" box="[1109,1399,1350,1377]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B966E017E10CFF80FBFD7A2D80B5FABB" bold="true" box="[1109,1200,1350,1377]" italics="true" pageId="27" pageNumber="28">grandis</emphasis>
Grouvelle, 1908
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<keyLead id="30E39F30E10CFF80FF6D7A068172FA76" pageId="27" pageNumber="28">
Body length not more than
<quantity id="4CEA91E0E10CFF80FDB37A068683FA5D" box="[539,646,1389,1415]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="27" pageNumber="28" unit="mm" value="3.3">3.30 mm</quantity>
; lateral margins of pronotum with distinct teeth, 1
<superScript id="7C67914DE10CFF80FB657A0080DDFAA3" attach="left" box="[1229,1240,1387,1401]" fontSize="6" pageId="27" pageNumber="28">st</superScript>
antennomere less than three times as long as 2
<superScript id="7C67914DE10CFF80FDF37AFA8669FA45" attach="none" box="[603,620,1425,1439]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
....................................................................................................... 4
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<keyLead id="30E39F30E10CFF80FF157ABD8172F9CC" pageId="27" pageNumber="28">
4. Interstices of lateral striae of elytra with small, distinct spines; pronotum with large lateral teeth, bases of tooth II and III connate.............................................................................
<taxonomicName id="4C124786E10CFF80FBE07A978172F9CC" authority="Grouvelle, 1882" authorityName="Grouvelle" authorityYear="1882" box="[1096,1399,1532,1558]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="spinosus">
<emphasis id="B966E017E10CFF80FBE07A9780B5F9CC" bold="true" box="[1096,1200,1532,1558]" italics="true" pageId="27" pageNumber="28">spinosus</emphasis>
<bibRefCitation id="EF8341F4E10CFF80FB107A978172F9CC" author="Grouvelle A." box="[1208,1399,1532,1558]" pageId="27" pageNumber="28" pagination="275 - 296" refId="ref14966" refString="Grouvelle A. 1882. Cucujides nouveaux du Musee Civique de Genes. Annali del Museo civico di storia naturale Giacomo Doria 18: 275 - 296, pl. 7." type="journal article" year="1882">Grouvelle, 1882</bibRefCitation>
</taxonomicName>
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</paragraph>
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<keyLead id="30E39F30E10CFF80FF6D79498172F9B8" pageId="27" pageNumber="28"> Interstices of lateral striae of elytra without spines, often with tubercles; lateral teeth of pronotum always well separated .......................................................................................................................... 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E624A0E10CFF80FF1579E08172F916" pageId="27" pageNumber="28">
<paragraph id="8BAD3C05E10CFF80FF1579E08172F97C" blockId="27.[189,1399,1674,1740]" box="[189,1399,1674,1702]" pageId="27" pageNumber="28">
<keyLead id="30E39F30E10CFF80FF1579E08172F97C" box="[189,1399,1674,1702]" pageId="27" pageNumber="28">
5. 1
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antennomere less than two times as long as 2
<superScript id="7C67914DE10CFF80FCA579E1871BF942" attach="none" box="[781,798,1674,1688]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
............................................................................... 6
</keyLead>
</paragraph>
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1
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antennomere more than two times as long as 2
<superScript id="7C67914DE10CFF80FCB579DB872BF964" attach="left" box="[797,814,1712,1726]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
............................................................................ 8
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<keyLead id="30E39F30E10CFF80FF15799E8172F8CA" box="[189,1399,1780,1808]" pageId="27" pageNumber="28">
6. 3
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antennomere longer than 2
<superScript id="7C67914DE10CFF80FD94799F8648F8D8" attach="left" box="[572,589,1780,1794]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
............................................................
<taxonomicName id="4C124786E10CFF80FBB9799E8172F8CA" authority="Grouvelle, 1908" authorityName="Grouvelle" authorityYear="1908" box="[1041,1399,1781,1808]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="marginicollis">
<emphasis id="B966E017E10CFF80FBB9799E80B4F8CA" bold="true" box="[1041,1201,1781,1808]" italics="true" pageId="27" pageNumber="28">marginicollis</emphasis>
Grouvelle, 1908
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</paragraph>
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<keyLead id="30E39F30E10CFF80FF6D78778172F8EC" box="[197,1399,1818,1846]" pageId="27" pageNumber="28">
3
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antennomere as long as 2
<superScript id="7C67914DE10CFF80FD807871863CF8F2" attach="left" box="[552,569,1818,1832]" fontSize="6" pageId="27" pageNumber="28">nd</superScript>
............................................................................................................. 7
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7. Striae on elytral disc narrower than interstices; lateral teeth of pronotum with narrow bases.............. ..................................................................................................................
<taxonomicName id="4C124786E10CFF80FB8478ED8172F845" authority="Grouvelle, 1919" authorityName="Grouvelle" authorityYear="1919" box="[1068,1399,1925,1952]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="personatus">
<emphasis id="B966E017E10CFF80FB8478ED80B5F87A" bold="true" box="[1068,1200,1926,1952]" italics="true" pageId="27" pageNumber="28">personatus</emphasis>
<bibRefCitation id="EF8341F4E10CFF80FB1078EE8172F845" author="Grouvelle A." box="[1208,1399,1925,1951]" pageId="27" pageNumber="28" pagination="5 - 38" refId="ref15275" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus Latr. In: Memoires Entomologiques. Etudes sur les coleopteres 2: 5 - 38. Societe Entomologique de France, Paris." type="journal article" year="1919">Grouvelle, 1919</bibRefCitation>
</taxonomicName>
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</paragraph>
<paragraph id="8BAD3C05E10CFF80FF6D78C08172F831" blockId="27.[189,1399,1887,2027]" pageId="27" pageNumber="28">
<keyLead id="30E39F30E10CFF80FF6D78C08172F831" pageId="27" pageNumber="28">
Striae on elytral disc wider than interstices, lateral teeth of pronotum with wide bases....................... .......................................................................................................................................
<taxonomicName id="4C124786E10CFF80FB6B78BB8111F831" authority="Karner, 2012" authorityName="Karner" authorityYear="2012" box="[1219,1300,2000,2027]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="leleupi" status="sp. nov.">
<emphasis id="B966E017E10CFF80FB6B78BB8111F831" bold="true" box="[1219,1300,2000,2027]" italics="true" pageId="27" pageNumber="28">leleupi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555D6CE10CFF80FAB478BA8172F831" box="[1308,1399,2001,2027]" pageId="27" pageNumber="28" rank="species">sp. nov.</taxonomicNameLabel>
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<keyStep id="30E624A0E10BFF87FF157E678170FE96" pageId="28" pageNumber="29">
<paragraph id="8BAD3C05E10BFF87FF157E678172FEFC" blockId="28.[189,1399,268,333]" box="[189,1399,268,295]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF157E678172FEFC" box="[189,1399,268,295]" pageId="28" pageNumber="29">8. Maximum pronotal width near middle (excluding lateral teeth) ......................................................... 9</keyLead>
</paragraph>
<paragraph id="8BAD3C05E10BFF87FF6D7E598170FE96" blockId="28.[189,1399,268,333]" box="[197,1397,306,333]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF6D7E598170FE96" box="[197,1397,306,333]" pageId="28" pageNumber="29"> Maximum pronotal width in anterior third (excluding lateral teeth)..................................................11</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E624A0E10BFF87FF157E1D8173FE6C" pageId="28" pageNumber="29">
<paragraph id="8BAD3C05E10BFF87FF157E1D8172FE4A" blockId="28.[189,1399,374,438]" box="[189,1399,374,401]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF157E1D8172FE4A" box="[189,1399,374,401]" pageId="28" pageNumber="29">
9. Lateral teeth of pronotum slender...............................................................
<taxonomicName id="4C124786E10BFF87FB9D7E1D8172FE4A" authority="Grouvelle, 1889" authorityName="Grouvelle" authorityYear="1889" box="[1077,1399,374,401]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="hacquardi">
<emphasis id="B966E017E10BFF87FB9D7E1D80B4FE4B" bold="true" box="[1077,1201,374,401]" italics="true" pageId="28" pageNumber="29">hacquardi</emphasis>
Grouvelle, 1889
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAD3C05E10BFF87FF6D7EF78173FE6C" blockId="28.[189,1399,374,438]" box="[197,1398,412,438]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF6D7EF78173FE6C" box="[197,1398,412,438]" pageId="28" pageNumber="29"> Lateral teeth of pronotum triangular, with wide bases ...................................................................... 10</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E624A0E10BFF87FF157E8B8172FD62" pageId="28" pageNumber="29">
<paragraph id="8BAD3C05E10BFF87FF157E8B8172FD9C" blockId="28.[189,1399,480,696]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF157E8B8172FD9C" pageId="28" pageNumber="29">
10. Elytral striae distinctly wider than interstices; humeral swelling, a transverse band in the middle of elytra, elytral suture along posterior two thirds and elytral apex dark; parameres short, stout, fused with basal piece................................................................................................
<taxonomicName id="4C124786E10BFF87FBF77D478172FD9C" authority="Grouvelle, 1892" authorityName="Grouvelle" authorityYear="1892" box="[1119,1399,556,582]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="simoni">
<emphasis id="B966E017E10BFF87FBF77D4780B4FD9C" bold="true" box="[1119,1201,556,582]" italics="true" pageId="28" pageNumber="29">simoni</emphasis>
<bibRefCitation id="EF8341F4E10BFF87FB107D478172FD9C" author="Grouvelle A." box="[1208,1399,556,582]" pageId="28" pageNumber="29" pagination="285 - 288" refId="ref14998" refString="Grouvelle A. 1892. Voyage de M. E. Simon a l'ile de Luzon, Philippines (Mars et Avril 1890), 5 e memoire. Cucujides. Annales de la Societe Entomologique de France 61: 285 - 288." type="journal article" year="1892">Grouvelle, 1892</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAD3C05E10BFF87FF6D7D398172FD62" blockId="28.[189,1399,480,696]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF6D7D398172FD62" pageId="28" pageNumber="29">
Elytral striae slightly wider than interstices; elytra with dark, transversal band at beginning of posterior portion, sometimes with darkened suture near elytral apex; parameres spatula-shaped, distinctly separated from basal piece...............................................................................
<taxonomicName id="4C124786E10BFF87FB677DF68110FD62" authority="Karner, 2012" authorityName="Karner" authorityYear="2012" box="[1231,1301,669,696]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="luchti" status="sp. nov.">
<emphasis id="B966E017E10BFF87FB677DF68110FD62" bold="true" box="[1231,1301,669,696]" italics="true" pageId="28" pageNumber="29">luchti</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2555D6CE10BFF87FAB47DF58172FD62" box="[1308,1399,670,696]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E624A0E10BFF87FF157D898173FC92" pageId="28" pageNumber="29">
<paragraph id="8BAD3C05E10BFF87FF157D898172FCF8" blockId="28.[189,1399,738,840]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF157D898172FCF8" pageId="28" pageNumber="29">
11. Elytrae with darkened humeral area, wide dark band across middle and darkened suture near apex; paramera slender............................................................................................
<taxonomicName id="4C124786E10BFF87FBF87C6C8172FCF8" authority="Grouvelle, 1914" authorityName="Grouvelle" authorityYear="1914" box="[1104,1399,775,802]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="laetulus">
<emphasis id="B966E017E10BFF87FBF87C6C80B5FCF8" bold="true" box="[1104,1200,775,802]" italics="true" pageId="28" pageNumber="29">laetulus</emphasis>
<bibRefCitation id="EF8341F4E10BFF87FB107C638172FCF8" author="Grouvelle A. &amp; de Peyerimhoff P." box="[1208,1399,776,802]" pageId="28" pageNumber="29" pagination="141 - 159" refId="ref15196" refString="Grouvelle A. &amp; de Peyerimhoff P. 1914. No. II. - Coleoptera: Cucujidae, Cryptophagidae: avec une description de la larve et de la nymphe de Prostominia convexiuscula Grouvelle [Cucujidae]. Transactions of the Linnean Society of London. 2 nd Series: Zoology 17: 141 - 159. http: // dx. doi. org / 10.1111 / j. 1096 - 3642.1914. tb 00591. x" type="journal article" year="1914">Grouvelle, 1914</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAD3C05E10BFF87FF6D7C458173FC92" blockId="28.[189,1399,738,840]" box="[197,1398,814,840]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF6D7C458173FC92" box="[197,1398,814,840]" pageId="28" pageNumber="29"> Elytrae with round macula and darkened suture near apex .............................................................. 12</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E624A0E10BFF87FF157C198172FC24" pageId="28" pageNumber="29">
<paragraph id="8BAD3C05E10BFF87FF157C198172FC69" blockId="28.[189,1399,882,1022]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF157C198172FC69" pageId="28" pageNumber="29">
12. Lateral margin of pronotum with 4 distinct teeth between small posterior tooth and anterior group of small teeth..........................................................................................
<taxonomicName id="4C124786E10BFF87FC5F7CFC8172FC69" authority="Motschulsky, 1858" authorityName="Motschulsky" authorityYear="1858" box="[1015,1399,919,947]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="trimaculatus">
<emphasis id="B966E017E10BFF87FC5F7CFC8095FC68" bold="true" box="[1015,1168,919,946]" italics="true" pageId="28" pageNumber="29">trimaculatus</emphasis>
Motschulsky, 1858
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAD3C05E10BFF87FF6D7CD58172FC24" blockId="28.[189,1399,882,1022]" pageId="28" pageNumber="29">
<keyLead id="30E39F30E10BFF87FF6D7CD58172FC24" pageId="28" pageNumber="29">
Lateral margin of pronotum with 5 distinct teeth between small posterior tooth and anterior group of small teeth...................................................................................................
<taxonomicName id="4C124786E10BFF87FB907C888172FC24" authority="Grouvelle, 1919" authorityName="Grouvelle" authorityYear="1919" box="[1080,1399,995,1022]" class="Insecta" family="Silvanidae" genus="Psammoecus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="parallelus">
<emphasis id="B966E017E10BFF87FB907C8880B4FC24" bold="true" box="[1080,1201,995,1022]" italics="true" pageId="28" pageNumber="29">parallelus</emphasis>
<bibRefCitation id="EF8341F4E10BFF87FB107C888172FC24" author="Grouvelle A." box="[1208,1399,995,1022]" pageId="28" pageNumber="29" pagination="5 - 38" refId="ref15275" refString="Grouvelle A. 1919. Descriptions d'especes nouvelles du genre Psammoecus Latr. In: Memoires Entomologiques. Etudes sur les coleopteres 2: 5 - 38. Societe Entomologique de France, Paris." type="journal article" year="1919">Grouvelle, 1919</bibRefCitation>
</taxonomicName>
</keyLead>
</paragraph>
</keyStep>
</key>
</subSubSection>
</treatment>
</subSection>
</document>