treatments-xml/data/03/BB/87/03BB87BEFFDEAE3EFED19535FEEAFB18.xml

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<mods:title id="104D779AC313BB94177A85650F0A5E0B">The caddisfly genus Protoptila in Costa Rica (Trichoptera: Glossosomatidae)</mods:title>
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<mods:namePart id="48FF32E5FF35FBF7866C13EA1FFF9F8B">Holzenthal, Ralph W.</mods:namePart>
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<mods:namePart id="AFDEA77763107E216A19A9501245FDF9">Blahnik, Roger J.</mods:namePart>
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<mods:date id="750427CCCFAFF2F46959D1BBCF87F2BE">2006</mods:date>
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<treatment id="03BB87BEFFDEAE3EFED19535FEEAFB18" ID-DOI="http://doi.org/10.5281/zenodo.6260775" ID-GBIF-Taxon="156789594" ID-Zenodo-Dep="6260775" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18" httpUri="http://treatment.plazi.org/id/03BB87BEFFDEAE3EFED19535FEEAFB18" lastPageId="12" lastPageNumber="12" pageId="8" pageNumber="9">
<subSubSection id="C3086523FFDEAE3AFED19535FDEEF9DE" box="[264,619,1558,1584]" pageId="8" pageNumber="9" type="nomenclature">
<paragraph id="8BAD36A8FFDEAE3AFED19535FDEEF9DE" blockId="8.[264,619,1558,1619]" box="[264,619,1558,1584]" pageId="8" pageNumber="9">
<heading id="D0E581C4FFDEAE3AFED19535FDEEF9DE" bold="true" box="[264,619,1558,1584]" fontSize="11" level="1" pageId="8" pageNumber="9" reason="1">
<emphasis id="B966EABAFFDEAE3AFED19535FDEEF9DE" bold="true" box="[264,619,1558,1584]" pageId="8" pageNumber="9">
<taxonomicName id="4C124D2BFFDEAE3AFED19535FE49F9DE" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[264,460,1558,1584]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bribri" status="sp. nov.">
<emphasis id="B966EABAFFDEAE3AFED19535FE49F9DE" bold="true" box="[264,460,1558,1584]" italics="true" pageId="8" pageNumber="9">Protoptila bribri</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFDEAE3AFE029535FDEEF9DE" box="[475,619,1558,1584]" pageId="8" pageNumber="9" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3086523FFDEAE3BFED1951AFA92FBC0" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9" type="description">
<paragraph id="8BAD36A8FFDEAE3AFED1951AFECEF9BD" blockId="8.[264,619,1558,1619]" box="[264,331,1593,1619]" pageId="8" pageNumber="9">
<figureCitation id="13292A2DFFDEAE3AFED1951AFECEF9BD" box="[264,331,1593,1619]" captionStart="FIGURE 5" captionStartId="10.[264,368,1790,1814]" captionTargetBox="[316,1493,285,1774]" captionTargetId="figure@10.[316,1279,319,1774]" captionTargetPageId="10" captionText="FIGURE 5. Protoptila bribri, new species. Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral (inset: apex of phallicata, dorsolateral); D—phallicata apex, dorsal." httpUri="https://zenodo.org/record/2646275/files/figure.svg" pageId="8" pageNumber="9">Fig. 5</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFDEAE3BFED195AAFCD9FED8" blockId="8.[264,1323,1673,1899]" lastBlockId="9.[264,1325,284,1870]" lastPageId="9" lastPageNumber="10" pageId="8" pageNumber="9">
This species is most similar to 
<taxonomicName id="4C124D2BFFDEAE3AFD5095AAFCDAF94C" authority="orotina" box="[649,863,1673,1699]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="orotina" subSpecies="orotina">
<emphasis id="B966EABAFFDEAE3AFD5095AAFD1DF94C" box="[649,664,1673,1698]" italics="true" pageId="8" pageNumber="9">P</emphasis>
. 
<emphasis id="B966EABAFFDEAE3AFD7395AAFCDAF94C" box="[682,863,1673,1698]" italics="true" pageId="8" pageNumber="9">orotina orotina</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C124D2BFFDEAE3AFCAB95AAFB55F94D" authority="Flint 1974" authorityName="Flint" authorityYear="1974" box="[882,1232,1673,1699]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="subSpecies" species="orotina" subSpecies="raposa">
<emphasis id="B966EABAFFDEAE3AFCAB95AAFC05F94C" box="[882,896,1673,1698]" italics="true" pageId="8" pageNumber="9">P</emphasis>
. 
<emphasis id="B966EABAFFDEAE3AFC4A95AAFBC0F94C" box="[915,1093,1673,1698]" italics="true" pageId="8" pageNumber="9">orotina raposa</emphasis>
<bibRefCitation id="EF834B59FFDEAE3AFB8995AAFB55F94D" author="Flint, O. S., Jr." box="[1104,1232,1673,1699]" pageId="8" pageNumber="9" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint 1974</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C124D2BFFDEAE3AFACC95AAFECFF924" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="ixtala">
<emphasis id="B966EABAFFDEAE3AFACC95AAFECFF924" italics="true" pageId="8" pageNumber="9">P. ixtala</emphasis>
</taxonomicName>
. All of these species have a similarly developed phallicata, which is elongate and very angularly bent in the middle. The different species differ most diagnostically in the development of the apex of this structure. In 
<taxonomicName id="4C124D2BFFDEAE3AFCF49422FC08F8F4" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[813,909,1793,1818]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bribri">
<emphasis id="B966EABAFFDEAE3AFCF49422FC08F8F4" box="[813,909,1793,1818]" italics="true" pageId="8" pageNumber="9">P. bribri</emphasis>
</taxonomicName>
it is greatly enlarged and inflated with a sclerotized ridge on its dorsal margin. Additionally, there are minor differences among the species in the shapes of the apical section of tergum X, the parameres, and the apex of sternum VIII, as shown in the illustrations.
</paragraph>
<paragraph id="8BAD36A8FFDFAE3BFEE19267FC79FE68" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Adult. Length of forewing: male 2.4–3.0 mm; female 
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. Color brown (in alcohol), with distinct transverse bar of whitish setae along cord.
</paragraph>
<paragraph id="8BAD36A8FFDFAE3BFEE192B7FA92FBC0" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">Male genitalia. Sternum VI process prominent, slightly curved, longer than wide, apex subacute. Tergum VIII posterior margin with row of relatively short setae; sternum VIII strongly produced ventrally, not bifurcate apically, strongly narrowed, apex slightly broader, weakly emarginate. Segment IX with anterior margin broadly rounded; sternum IX medially produced posteriorly, long, narrow, apex slightly wider, angularly emarginate; posterolateral margin of segment IX without processes. Preanal appendages absent. Tergum X divided mesally to form sclerotized lateral parts with basal and apical sections; basal section subquadrate, longer than wide, apical section elongate, angularly bent from base, apex with acute apicodorsal process and numerous minute spine­like projections. Phallobase dorsally with large, semicircular, laterally compressed apodeme; ventrally with short, rod­like, articulated appendages with apical setae, appendages fitting into sclerotized pockets on ventral margin of phallobase; posteroventral margin of phallobase without paired basal or dorsolateral processes; paramere elongate, membranous, paramere spine elongate, fishhook­shaped, bulbous basally; phallicata very elongate, narrow throughout length, angularly bent near middle, apex much enlarged, with diverging apicodorsal sclerotized ridges; endophallus reduced, not noticeably everted; phallotremal sclerite lightly sclerotized, an incomplete basal ring, with curved, apicomesal projection.</paragraph>
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<paragraph id="8BAD36A8FFDFAE3BFEE1971FFDE4FB48" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<materialsCitation id="3B7A3CF5FFDFAE3BFEE1971FFDE4FB48" ID-GBIF-Occurrence="2236573127" collectingDate="1986-03-09" collectionCode="UMSP" collectorName="S Brasilia &amp; Holzenthal &amp; Fasth" country="Costa Rica" elevation="390" latitude="10.971944" location="Rio Pizote" longLatPrecision="21" longitude="-85.345" pageId="9" pageNumber="10" specimenCode="UMSP000208539" specimenCount="1" stateProvince="Alajuela" typeStatus="holotype">
<emphasis id="B966EABAFFDFAE3BFEE1971FFE76FBB8" bold="true" box="[312,499,1084,1110]" pageId="9" pageNumber="10">
<typeStatus id="54A9880AFFDFAE3BFEE1971FFE2FFBB8" box="[312,426,1084,1110]" pageId="9" pageNumber="10" type="holotype">Holotype</typeStatus>
male
</emphasis>
: 
<emphasis id="B966EABAFFDFAE3BFDD1971FFD41FBB8" bold="true" box="[520,708,1084,1110]" pageId="9" pageNumber="10">
<collectingCountry id="F3057638FFDFAE3BFDD1971FFD41FBB8" box="[520,708,1084,1110]" name="Costa Rica" pageId="9" pageNumber="10">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFDFAE3BFD00971FFCC6FBB8" bold="true" box="[729,835,1084,1110]" pageId="9" pageNumber="10">
<collectingRegion id="49D6F84AFFDFAE3BFD00971FFCC6FBB8" box="[729,835,1084,1110]" country="Costa Rica" name="Alajuela" pageId="9" pageNumber="10">Alajuela</collectingRegion>
</emphasis>
: 
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, ca. 
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(air) 
<collectorName id="26E7537EFFDFAE3BFB72971FFAA3FBB8" box="[1195,1318,1084,1110]" pageId="9" pageNumber="10">S Brasilia</collectorName>
, 
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, 
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, el. 
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<quantity id="4CEA9B4DFFDFAE3BFD419747FD6AFB90" box="[664,751,1124,1150]" metricMagnitude="2" metricUnit="m" metricValue="3.9" pageId="9" pageNumber="10" unit="m" value="390.0">390 m</quantity>
</elevation>
, 
<date id="FFAC1068FFDFAE3BFCDC9747FCFDFB90" box="[773,888,1124,1150]" pageId="9" pageNumber="10" value="1986-03-09">
<collectingDate id="EFE8E980FFDFAE3BFCDC9747FCFDFB90" box="[773,888,1124,1150]" pageId="9" pageNumber="10" value="1986-03-09">9.iii.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFC579747FB8AFB90" box="[910,1039,1124,1150]" pageId="9" pageNumber="10">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFDFAE3BFB919747FB02FB90" box="[1096,1159,1124,1150]" pageId="9" pageNumber="10">Fasth</collectorName>
(in alcohol) (
<specimenCode id="DBB49ED3FFDFAE3BFEC997AFFE62FB48" box="[272,487,1164,1190]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="9" pageNumber="10" type="University or college">UMSP000208539</specimenCode>
) (
<collectionCode id="ED03AE6DFFDFAE3BFE2697AFFDD0FB48" box="[511,597,1164,1190]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="9" pageNumber="10" type="University or college">UMSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAD36A8FFDFAE3BFED19797FAAEFB18" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<materialsCitation id="3B7A3CF5FFDFAE3BFED19797FBF3FB20" ID-GBIF-Occurrence="2236573161" box="[264,1142,1204,1230]" collectionCode="NMNH" collectorName="Same" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B966EABAFFDFAE3BFED19797FE07FB20" bold="true" box="[264,386,1204,1230]" pageId="9" pageNumber="10">
<typeStatus id="54A9880AFFDFAE3BFED19797FE07FB20" box="[264,386,1204,1230]" pageId="9" pageNumber="10" type="paratype">Paratypes</typeStatus>
</emphasis>
: 
<collectorName id="26E7537EFFDFAE3BFE459797FE59FB20" box="[412,476,1204,1230]" pageId="9" pageNumber="10">Same</collectorName>
data as 
<typeStatus id="54A9880AFFDFAE3BFD809797FD38FB20" box="[601,701,1204,1230]" pageId="9" pageNumber="10" type="holotype">holotype</typeStatus>
— 
<specimenCount id="9D14FD21FFDFAE3BFD249797FCD0FB20" box="[765,853,1204,1230]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFBD69797FBE8FB20" box="[1039,1133,1204,1230]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDFAE3BFB559797FBCAFB18" ID-GBIF-Occurrence="2236573122" collectingDate="1986-03-12" collectionCode="UMSP" collectorName="Holzenthal &amp; Fasth" pageId="9" pageNumber="10" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
same, except 
<date id="FFAC1068FFDFAE3BFED197FFFE02FB18" box="[264,391,1244,1270]" pageId="9" pageNumber="10" value="1986-03-12">
<collectingDate id="EFE8E980FFDFAE3BFED197FFFE02FB18" box="[264,391,1244,1270]" pageId="9" pageNumber="10" value="1986-03-12">12.iii.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFE4497FFFD9EFB18" box="[413,539,1244,1270]" pageId="9" pageNumber="10">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFDFAE3BFD8C97FFFD16FB18" box="[597,659,1244,1270]" pageId="9" pageNumber="10">Fasth</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFD0D97FFFCB2FB18" box="[724,823,1244,1270]" pageId="9" pageNumber="10" type="male">2 males</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFC2A97FFFBC2FB18" box="[1011,1095,1244,1270]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="9" pageNumber="10" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
; 
<emphasis id="B966EABAFFDFAE3BFBBF97FFFAA7FB18" bold="true" box="[1126,1314,1244,1270]" pageId="9" pageNumber="10">
<collectingCountry id="F3057638FFDFAE3BFBBF97FFFAA7FB18" box="[1126,1314,1244,1270]" name="Costa Rica" pageId="9" pageNumber="10">COSTA RICA</collectingCountry>
</emphasis>
:
</paragraph>
</subSubSection>
<subSubSection id="C3086523FFDFAE3EFED19627FEEAFB18" lastPageId="12" lastPageNumber="13" pageId="9" pageNumber="10" type="description">
<paragraph id="8BAD36A8FFDFAE3BFED19627FC69F910" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
<materialsCitation id="3B7A3CF5FFDFAE3BFED19627FE51FAF0" ID-GBIF-Occurrence="2236573135" box="[264,468,1284,1310]" collectorName="Rio" location="Guanacaste" pageId="9" pageNumber="10" specimenCount="1" stateProvince="Guanacaste">
<emphasis id="B966EABAFFDFAE3BFED19627FE12FAF0" bold="true" box="[264,407,1284,1310]" pageId="9" pageNumber="10">
<collectingRegion id="49D6F84AFFDFAE3BFED19627FE12FAF0" box="[264,407,1284,1310]" country="Costa Rica" name="Guanacaste" pageId="9" pageNumber="10">Guanacaste</collectingRegion>
</emphasis>
: 
<collectorName id="26E7537EFFDFAE3BFE739627FE51FAF0" box="[426,468,1284,1310]" pageId="9" pageNumber="10">Río</collectorName>
</materialsCitation>
<materialsCitation id="3B7A3CF5FFDFAE3BFE079627FC64FAA8" ID-GBIF-Occurrence="2236573155" collectingDate="1986-03-11" collectionCode="UMSP" collectorName="Holzenthal &amp; Fasth" elevation="260" latitude="11.058889" location="4.2 km W Santa Cecilia" longLatPrecision="21" longitude="-85.44806" pageId="9" pageNumber="10" specimenCount="1" specimenCount-male="1" stateProvince="Maine">
<collectingRegion id="49D6F84AFFDFAE3BFE079627FDA7FAF0" box="[478,546,1284,1310]" country="United States of America" name="Maine" pageId="9" pageNumber="10">Mena</collectingRegion>
, 
<location id="8ECD6073FFDFAE3BFDE89627FCD6FAF0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDFAE3BFDE89627FCD6FAF0" box="[561,851,1284,1310]" latitude="11.058889" longLatPrecision="21" longitude="-85.44806" name="4.2 km W Santa Cecilia" pageId="9" pageNumber="10" stateProvince="Maine">
<quantity id="4CEA9B4DFFDFAE3BFDE89627FD01FAF0" box="[561,644,1284,1310]" metricMagnitude="3" metricUnit="m" metricValue="4.2" pageId="9" pageNumber="10" unit="km" value="4.2">4.2 km</quantity>
W Santa Cecilia
</location>
, 
<geoCoordinate id="EE26506FFFDFAE3BFCBA9627FC75FAF0" box="[867,1008,1284,1310]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="11.058889">11°03’32”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDFAE3BFBD99627FB24FAF0" box="[1024,1185,1284,1310]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-85.44806">085°26’53”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDFAE3BFB0E9627FAA3FAF0" box="[1239,1318,1284,1310]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="10" unit="m" value="260.0">
<elevation id="003FD19BFFDFAE3BFB0E9627FAA3FAF0" box="[1239,1318,1284,1310]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="10" unit="m" value="260.0">260 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDFAE3BFED1960FFE03FAA8" box="[264,390,1324,1350]" pageId="9" pageNumber="10" value="1986-03-11">
<collectingDate id="EFE8E980FFDFAE3BFED1960FFE03FAA8" box="[264,390,1324,1350]" pageId="9" pageNumber="10" value="1986-03-11">11.iii.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFE48960FFD8AFAA8" box="[401,527,1324,1350]" pageId="9" pageNumber="10">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFDFAE3BFDED960FFDF7FAA8" box="[564,626,1324,1350]" pageId="9" pageNumber="10">Fasth</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFD45960FFD6FFAA8" box="[668,746,1324,1350]" pageId="9" pageNumber="10" type="male">1 male</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFC5D960FFC5CFAA8" box="[900,985,1324,1350]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="9" pageNumber="10" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDFAE3BFC35960FFC2CFA78" ID-GBIF-Occurrence="2236573121" collectingDate="1986-06-19" collectionCode="UMSP" collectorName="Holzenthal &amp; Heyn &amp; Armitage" elevation="30" latitude="10.440001" location="Rio Puerto Viejo" longLatPrecision="21" longitude="-84.01195" municipality="Estacion Biologia La Selva" pageId="9" pageNumber="10" specimenCount="22" specimenCount-male="22" stateProvince="Heredia">
<emphasis id="B966EABAFFDFAE3BFC35960FFBCBFAA8" bold="true" box="[1004,1102,1324,1350]" pageId="9" pageNumber="10">
<collectingRegion id="49D6F84AFFDFAE3BFC35960FFBCBFAA8" box="[1004,1102,1324,1350]" country="Costa Rica" name="Heredia" pageId="9" pageNumber="10">Heredia</collectingRegion>
</emphasis>
: 
<collectingMunicipality id="6BC9ACD2FFDFAE3BFB84960FFEF7FA80" pageId="9" pageNumber="10">Estación Biología La Selva</collectingMunicipality>
, 
<location id="8ECD6073FFDFAE3BFE589677FDCFFA80" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDFAE3BFE589677FDCFFA80" box="[385,586,1364,1390]" latitude="10.440001" longLatPrecision="21" longitude="-84.01195" municipality="Estacion Biologia La Selva" name="Rio Puerto Viejo" pageId="9" pageNumber="10" stateProvince="Heredia">Río Puerto Viejo</location>
, 
<geoCoordinate id="EE26506FFFDFAE3BFD819677FD62FA80" box="[600,743,1364,1390]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="10.440001">10°26’24”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDFAE3BFD2C9677FC12FA80" box="[757,919,1364,1390]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-84.01195">084°00’43”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDFAE3BFC129677FB8FFA80" box="[971,1034,1364,1390]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="m" value="30.0">
<elevation id="003FD19BFFDFAE3BFC129677FB8FFA80" box="[971,1034,1364,1390]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="9" pageNumber="10" unit="m" value="30.0">30 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDFAE3BFBC09677FB13FA80" box="[1049,1174,1364,1390]" pageId="9" pageNumber="10" value="1986-06-19">
<collectingDate id="EFE8E980FFDFAE3BFBC09677FB13FA80" box="[1049,1174,1364,1390]" pageId="9" pageNumber="10" value="1986-06-19">19.vi.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFB7C9677FAA3FA80" box="[1189,1318,1364,1390]" pageId="9" pageNumber="10">Holzenthal</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFED1965FFECCFA78" box="[264,329,1404,1430]" pageId="9" pageNumber="10">Heyn</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFEBB965FFE4BFA78" box="[354,462,1404,1430]" pageId="9" pageNumber="10">Armitage</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFDCD965FFD0DFA78" box="[532,648,1404,1430]" pageId="9" pageNumber="10" type="male">22 males</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFC95965FFC24FA78" box="[844,929,1404,1430]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="9" pageNumber="10" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDFAE3BFC1B965FFE61FA08" ID-GBIF-Occurrence="2236573132" collectingDate="1986-06-20" collectingDateMax="1986-06-21" collectingDateMin="1986-06-20" collectionCode="NMNH" collectorName="Holzenthal &amp; Heyn &amp; Armitage" elevation="50" latitude="10.436945" location="Quebrada" longLatPrecision="21" longitude="-84.01" pageId="9" pageNumber="10" specimenCount="9" specimenCount-male="9" stateProvince="Heredia">
<location id="8ECD6073FFDFAE3BFC1B965FFBB4FA78" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDFAE3BFC1B965FFBB4FA78" box="[962,1073,1404,1430]" latitude="10.436945" longLatPrecision="21" longitude="-84.01" name="Quebrada" pageId="9" pageNumber="10" stateProvince="Heredia">Quebrada</location>
Sura, 
<geoCoordinate id="EE26506FFFDFAE3BFB4F965FFAA0FA78" box="[1174,1317,1404,1430]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="10.436945">10°26’13”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDFAE3BFED19687FE28FA50" box="[264,429,1444,1470]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-84.01">084°00’36”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDFAE3BFE359687FDB7FA50" box="[492,562,1444,1470]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="m" value="50.0">
<elevation id="003FD19BFFDFAE3BFE359687FDB7FA50" box="[492,562,1444,1470]" metricMagnitude="1" metricUnit="m" metricValue="5.0" pageId="9" pageNumber="10" unit="m" value="50.0">50 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDFAE3BFD9F9687FD68FA50" box="[582,749,1444,1470]" pageId="9" pageNumber="10" value="1986-06-20" valueMax="1986-06-21" valueMin="1986-06-20">
<collectingDate id="EFE8E980FFDFAE3BFD9F9687FD68FA50" box="[582,749,1444,1470]" pageId="9" pageNumber="10" value="1986-06-20" valueMax="1986-06-21" valueMin="1986-06-20">20­21.vi.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFCD89687FC00FA50" box="[769,901,1444,1470]" pageId="9" pageNumber="10">Holzenthal</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFC419687FC5FFA50" box="[920,986,1444,1470]" pageId="9" pageNumber="10">Heyn</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFC379687FBD8FA50" box="[1006,1117,1444,1470]" pageId="9" pageNumber="10">Armitage</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFB419687FB7FFA50" box="[1176,1274,1444,1470]" pageId="9" pageNumber="10" type="male">9 males</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFEA296EFFE5EFA08" box="[379,475,1484,1510]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDFAE3BFE2D96EFFDEFF9D8" ID-GBIF-Occurrence="2236573143" collectingDate="1987-03-23" collectingDateMax="1987-03-24" collectingDateMin="1987-03-23" collectionCode="UMSP" collectorName="Holzenthal &amp; Hamilton &amp; Heyn" county="Reserva Biologica Hitoy-Cerere" elevation="90" latitude="9.671111" location="Est. Miramar" longLatPrecision="21" longitude="-83.02806" municipality="Rio Cerere" pageId="9" pageNumber="10" specimenCount="107" specimenCount-male="107" stateProvince="Limon">
<emphasis id="B966EABAFFDFAE3BFE2D96EFFDC2FA08" bold="true" box="[500,583,1484,1510]" pageId="9" pageNumber="10">
<collectingRegion id="49D6F84AFFDFAE3BFE2D96EFFDC2FA08" box="[500,583,1484,1510]" country="Costa Rica" name="Limon" pageId="9" pageNumber="10">Limón</collectingRegion>
</emphasis>
: 
<collectingCounty id="62CC4E24FFDFAE3BFD8296EFFC65FA08" box="[603,992,1484,1510]" pageId="9" pageNumber="10">Reserva Biológica Hitoy­Cerere</collectingCounty>
, 
<collectingMunicipality id="6BC9ACD2FFDFAE3BFC3696EFFBF3FA08" box="[1007,1142,1484,1510]" pageId="9" pageNumber="10">Río Cerere</collectingMunicipality>
, 
<location id="8ECD6073FFDFAE3BFB5F96EFFAA2FA08" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDFAE3BFB5F96EFFAA2FA08" box="[1158,1319,1484,1510]" county="Reserva Biologica Hitoy-Cerere" latitude="9.671111" longLatPrecision="21" longitude="-83.02806" municipality="Rio Cerere" name="Est. Miramar" pageId="9" pageNumber="10" stateProvince="Limon">Est. Miramar</location>
, 
<geoCoordinate id="EE26506FFFDFAE3BFED196D7FE1FF9E0" box="[264,410,1524,1550]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="9.671111">09°40’16”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDFAE3BFE7196D7FDC8F9E0" box="[424,589,1524,1550]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-83.02806">083°01’41”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDFAE3BFDA696D7FD3AF9E0" box="[639,703,1524,1550]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="10" unit="m" value="90.0">
<elevation id="003FD19BFFDFAE3BFDA696D7FD3AF9E0" box="[639,703,1524,1550]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="9" pageNumber="10" unit="m" value="90.0">90 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDFAE3BFD1596D7FCF0F9E0" box="[716,885,1524,1550]" pageId="9" pageNumber="10" value="1987-03-23" valueMax="1987-03-24" valueMin="1987-03-23">
<collectingDate id="EFE8E980FFDFAE3BFD1596D7FCF0F9E0" box="[716,885,1524,1550]" pageId="9" pageNumber="10" value="1987-03-23" valueMax="1987-03-24" valueMin="1987-03-23">23­24.iii.1987</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFC5B96D7FB83F9E0" box="[898,1030,1524,1550]" pageId="9" pageNumber="10">Holzenthal</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFBCA96D7FB00F9E0" box="[1043,1157,1524,1550]" pageId="9" pageNumber="10">Hamilton</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFB4B96D7FB57F9E0" box="[1170,1234,1524,1550]" pageId="9" pageNumber="10">Heyn</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFAD896D7FEC9F9D8" pageId="9" pageNumber="10" type="male">107 males</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFDD5953FFDE7F9D8" box="[524,610,1564,1590]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="9" pageNumber="10" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDFAE3BFD58953FFEFCF968" ID-GBIF-Occurrence="2236573157" collectingDate="1986-02-01" collectionCode="NMNH" collectorName="SE Suretka &amp; Holzenthal &amp; Morse &amp; Fasth" elevation="48" latitude="9.553889" location="Rio Telire" longLatPrecision="21" longitude="-82.891945" pageId="9" pageNumber="10" specimenCount="4" specimenCount-male="4" stateProvince="Limon">
<location id="8ECD6073FFDFAE3BFD58953FFC87F9D8" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDFAE3BFD58953FFC87F9D8" box="[641,770,1564,1590]" latitude="9.553889" longLatPrecision="21" longitude="-82.891945" name="Rio Telire" pageId="9" pageNumber="10" stateProvince="Limon">Río Telire</location>
and small trib., 
<collectorName id="26E7537EFFDFAE3BFC35953FFBF9F9D8" box="[1004,1148,1564,1590]" pageId="9" pageNumber="10">SE Suretka</collectorName>
, 
<geoCoordinate id="EE26506FFFDFAE3BFB4A953FFAA3F9D8" box="[1171,1318,1564,1590]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="9.553889">09°33’14”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDFAE3BFED19567FE29F9B0" box="[264,428,1604,1630]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-82.891945">082°53’31”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDFAE3BFE339567FDAAF9B0" box="[490,559,1604,1630]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="10" unit="m" value="48.0">
<elevation id="003FD19BFFDFAE3BFE339567FDAAF9B0" box="[490,559,1604,1630]" metricMagnitude="1" metricUnit="m" metricValue="4.8" pageId="9" pageNumber="10" unit="m" value="48.0">48 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDFAE3BFD989567FD29F9B0" box="[577,684,1604,1630]" pageId="9" pageNumber="10" value="1986-02-01">
<collectingDate id="EFE8E980FFDFAE3BFD989567FD29F9B0" box="[577,684,1604,1630]" pageId="9" pageNumber="10" value="1986-02-01">1.ii.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFD669567FCC6F9B0" box="[703,835,1604,1630]" pageId="9" pageNumber="10">Holzenthal</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFC8F9567FC21F9B0" box="[854,932,1604,1630]" pageId="9" pageNumber="10">Morse</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFC6F9567FC73F9B0" box="[950,1014,1604,1630]" pageId="9" pageNumber="10">Fasth</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFBF69567FB15F9B0" box="[1071,1168,1604,1630]" pageId="9" pageNumber="10" type="male">4 males</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFEC9954FFEF5F968" box="[272,368,1644,1670]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="9" pageNumber="10" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDFAE3BFE52954FFB0CF940" ID-GBIF-Occurrence="2236573156" collectingDate="1987-03-25" collectionCode="UMSP" collectorName="W Bribri &amp; Holzenthal &amp; Hamilton &amp; Heyn" elevation="60" latitude="9.62" location="Rio Uatsi" longLatPrecision="21" longitude="-82.9" pageId="9" pageNumber="10" specimenCount="98" specimenCount-male="98" stateProvince="Limon">
<location id="8ECD6073FFDFAE3BFE52954FFD80F968" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDFAE3BFE52954FFD80F968" box="[395,517,1644,1670]" latitude="9.62" longLatPrecision="21" longitude="-82.9" name="Rio Uatsi" pageId="9" pageNumber="10" stateProvince="Limon">Río Uatsi</location>
, ca. 
<quantity id="4CEA9B4DFFDFAE3BFD9F954FFD0CF968" box="[582,649,1644,1670]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="9" pageNumber="10" unit="km" value="8.0">8 km</quantity>
(air) 
<collectorName id="26E7537EFFDFAE3BFD0E954FFCCFF968" box="[727,842,1644,1670]" pageId="9" pageNumber="10">W Bribri</collectorName>
, 
<geoCoordinate id="EE26506FFFDFAE3BFC85954FFC6AF968" box="[860,1007,1644,1670]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="9.62">09°37’12”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDFAE3BFBD8954FFB22F968" box="[1025,1191,1644,1670]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-82.9">082°54’00”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDFAE3BFB3A954FFAACF968" box="[1251,1321,1644,1670]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="10" unit="m" value="60.0">
<elevation id="003FD19BFFDFAE3BFB3A954FFAACF968" box="[1251,1321,1644,1670]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="9" pageNumber="10" unit="m" value="60.0">60 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDFAE3BFED195B7FE0CF940" box="[264,393,1684,1710]" pageId="9" pageNumber="10" value="1987-03-25">
<collectingDate id="EFE8E980FFDFAE3BFED195B7FE0CF940" box="[264,393,1684,1710]" pageId="9" pageNumber="10" value="1987-03-25">25.iii.1987</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFE4195B7FD99F940" box="[408,540,1684,1710]" pageId="9" pageNumber="10">Holzenthal</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFDF395B7FD19F940" box="[554,668,1684,1710]" pageId="9" pageNumber="10">Hamilton</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFD7395B7FD6CF940" box="[682,745,1684,1710]" pageId="9" pageNumber="10">Heyn</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFCC395B7FC03F940" box="[794,902,1684,1710]" pageId="9" pageNumber="10" type="male">98 males</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFBF295B7FB04F940" box="[1067,1153,1684,1710]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="9" pageNumber="10" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDFAE3BFB4195B7FC62F910" ID-GBIF-Occurrence="2236573140" collectingDate="1987-03-26" collectionCode="INBIO" collectorName="Holzenthal &amp; Hamilton &amp; Heyn" elevation="150" latitude="9.888056" location="Rio Banano" longLatPrecision="21" longitude="-83.16694" pageId="9" pageNumber="10" specimenCount="28" specimenCount-female="11" specimenCount-male="17" stateProvince="Limon">
<location id="8ECD6073FFDFAE3BFB4195B7FAACF940" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDFAE3BFB4195B7FAACF940" box="[1176,1321,1684,1710]" latitude="9.888056" longLatPrecision="21" longitude="-83.16694" name="Rio Banano" pageId="9" pageNumber="10" stateProvince="Limon">Río Banano</location>
, 
<locationDeviation id="9A1F2F8AFFDFAE3BFED1959FFD91F938" box="[264,532,1724,1750]" location="Bomba" pageId="9" pageNumber="10">
<quantity id="4CEA9B4DFFDFAE3BFED1959FFEDCF938" box="[264,345,1724,1750]" metricMagnitude="4" metricUnit="m" metricValue="1.6" pageId="9" pageNumber="10" unit="km" value="16.0">16 km</quantity>
WSW Bomba
</locationDeviation>
, 
<geoCoordinate id="EE26506FFFDFAE3BFDFC959FFD3DF938" box="[549,696,1724,1750]" direction="north" orientation="latitude" pageId="9" pageNumber="10" precision="15" value="9.888056">09°53’17”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDFAE3BFD13959FFCEAF938" box="[714,879,1724,1750]" direction="west" orientation="longitude" pageId="9" pageNumber="10" precision="15" value="-83.16694">083°10’01”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDFAE3BFC72959FFC7AF938" box="[939,1023,1724,1750]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="m" value="150.0">
<elevation id="003FD19BFFDFAE3BFC72959FFC7AF938" box="[939,1023,1724,1750]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="9" pageNumber="10" unit="m" value="150.0">150 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDFAE3BFBC9959FFB17F938" box="[1040,1170,1724,1750]" pageId="9" pageNumber="10" value="1987-03-26">
<collectingDate id="EFE8E980FFDFAE3BFBC9959FFB17F938" box="[1040,1170,1724,1750]" pageId="9" pageNumber="10" value="1987-03-26">26.iii.1987</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDFAE3BFB7C959FFAACF938" box="[1189,1321,1724,1750]" pageId="9" pageNumber="10">Holzenthal</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFED195C7FEFFF910" box="[264,378,1764,1790]" pageId="9" pageNumber="10">Hamilton</collectorName>
, 
<collectorName id="26E7537EFFDFAE3BFE5F95C7FE43F910" box="[390,454,1764,1790]" pageId="9" pageNumber="10">Heyn</collectorName>
— 
<specimenCount id="9D14FD21FFDFAE3BFE2895C7FDDBF910" box="[497,606,1764,1790]" pageId="9" pageNumber="10" type="male">17 males</specimenCount>
, 
<specimenCount id="9D14FD21FFDFAE3BFDB095C7FD6DF910" box="[617,744,1764,1790]" pageId="9" pageNumber="10" type="female">11 females</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFDFAE3BFC5E95C7FC5AF910" box="[903,991,1764,1790]" pageId="9" pageNumber="10">INBIO</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD36A8FFDFAE3BFEE1942FFD42F8A0" blockId="9.[264,1325,284,1870]" pageId="9" pageNumber="10">
Etymology. 
<emphasis id="B966EABAFFDFAE3BFE10942FFD94F8CB" box="[457,529,1804,1829]" italics="true" pageId="9" pageNumber="10">Bribri</emphasis>
comes from the name of the indigenous people of the Talamanca mountains of 
<collectingCountry id="F3057638FFDFAE3BFE709417FDACF8A0" box="[425,553,1844,1870]" name="Costa Rica" pageId="9" pageNumber="10">Costa Rica</collectingCountry>
and 
<collectingCountry id="F3057638FFDFAE3BFDBB9417FD46F8A0" box="[610,707,1844,1870]" name="Panama" pageId="9" pageNumber="10">Panama</collectingCountry>
.
</paragraph>
<caption id="DF6D6620FFDCAE38FED195DDFED5F8B5" httpUri="https://zenodo.org/record/2646275/files/figure.svg" pageId="10" pageNumber="11" startId="10.[264,368,1790,1814]" targetBox="[316,1493,285,1774]" targetPageId="10">
<paragraph id="8BAD36A8FFDCAE38FED195DDFED5F8B5" blockId="10.[264,1323,1790,1883]" pageId="10" pageNumber="11">
<emphasis id="B966EABAFFDCAE38FED195DDFE0BF8F8" bold="true" box="[264,398,1790,1814]" pageId="10" pageNumber="11">FIGURE 5.</emphasis>
<taxonomicName id="4C124D2BFFDCAE38FE4095DCFDCAF8F8" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[409,591,1791,1814]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="10" pageNumber="11" phylum="Arthropoda" rank="species" species="bribri" status="sp. nov.">
<emphasis id="B966EABAFFDCAE38FE4095DCFDCAF8F8" box="[409,591,1791,1814]" italics="true" pageId="10" pageNumber="11">Protoptila bribri</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFDCAE38FDB895DDFD6DF8FB" box="[609,744,1790,1813]" pageId="10" pageNumber="11" rank="species">new species</taxonomicNameLabel>
. Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral (inset: apex of phallicata, dorsolateral); D—phallicata apex, dorsal.
</paragraph>
</caption>
<paragraph id="8BAD36A8FFDDAE39FED1923FFD9CFEDB" blockId="11.[264,537,283,350]" box="[264,537,283,310]" pageId="11" pageNumber="12">
<taxonomicName id="4C124D2BFFDDAE39FED1923FFD9CFEDB" authority="Flint" authorityName="Flint" box="[264,537,283,310]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="burica">
<emphasis id="B966EABAFFDDAE39FED1923FFD9CFEDB" bold="true" box="[264,537,283,310]" pageId="11" pageNumber="12">
<emphasis id="B966EABAFFDDAE39FED1923FFE53FED8" bold="true" box="[264,470,284,310]" italics="true" pageId="11" pageNumber="12">Protoptila burica</emphasis>
Flint
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FED19267FECEFEB0" blockId="11.[264,537,283,350]" box="[264,331,324,350]" pageId="11" pageNumber="12">
<figureCitation id="13292A2DFFDDAE39FED19267FECEFEB0" box="[264,331,324,350]" captionStart="FIGURE 6" captionStartId="13.[264,368,1407,1431]" captionTargetBox="[94,1286,286,1395]" captionTargetId="figure@13.[301,1286,319,1395]" captionTargetPageId="13" captionText="FIGURE 6. Protoptila burica, Flint. Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral." httpUri="https://zenodo.org/record/2646281/files/figure.svg" pageId="11" pageNumber="12">Fig. 6</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FED192BAFE23FE5E" blockId="11.[264,422,409,432]" box="[264,422,409,432]" pageId="11" pageNumber="12">
<bibRefCitation id="EF834B59FFDDAE39FED192BAFEFFFE5E" author="Flint, O. S., Jr." box="[264,378,409,432]" pageId="11" pageNumber="12" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint 1974</bibRefCitation>
: 17.
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FED192CFFDC1FD90" blockId="11.[264,1325,492,1158]" pageId="11" pageNumber="12">
<bibRefCitation id="EF834B59FFDDAE39FED192CFFE19FDE8" author="Flint, O. S., Jr." box="[264,412,492,518]" pageId="11" pageNumber="12" pagination="1 - 24" refId="ref12774" refString="Flint, O. S., Jr. (1967) Studies of Neotropical caddis flies, IV: new species from Mexico and Central America. Proceedings of the United States National Museum, 123, 1 - 24." type="journal article" year="1967">Flint (1967)</bibRefCitation>
noted the similarity of this species to 
<taxonomicName id="4C124D2BFFDDAE39FC4A92CFFB26FDE8" authority="Flint 1967" authorityName="Flint" authorityYear="1967" box="[915,1187,492,518]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cristata">
<emphasis id="B966EABAFFDDAE39FC4A92CFFC27FDEB" box="[915,930,492,517]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. 
<emphasis id="B966EABAFFDDAE39FC6E92CFFB94FDEB" box="[951,1041,492,517]" italics="true" pageId="11" pageNumber="12">cristata</emphasis>
<bibRefCitation id="EF834B59FFDDAE39FBC692CFFB26FDE8" author="Flint, O. S., Jr." box="[1055,1187,492,518]" pageId="11" pageNumber="12" pagination="1 - 24" refId="ref12774" refString="Flint, O. S., Jr. (1967) Studies of Neotropical caddis flies, IV: new species from Mexico and Central America. Proceedings of the United States National Museum, 123, 1 - 24." type="journal article" year="1967">Flint 1967</bibRefCitation>
</taxonomicName>
. The most distinctive and diagnostic feature of 
<taxonomicName id="4C124D2BFFDDAE39FD609137FCA0FDC3" box="[697,805,532,558]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="burica">
<emphasis id="B966EABAFFDDAE39FD609137FD4DFDC3" box="[697,712,532,557]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. 
<emphasis id="B966EABAFFDDAE39FD019137FCA0FDC3" box="[728,805,532,557]" italics="true" pageId="11" pageNumber="12">burica</emphasis>
</taxonomicName>
is the shape of the apical section of tergum X, which is branched apically into an elongate, narrow, curved dorsal projection, and three shorter ventral projections.
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FEE191AFFCE0FB68" blockId="11.[264,1325,492,1158]" pageId="11" pageNumber="12">
Material examined. 
<emphasis id="B966EABAFFDDAE39FDFA91AFFD58FD48" bold="true" box="[547,733,652,678]" pageId="11" pageNumber="12">
<collectingCountry id="F3057638FFDDAE39FDFA91AFFD58FD48" box="[547,733,652,678]" name="Costa Rica" pageId="11" pageNumber="12">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFDDAE39FD2B91AFFCDDFD48" bold="true" box="[754,856,652,678]" pageId="11" pageNumber="12">
<collectingRegion id="49D6F84AFFDDAE39FD2B91AFFCDDFD48" box="[754,856,652,678]" country="Costa Rica" name="Cartago" pageId="11" pageNumber="12">Cartago</collectingRegion>
</emphasis>
: Río Chitaría, rt 10, 10 km NW Río Reventazón, 
<geoCoordinate id="EE26506FFFDDAE39FE739197FDB9FD20" box="[426,572,692,718]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="9.92">09°55’12”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDDAE39FD889197FD70FD20" box="[593,757,692,718]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-83.60389">083°36’14”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDDAE39FCEE9197FC09FD20" box="[823,908,692,718]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="12" unit="m" value="740.0">740 m</quantity>
, 
<date id="FFAC1068FFDDAE39FC789197FBA6FD20" box="[929,1059,692,718]" pageId="11" pageNumber="12" value="1991-03-21">21.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFDDAE39FE7891FFFE70FD18" box="[417,501,732,758]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
; 
<materialsCitation id="3B7A3CF5FFDDAE39FDDD91FFFB33FCF0" ID-GBIF-Occurrence="2236573129" collectingDate="1986-02-15" county="Quebrada Pita" elevation="15" latitude="8.641944" location="Holzenthal" longLatPrecision="21" longitude="-83.193054" municipality="Golfito" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas">
<emphasis id="B966EABAFFDDAE39FDDD91FFFD16FD18" bold="true" box="[516,659,732,758]" pageId="11" pageNumber="12">
<collectingRegion id="49D6F84AFFDDAE39FDDD91FFFD16FD18" box="[516,659,732,758]" country="Costa Rica" name="Puntarenas" pageId="11" pageNumber="12">Puntarenas</collectingRegion>
</emphasis>
: 
<collectingCounty id="62CC4E24FFDDAE39FD7D91FFFCD5FD18" box="[676,848,732,758]" pageId="11" pageNumber="12">Quebrada Pita</collectingCounty>
, ca. 
<quantity id="4CEA9B4DFFDDAE39FC5191FFFC40FD18" box="[904,965,732,758]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="11" pageNumber="12" unit="km" value="3.0">3 km</quantity>
(air) W 
<collectingMunicipality id="6BC9ACD2FFDDAE39FBF691FFFB02FD18" box="[1071,1159,732,758]" pageId="11" pageNumber="12">Golfito</collectingMunicipality>
, 
<geoCoordinate id="EE26506FFFDDAE39FB4D91FFFAA3FD18" box="[1172,1318,732,758]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="8.641944">08°38’31”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDDAE39FED19027FE2EFCF0" box="[264,427,772,798]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-83.193054">083°11’35”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDDAE39FE369027FDBDFCF0" box="[495,568,772,798]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="m" value="15.0">
<elevation id="003FD19BFFDDAE39FE369027FDBDFCF0" box="[495,568,772,798]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="m" value="15.0">15 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDDAE39FD979027FD42FCF0" box="[590,711,772,798]" pageId="11" pageNumber="12" value="1986-02-15">
<collectingDate id="EFE8E980FFDDAE39FD979027FD42FCF0" box="[590,711,772,798]" pageId="11" pageNumber="12" value="1986-02-15">15.ii.1986</collectingDate>
</date>
, 
<location id="8ECD6073FFDDAE39FD049027FCE5FCF0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FD049027FCE5FCF0" box="[733,864,772,798]" county="Quebrada Pita" latitude="8.641944" longLatPrecision="21" longitude="-83.193054" municipality="Golfito" name="Holzenthal" pageId="11" pageNumber="12" stateProvince="Puntarenas">Holzenthal</location>
, 
<location id="8ECD6073FFDDAE39FCAF9027FC41FCF0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FCAF9027FC41FCF0" box="[886,964,772,798]" county="Quebrada Pita" latitude="8.641944" longLatPrecision="21" longitude="-83.193054" municipality="Golfito" name="Morse" pageId="11" pageNumber="12" stateProvince="Puntarenas">Morse</location>
, 
<location id="8ECD6073FFDDAE39FC039027FB9CFCF0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FC039027FB9CFCF0" box="[986,1049,772,798]" county="Quebrada Pita" latitude="8.641944" longLatPrecision="21" longitude="-83.193054" municipality="Golfito" name="Fasth" pageId="11" pageNumber="12" stateProvince="Puntarenas">Fasth</location>
— 
<specimenCount id="9D14FD21FFDDAE39FB809027FB33FCF0" box="[1113,1206,772,798]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDDAE39FB159027FD7BFC80" ID-GBIF-Occurrence="2236573154" collectingDate="1990-05-18" collectingDateMax="1990-05-20" collectingDateMin="1990-05-18" collectorName="Holzenthal &amp; Blahnik" elevation="35" latitude="9.775" location="Quebrada Bonita" longLatPrecision="21" longitude="-84.604996" municipality="Reserva Biologica Carara" pageId="11" pageNumber="12" specimenCount="2" specimenCount-male="2" stateProvince="Puntarenas">
<collectingMunicipality id="6BC9ACD2FFDDAE39FB159027FE50FCA8" pageId="11" pageNumber="12">Reserva Biológica Carara</collectingMunicipality>
, 
<location id="8ECD6073FFDDAE39FE3A900FFD35FCA8" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FE3A900FFD35FCA8" box="[483,688,812,838]" latitude="9.775" longLatPrecision="21" longitude="-84.604996" municipality="Reserva Biologica Carara" name="Quebrada Bonita" pageId="11" pageNumber="12" stateProvince="Puntarenas">Quebrada Bonita</location>
, 
<geoCoordinate id="EE26506FFFDDAE39FD67900FFCD4FCA8" box="[702,849,812,838]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="9.775">09°46’30”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDDAE39FC86900FFB81FCA8" box="[863,1028,812,838]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-84.604996">084°36’18”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDDAE39FBE0900FFBFFFCA8" box="[1081,1146,812,838]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="m" value="35.0">
<elevation id="003FD19BFFDDAE39FBE0900FFBFFFCA8" box="[1081,1146,812,838]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="m" value="35.0">35 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDDAE39FB50900FFAA3FCA8" box="[1161,1318,812,838]" pageId="11" pageNumber="12" value="1990-05-18" valueMax="1990-05-20" valueMin="1990-05-18">
<collectingDate id="EFE8E980FFDDAE39FB50900FFAA3FCA8" box="[1161,1318,812,838]" pageId="11" pageNumber="12" value="1990-05-18" valueMax="1990-05-20" valueMin="1990-05-18">18­20.v.1990</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDDAE39FED19077FE0DFC80" box="[264,392,852,878]" pageId="11" pageNumber="12">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFDDAE39FE779077FD8EFC80" box="[430,523,852,878]" pageId="11" pageNumber="12">Blahnik</collectorName>
— 
<specimenCount id="9D14FD21FFDDAE39FDEE9077FD14FC80" box="[567,657,852,878]" pageId="11" pageNumber="12" type="male">2 males</specimenCount>
(pinned)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFDDAE39FCD39077FC73FC80" ID-GBIF-Occurrence="2236573166" box="[778,1014,852,878]" collectingDate="1990-05-18" collectingDateMax="1990-05-20" collectingDateMin="1990-05-18" collectorName="Holzenthal &amp; Blahnik" elevation="35" isTailEnumeration="true" latitude="9.775" location="Quebrada Bonita" longLatPrecision="21" longitude="-84.604996" municipality="Reserva Biologica Carara" pageId="11" pageNumber="12" specimenCount="4" specimenCount-male="4" stateProvince="Puntarenas">
<specimenCount id="9D14FD21FFDDAE39FCD39077FCE1FC80" box="[778,868,852,878]" pageId="11" pageNumber="12" type="male">4 males</specimenCount>
(in alcohol)
</materialsCitation>
, 
<specimenCount id="9D14FD21FFDDAE39FBDB9077FB03FC80" box="[1026,1158,852,878]" pageId="11" pageNumber="12" type="female">22 females</specimenCount>
; 
<materialsCitation id="3B7A3CF5FFDDAE39FB489077FC17FC78" ID-GBIF-Occurrence="2236573130" collectingDate="1991-03-11" county="Holzenthal" location="Huisman" municipality="Munoz" pageId="11" pageNumber="12" specimenCount="42" specimenCount-male="42" stateProvince="Puntarenas">
same, except 
<date id="FFAC1068FFDDAE39FED1905FFE0DFC78" box="[264,392,892,918]" pageId="11" pageNumber="12" value="1991-03-11">
<collectingDate id="EFE8E980FFDDAE39FED1905FFE0DFC78" box="[264,392,892,918]" pageId="11" pageNumber="12" value="1991-03-11">11.iii.1991</collectingDate>
</date>
, 
<collectingCounty id="62CC4E24FFDDAE39FE4F905FFD9FFC78" box="[406,538,892,918]" pageId="11" pageNumber="12">Holzenthal</collectingCounty>
, 
<collectingMunicipality id="6BC9ACD2FFDDAE39FDFE905FFDF8FC78" box="[551,637,892,918]" pageId="11" pageNumber="12">Muñoz</collectingMunicipality>
, 
<location id="8ECD6073FFDDAE39FD52905FFD70FC78" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FD52905FFD70FC78" box="[651,757,892,918]" county="Holzenthal" municipality="Munoz" name="Huisman" pageId="11" pageNumber="12" stateProvince="Puntarenas">Huisman</location>
— 
<specimenCount id="9D14FD21FFDDAE39FCFD905FFC17FC78" box="[804,914,892,918]" pageId="11" pageNumber="12" type="male">42 males</specimenCount>
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDDAE39FC79905FFCE4FC50" ID-GBIF-Occurrence="2236573152" elevation="230" latitude="9.68" location="Rio Negro" longLatPrecision="21" longitude="-84.39389" pageId="11" pageNumber="12" specimenCount="1" stateProvince="San Jose">
<emphasis id="B966EABAFFDDAE39FC79905FFB88FC78" bold="true" box="[928,1037,892,918]" pageId="11" pageNumber="12">
<collectingRegion id="49D6F84AFFDDAE39FC79905FFB88FC78" box="[928,1037,892,918]" country="Costa Rica" name="San Jose" pageId="11" pageNumber="12">San José</collectingRegion>
</emphasis>
: 
<location id="8ECD6073FFDDAE39FBC6905FFB1BFC78" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FBC6905FFB1BFC78" box="[1055,1182,892,918]" latitude="9.68" longLatPrecision="21" longitude="-84.39389" name="Rio Negro" pageId="11" pageNumber="12" stateProvince="San Jose">Río Negro</location>
, 
<quantity id="4CEA9B4DFFDDAE39FB75905FFB7AFC78" box="[1196,1279,892,918]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="11" pageNumber="12" unit="km" value="3.5">3.5 km</quantity>
SE jct. rt. 239, 
<geoCoordinate id="EE26506FFFDDAE39FE4E9087FDACFC50" box="[407,553,932,958]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="9.68">09°40’48”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDDAE39FDE19087FD59FC50" box="[568,732,932,958]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-84.39389">084°23’38”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDDAE39FCC89087FCE4FC50" box="[785,865,932,958]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="12" unit="m" value="230.0">
<elevation id="003FD19BFFDDAE39FCC89087FCE4FC50" box="[785,865,932,958]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="11" pageNumber="12" unit="m" value="230.0">230 m</elevation>
</quantity>
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFDDAE39FCB69087FD23FBD8" ID-GBIF-Occurrence="2236573151" collectingDate="1986-03-21" collectingDateMax="1991-03-13" collectingDateMin="1986-03-21" collectionCode="NMNH, INBIO" collectorName="Holzenthal &amp; Fasth" county="Reserva Biologica Carara" elevation="160" latitude="9.768888" location="Sur" longLatPrecision="21" longitude="-84.53111" municipality="Rio" pageId="11" pageNumber="12" specimenCount="18" specimenCount-female="1" specimenCount-male="17" stateProvince="San Jose">
<date id="FFAC1068FFDDAE39FCB69087FC74FC50" box="[879,1009,932,958]" pageId="11" pageNumber="12" value="1986-03-21">
<collectingDate id="EFE8E980FFDDAE39FCB69087FC74FC50" box="[879,1009,932,958]" pageId="11" pageNumber="12" value="1986-03-21">21.iii.1986</collectingDate>
</date>
, 
<collectorName id="26E7537EFFDDAE39FBD99087FB05FC50" box="[1024,1152,932,958]" pageId="11" pageNumber="12">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFDDAE39FB729087FB6FFC50" box="[1195,1258,932,958]" pageId="11" pageNumber="12">Fasth</collectorName>
— 
<specimenCount id="9D14FD21FFDDAE39FAC59087FEC9FC08" pageId="11" pageNumber="12" type="male">8 males</specimenCount>
(
<collectionCode id="ED03AE6DFFDDAE39FE8290EFFE3EFC08" box="[347,443,972,998]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="11" pageNumber="12" type="Museum">NMNH</collectionCode>
), 
<specimenCount id="9D14FD21FFDDAE39FE1690EFFDA5FC08" box="[463,544,972,998]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
, 
<specimenCount id="9D14FD21FFDDAE39FDF590EFFD14FC08" box="[556,657,972,998]" pageId="11" pageNumber="12" type="female">1 female</specimenCount>
(
<collectionCode id="ED03AE6DFFDDAE39FD7990EFFD7DFC08" box="[672,760,972,998]" pageId="11" pageNumber="12">INBIO</collectionCode>
); 
<collectingCounty id="62CC4E24FFDDAE39FCD590EFFBBEFC08" box="[780,1083,972,998]" pageId="11" pageNumber="12">Reserva Biológica Carara</collectingCounty>
, 
<collectingMunicipality id="6BC9ACD2FFDDAE39FB9E90EFFBF4FC08" box="[1095,1137,972,998]" pageId="11" pageNumber="12">Río</collectingMunicipality>
del 
<location id="8ECD6073FFDDAE39FB7A90EFFB48FC08" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FB7A90EFFB48FC08" box="[1187,1229,972,998]" county="Reserva Biologica Carara" latitude="9.768888" longLatPrecision="21" longitude="-84.53111" municipality="Rio" name="Sur" pageId="11" pageNumber="12" stateProvince="San Jose">Sur</location>
, 
<quantity id="4CEA9B4DFFDDAE39FB0090EFFAAEFC08" box="[1241,1323,972,998]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="11" pageNumber="12" unit="km" value="1.5">1.5 km</quantity>
(rd) S 
<location id="8ECD6073FFDDAE39FE8790D7FE2AFBE0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FE8790D7FE2AFBE0" box="[350,431,1012,1038]" county="Reserva Biologica Carara" latitude="9.768888" longLatPrecision="21" longitude="-84.53111" municipality="Rio" name="Carara" pageId="11" pageNumber="12" stateProvince="San Jose">Carara</location>
, 
<geoCoordinate id="EE26506FFFDDAE39FE1990D7FDD7FBE0" box="[448,594,1012,1038]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="9.768888">09°46’08”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDDAE39FDBD90D7FC8CFBE0" box="[612,777,1012,1038]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDDAE39FC9D90D7FC12FBE0" box="[836,919,1012,1038]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="160.0">
<elevation id="003FD19BFFDDAE39FC9D90D7FC12FBE0" box="[836,919,1012,1038]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="11" pageNumber="12" unit="m" value="160.0">160 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDDAE39FC7190D7FBACFBE0" box="[936,1065,1012,1038]" pageId="11" pageNumber="12" value="1991-03-13">
<collectingDate id="EFE8E980FFDDAE39FC7190D7FBACFBE0" box="[936,1065,1012,1038]" pageId="11" pageNumber="12" value="1991-03-13">13.iii.1991</collectingDate>
</date>
, 
<location id="8ECD6073FFDDAE39FBE290D7FB3AFBE0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FBE290D7FB3AFBE0" box="[1083,1215,1012,1038]" county="Reserva Biologica Carara" latitude="9.768888" longLatPrecision="21" longitude="-84.53111" municipality="Rio" name="Holzenthal" pageId="11" pageNumber="12" stateProvince="San Jose">Holzenthal</location>
, 
<location id="8ECD6073FFDDAE39FB0990D7FAA3FBE0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FB0990D7FAA3FBE0" box="[1232,1318,1012,1038]" county="Reserva Biologica Carara" latitude="9.768888" longLatPrecision="21" longitude="-84.53111" municipality="Rio" name="Munoz" pageId="11" pageNumber="12" stateProvince="San Jose">Muñoz</location>
, 
<location id="8ECD6073FFDDAE39FED1973FFEF7FBD8" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FED1973FFEF7FBD8" box="[264,370,1052,1078]" county="Reserva Biologica Carara" latitude="9.768888" longLatPrecision="21" longitude="-84.53111" municipality="Rio" name="Huisman" pageId="11" pageNumber="12" stateProvince="San Jose">Huisman</location>
— 
<specimenCount id="9D14FD21FFDDAE39FE70973FFD8CFBD8" box="[425,521,1052,1078]" pageId="11" pageNumber="12" type="male">8 males</specimenCount>
(in alcohol)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFDDAE39FD61973FFC27FBD8" ID-GBIF-Occurrence="2236573124" box="[696,930,1052,1078]" collectingDate="1986-03-21" collectingDateMax="1991-03-13" collectingDateMin="1986-03-21" collectionCode="NMNH, INBIO" collectorName="Holzenthal &amp; Fasth" county="Reserva Biologica Carara" elevation="160" isTailEnumeration="true" latitude="9.768888" location="Sur" longLatPrecision="21" longitude="-84.53111" municipality="Rio" pageId="11" pageNumber="12" specimenCount="7" specimenCount-female="7" stateProvince="San Jose">
<specimenCount id="9D14FD21FFDDAE39FD61973FFCABFBD8" box="[696,814,1052,1078]" pageId="11" pageNumber="12" type="female">7 females</specimenCount>
(pinned)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFDDAE39FC6A973FFDDCFB68" ID-GBIF-Occurrence="2236573162" collectingDate="1991-03-14" county="Reserva Biologica Carara" elevation="200" latitude="9.778055" location="Holzenthal" longLatPrecision="21" longitude="-84.53111" municipality="Rio Carara in Carara" pageId="11" pageNumber="12" specimenCount="1" specimenCount-male="1" stateProvince="San Jose">
<collectingCounty id="62CC4E24FFDDAE39FC6A973FFB6BFBD8" box="[947,1262,1052,1078]" pageId="11" pageNumber="12">Reserva Biológica Carara</collectingCounty>
, 
<collectingMunicipality id="6BC9ACD2FFDDAE39FAD9973FFE4AFBB0" pageId="11" pageNumber="12">Río Carara in Carara</collectingMunicipality>
, 
<geoCoordinate id="EE26506FFFDDAE39FE049767FDEBFBB0" box="[477,622,1092,1118]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="9.778055">09°46’41”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDDAE39FDA59767FCA4FBB0" box="[636,801,1092,1118]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDDAE39FC8D9767FC26FBB0" box="[852,931,1092,1118]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="200.0">
<elevation id="003FD19BFFDDAE39FC8D9767FC26FBB0" box="[852,931,1092,1118]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="11" pageNumber="12" unit="m" value="200.0">200 m</elevation>
</quantity>
, 
<date id="FFAC1068FFDDAE39FC699767FBB4FBB0" box="[944,1073,1092,1118]" pageId="11" pageNumber="12" value="1991-03-14">
<collectingDate id="EFE8E980FFDDAE39FC699767FBB4FBB0" box="[944,1073,1092,1118]" pageId="11" pageNumber="12" value="1991-03-14">14.iii.1991</collectingDate>
</date>
, 
<location id="8ECD6073FFDDAE39FBE69767FB46FBB0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FBE69767FB46FBB0" box="[1087,1219,1092,1118]" county="Reserva Biologica Carara" latitude="9.778055" longLatPrecision="21" longitude="-84.53111" municipality="Rio Carara in Carara" name="Holzenthal" pageId="11" pageNumber="12" stateProvince="San Jose">Holzenthal</location>
, 
<location id="8ECD6073FFDDAE39FB099767FAA3FBB0" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FB099767FAA3FBB0" box="[1232,1318,1092,1118]" county="Reserva Biologica Carara" latitude="9.778055" longLatPrecision="21" longitude="-84.53111" municipality="Rio Carara in Carara" name="Munoz" pageId="11" pageNumber="12" stateProvince="San Jose">Muñoz</location>
, 
<location id="8ECD6073FFDDAE39FED1974FFEF7FB68" LSID="urn:lsid:plazi:treatment:03BB87BEFFDEAE3EFED19535FEEAFB18:8ECD6073FFDDAE39FED1974FFEF7FB68" box="[264,370,1132,1158]" county="Reserva Biologica Carara" latitude="9.778055" longLatPrecision="21" longitude="-84.53111" municipality="Rio Carara in Carara" name="Huisman" pageId="11" pageNumber="12" stateProvince="San Jose">Huisman</location>
— 
<specimenCount id="9D14FD21FFDDAE39FE44974FFE69FB68" box="[413,492,1132,1158]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
(pinned)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFDDAE39FDBC974FFD4BFB68" ID-GBIF-Occurrence="2236573131" box="[613,718,1132,1158]" collectingDate="1991-03-14" county="Reserva Biologica Carara" elevation="200" isTailEnumeration="true" latitude="9.778055" location="Holzenthal" longLatPrecision="21" longitude="-84.53111" municipality="Rio Carara in Carara" pageId="11" pageNumber="12" specimenCount="18" specimenCount-male="18" stateProvince="San Jose">
<specimenCount id="9D14FD21FFDDAE39FDBC974FFD4BFB68" box="[613,718,1132,1158]" pageId="11" pageNumber="12" type="male">18 males</specimenCount>
</materialsCitation>
(in alcohol).
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FED197FAFD83FB1D" blockId="11.[264,518,1241,1307]" box="[264,518,1241,1267]" pageId="11" pageNumber="12">
<taxonomicName id="4C124D2BFFDDAE39FED197FAFD83FB1D" authority="Flint" authorityName="Flint" box="[264,518,1241,1267]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cana">
<emphasis id="B966EABAFFDDAE39FED197FAFD83FB1D" bold="true" box="[264,518,1241,1267]" pageId="11" pageNumber="12">
<emphasis id="B966EABAFFDDAE39FED197FAFE47FB1D" bold="true" box="[264,450,1241,1267]" italics="true" pageId="11" pageNumber="12">Protoptila cana</emphasis>
Flint
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FED19622FECEFAF5" blockId="11.[264,518,1241,1307]" box="[264,331,1281,1307]" pageId="11" pageNumber="12">
<figureCitation id="13292A2DFFDDAE39FED19622FECEFAF5" box="[264,331,1281,1307]" captionStart="FIGURE 7" captionStartId="14.[264,367,1720,1744]" captionTargetBox="[283,1307,681,1693]" captionTargetId="figure@14.[280,1307,681,1693]" captionTargetPageId="14" captionText="FIGURE 7. Protoptila cana, Flint. Male genitalia: A—lateral (inset: apex of segment X, caudal); B—segment VIII, ventral; C—phallic complex, lateral." httpUri="https://zenodo.org/record/2646283/files/figure.png" pageId="11" pageNumber="12">Fig. 7</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FED19672FE23FA86" blockId="11.[264,422,1361,1384]" box="[264,422,1361,1384]" pageId="11" pageNumber="12">
<bibRefCitation id="EF834B59FFDDAE39FED19672FEFFFA86" author="Flint, O. S., Jr." box="[264,378,1361,1384]" pageId="11" pageNumber="12" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint 1974</bibRefCitation>
: 18.
</paragraph>
<paragraph id="8BAD36A8FFDDAE39FED19682FBDFF945" blockId="11.[264,1323,1441,1867]" pageId="11" pageNumber="12">
This species is very similar to 
<taxonomicName id="4C124D2BFFDDAE39FD5F9682FC80FA54" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[646,773,1441,1467]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="chitaria" status="sp. nov.">
<emphasis id="B966EABAFFDDAE39FD5F9682FD10FA54" box="[646,661,1441,1466]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. 
<emphasis id="B966EABAFFDDAE39FD719682FC80FA54" box="[680,773,1441,1466]" italics="true" pageId="11" pageNumber="12">chitaria</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFDDAE39FCCE9682FCD9FA55" box="[791,860,1441,1467]" pageId="11" pageNumber="12" rank="species">n. sp.</taxonomicNameLabel>
, and the two undoubtedly are closely related. The most distinctive difference between them is the shape of the apex of the ventral margin of segment IX, which is closely associated with and partially fused to the ventral apex of sternum VIII. In 
<taxonomicName id="4C124D2BFFDDAE39FD5A953AFD5CF9DC" box="[643,729,1561,1587]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="cana">
<emphasis id="B966EABAFFDDAE39FD5A953AFD17F9DC" box="[643,658,1561,1586]" italics="true" pageId="11" pageNumber="12">P</emphasis>
. 
<emphasis id="B966EABAFFDDAE39FD79953AFD5CF9DC" box="[672,729,1561,1586]" italics="true" pageId="11" pageNumber="12">cana</emphasis>
</taxonomicName>
this apex is more elongate than the ventral apex of sternum VIII and nearly straight, whereas in 
<taxonomicName id="4C124D2BFFDDAE39FCC69562FC13F9B4" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[799,918,1601,1626]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="11" pageNumber="12" phylum="Arthropoda" rank="species" species="chitaria">
<emphasis id="B966EABAFFDDAE39FCC69562FC13F9B4" box="[799,918,1601,1626]" italics="true" pageId="11" pageNumber="12">P. chitaria</emphasis>
</taxonomicName>
it is shorter than the ventral apex of sternum VIII and curved apically. The apices of the apical sections of tergum X are also shaped slightly different in the two species, as shown in the illustrations.
</paragraph>
<paragraph id="8BAD36A8FFDDAE3EFEE1959AFEEAFB18" blockId="11.[264,1323,1441,1867]" lastBlockId="12.[264,1325,284,1270]" lastPageId="12" lastPageNumber="13" pageId="11" pageNumber="12">
Material examined. 
<emphasis id="B966EABAFFDDAE39FDFA959AFD5AF93D" bold="true" box="[547,735,1721,1747]" pageId="11" pageNumber="12">
<collectingCountry id="F3057638FFDDAE39FDFA959AFD5AF93D" box="[547,735,1721,1747]" name="Costa Rica" pageId="11" pageNumber="12">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFDDAE39FD2F959AFCE5F93D" bold="true" box="[758,864,1721,1747]" pageId="11" pageNumber="12">
<collectingRegion id="49D6F84AFFDDAE39FD2F959AFCE5F93D" box="[758,864,1721,1747]" country="Costa Rica" name="Alajuela" pageId="11" pageNumber="12">Alajuela</collectingRegion>
</emphasis>
: 
<quantity id="4CEA9B4DFFDDAE39FCAE959AFC55F93D" box="[887,976,1721,1747]" metricMagnitude="3" metricUnit="m" metricValue="7.6" pageId="11" pageNumber="12" unit="km" value="7.6">7.6 km</quantity>
E Artenas, Rio Grande de Tarcoles river bed, 
<date id="FFAC1068FFDDAE39FE3595C2FDF1F915" box="[492,628,1761,1787]" pageId="11" pageNumber="12" value="1987-12-15">15.xii.1987</date>
, Burdick — 
<specimenCount id="9D14FD21FFDDAE39FCD795C2FCDBF915" box="[782,862,1761,1787]" pageId="11" pageNumber="12" type="male">1 male</specimenCount>
(in alcohol); 
<emphasis id="B966EABAFFDDAE39FBD895C2FBE2F915" bold="true" box="[1025,1127,1761,1787]" pageId="11" pageNumber="12">
<collectingRegion id="49D6F84AFFDDAE39FBD895C2FBE2F915" box="[1025,1127,1761,1787]" country="Costa Rica" name="Cartago" pageId="11" pageNumber="12">Cartago</collectingRegion>
</emphasis>
: Río Chitaría, rt 10, 10 km NW Río Reventazón, 
<geoCoordinate id="EE26506FFFDDAE39FD6D942AFCC2F8CD" box="[692,839,1801,1827]" direction="north" orientation="latitude" pageId="11" pageNumber="12" precision="15" value="9.92">09°55’12”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDDAE39FC80942AFC7BF8CD" box="[857,1022,1801,1827]" direction="west" orientation="longitude" pageId="11" pageNumber="12" precision="15" value="-83.60389">083°36’14”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDDAE39FBE4942AFB14F8CD" box="[1085,1169,1801,1827]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="11" pageNumber="12" unit="m" value="740.0">740 m</quantity>
, 
<date id="FFAC1068FFDDAE39FB7D942AFAA0F8CD" box="[1188,1317,1801,1827]" pageId="11" pageNumber="12" value="1991-03-21">21.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFDDAE39FD739412FC8AF8A5" box="[682,783,1841,1867]" pageId="11" pageNumber="12" type="male">2 males</specimenCount>
; Río Reventazón, CATIE along Sendero E spaveles, 
<geoCoordinate id="EE26506FFFDAAE3EFEA5923FFD88FED8" box="[380,525,284,310]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="9.893055">09°53’35”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFDC5923FFD45FED8" box="[540,704,284,310]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-83.651115">083°39’04”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFD2D923FFCC6FED8" box="[756,835,284,310]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="m" value="500.0">500 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFC88923FFC56FED8" box="[849,979,284,310]" pageId="12" pageNumber="13" value="1991-03-22">22.iii.1991</date>
, Muñoz­Quesada — 
<specimenCount id="9D14FD21FFDAAE3EFB0B923FFAA3FED8" box="[1234,1318,284,310]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
, 
<specimenCount id="9D14FD21FFDAAE3EFED19267FE09FEB0" box="[264,396,324,350]" pageId="12" pageNumber="13" type="female">23 females</specimenCount>
; 
<emphasis id="B966EABAFFDAAE3EFE4E9260FDAFFEB3" bold="true" box="[407,554,323,349]" pageId="12" pageNumber="13">
<collectingRegion id="49D6F84AFFDAAE3EFE4E9260FDAFFEB3" box="[407,554,323,349]" country="Costa Rica" name="Guanacaste" pageId="12" pageNumber="13">Guanacaste</collectingRegion>
</emphasis>
: Quebrada Garcia, 
<quantity id="4CEA9B4DFFDAAE3EFCD69267FCEAFEB0" box="[783,879,324,350]" metricMagnitude="4" metricUnit="m" metricValue="1.06" pageId="12" pageNumber="13" unit="km" value="10.6">10.6 km</quantity>
ENE Quebrada Grande, 
<geoCoordinate id="EE26506FFFDAAE3EFB4D9267FAA3FEB0" box="[1172,1318,324,350]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="10.861945">10°51’43”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFED1924FFE29FE68" box="[264,428,364,390]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-85.428055">085°25’41”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFE38924FFDB4FE68" box="[481,561,364,390]" metricMagnitude="2" metricUnit="m" metricValue="4.7" pageId="12" pageNumber="13" unit="m" value="470.0">470 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFDE7924FFD34FE68" box="[574,689,364,390]" pageId="12" pageNumber="13" value="1986-03-08">8.iii.1986</date>
, Holzenthal &amp; Fasth — 
<specimenCount id="9D14FD21FFDAAE3EFC02924FFBBEFE68" box="[987,1083,364,390]" pageId="12" pageNumber="13" type="male">7 males</specimenCount>
; Río Tizate, 
<quantity id="4CEA9B4DFFDAAE3EFB0F924FFAAEFE68" box="[1238,1323,364,390]" metricMagnitude="3" metricUnit="m" metricValue="7.2" pageId="12" pageNumber="13" unit="km" value="7.2">7.2 km</quantity>
NE Cañas Dulces, 
<geoCoordinate id="EE26506FFFDAAE3EFE3592B7FDFBFE40" box="[492,638,404,430]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="10.773055">10°46’23”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFD5492B7FCB7FE40" box="[653,818,404,430]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-85.44889">085°26’56”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFCBF92B7FC31FE40" box="[870,948,404,430]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="12" pageNumber="13" unit="m" value="275.0">275 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFC1A92B7FBC6FE40" box="[963,1091,404,430]" pageId="12" pageNumber="13" value="1986-06-28">28.vi.1986</date>
, Holzenthal, Heyn, Armitage — 
<specimenCount id="9D14FD21FFDAAE3EFE7A929FFE78FE38" box="[419,509,444,470]" pageId="12" pageNumber="13" type="male">4 males</specimenCount>
(in alcohol); Parque Nacional 
<collectingRegion id="49D6F84AFFDAAE3EFCBF929FFC75FE38" box="[870,1008,444,470]" country="Costa Rica" name="Guanacaste" pageId="12" pageNumber="13">Guanacaste</collectingRegion>
, Quebrada Alcornoque, El Hacha, 
<geoCoordinate id="EE26506FFFDAAE3EFEB892C7FE76FE10" box="[353,499,484,510]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="11.008889">11°00’32”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFE2792C7FD26FE10" box="[510,675,484,510]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-85.57694">085°34’37”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFD0B92C7FC9AFE10" box="[722,799,484,510]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="m" value="250.0">250 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFCF292C7FC36FE10" box="[811,947,484,510]" pageId="12" pageNumber="13" value="1987-07-26">26.vii.1987</date>
, Holzenthal, Morse, Clausen — 
<specimenCount id="9D14FD21FFDAAE3EFED1912FFEEFFDC8" box="[264,362,524,550]" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
, 
<specimenCount id="9D14FD21FFDAAE3EFEA5912FFE62FDC8" box="[380,487,524,550]" pageId="12" pageNumber="13" type="female">1 female</specimenCount>
(in alcohol) (INBIO); 
<emphasis id="B966EABAFFDAAE3EFCD19128FCDEFDCB" bold="true" box="[776,859,523,549]" pageId="12" pageNumber="13">
<collectingRegion id="49D6F84AFFDAAE3EFCD19128FCDEFDCB" box="[776,859,523,549]" country="Costa Rica" name="Limon" pageId="12" pageNumber="13">Limón</collectingRegion>
</emphasis>
: Río Uatsi, ca. 
<quantity id="4CEA9B4DFFDAAE3EFBF1912FFBECFDC8" box="[1064,1129,524,550]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="12" pageNumber="13" unit="km" value="8.0">8 km</quantity>
(air) W Bribri, 
<geoCoordinate id="EE26506FFFDAAE3EFED19117FE1EFDA0" box="[264,411,564,590]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="9.62">09°37’12”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFE7E9117FDC8FDA0" box="[423,589,564,590]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-82.9">082°54’00”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFDA79117FD38FDA0" box="[638,701,564,590]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="m" value="60.0">60 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFD139117FCC9FDA0" box="[714,844,564,590]" pageId="12" pageNumber="13" value="1987-03-25">25.iii.1987</date>
, Holzenthal, Hamilton, Heyn — 
<specimenCount id="9D14FD21FFDAAE3EFB0C9117FAADFDA0" box="[1237,1320,564,590]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
; 
<emphasis id="B966EABAFFDAAE3EFED19178FE12FD9B" bold="true" box="[264,407,603,629]" pageId="12" pageNumber="13">
<collectingRegion id="49D6F84AFFDAAE3EFED19178FE12FD9B" box="[264,407,603,629]" country="Costa Rica" name="Puntarenas" pageId="12" pageNumber="13">Puntarenas</collectingRegion>
</emphasis>
: Río Ceibo, route 2 ca. 
<quantity id="4CEA9B4DFFDAAE3EFD2A917FFCBEFD98" box="[755,827,604,630]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="12" pageNumber="13" unit="km" value="6.0">6 km</quantity>
W rd to Buenos Aires, 
<geoCoordinate id="EE26506FFFDAAE3EFB4D917FFAA2FD98" box="[1172,1319,604,630]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="9.148889">09°08’56”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFED191A7FE28FD70" box="[264,429,644,670]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-83.376945">083°22’37”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFE3791A7FDC1FD70" box="[494,580,644,670]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="12" pageNumber="13" unit="m" value="250.0">250 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFD8091A7FD57FD70" box="[601,722,644,670]" pageId="12" pageNumber="13" value="1986-02-20">20.ii.1986</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFDAAE3EFB8691A7FB43FD70" box="[1119,1222,644,670]" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
; Parque Nacional Corcovado, Est. Sirena, Río Camaronal, 
<geoCoordinate id="EE26506FFFDAAE3EFCB3918FFC79FD28" box="[874,1020,684,710]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="8.481944">08°28’55”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFBD5918FFB35FD28" box="[1036,1200,684,710]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-83.58889">083°35’20”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFB3E918FFAADFD28" box="[1255,1320,684,710]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="12" pageNumber="13" unit="m" value="30.0">30 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFED191F7FE03FD00" box="[264,390,724,750]" pageId="12" pageNumber="13" value="1989-04-13">13.iv.1989</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFDAAE3EFD0491F7FCB8FD00" box="[733,829,724,750]" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
(in alcohol); same, except 
<geoCoordinate id="EE26506FFFDAAE3EFB4C91F7FAADFD00" box="[1173,1320,724,750]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="8.481111">08°28’52”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFED191DFFE29FCF8" box="[264,428,764,790]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-83.593895">083°35’38”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFE3E91DFFD98FCF8" box="[487,541,764,790]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="12" pageNumber="13" unit="m" value="5.0">5 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFDF791DFFD57FCF8" box="[558,722,764,790]" pageId="12" pageNumber="13" value="1989-04-12" valueMax="1989-04-13" valueMin="1989-04-12">12­13.iv.1989</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFDAAE3EFBF091DFFB0CFCF8" box="[1065,1161,764,790]" pageId="12" pageNumber="13" type="male">8 males</specimenCount>
(in alcohol); Reserva Biológica Carara, Quebrada Bonita, 
<geoCoordinate id="EE26506FFFDAAE3EFCE89007FC41FCD0" box="[817,964,804,830]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="9.775">09°46’30”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFC0D9007FBFCFCD0" box="[980,1145,804,830]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-84.604996">084°36’18”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFB6B9007FB70FCD0" box="[1202,1269,804,830]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="13" unit="m" value="35.0">35 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFADF9007FEFBFC88" pageId="12" pageNumber="13" value="1990-05-18" valueMax="1990-05-20" valueMin="1990-05-18">18­ 20.v.1990</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFDAAE3EFD10906FFCA3FC88" box="[713,806,844,870]" pageId="12" pageNumber="13" type="male">2 males</specimenCount>
(pinned), 
<specimenCount id="9D14FD21FFDAAE3EFC7C906FFB86FC88" box="[933,1027,844,870]" pageId="12" pageNumber="13" type="male">6 males</specimenCount>
(in alcohol), 
<specimenCount id="9D14FD21FFDAAE3EFB73906FFAA9FC88" box="[1194,1324,844,870]" pageId="12" pageNumber="13" type="female">22 females</specimenCount>
(pinned); same, except 
<date id="FFAC1068FFDAAE3EFDFB9057FD27FC60" box="[546,674,884,910]" pageId="12" pageNumber="13" value="1991-03-11">11.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFDAAE3EFB9D9057FB20FC60" box="[1092,1189,884,910]" pageId="12" pageNumber="13" type="male">6 males</specimenCount>
; 
<emphasis id="B966EABAFFDAAE3EFB6A9057FAA6FC60" bold="true" box="[1203,1315,884,910]" pageId="12" pageNumber="13">
<collectingRegion id="49D6F84AFFDAAE3EFB6A9057FAA6FC60" box="[1203,1315,884,910]" country="Costa Rica" name="San Jose" pageId="12" pageNumber="13">San José</collectingRegion>
</emphasis>
: Río Negro, 
<quantity id="4CEA9B4DFFDAAE3EFE4290BFFE77FC58" box="[411,498,924,950]" metricMagnitude="3" metricUnit="m" metricValue="3.5" pageId="12" pageNumber="13" unit="km" value="3.5">3.5 km</quantity>
SE jct. rt. 239, 
<geoCoordinate id="EE26506FFFDAAE3EFD1890BFFCD6FC58" box="[705,851,924,950]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="9.68">09°40’48”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFCBD90BFFB8CFC58" box="[868,1033,924,950]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-84.39389">084°23’38”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFB9B90BFFB11FC58" box="[1090,1172,924,950]" metricMagnitude="2" metricUnit="m" metricValue="2.3" pageId="12" pageNumber="13" unit="m" value="230.0">230 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFB7D90BFFAA3FC58" box="[1188,1318,924,950]" pageId="12" pageNumber="13" value="1986-03-21">21.iii.1986</date>
, Holzenthal &amp; Fasth — 
<specimenCount id="9D14FD21FFDAAE3EFDC390E7FDECFC30" box="[538,617,964,990]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(in alcohol); Reserva Biológica Carara, Río del Sur, 
<quantity id="4CEA9B4DFFDAAE3EFB0090E7FAAEFC30" box="[1241,1323,964,990]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="12" pageNumber="13" unit="km" value="1.5">1.5 km</quantity>
(rd) S Carara, 
<geoCoordinate id="EE26506FFFDAAE3EFE1990CFFDD7FBE8" box="[448,594,1004,1030]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="9.768888">09°46’08”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFDBD90CFFC8CFBE8" box="[612,777,1004,1030]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFC9D90CFFC12FBE8" box="[836,919,1004,1030]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="12" pageNumber="13" unit="m" value="160.0">160 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFC7190CFFBACFBE8" box="[936,1065,1004,1030]" pageId="12" pageNumber="13" value="1991-03-13">13.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFDAAE3EFE799737FE74FBC0" box="[416,497,1044,1070]" pageId="12" pageNumber="13" type="male">1 male</specimenCount>
(pinned), 
<specimenCount id="9D14FD21FFDAAE3EFDB59737FD42FBC0" box="[620,711,1044,1070]" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
(in alcohol); Reserva Biológica Carara, Río Carara in Carara, 
<geoCoordinate id="EE26506FFFDAAE3EFE40971FFDAEFBB8" box="[409,555,1084,1110]" direction="north" orientation="latitude" pageId="12" pageNumber="13" precision="15" value="9.778055">09°46’41”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFDAAE3EFD9A971FFD62FBB8" box="[579,743,1084,1110]" direction="west" orientation="longitude" pageId="12" pageNumber="13" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFDAAE3EFCF4971FFC03FBB8" box="[813,902,1084,1110]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="12" pageNumber="13" unit="m" value="200.0">200 m</quantity>
, 
<date id="FFAC1068FFDAAE3EFC45971FFB9BFBB8" box="[924,1054,1084,1110]" pageId="12" pageNumber="13" value="1991-03-14">14.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFDAAE3EFE709747FD8CFB90" box="[425,521,1124,1150]" pageId="12" pageNumber="13" type="male">3 males</specimenCount>
(in alcohol) (NMNH). 
<emphasis id="B966EABAFFDAAE3EFCEB9747FBBCFB90" bold="true" box="[818,1081,1124,1150]" pageId="12" pageNumber="13">
<collectingCountry id="F3057638FFDAAE3EFCEB9747FC32FB90" box="[818,951,1124,1150]" name="Panama" pageId="12" pageNumber="13">PANAMA</collectingCountry>
: 
<collectingCountry id="F3057638FFDAAE3EFC109747FBB1FB90" box="[969,1076,1124,1150]" name="Panama" pageId="12" pageNumber="13">Panama</collectingCountry>
:
</emphasis>
Canal Zone, Barro Colorado Island, Snyder­Molino trail, marker 3, 
<date id="FFAC1068FFDAAE3EFC8A97AFFBA5FB48" box="[851,1056,1164,1190]" pageId="12" pageNumber="13" value="1988-07-13" valueMax="1988-09-20" valueMin="1988-07-13">13.vii­20.ix.1988</date>
, Wolda — 
<specimenCount id="9D14FD21FFDAAE3EFB7497AFFA88FB48" box="[1197,1293,1164,1190]" pageId="12" pageNumber="13" type="male">9 males</specimenCount>
, 
<specimenCount id="9D14FD21FFDAAE3EFAC597AFFEE6FB20" pageId="12" pageNumber="13" type="female">7 females</specimenCount>
(in alcohol); same, except 
<date id="FFAC1068FFDAAE3EFD6C9797FC05FB20" box="[693,896,1204,1230]" pageId="12" pageNumber="13" value="1990-06-27" valueMax="1990-11-13" valueMin="1990-06-27">27.vi.­13.xi.1990</date>
, Wolda, — 
<specimenCount id="9D14FD21FFDAAE3EFBC19797FBFCFB20" box="[1048,1145,1204,1230]" pageId="12" pageNumber="13" type="male">5 males</specimenCount>
, 
<specimenCount id="9D14FD21FFDAAE3EFB509797FB7BFB20" box="[1161,1278,1204,1230]" pageId="12" pageNumber="13" type="female">6 females</specimenCount>
(in alcohol).
</paragraph>
</subSubSection>
</treatment>
</document>