treatments-xml/data/03/BB/87/03BB87BEFFCEAE16FED19615FC9BFE5D.xml

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<mods:title id="AE35AE080B933F6231E12ADAF3A55111">The caddisfly genus Protoptila in Costa Rica (Trichoptera: Glossosomatidae)</mods:title>
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<mods:namePart id="493FAF651BD5B9714E259BB3ABDE7BE2">Blahnik, Roger J.</mods:namePart>
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<paragraph id="8BAD36A8FFCEAE2AFED19615FD07FABE" blockId="24.[264,642,1334,1395]" box="[264,642,1334,1360]" pageId="24" pageNumber="25">
<heading id="D0E581C4FFCEAE2AFED19615FD07FABE" bold="true" box="[264,642,1334,1360]" fontSize="11" level="1" pageId="24" pageNumber="25" reason="1">
<taxonomicName id="4C124D2BFFCEAE2AFED19615FD07FABE" ID-CoL="4N6SX" authority="orotina Flint" authorityName="Flint" box="[264,642,1334,1360]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="orotina" subSpecies="orotina">
<emphasis id="B966EABAFFCEAE2AFED19615FD07FABE" bold="true" box="[264,642,1334,1360]" pageId="24" pageNumber="25">
<emphasis id="B966EABAFFCEAE2AFED19615FDBAFABE" bold="true" box="[264,575,1334,1360]" italics="true" pageId="24" pageNumber="25">Protoptila orotina orotina</emphasis>
Flint
</emphasis>
</taxonomicName>
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<paragraph id="8BAD36A8FFCEAE2AFED1967AFEDEFA9D" blockId="24.[264,642,1334,1395]" box="[264,347,1369,1395]" pageId="24" pageNumber="25">
<figureCitation id="13292A2DFFCEAE2AFED1967AFEDEFA9D" box="[264,347,1369,1395]" captionStart="FIGURE 14" captionStartId="25.[264,368,1495,1519]" captionTargetBox="[94,1298,286,1467]" captionTargetId="figure@25.[295,1298,329,1467]" captionTargetPageId="25" captionText="FIGURE 14. Protoptila orotina orotina, Flint. Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral." httpUri="https://zenodo.org/record/2646297/files/figure.svg" pageId="24" pageNumber="25">Fig. 14</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFCEAE2AFED19687FE23FA55" blockId="24.[264,422,1444,1467]" box="[264,422,1444,1467]" pageId="24" pageNumber="25">
<bibRefCitation id="EF834B59FFCEAE2AFED19687FEFFFA55" author="Flint, O. S., Jr." box="[264,378,1444,1467]" pageId="24" pageNumber="25" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint 1974</bibRefCitation>
: 13.
</paragraph>
<paragraph id="8BAD36A8FFCEAE2AFED196CFFDD2F880" blockId="24.[264,1323,1516,1902]" pageId="24" pageNumber="25">
This species is most similar to 
<taxonomicName id="4C124D2BFFCEAE2AFD4B96CFFCEDF9EB" authorityName="Flint" authorityYear="1974" box="[658,872,1516,1542]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="orotina" subSpecies="raposa">
<emphasis id="B966EABAFFCEAE2AFD4B96CFFD24F9EB" box="[658,673,1516,1541]" italics="true" pageId="24" pageNumber="25">P</emphasis>
. 
<emphasis id="B966EABAFFCEAE2AFD6D96CFFCEDF9EB" box="[692,872,1516,1541]" italics="true" pageId="24" pageNumber="25">orotina raposa</emphasis>
</taxonomicName>
, 
<emphasis id="B966EABAFFCEAE2AFCA596CFFB16F9EB" box="[892,1171,1516,1541]" italics="true" pageId="24" pageNumber="25">
<taxonomicName id="4C124D2BFFCEAE2AFCA596CFFC66F9EB" box="[892,995,1516,1541]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="ixtala">P. ixtala</taxonomicName>
, and 
<taxonomicName id="4C124D2BFFCEAE2AFBF496CFFB16F9EB" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[1069,1171,1516,1541]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="bribri">P. bribri</taxonomicName>
</emphasis>
. All have a similarly developed phallicata, which is elongate and very angularly bent in the middle. The different species differ most in the development of the apex of this structure. In 
<taxonomicName id="4C124D2BFFCEAE2AFACC951FFE41F993" authority="orotina" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="subSpecies" species="orotina" subSpecies="orotina">
<emphasis id="B966EABAFFCEAE2AFACC951FFE41F993" italics="true" pageId="24" pageNumber="25">P. orotina orotina</emphasis>
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it is somewhat enlarged apically and has a subacute apicoventral projection. Diagnostic differences in the two subspecies of 
<taxonomicName id="4C124D2BFFCEAE2AFC1795AFFBC6F94B" authorityName="Flint" authorityYear="1974" box="[974,1091,1676,1702]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="24" pageNumber="25" phylum="Arthropoda" rank="species" species="orotina">
<emphasis id="B966EABAFFCEAE2AFC1795AFFC59F94B" box="[974,988,1676,1701]" italics="true" pageId="24" pageNumber="25">P</emphasis>
. 
<emphasis id="B966EABAFFCEAE2AFC3795AFFBC6F94B" box="[1006,1091,1676,1701]" italics="true" pageId="24" pageNumber="25">orotina</emphasis>
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were illustrated by 
<bibRefCitation id="EF834B59FFCEAE2AFED19597FE1EF920" author="Flint, O. S., Jr." box="[264,411,1716,1742]" pageId="24" pageNumber="25" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint (1974)</bibRefCitation>
. Considering that active selection appears to be occurring within this group for structures at the apex of the phallus, it seems possible that the two subspecies may actually represent different species. There are also minor differences among the species in the shapes of the apical section of tergum X, the parameres, and the apex of sternum VIII, as shown in the illustrations.
</paragraph>
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<paragraph id="8BAD36A8FFCFAE2BFED196F4FD0AF9FC" blockId="25.[264,1322,1495,1554]" pageId="25" pageNumber="26">
<emphasis id="B966EABAFFCFAE2BFED196F4FE1DFA01" bold="true" box="[264,408,1495,1519]" pageId="25" pageNumber="26">FIGURE 14.</emphasis>
<taxonomicName id="4C124D2BFFCFAE2BFE7996FBFD7AFA01" authority="orotina, Flint." authorityName="Flint." box="[416,767,1496,1519]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="25" pageNumber="26" phylum="Arthropoda" rank="subSpecies" species="orotina" subSpecies="orotina">
<emphasis id="B966EABAFFCFAE2BFE7996FBFD32FA01" box="[416,695,1496,1519]" italics="true" pageId="25" pageNumber="26">Protoptila orotina orotina</emphasis>
, Flint.
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Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral.
</paragraph>
</caption>
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Material examined. 
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: 
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: Río Tizate, 
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NE Cañas Dulces, 
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, 
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, el. 
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, 
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<collectingDate id="EFE8E980FFCFAE2BFCE9954BFC35F96C" box="[816,944,1640,1666]" pageId="25" pageNumber="26" value="1986-06-28">28.vi.1986</collectingDate>
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, Holzenthal, Heyn, Armitage — 
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</materialsCitation>
; 
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<collectingRegion id="49D6F84AFFCFAE2BFEB095B3FE39F944" box="[361,444,1680,1706]" country="Costa Rica" name="Limon" pageId="25" pageNumber="26">Limón</collectingRegion>
</emphasis>
: 
<collectingCounty id="62CC4E24FFCFAE2BFE1495B3FCC9F944" box="[461,844,1680,1706]" pageId="25" pageNumber="26">Reserva Biológica Hitoy­Cerere</collectingCounty>
, 
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, 
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. Miramar, 
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, 
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, el. 
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<elevation id="003FD19BFFCFAE2BFE04959BFD99F93C" box="[477,540,1720,1746]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="25" pageNumber="26" unit="m" value="90.0">90 m</elevation>
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, 
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<collectingDate id="EFE8E980FFCFAE2BFDF0959BFD54F93C" box="[553,721,1720,1746]" pageId="25" pageNumber="26" value="1987-03-23" valueMax="1987-03-24" valueMin="1987-03-23">23­24.iii.1987</collectingDate>
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, Holzenthal, Hamilton, Heyn — 
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, 
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</materialsCitation>
; 
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<collectingRegion id="49D6F84AFFCFAE2BFED195C3FE12F914" box="[264,407,1760,1786]" country="Costa Rica" name="Puntarenas" pageId="25" pageNumber="26">Puntarenas</collectingRegion>
</emphasis>
: 
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, route 2 ca. 
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W rd to 
<location id="8ECD6073FFCFAE2BFC1695C3FBF9F914" LSID="urn:lsid:plazi:treatment:03BB87BEFFCEAE16FED19615FC9BFE5D:8ECD6073FFCFAE2BFC1695C3FBF9F914" box="[975,1148,1760,1786]" country="Costa Rica" county="Reserva Biologica Hitoy-Cerere" latitude="9.671111" longLatPrecision="21" longitude="-84.53111" municipality="Rio Cerere" name="Buenos Aires" pageId="25" pageNumber="26" stateProvince="Guanacaste">Buenos Aires</location>
, 
<geoCoordinate id="EE26506FFFCFAE2BFB4A95C3FAA0F914" box="[1171,1317,1760,1786]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="9.148889">09°08’56”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCFAE2BFED1942BFE28F8CC" box="[264,429,1800,1826]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="15" value="-83.376945">083°22’37”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCFAE2BFE07942BFDA9F8CC" box="[478,556,1800,1826]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="25" pageNumber="26" unit="m" value="250.0">
<elevation id="003FD19BFFCFAE2BFE07942BFDA9F8CC" box="[478,556,1800,1826]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="25" pageNumber="26" unit="m" value="250.0">250 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCFAE2BFDE0942BFD36F8CC" box="[569,691,1800,1826]" pageId="25" pageNumber="26" value="1986-02-20">
<collectingDate id="EFE8E980FFCFAE2BFDE0942BFD36F8CC" box="[569,691,1800,1826]" pageId="25" pageNumber="26" value="1986-02-20">20.ii.1986</collectingDate>
</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFCFAE2BFBC1942BFB15F8CC" box="[1048,1168,1800,1826]" pageId="25" pageNumber="26" type="male">103 males</specimenCount>
(in alcohol)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFCFAE28FED19413FC01FED8" ID-GBIF-Occurrence="2417205427" collectingDate="1986-02-20" collectingDateMax="1991-03-14" collectingDateMin="1986-02-20" country="Costa Rica" county="Reserva Biologica Hitoy-Cerere" elevation="275" lastPageId="26" lastPageNumber="27" latitude="9.671111" location="Est" longLatPrecision="21" longitude="-84.53111" municipality="Rio Cerere" pageId="25" pageNumber="26" specimenCount="130" specimenCount-female="1" specimenCount-male="129" stateProvince="Guanacaste">
Río Rincón, 
<quantity id="4CEA9B4DFFCFAE2BFE739413FD86F8A4" box="[426,515,1840,1866]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="25" pageNumber="26" unit="km" value="6.5">
<locationDeviation id="9A1F2F8AFFCFAE2BFE739413FD86F8A4" box="[426,515,1840,1866]" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="25" pageNumber="26" unit="km" value="6.5">6.5 km</locationDeviation>
</quantity>
(air) S Rincón, 
<geoCoordinate id="EE26506FFFCFAE2BFD009413FCEEF8A4" box="[729,875,1840,1866]" direction="north" orientation="latitude" pageId="25" pageNumber="26" precision="15" value="8.638056">08°38’17”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCFAE2BFCA49413FBA7F8A4" box="[893,1058,1840,1866]" direction="west" orientation="longitude" pageId="25" pageNumber="26" precision="15" value="-83.48">083°28’48”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCFAE2BFB869413FB26F8A4" box="[1119,1187,1840,1866]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="m" value="20.0">
<elevation id="003FD19BFFCFAE2BFB869413FB26F8A4" box="[1119,1187,1840,1866]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="25" pageNumber="26" unit="m" value="20.0">20 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCFAE2BFB6F9413FAA3F8A4" box="[1206,1318,1840,1866]" pageId="25" pageNumber="26" value="1987-04-07">
<collectingDate id="EFE8E980FFCFAE2BFB6F9413FAA3F8A4" box="[1206,1318,1840,1866]" pageId="25" pageNumber="26" value="1987-04-07">7.iv.1987</collectingDate>
</date>
, Holzenthal, Hamilton, Heyn — 
<specimenCount id="9D14FD21FFCCAE28FD57923FFD6EFED8" box="[654,747,284,310]" pageId="26" pageNumber="27" type="male">8 males</specimenCount>
(in alcohol)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFCCAE28FC4D923FFC4BFE68" ID-GBIF-Occurrence="2417205423" collectingDate="1986-02-20" collectingDateMax="1991-03-14" collectingDateMin="1986-02-20" country="Costa Rica" county="Reserva Biologica Hitoy-Cerere" elevation="275" lastPageId="26" lastPageNumber="27" latitude="9.671111" location="Est" longLatPrecision="21" longitude="-84.53111" municipality="Rio Cerere" pageId="26" pageNumber="26" specimenCount="130" specimenCount-female="1" specimenCount-male="129" stateProvince="Guanacaste">
Río Singrí, ca 
<quantity id="4CEA9B4DFFCCAE28FB9E923FFB03FED8" box="[1095,1158,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="km" value="2.0">
<locationDeviation id="9A1F2F8AFFCCAE28FB9E923FFB03FED8" box="[1095,1158,284,310]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="km" value="2.0">2 km</locationDeviation>
</quantity>
(air) S Finca Helechales, 
<geoCoordinate id="EE26506FFFCCAE28FE439267FDA9FEB0" box="[410,556,324,350]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="9.056945">09°03’25”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCCAE28FDE29267FD65FEB0" box="[571,736,324,350]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-83.08194">083°04’55”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCCAE28FCCC9267FCE0FEB0" box="[789,869,324,350]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="26" pageNumber="27" unit="m" value="720.0">
<elevation id="003FD19BFFCCAE28FCCC9267FCE0FEB0" box="[789,869,324,350]" metricMagnitude="2" metricUnit="m" metricValue="7.2" pageId="26" pageNumber="27" unit="m" value="720.0">720 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCCAE28FCAD9267FC68FEB0" box="[884,1005,324,350]" pageId="26" pageNumber="27" value="1986-02-21">
<collectingDate id="EFE8E980FFCCAE28FCAD9267FC68FEB0" box="[884,1005,324,350]" pageId="26" pageNumber="27" value="1986-02-21">21.ii.1986</collectingDate>
</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFCCAE28FEEF924FFE1CFE68" box="[310,409,364,390]" pageId="26" pageNumber="27" type="male">9 males</specimenCount>
(in alcohol), 
<specimenCount id="9D14FD21FFCCAE28FD88924FFD31FE68" box="[593,692,364,390]" pageId="26" pageNumber="27" type="male">2 males</specimenCount>
(in alcohol) (INBIO)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFCCAE28FC3A924FFDFAFE38" ID-GBIF-Occurrence="2417205428" collectingDate="1986-02-20" collectingDateMax="1991-03-14" collectingDateMin="1986-02-20" country="Costa Rica" county="Reserva Biologica Hitoy-Cerere" elevation="275" lastPageId="26" lastPageNumber="27" latitude="9.671111" location="Est" longLatPrecision="21" longitude="-84.53111" municipality="Rio Cerere" pageId="26" pageNumber="26" specimenCount="130" specimenCount-female="1" specimenCount-male="129" stateProvince="Guanacaste">
Reserva Biológica Carara, Quebrada Bonita, 
<geoCoordinate id="EE26506FFFCCAE28FE3792B7FD05FE40" box="[494,640,404,430]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="9.775">09°46’30”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCCAE28FD4D92B7FCBDFE40" box="[660,824,404,430]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-84.604996">084°36’18”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCCAE28FCA192B7FC3AFE40" box="[888,959,404,430]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="27" unit="m" value="35.0">
<elevation id="003FD19BFFCCAE28FCA192B7FC3AFE40" box="[888,959,404,430]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="26" pageNumber="27" unit="m" value="35.0">35 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCCAE28FC0A92B7FBEAFE40" box="[979,1135,404,430]" pageId="26" pageNumber="27" value="1990-05-18" valueMax="1990-05-20" valueMin="1990-05-18">
<collectingDate id="EFE8E980FFCCAE28FC0A92B7FBEAFE40" box="[979,1135,404,430]" pageId="26" pageNumber="27" value="1990-05-18" valueMax="1990-05-20" valueMin="1990-05-18">18­20.v.1990</collectingDate>
</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFCCAE28FE48929FFE69FE38" box="[401,492,444,470]" pageId="26" pageNumber="27" type="male">2 males</specimenCount>
(in alcohol)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFCCAE28FD559298FD2FFDC8" ID-GBIF-Occurrence="2417205424" collectingDate="1986-02-20" collectingDateMax="1991-03-14" collectingDateMin="1986-02-20" country="Costa Rica" county="Reserva Biologica Hitoy-Cerere" elevation="275" lastPageId="26" lastPageNumber="27" latitude="9.671111" location="Est" longLatPrecision="21" longitude="-84.53111" municipality="Rio Cerere" pageId="26" pageNumber="26" specimenCount="130" specimenCount-female="1" specimenCount-male="129" stateProvince="Guanacaste">
<emphasis id="B966EABAFFCCAE28FD559298FD7CFE3B" bold="true" box="[652,761,443,469]" pageId="26" pageNumber="27">
<collectingRegion id="49D6F84AFFCCAE28FD559298FD7CFE3B" box="[652,761,443,469]" country="Costa Rica" name="San Jose" pageId="26" pageNumber="27">San José</collectingRegion>
</emphasis>
: Reserva Biológica Carara, Río del Sur, 
<quantity id="4CEA9B4DFFCCAE28FB01929FFAAFFE38" box="[1240,1322,444,470]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="km" value="1.5">
<locationDeviation id="9A1F2F8AFFCCAE28FB01929FFAAFFE38" box="[1240,1322,444,470]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="26" pageNumber="27" unit="km" value="1.5">1.5 km</locationDeviation>
</quantity>
(rd) S Carara, 
<geoCoordinate id="EE26506FFFCCAE28FE1992C7FDD7FE10" box="[448,594,484,510]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="9.768888">09°46’08”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCCAE28FDBD92C7FC8CFE10" box="[612,777,484,510]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCCAE28FC9D92C7FC12FE10" box="[836,919,484,510]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="27" unit="m" value="160.0">
<elevation id="003FD19BFFCCAE28FC9D92C7FC12FE10" box="[836,919,484,510]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="26" pageNumber="27" unit="m" value="160.0">160 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCCAE28FC7192C7FBACFE10" box="[936,1065,484,510]" pageId="26" pageNumber="27" value="1991-03-13">
<collectingDate id="EFE8E980FFCCAE28FC7192C7FBACFE10" box="[936,1065,484,510]" pageId="26" pageNumber="27" value="1991-03-13">13.iii.1991</collectingDate>
</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFCCAE28FE76912FFD82FDC8" box="[431,519,524,550]" pageId="26" pageNumber="27" type="male">1 male</specimenCount>
(in alcohol)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFCCAE28FD66912FFE38FD98" ID-GBIF-Occurrence="2417205426" collectingDate="1986-02-20" collectingDateMax="1991-03-14" collectingDateMin="1986-02-20" country="Costa Rica" county="Reserva Biologica Hitoy-Cerere" elevation="275" lastPageId="26" lastPageNumber="27" latitude="9.671111" location="Est" longLatPrecision="21" longitude="-84.53111" municipality="Rio Cerere" pageId="26" pageNumber="26" specimenCount="130" specimenCount-female="1" specimenCount-male="129" stateProvince="Guanacaste">
Reserva Biológica Carara, Río Carara in Carara, 
<geoCoordinate id="EE26506FFFCCAE28FED19117FE1FFDA0" box="[264,410,564,590]" direction="north" orientation="latitude" pageId="26" pageNumber="27" precision="15" value="9.778055">09°46’41”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCCAE28FE729117FDD5FDA0" box="[427,592,564,590]" direction="west" orientation="longitude" pageId="26" pageNumber="27" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCCAE28FD539117FD59FDA0" box="[650,732,564,590]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="m" value="200.0">
<elevation id="003FD19BFFCCAE28FD539117FD59FDA0" box="[650,732,564,590]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="26" pageNumber="27" unit="m" value="200.0">200 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCCAE28FD349117FCEAFDA0" box="[749,879,564,590]" pageId="26" pageNumber="27" value="1991-03-14">
<collectingDate id="EFE8E980FFCCAE28FD349117FCEAFDA0" box="[749,879,564,590]" pageId="26" pageNumber="27" value="1991-03-14">14.iii.1991</collectingDate>
</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFCCAE28FAC59117FEC4FD98" pageId="26" pageNumber="27" type="male">1 male</specimenCount>
(NMNH).
</materialsCitation>
</paragraph>
<caption id="DF6D6620FFCCAE28FED195D8FCF5F8DB" ID-DOI="http://doi.org/10.5281/zenodo.3492053" ID-Zenodo-Dep="3492053" httpUri="https://zenodo.org/record/3492053/files/figure.png" pageId="26" pageNumber="27" startId="26.[264,367,1787,1811]" targetBox="[340,1248,695,1755]" targetPageId="26">
<paragraph id="8BAD36A8FFCCAE28FED195D8FCF5F8DB" blockId="26.[264,1322,1787,1845]" pageId="26" pageNumber="27">
<emphasis id="B966EABAFFCCAE28FED195D8FE12F8FD" bold="true" box="[264,407,1787,1811]" pageId="26" pageNumber="27">FIGURE 15.</emphasis>
<taxonomicName id="4C124D2BFFCCAE28FE4795DFFD31F8FD" authority=", Flint." authorityName="Flint." box="[414,692,1788,1811]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="26" pageNumber="27" phylum="Arthropoda" rank="species" species="spirifera">
<emphasis id="B966EABAFFCCAE28FE4795DFFDE8F8FD" box="[414,621,1788,1811]" italics="true" pageId="26" pageNumber="27">Protoptila spirifera</emphasis>
, Flint.
</taxonomicName>
Male genitalia: A—lateral; B—segment VIII, ventral; C— phallic complex, lateral (inset: apex of phallicata, dorsal).
</paragraph>
</caption>
<paragraph id="8BAD36A8FFCDAE29FED1923FFDB7FEDB" blockId="27.[264,562,283,345]" box="[264,562,283,310]" pageId="27" pageNumber="28">
<taxonomicName id="4C124D2BFFCDAE29FED1923FFDB7FEDB" authority="Flint" authorityName="Flint" box="[264,562,283,310]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="spirifera">
<emphasis id="B966EABAFFCDAE29FED1923FFDB7FEDB" bold="true" box="[264,562,283,310]" pageId="27" pageNumber="28">
<emphasis id="B966EABAFFCDAE29FED1923FFE6AFED8" bold="true" box="[264,495,284,310]" italics="true" pageId="27" pageNumber="28">Protoptila spirifera</emphasis>
Flint
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FED1921CFEDFFEB7" blockId="27.[264,562,283,345]" box="[264,346,319,345]" pageId="27" pageNumber="28">
<figureCitation id="13292A2DFFCDAE29FED1921CFEDFFEB7" box="[264,346,319,345]" captionStart="FIGURE 15" captionStartId="26.[264,367,1787,1811]" captionTargetBox="[340,1248,695,1755]" captionTargetId="figure@26.[340,1248,689,1765]" captionTargetPageId="26" captionText="FIGURE 15. Protoptila spirifera, Flint. Male genitalia: A—lateral; B—segment VIII, ventral; C— phallic complex, lateral (inset: apex of phallicata, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.3492053" httpUri="https://zenodo.org/record/3492053/files/figure.png" pageId="27" pageNumber="28">Fig. 15</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FED192ADFE23FE4B" blockId="27.[264,422,398,421]" box="[264,422,398,421]" pageId="27" pageNumber="28">
<bibRefCitation id="EF834B59FFCDAE29FED192ADFEFFFE4B" author="Flint, O. S., Jr." box="[264,378,398,421]" pageId="27" pageNumber="28" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint 1974</bibRefCitation>
: 14.
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FED192F4FBC3FD7F" blockId="27.[264,1323,471,737]" pageId="27" pageNumber="28">
The species was named for the spiral paramere spines and seems to be a member of the 
<emphasis id="B966EABAFFCDAE29FED192DCFE0BFDF6" box="[264,398,511,536]" italics="true" pageId="27" pageNumber="28">condylifera</emphasis>
complex of 
<bibRefCitation id="EF834B59FFCDAE29FDF592DCFD45FDF7" author="Flint, O. S., Jr." box="[556,704,511,537]" pageId="27" pageNumber="28" pagination="1 - 113" refId="ref12856" refString="Flint, O. S., Jr. (1991) Studies in Neotropical caddisflies, XLV: the taxonomy, phenology, and faunistics of the Trichoptera of Antioquia, Colombia. Smithsonian Contributions to Zoology, 520, 1 - 113." type="journal article" year="1991">Flint (1991)</bibRefCitation>
. As in those species the phallicata has a relatively long, narrow, arched “neck” and inflated apex and the ventral apex of segment VIII is long and narrow and not at all bifid apically. It differs from its relatives in the shape of the apical section of tergum X, which has hook­like apicomesal processes.
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FEE191BCFD68FD0F" blockId="27.[264,1323,471,737]" pageId="27" pageNumber="28">
Material examined. 
<emphasis id="B966EABAFFCDAE29FDC591BDFCD0FD56" bold="true" box="[540,853,670,696]" pageId="27" pageNumber="28">
<collectingCountry id="F3057638FFCDAE29FDC591BDFD53FD56" box="[540,726,670,696]" name="Costa Rica" pageId="27" pageNumber="28">COSTA RICA</collectingCountry>
: 
<collectingRegion id="49D6F84AFFCDAE29FD3C91BDFCD5FD56" box="[741,848,670,696]" country="Costa Rica" name="Cartago" pageId="27" pageNumber="28">Cartago</collectingRegion>
:
</emphasis>
Ojo de Agua, rt. 2, km 75, 
<date id="FFAC1068FFCDAE29FB7F91BCFAA3FD57" box="[1190,1318,671,697]" pageId="27" pageNumber="28" value="1967-06-30">30.vi.1967</date>
, Flint &amp; Ortiz — male 
<typeStatus id="54A9880AFFCDAE29FDD591E4FDF4FD0F" box="[524,625,711,737]" pageId="27" pageNumber="28" type="paratype">paratype</typeStatus>
(NMNH).
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FED1901DFD2CFCB6" blockId="27.[264,681,830,891]" box="[264,681,830,856]" pageId="27" pageNumber="28">
<emphasis id="B966EABAFFCDAE29FED1901DFD2CFCB6" bold="true" box="[264,681,830,856]" pageId="27" pageNumber="28">
<taxonomicName id="4C124D2BFFCDAE29FED1901DFD8FFCB6" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[264,522,830,856]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="strepsicera" status="sp. nov.">
<emphasis id="B966EABAFFCDAE29FED1901DFD8FFCB6" bold="true" box="[264,522,830,856]" italics="true" pageId="27" pageNumber="28">Protoptila strepsicera</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFCDAE29FDC0901DFD2CFCB6" box="[537,681,830,856]" pageId="27" pageNumber="28" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FED19042FEDFFC95" blockId="27.[264,681,830,891]" box="[264,346,865,891]" pageId="27" pageNumber="28">
<figureCitation id="13292A2DFFCDAE29FED19042FEDFFC95" box="[264,346,865,891]" captionStart="FIGURE 16" captionStartId="28.[264,368,1567,1591]" captionTargetBox="[283,1317,522,1544]" captionTargetId="figure@28.[279,1317,515,1554]" captionTargetPageId="28" captionText="FIGURE 16. Protoptila strepsicera, new species. Male genitalia: A—lateral (inset: segment X, dorsal); B—segment VIII, ventral; C—phallic complex, lateral." figureDoi="http://doi.org/10.5281/zenodo.3492055" httpUri="https://zenodo.org/record/3492055/files/figure.png" pageId="27" pageNumber="28">Fig. 16</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FED19092FD61FB0D" blockId="27.[264,1325,945,1891]" pageId="27" pageNumber="28">
This species is most similar to 
<taxonomicName id="4C124D2BFFCDAE29FDB69092FD52FC24" box="[623,727,945,971]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tojana">
<emphasis id="B966EABAFFCDAE29FDB69092FDFBFC24" box="[623,638,945,970]" italics="true" pageId="27" pageNumber="28">P</emphasis>
. 
<emphasis id="B966EABAFFCDAE29FD559092FD52FC24" box="[652,727,945,970]" italics="true" pageId="27" pageNumber="28">tojana</emphasis>
</taxonomicName>
, especially in the structure and development of the apical sections of tergum X and the shape and development the ventral apex of sternum IX. However, the species differ greatly in the processes emanating from the posterodorsal margin of sternum VIII. In 
<taxonomicName id="4C124D2BFFCDAE29FD8E970AFD47FBAC" box="[599,706,1065,1090]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="tojana">
<emphasis id="B966EABAFFCDAE29FD8E970AFD47FBAC" box="[599,706,1065,1090]" italics="true" pageId="27" pageNumber="28">P. tojana</emphasis>
</taxonomicName>
they are very narrow and sometimes down­curved apically (straight in the specimen illustrated by 
<bibRefCitation id="EF834B59FFCDAE29FCEC9772FC3CFB85" author="Flint, O. S., Jr." box="[821,953,1105,1131]" pageId="27" pageNumber="28" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint, 1974</bibRefCitation>
), whereas in 
<taxonomicName id="4C124D2BFFCDAE29FB8A9772FB74FB84" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[1107,1265,1105,1130]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="strepsicera">
<emphasis id="B966EABAFFCDAE29FB8A9772FB74FB84" box="[1107,1265,1105,1130]" italics="true" pageId="27" pageNumber="28">P. strepsicera</emphasis>
</taxonomicName>
they are very prominent, enlarged basally and tapering apically. In 
<taxonomicName id="4C124D2BFFCDAE29FBEC975AFB58FB7C" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[1077,1245,1145,1170]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="27" pageNumber="28" phylum="Arthropoda" rank="species" species="strepsicera">
<emphasis id="B966EABAFFCDAE29FBEC975AFB58FB7C" box="[1077,1245,1145,1170]" italics="true" pageId="27" pageNumber="28">P. strepsicera</emphasis>
</taxonomicName>
these structures are also somewhat spirally curved, almost as in the horns of some antelopes, making the species very easy to identify.
</paragraph>
<paragraph id="8BAD36A8FFCDAE29FEE197D2FEF8FADD" blockId="27.[264,1325,945,1891]" pageId="27" pageNumber="28">Adult. Length of forewing: male 2.7–3.1. Color brown (in alcohol), wing markings indistinct.</paragraph>
<paragraph id="8BAD36A8FFCDAE2EFEE19662FB2DFE38" blockId="27.[264,1325,945,1891]" lastBlockId="28.[264,1323,284,470]" lastPageId="28" lastPageNumber="29" pageId="27" pageNumber="28">Male genitalia. Sternum VI process prominent, slightly curved, subtriangular, apex subacute. Tergum VIII posterior margin with row of elongate setae; sternum VIII produced ventrally, not bifurcate apically, broadly rounded, sclerotized and setose basally, membranous apically, apparently fused to sternum IX apicoventrally; dorsolateral margin of sternum VIII basally with very long, prominent, sinuous, apically acute process. Segment IX with anterior margin broadly rounded; sternum IX medially produced, in lateral view forming short, acute, strongly upturned process, weakly bifid mesally when viewed ventrally; posterolateral margin of segment IX without processes. Preanal appendages absent. Tergum X divided mesally to form sclerotized lateral parts with basal and apical sections; basal section subquadrate, about as long as wide, apical section short, divided apically to form acuminate dorsal projection and more elongate, sinuously upturned, apically acuminate ventral projection. Phallobase dorsally with large, semicircular, laterally compressed apodeme; ventrally with short, rod­like, articulated appendages with apical setae, appendages fitting into sclerotized pockets on ventral margin of phallobase; posteroventral margin of phallobase with short, paired, upturned, apically acute basal processes and also dorsolateral processes; dorsolateral processes short, apically truncate; paramere absent; phallicata very short, ventrally curved, with pair of short apicoventral spine­like processes; endophallus membranous, enlarged when everted; phallotremal sclerite spine­like, sclerotized apically, somewhat divided basally.</paragraph>
</subSubSection>
<caption id="DF6D6620FFCAAE2EFED1953CFC2BF9B7" ID-DOI="http://doi.org/10.5281/zenodo.3492055" ID-Zenodo-Dep="3492055" httpUri="https://zenodo.org/record/3492055/files/figure.png" pageId="28" pageNumber="29" startId="28.[264,368,1567,1591]" targetBox="[283,1317,522,1544]" targetPageId="28">
<paragraph id="8BAD36A8FFCAAE2EFED1953CFC2BF9B7" blockId="28.[264,1322,1567,1625]" pageId="28" pageNumber="29">
<emphasis id="B966EABAFFCAAE2EFED1953CFE1EF9D9" bold="true" box="[264,411,1567,1591]" pageId="28" pageNumber="29">FIGURE 16.</emphasis>
<taxonomicName id="4C124D2BFFCAAE2EFE7C9503FD14F9D9" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[421,657,1568,1591]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="28" pageNumber="29" phylum="Arthropoda" rank="species" species="strepsicera" status="sp. nov.">
<emphasis id="B966EABAFFCAAE2EFE7C9503FD14F9D9" box="[421,657,1568,1591]" italics="true" pageId="28" pageNumber="29">Protoptila strepsicera</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFCAAE2EFD7A9503FCAFF9D9" box="[675,810,1568,1591]" pageId="28" pageNumber="29" rank="species">new species</taxonomicNameLabel>
. Male genitalia: A—lateral (inset: segment X, dorsal); B—segment VIII, ventral; C—phallic complex, lateral.
</paragraph>
</caption>
<subSubSection id="C3086523FFCAAE11FEE195AEFAAFFE40" lastPageId="35" lastPageNumber="36" pageId="28" pageNumber="29" type="materials_examined">
<paragraph id="8BAD36A8FFCAAE2EFEE195AEFCA3F919" blockId="28.[264,1323,1677,1903]" pageId="28" pageNumber="29">
<materialsCitation id="3B7A3CF5FFCAAE2EFEE195AEFCA3F919" ID-GBIF-Occurrence="2236573139" collectingDate="1987-03-23" collectingDateMax="1987-03-24" collectingDateMin="1987-03-23" collectionCode="UMSP" country="Costa Rica" elevation="90" latitude="9.671111" location="Rio Cerere" longLatPrecision="21" longitude="-83.02806" municipality="Reserva Biologica Hitoy-Cerere" pageId="28" pageNumber="29" specimenCode="UMSP000208540" specimenCount="1" stateProvince="Limon" typeStatus="holotype">
<emphasis id="B966EABAFFCAAE2EFEE195AEFE7DF949" bold="true" box="[312,504,1677,1703]" pageId="28" pageNumber="29">
<typeStatus id="54A9880AFFCAAE2EFEE195AEFE2FF949" box="[312,426,1677,1703]" pageId="28" pageNumber="29" type="holotype">Holotype</typeStatus>
male
</emphasis>
: 
<emphasis id="B966EABAFFCAAE2EFDCB95AEFD54F949" bold="true" box="[530,721,1677,1703]" pageId="28" pageNumber="29">
<collectingCountry id="F3057638FFCAAE2EFDCB95AEFD54F949" box="[530,721,1677,1703]" name="Costa Rica" pageId="28" pageNumber="29">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFCAAE2EFD3295AEFCBBF949" bold="true" box="[747,830,1677,1703]" pageId="28" pageNumber="29">
<collectingRegion id="49D6F84AFFCAAE2EFD3295AEFCBBF949" box="[747,830,1677,1703]" country="Costa Rica" name="Limon" pageId="28" pageNumber="29">Limón</collectingRegion>
</emphasis>
: 
<collectingMunicipality id="6BC9ACD2FFCAAE2EFC8195AEFB6FF949" box="[856,1258,1677,1703]" pageId="28" pageNumber="29">Reserva Biológica Hitoy­Cerere</collectingMunicipality>
, 
<location id="8ECD6073FFCAAE2EFB2695AEFEDCF921" LSID="urn:lsid:plazi:treatment:03BB87BEFFCEAE16FED19615FC9BFE5D:8ECD6073FFCAAE2EFB2695AEFEDCF921" country="Costa Rica" latitude="9.671111" longLatPrecision="21" longitude="-83.02806" municipality="Reserva Biologica Hitoy-Cerere" name="Rio Cerere" pageId="28" pageNumber="29" stateProvince="Limon">Río Cerere</location>
, Est. Miramar, 
<geoCoordinate id="EE26506FFFCAAE2EFDC09596FD2FF921" box="[537,682,1717,1743]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="15" value="9.671111">09°40’16”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCAAE2EFD629596FCE5F921" box="[699,864,1717,1743]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="15" value="-83.02806">083°01’41”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCAAE2EFC4E9596FC5CF921" box="[919,985,1717,1743]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="m" value="90.0">
<elevation id="003FD19BFFCAAE2EFC4E9596FC5CF921" box="[919,985,1717,1743]" metricMagnitude="1" metricUnit="m" metricValue="9.0" pageId="28" pageNumber="29" unit="m" value="90.0">90 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCAAE2EFC309596FB17F921" box="[1001,1170,1717,1743]" pageId="28" pageNumber="29" value="1987-03-23" valueMax="1987-03-24" valueMin="1987-03-23">
<collectingDate id="EFE8E980FFCAAE2EFC309596FB17F921" box="[1001,1170,1717,1743]" pageId="28" pageNumber="29" value="1987-03-23" valueMax="1987-03-24" valueMin="1987-03-23">23­24.iii.1987</collectingDate>
</date>
, Holzenthal, Hamilton, Heyn (
<specimenCode id="DBB49ED3FFCAAE2EFE0D95FEFD2EF919" box="[468,683,1757,1783]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="28" pageNumber="29" type="University or college">UMSP000208540</specimenCode>
) (
<collectionCode id="ED03AE6DFFCAAE2EFD1A95FEFC9CF919" box="[707,793,1757,1783]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="28" pageNumber="29" type="University or college">UMSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAD36A8FFCAAE2EFEE19426FB6BF881" blockId="28.[264,1323,1677,1903]" pageId="28" pageNumber="29">
<emphasis id="B966EABAFFCAAE2EFEE19426FE30F8F1" bold="true" box="[312,437,1797,1823]" pageId="28" pageNumber="29">
<typeStatus id="54A9880AFFCAAE2EFEE19426FE30F8F1" box="[312,437,1797,1823]" pageId="28" pageNumber="29" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as 
<typeStatus id="54A9880AFFCAAE2EFDAD9426FD5EF8F1" box="[628,731,1797,1823]" pageId="28" pageNumber="29" type="holotype">holotype</typeStatus>
— 
<materialsCitation id="3B7A3CF5FFCAAE2EFCD69426FC5AF8F1" ID-GBIF-Occurrence="2236573138" box="[783,991,1797,1823]" collectionCode="UMSP" pageId="28" pageNumber="29" specimenCount="6" specimenCount-male="6" typeStatus="paratype">
<specimenCount id="9D14FD21FFCAAE2EFCD69426FCE8F8F1" box="[783,877,1797,1823]" pageId="28" pageNumber="29" type="male">6 males</specimenCount>
(
<collectionCode id="ED03AE6DFFCAAE2EFC589426FC52F8F1" box="[897,983,1797,1823]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="28" pageNumber="29" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFCAAE2EFC369426FB4DF8F1" ID-GBIF-Occurrence="2236573147" box="[1007,1224,1797,1823]" collectionCode="NMNH" pageId="28" pageNumber="29" specimenCount="2" specimenCount-male="2" typeStatus="paratype">
<specimenCount id="9D14FD21FFCAAE2EFC369426FBC9F8F1" box="[1007,1100,1797,1823]" pageId="28" pageNumber="29" type="male">2 males</specimenCount>
(
<collectionCode id="ED03AE6DFFCAAE2EFBB99426FB45F8F1" box="[1120,1216,1797,1823]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="28" pageNumber="29" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
, 
<materialsCitation id="3B7A3CF5FFCAAE2EFB019426FEF5F8A9" ID-GBIF-Occurrence="2236573164" collectionCode="INBIO" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<specimenCount id="9D14FD21FFCAAE2EFB019426FAAEF8F1" box="[1240,1323,1797,1823]" pageId="28" pageNumber="29" type="male">1 male</specimenCount>
(
<collectionCode id="ED03AE6DFFCAAE2EFEC9940EFEEDF8A9" box="[272,360,1837,1863]" pageId="28" pageNumber="29">INBIO</collectionCode>
)
</materialsCitation>
; 
<materialsCitation id="3B7A3CF5FFCAAE2EFE5D940EFB85F881" ID-GBIF-Occurrence="2236573149" collectingDate="1987-03-25" collectorName="W Bribri &amp; Holzenthal &amp; Hamilton &amp; Heyn" country="Costa Rica" elevation="60" latitude="9.62" location="Rio Uatsi" longLatPrecision="21" longitude="-82.9" pageId="28" pageNumber="29" specimenCount="1" stateProvince="Limon" typeStatus="paratype">
<emphasis id="B966EABAFFCAAE2EFE5D940EFDC5F8A9" bold="true" box="[388,576,1837,1863]" pageId="28" pageNumber="29">
<collectingCountry id="F3057638FFCAAE2EFE5D940EFDC5F8A9" box="[388,576,1837,1863]" name="Costa Rica" pageId="28" pageNumber="29">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFCAAE2EFD8F940EFD2CF8A9" bold="true" box="[598,681,1837,1863]" pageId="28" pageNumber="29">
<collectingRegion id="49D6F84AFFCAAE2EFD8F940EFD2CF8A9" box="[598,681,1837,1863]" country="Costa Rica" name="Limon" pageId="28" pageNumber="29">Limón</collectingRegion>
</emphasis>
: 
<location id="8ECD6073FFCAAE2EFD19940EFCBFF8A9" LSID="urn:lsid:plazi:treatment:03BB87BEFFCEAE16FED19615FC9BFE5D:8ECD6073FFCAAE2EFD19940EFCBFF8A9" box="[704,826,1837,1863]" country="Costa Rica" latitude="9.62" longLatPrecision="21" longitude="-82.9" name="Rio Uatsi" pageId="28" pageNumber="29" stateProvince="Limon">Río Uatsi</location>
, ca. 
<quantity id="4CEA9B4DFFCAAE2EFCA5940EFC3AF8A9" box="[892,959,1837,1863]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="28" pageNumber="29" unit="km" value="8.0">8 km</quantity>
(air) 
<collectorName id="26E7537EFFCAAE2EFBD7940EFB04F8A9" box="[1038,1153,1837,1863]" pageId="28" pageNumber="29">W Bribri</collectorName>
, 
<geoCoordinate id="EE26506FFFCAAE2EFB4D940EFAA0F8A9" box="[1172,1317,1837,1863]" direction="north" orientation="latitude" pageId="28" pageNumber="29" precision="15" value="9.62">09°37’12”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCAAE2EFED19476FE29F881" box="[264,428,1877,1903]" direction="west" orientation="longitude" pageId="28" pageNumber="29" precision="15" value="-82.9">082°54’00”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCAAE2EFE059476FD9EF881" box="[476,539,1877,1903]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="29" unit="m" value="60.0">
<elevation id="003FD19BFFCAAE2EFE059476FD9EF881" box="[476,539,1877,1903]" metricMagnitude="1" metricUnit="m" metricValue="6.0" pageId="28" pageNumber="29" unit="m" value="60.0">60 m</elevation>
</quantity>
, 
<date id="FFAC1068FFCAAE2EFDFF9476FD2DF881" box="[550,680,1877,1903]" pageId="28" pageNumber="29" value="1987-03-25">
<collectingDate id="EFE8E980FFCAAE2EFDFF9476FD2DF881" box="[550,680,1877,1903]" pageId="28" pageNumber="29" value="1987-03-25">25.iii.1987</collectingDate>
</date>
, 
<collectorName id="26E7537EFFCAAE2EFD6D9476FCBDF881" box="[692,824,1877,1903]" pageId="28" pageNumber="29">Holzenthal</collectorName>
, 
<collectorName id="26E7537EFFCAAE2EFC9A9476FC30F881" box="[835,949,1877,1903]" pageId="28" pageNumber="29">Hamilton</collectorName>
, 
<collectorName id="26E7537EFFCAAE2EFC189476FB85F881" box="[961,1024,1877,1903]" pageId="28" pageNumber="29">Heyn</collectorName>
</materialsCitation>
— 
<materialsCitation id="3B7A3CF5FFCAAE2EFBF59476FB6CF881" ID-GBIF-Occurrence="2236573148" box="[1068,1257,1877,1903]" collectionCode="UMSP" country="Costa Rica" pageId="28" pageNumber="29" specimenCount="1" specimenCount-male="1" stateProvince="Limon" typeStatus="paratype">
<specimenCount id="9D14FD21FFCAAE2EFBF59476FBFEF881" box="[1068,1147,1877,1903]" pageId="28" pageNumber="29" type="male">1 male</specimenCount>
(
<collectionCode id="ED03AE6DFFCAAE2EFB529476FB64F881" box="[1163,1249,1877,1903]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="28" pageNumber="29" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFEE1923FFBABFEB0" blockId="29.[264,1323,284,350]" pageId="29" pageNumber="30">
Etymology. The name 
<taxonomicName id="4C124D2BFFCBAE2FFD9C923FFD43FEDB" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[581,710,284,309]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="strepsicera">
<emphasis id="B966EABAFFCBAE2FFD9C923FFD43FEDB" box="[581,710,284,309]" italics="true" pageId="29" pageNumber="30">strepsicera</emphasis>
</taxonomicName>
comes from Latin for “twisted horn,” and refers to the processes from the lateral margin of sternum VIII of this species.
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED1929FFDCFFE3B" blockId="29.[264,586,443,505]" box="[264,586,443,470]" pageId="29" pageNumber="30">
<taxonomicName id="4C124D2BFFCBAE2FFED1929FFDCFFE3B" authority="Flint" authorityName="Flint" box="[264,586,443,470]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="talamanca">
<emphasis id="B966EABAFFCBAE2FFED1929FFDCFFE3B" bold="true" box="[264,586,443,470]" pageId="29" pageNumber="30">
<emphasis id="B966EABAFFCBAE2FFED1929FFD82FE38" bold="true" box="[264,519,444,470]" italics="true" pageId="29" pageNumber="30">Protoptila talamanca</emphasis>
Flint
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED192FCFEDFFE17" blockId="29.[264,586,443,505]" box="[264,346,479,505]" pageId="29" pageNumber="30">
<figureCitation id="13292A2DFFCBAE2FFED192FCFEDFFE17" box="[264,346,479,505]" captionStart="FIGURE 17" captionStartId="30.[264,367,1787,1811]" captionTargetBox="[264,1493,285,1760]" captionTargetId="figure@30.[264,1323,329,1760]" captionTargetPageId="30" captionText="FIGURE 17. Protoptila talamanca, Flint. Male genitalia: A—lateral; B—segments VIII and IX, ventral; C—phallic complex, lateral." figureDoi="http://doi.org/10.5281/zenodo.3492057" httpUri="https://zenodo.org/record/3492057/files/figure.png" pageId="29" pageNumber="30">Fig. 17</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED1910DFE23FDAB" blockId="29.[264,422,558,581]" box="[264,422,558,581]" pageId="29" pageNumber="30">
<bibRefCitation id="EF834B59FFCBAE2FFED1910DFEFFFDAB" author="Flint, O. S., Jr." box="[264,378,558,581]" pageId="29" pageNumber="30" pagination="1 - 30" refId="ref12816" refString="Flint, O. S., Jr. (1974) Studies of Neotropical caddisflies, XVIII: new species of Rhyacophilidae and Glossosomatidae (Trichoptera). Smithsonian Contributions to Zoology, 169, 1 - 30." type="journal article" year="1974">Flint 1974</bibRefCitation>
: 14.
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED19154FBA3FCB7" blockId="29.[264,1324,631,1217]" pageId="29" pageNumber="30">This is another distinctive species, unlikely to be confused with any other. Its most unusual character is the development of the ventral margin of segment IX, which possesses a pair of rod­like, apically acute processes, each of which is branched preapically to form a secondary tine­like process. Another distinctive character is the shape and structure of the phallicata, which has its apical half enlarged. The dorsal margin of this apex has two apically diverging lateral ridges, separated by a concave depression.</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFEE19044FEEAFB2F" blockId="29.[264,1324,631,1217]" pageId="29" pageNumber="30">
Material examined. 
<emphasis id="B966EABAFFCBAE2FFDFC9045FD5EFC6E" bold="true" box="[549,731,870,896]" pageId="29" pageNumber="30">
<collectingCountry id="F3057638FFCBAE2FFDFC9045FD5EFC6E" box="[549,731,870,896]" name="Costa Rica" pageId="29" pageNumber="30">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFCBAE2FFD329045FCD0FC6E" bold="true" box="[747,853,870,896]" pageId="29" pageNumber="30">
<collectingRegion id="49D6F84AFFCBAE2FFD329045FCD0FC6E" box="[747,853,870,896]" country="Costa Rica" name="Alajuela" pageId="29" pageNumber="30">Alajuela</collectingRegion>
</emphasis>
: Reserva Forestal San Ramón, Río San Lorencito &amp; tribs., 
<geoCoordinate id="EE26506FFFCBAE2FFE2190ACFD0FFC47" box="[504,650,911,937]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="15" value="10.216111">10°12’58”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCBAE2FFD4290ACFCC5FC47" box="[667,832,911,937]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="15" value="-84.60694">084°36’25”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCBAE2FFCA090ACFC4EFC47" box="[889,971,911,937]" metricMagnitude="2" metricUnit="m" metricValue="9.8" pageId="29" pageNumber="30" unit="m" value="980.0">980 m</quantity>
, 
<date id="FFAC1068FFCBAE2FFC0290ACFB14FC47" box="[987,1169,911,937]" pageId="29" pageNumber="30" value="1987-03-30" valueMax="1987-04-01" valueMin="1987-03-30">30.iii­1.iv.1987</date>
, Holzenthal, Hamilton, Heyn — 
<specimenCount id="9D14FD21FFCBAE2FFE2D9094FDD5FC3F" box="[500,592,951,977]" pageId="29" pageNumber="30" type="male">7 males</specimenCount>
(in alcohol) (UMSP), 
<specimenCount id="9D14FD21FFCBAE2FFC849094FC3DFC3F" box="[861,952,951,977]" pageId="29" pageNumber="30" type="male">2 males</specimenCount>
(in alcohol) (NMNH), 
<specimenCount id="9D14FD21FFCBAE2FFB169094FAAFFC3F" box="[1231,1322,951,977]" pageId="29" pageNumber="30" type="male">2 males</specimenCount>
(in alcohol) (INBIO); same, except 
<date id="FFAC1068FFCBAE2FFD1590FCFCE3FC17" box="[716,870,991,1017]" pageId="29" pageNumber="30" value="1991-03-06" valueMax="1991-03-10" valueMin="1991-03-06">6­10.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFCBAE2FFAC590FCFECAFBCF" pageId="29" pageNumber="30" type="male">9 males</specimenCount>
, 
<specimenCount id="9D14FD21FFCBAE2FFE849724FE55FBCF" box="[349,464,1031,1057]" pageId="29" pageNumber="30" type="female">4 females</specimenCount>
(in alcohol); 
<emphasis id="B966EABAFFCBAE2FFDAF9725FD59FBCE" bold="true" box="[630,732,1030,1056]" pageId="29" pageNumber="30">
<collectingRegion id="49D6F84AFFCBAE2FFDAF9725FD59FBCE" box="[630,732,1030,1056]" country="Costa Rica" name="Cartago" pageId="29" pageNumber="30">Cartago</collectingRegion>
</emphasis>
: Reserva Tapantí, waterfall, ca. 
<quantity id="4CEA9B4DFFCBAE2FFBB89724FB1BFBCF" box="[1121,1182,1031,1057]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="29" pageNumber="30" unit="km" value="1.0">1 km</quantity>
(road) NW tunnel, 
<geoCoordinate id="EE26506FFFCBAE2FFEB8970CFE77FBA7" box="[353,498,1071,1097]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="15" value="9.690001">09°41’24”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCBAE2FFDD9970CFD21FBA7" box="[512,676,1071,1097]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="15" value="-83.76">083°45’36”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCBAE2FFD0F970CFCB6FBA7" box="[726,819,1071,1097]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="29" pageNumber="30" unit="m" value="1600.0">1600 m</quantity>
, 
<date id="FFAC1068FFCBAE2FFCE6970CFC44FBA7" box="[831,961,1071,1097]" pageId="29" pageNumber="30" value="1991-03-24">24.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFCBAE2FFEEB9774FE0CFB9F" box="[306,393,1111,1137]" pageId="29" pageNumber="30" type="male">1 male</specimenCount>
, 
<specimenCount id="9D14FD21FFCBAE2FFE439774FD95FB9F" box="[410,528,1111,1137]" pageId="29" pageNumber="30" type="female">3 females</specimenCount>
(in alcohol); Río Reventazón, CATIE along Sendero Espaveles, 
<geoCoordinate id="EE26506FFFCBAE2FFED1975CFE1FFB77" box="[264,410,1151,1177]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="15" value="9.893055">09°53’35”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCBAE2FFE69975CFDD0FB77" box="[432,597,1151,1177]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="15" value="-83.651115">083°39’04”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCBAE2FFD40975CFD75FB77" box="[665,752,1151,1177]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="m" value="500.0">500 m</quantity>
, 
<date id="FFAC1068FFCBAE2FFCDF975CFC02FB77" box="[774,903,1151,1177]" pageId="29" pageNumber="30" value="1991-03-22">22.iii.1991</date>
, Muñoz­Quesada — 
<specimenCount id="9D14FD21FFCBAE2FFB46975CFB7DFB77" box="[1183,1272,1151,1177]" pageId="29" pageNumber="30" type="male">1 male</specimenCount>
(in alcohol).
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED1963DFDD2FAD6" blockId="29.[264,599,1310,1371]" box="[264,599,1310,1336]" pageId="29" pageNumber="30">
<taxonomicName id="4C124D2BFFCBAE2FFED1963DFDD2FAD6" authority="Bueno-Soria" authorityName="Bueno-Soria" box="[264,599,1310,1336]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="29" pageNumber="30" phylum="Arthropoda" rank="species" species="tica">
<emphasis id="B966EABAFFCBAE2FFED1963DFDD2FAD6" bold="true" box="[264,599,1310,1336]" pageId="29" pageNumber="30">
<emphasis id="B966EABAFFCBAE2FFED1963DFE36FAD6" bold="true" box="[264,435,1310,1336]" italics="true" pageId="29" pageNumber="30">Protoptila tica</emphasis>
Bueno­Soria
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED19662FEDFFAB5" blockId="29.[264,599,1310,1371]" box="[264,346,1345,1371]" pageId="29" pageNumber="30">
<figureCitation id="13292A2DFFCBAE2FFED19662FEDFFAB5" box="[264,346,1345,1371]" captionStart="FIGURE 18" captionStartId="31.[264,368,1404,1428]" captionTargetBox="[94,1323,286,1369]" captionTargetId="figure@31.[264,1323,300,1380]" captionTargetPageId="31" captionText="FIGURE 18. Protoptila tica, Bueno­Soria. Male genitalia: A—lateral; B—segments VIII and IX, ventral; C—phallic complex, lateral." httpUri="https://zenodo.org/record/2646305/files/figure.svg" pageId="29" pageNumber="30">Fig. 18</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED196B2FD8DFA46" blockId="29.[264,520,1425,1448]" box="[264,520,1425,1448]" pageId="29" pageNumber="30">
<bibRefCitation id="EF834B59FFCBAE2FFED196B2FE4BFA46" author="Bueno-Soria, J." box="[264,462,1425,1448]" pageId="29" pageNumber="30" pagination="392 - 394" refId="ref12694" refString="Bueno-Soria, J. (1984) Three new species of the genus Protoptila from Mexico and Costa Rica (Trichoptera: Glossosomatidae). Proceedings of the Biological Society of Washington, 97, 392 - 394." type="journal article" year="1984">Bueno­Soria 1984</bibRefCitation>
: 392.
</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFED196FAFC96F985" blockId="29.[264,1323,1497,1803]" pageId="29" pageNumber="30">The species has no obvious relationship to any other described species. It is easily diagnosed by the absence of parameres and the unique structure of sternum VIII, which forms broad, widely separated apical lobes, each with a fringe of thickened apical setae and a distinct apicodorsal spine­like process.</paragraph>
<paragraph id="8BAD36A8FFCBAE2FFEE1955AFCA2F8E5" blockId="29.[264,1323,1497,1803]" pageId="29" pageNumber="30">
Material examined. 
<emphasis id="B966EABAFFCBAE2FFDF2955AFD60F97D" bold="true" box="[555,741,1657,1683]" pageId="29" pageNumber="30">
<collectingCountry id="F3057638FFCBAE2FFDF2955AFD60F97D" box="[555,741,1657,1683]" name="Costa Rica" pageId="29" pageNumber="30">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFCBAE2FFD21955AFC02F97D" bold="true" box="[760,903,1657,1683]" pageId="29" pageNumber="30">
<collectingRegion id="49D6F84AFFCBAE2FFD21955AFC02F97D" box="[760,903,1657,1683]" country="Costa Rica" name="Puntarenas" pageId="29" pageNumber="30">Puntarenas</collectingRegion>
</emphasis>
: Parque Nacional Corcovado, Río Camaronal, 
<geoCoordinate id="EE26506FFFCBAE2FFE419582FDAFF955" box="[408,554,1697,1723]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="15" value="8.481944">08°28’55”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCBAE2FFDE19582FD59F955" box="[568,732,1697,1723]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="15" value="-83.58889">083°35’20”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCBAE2FFCD69582FCCAF955" box="[783,847,1697,1723]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="29" pageNumber="30" unit="m" value="30.0">30 m</quantity>
, 
<date id="FFAC1068FFCBAE2FFC859582FC5FF955" box="[860,986,1697,1723]" pageId="29" pageNumber="30" value="1989-04-13">13.iv.1989</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFCBAE2FFAC59582FEC4F90D" pageId="29" pageNumber="30" type="male">1 male</specimenCount>
(in alcohol); same, except 
<geoCoordinate id="EE26506FFFCBAE2FFD7195EAFCBFF90D" box="[680,826,1737,1763]" direction="north" orientation="latitude" pageId="29" pageNumber="30" precision="15" value="8.481111">08°28’52”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFCBAE2FFC9795EAFC76F90D" box="[846,1011,1737,1763]" direction="west" orientation="longitude" pageId="29" pageNumber="30" precision="15" value="-83.593895">083°35’38”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFCBAE2FFBED95EAFBE9F90D" box="[1076,1132,1737,1763]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="29" pageNumber="30" unit="m" value="5.0">5 m</quantity>
, 
<date id="FFAC1068FFCBAE2FFB5995EAFAA3F90D" box="[1152,1318,1737,1763]" pageId="29" pageNumber="30" value="1989-04-12" valueMax="1989-04-13" valueMin="1989-04-12">12­13.iv.1989</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFCBAE2FFDEF95D2FD15F8E5" box="[566,656,1777,1803]" pageId="29" pageNumber="30" type="male">4 males</specimenCount>
(in alcohol).
</paragraph>
<caption id="DF6D6620FFC8AE2CFED195D8FD0AF8DA" ID-DOI="http://doi.org/10.5281/zenodo.3492057" ID-Zenodo-Dep="3492057" httpUri="https://zenodo.org/record/3492057/files/figure.png" pageId="30" pageNumber="31" startId="30.[264,367,1787,1811]" targetBox="[264,1493,285,1760]" targetPageId="30">
<paragraph id="8BAD36A8FFC8AE2CFED195D8FD0AF8DA" blockId="30.[264,1322,1787,1844]" pageId="30" pageNumber="31">
<emphasis id="B966EABAFFC8AE2CFED195D8FE1FF8FD" bold="true" box="[264,410,1787,1811]" pageId="30" pageNumber="31">FIGURE 17.</emphasis>
<taxonomicName id="4C124D2BFFC8AE2CFE7A95D8FD56F8FC" authority=", Flint." authorityName="Flint." box="[419,723,1787,1810]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="30" pageNumber="31" phylum="Arthropoda" rank="species" species="talamanca">
<emphasis id="B966EABAFFC8AE2CFE7A95D8FD0FF8FC" box="[419,650,1787,1810]" italics="true" pageId="30" pageNumber="31">Protoptila talamanca</emphasis>
, Flint.
</taxonomicName>
Male genitalia: A—lateral; B—segments VIII and IX, ventral; C—phallic complex, lateral.
</paragraph>
</caption>
<caption id="DF6D6620FFC9AE2DFED1965FFD0AFA58" httpUri="https://zenodo.org/record/2646305/files/figure.svg" pageId="31" pageNumber="32" startId="31.[264,368,1404,1428]" targetBox="[94,1323,286,1369]" targetPageId="31">
<paragraph id="8BAD36A8FFC9AE2DFED1965FFD0AFA58" blockId="31.[264,1322,1404,1462]" pageId="31" pageNumber="32">
<emphasis id="B966EABAFFC9AE2DFED1965FFE1CFA7A" bold="true" box="[264,409,1404,1428]" pageId="31" pageNumber="32">FIGURE 18.</emphasis>
<taxonomicName id="4C124D2BFFC9AE2DFE78965FFD5EFA7D" authority=", Bueno-Soria." authorityName="Bueno-Soria." box="[417,731,1404,1427]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tica">
<emphasis id="B966EABAFFC9AE2DFE78965FFDB8FA7D" box="[417,573,1404,1427]" italics="true" pageId="31" pageNumber="32">Protoptila tica</emphasis>
, Bueno­Soria.
</taxonomicName>
Male genitalia: A—lateral; B—segments VIII and IX, ventral; C—phallic complex, lateral.
</paragraph>
</caption>
<paragraph id="8BAD36A8FFC9AE2DFED1952FFDB3F9C8" blockId="31.[264,566,1548,1609]" box="[264,566,1548,1574]" pageId="31" pageNumber="32">
<taxonomicName id="4C124D2BFFC9AE2DFED1952FFDB3F9C8" authority="Mosely" authorityName="Mosely" box="[264,566,1548,1574]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tojana">
<emphasis id="B966EABAFFC9AE2DFED1952FFDB3F9C8" bold="true" box="[264,566,1548,1574]" pageId="31" pageNumber="32">
<emphasis id="B966EABAFFC9AE2DFED1952FFE51F9C8" bold="true" box="[264,468,1548,1574]" italics="true" pageId="31" pageNumber="32">Protoptila tojana</emphasis>
Mosely
</emphasis>
</taxonomicName>
</paragraph>
<paragraph id="8BAD36A8FFC9AE2DFED1950CFEDFF9A7" blockId="31.[264,566,1548,1609]" box="[264,346,1583,1609]" pageId="31" pageNumber="32">
<figureCitation id="13292A2DFFC9AE2DFED1950CFEDFF9A7" box="[264,346,1583,1609]" captionStart="FIGURE 19" captionStartId="33.[264,368,1034,1058]" captionTargetBox="[94,1315,286,1008]" captionTargetId="figure@33.[271,1315,326,1008]" captionTargetPageId="33" captionText="FIGURE 19. Protoptila tojana, Mosely. Male genitalia: A—lateral; B—phallic complex, lateral." httpUri="https://zenodo.org/record/2646307/files/figure.svg" pageId="31" pageNumber="32">Fig. 19</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFC9AE2DFED1955CFE54F978" blockId="31.[264,465,1663,1686]" box="[264,465,1663,1686]" pageId="31" pageNumber="32">
<bibRefCitation id="EF834B59FFC9AE2DFED1955CFE12F978" author="Mosely, M. E." box="[264,407,1663,1686]" pageId="31" pageNumber="32" pagination="317 - 346" refId="ref13051" refString="Mosely, M. E. (1954) The Protoptila group of the Glossosomatinae (Trichoptera: Rhyacophilidae). Bulletin of the British Museum (Natural History) Entomology, 3, 317 - 346." type="journal article" year="1954">Mosely 1954</bibRefCitation>
: 331.
</paragraph>
<paragraph id="8BAD36A8FFC9AE2DFED195E4FCE2F8B7" blockId="31.[264,1324,1735,1881]" pageId="31" pageNumber="32">
This species is most similar to 
<taxonomicName id="4C124D2BFFC9AE2DFDAE95E4FC93F90E" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[631,790,1735,1761]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="strepsicera">
<emphasis id="B966EABAFFC9AE2DFDAE95E4FD03F90E" box="[631,646,1735,1760]" italics="true" pageId="31" pageNumber="32">P</emphasis>
. 
<emphasis id="B966EABAFFC9AE2DFD4C95E4FC93F90E" box="[661,790,1735,1760]" italics="true" pageId="31" pageNumber="32">strepsicera</emphasis>
</taxonomicName>
and the two are undoubtedly closely related. Both species differ greatly in the processes emanating from the posterodorsal margin of sternum VIII. In 
<taxonomicName id="4C124D2BFFC9AE2DFE0E9434FDC4F8DE" box="[471,577,1815,1840]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="tojana">
<emphasis id="B966EABAFFC9AE2DFE0E9434FDC4F8DE" box="[471,577,1815,1840]" italics="true" pageId="31" pageNumber="32">P. tojana</emphasis>
</taxonomicName>
they are very narrow, whereas in 
<taxonomicName id="4C124D2BFFC9AE2DFC389434FB07F8DE" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[993,1154,1815,1840]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="species" species="strepsicera">
<emphasis id="B966EABAFFC9AE2DFC389434FB07F8DE" box="[993,1154,1815,1840]" italics="true" pageId="31" pageNumber="32">P. strepsicera</emphasis>
</taxonomicName>
they are very prominent and shaped like the horns of an antelope.
</paragraph>
<paragraph id="8BAD36A8FFF6AE12FEE1923FFEF5F910" blockId="32.[264,1325,283,1790]" pageId="32" pageNumber="33">
Material examined. 
<emphasis id="B966EABAFFF6AE12FDFD9238FD5CFEDB" bold="true" box="[548,729,283,309]" pageId="32" pageNumber="33">
<collectingCountry id="F3057638FFF6AE12FDFD9238FD5CFEDB" box="[548,729,283,309]" name="Costa Rica" pageId="32" pageNumber="33">COSTA RICA</collectingCountry>
</emphasis>
: 
<collectingRegion id="49D6F84AFFF6AE12FD30923FFCD4FED8" box="[745,849,284,310]" country="Costa Rica" name="Alajuela" pageId="32" pageNumber="33">Alajuela</collectingRegion>
: Río La Paz Pequeña, route 9, 
<quantity id="4CEA9B4DFFF6AE12FB62923FFA8BFED8" box="[1211,1294,284,310]" metricMagnitude="3" metricUnit="m" metricValue="7.8" pageId="32" pageNumber="33" unit="km" value="7.8">7.8 km</quantity>
N Vara Blanca, 
<geoCoordinate id="EE26506FFFF6AE12FE7D9267FDB2FEB0" box="[420,567,324,350]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="10.211111">10°12’40”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FD9B9267FD62FEB0" box="[578,743,324,350]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-84.166115">084°09’58”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FCCF9267FCF7FEB0" box="[790,882,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="32" pageNumber="33" unit="m" value="1230.0">1230 m</quantity>
, 
<date id="FFAC1068FFF6AE12FCA49267FC73FEB0" box="[893,1014,324,350]" pageId="32" pageNumber="33" value="1986-02-13">13.ii.1986</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFF6AE12FEF5924FFEFBFE68" box="[300,382,364,390]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
; 
<emphasis id="B966EABAFFF6AE12FE539248FDBAFE6B" bold="true" box="[394,575,363,389]" pageId="32" pageNumber="33">
<collectingCountry id="F3057638FFF6AE12FE539248FDBAFE6B" box="[394,575,363,389]" name="Costa Rica" pageId="32" pageNumber="33">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFF6AE12FD979248FD3DFE6B" bold="true" box="[590,696,363,389]" pageId="32" pageNumber="33">
<collectingRegion id="49D6F84AFFF6AE12FD979248FD3DFE6B" box="[590,696,363,389]" country="Costa Rica" name="Alajuela" pageId="32" pageNumber="33">Alajuela</collectingRegion>
</emphasis>
: Río La Paz Pequeña, route 9, 
<quantity id="4CEA9B4DFFF6AE12FBF8924FFBF6FE68" box="[1057,1139,364,390]" metricMagnitude="3" metricUnit="m" metricValue="7.8" pageId="32" pageNumber="33" unit="km" value="7.8">7.8 km</quantity>
N Vara Blanca, 
<geoCoordinate id="EE26506FFFF6AE12FED192B7FE1FFE40" box="[264,410,404,430]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="10.211111">10°12’40”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FE7192B7FDC8FE40" box="[424,589,404,430]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-84.166115">084°09’58”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FD5892B7FD5AFE40" box="[641,735,404,430]" metricMagnitude="3" metricUnit="m" metricValue="1.23" pageId="32" pageNumber="33" unit="m" value="1230.0">1230 m</quantity>
, 
<date id="FFAC1068FFF6AE12FD3492B7FCE3FE40" box="[749,870,404,430]" pageId="32" pageNumber="33" value="1986-02-13">13.ii.1986</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFF6AE12FB0892B7FAA3FE40" box="[1233,1318,404,430]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
; 
<emphasis id="B966EABAFFF6AE12FED19298FEEBFE3B" bold="true" box="[264,366,443,469]" pageId="32" pageNumber="33">
<collectingRegion id="49D6F84AFFF6AE12FED19298FEEBFE3B" box="[264,366,443,469]" country="Costa Rica" name="Cartago" pageId="32" pageNumber="33">Cartago</collectingRegion>
</emphasis>
: Río Chitaría, rt 10, 10 km NW Río Reventazón, 
<geoCoordinate id="EE26506FFFF6AE12FC67929FFBD5FE38" box="[958,1104,444,470]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="9.92">09°55’12”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FB84929FFA84FE38" box="[1117,1281,444,470]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.60389">083°36’14”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FED192C7FED2FE10" box="[264,343,484,510]" metricMagnitude="2" metricUnit="m" metricValue="7.4" pageId="32" pageNumber="33" unit="m" value="740.0">740 m</quantity>
, 
<date id="FFAC1068FFF6AE12FEBE92C7FE6DFE10" box="[359,488,484,510]" pageId="32" pageNumber="33" value="1991-03-21">21.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFF6AE12FC5392C7FC62FE10" box="[906,999,484,510]" pageId="32" pageNumber="33" type="male">2 males</specimenCount>
(in alcohol); 
<emphasis id="B966EABAFFF6AE12FB4992C0FAA7FE13" bold="true" box="[1168,1314,483,509]" pageId="32" pageNumber="33">
<collectingRegion id="49D6F84AFFF6AE12FB4992C0FAA7FE13" box="[1168,1314,483,509]" country="Costa Rica" name="Guanacaste" pageId="32" pageNumber="33">Guanacaste</collectingRegion>
</emphasis>
: Río Tempisquito, ca. 
<quantity id="4CEA9B4DFFF6AE12FDD7912FFDC8FDC8" box="[526,589,524,550]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="km" value="3.0">3 km</quantity>
S Route 1, 
<geoCoordinate id="EE26506FFFF6AE12FD3C912FFCF3FDC8" box="[741,886,524,550]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="10.790001">10°47’24”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FC5F912FFBAFFDC8" box="[902,1066,524,550]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-85.55195">085°33’07”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FBBB912FFB21FDC8" box="[1122,1188,524,550]" metricMagnitude="1" metricUnit="m" metricValue="7.5" pageId="32" pageNumber="33" unit="m" value="75.0">75 m</quantity>
, 
<date id="FFAC1068FFF6AE12FB6A912FFAA3FDC8" box="[1203,1318,524,550]" pageId="32" pageNumber="33" value="1986-03-06">6.iii.1986</date>
, Holzenthal &amp; Fasth — 
<specimenCount id="9D14FD21FFF6AE12FDE89117FD14FDA0" box="[561,657,564,590]" pageId="32" pageNumber="33" type="male">3 males</specimenCount>
(in alcohol) (INBIO); Río Tizate, 
<quantity id="4CEA9B4DFFF6AE12FB909117FB24FDA0" box="[1097,1185,564,590]" metricMagnitude="3" metricUnit="m" metricValue="7.2" pageId="32" pageNumber="33" unit="km" value="7.2">7.2 km</quantity>
NE Cañas Dulces, 
<geoCoordinate id="EE26506FFFF6AE12FEBE917FFE7CFD98" box="[359,505,604,630]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="10.773055">10°46’23”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FDDC917FFD2FFD98" box="[517,682,604,630]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-85.44889">085°26’56”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FD01917FFCA0FD98" box="[728,805,604,630]" metricMagnitude="2" metricUnit="m" metricValue="2.75" pageId="32" pageNumber="33" unit="m" value="275.0">275 m</quantity>
, 
<date id="FFAC1068FFF6AE12FCE9917FFC35FD98" box="[816,944,604,630]" pageId="32" pageNumber="33" value="1986-06-28">28.vi.1986</date>
, Holzenthal, Heyn, Armitage — 
<specimenCount id="9D14FD21FFF6AE12FED191A7FED9FD70" box="[264,348,644,670]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
; Parque Nacional Santa Rosa, Río Poza Salada, 
<geoCoordinate id="EE26506FFFF6AE12FC4191A7FBACFD70" box="[920,1065,644,670]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="10.798889">10°47’56”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FBEE91A7FB5EFD70" box="[1079,1243,644,670]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-85.65195">085°39’07”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FAD491A7FEA7FD28" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="32" pageNumber="33" unit="m" value="10.0">10 m</quantity>
, 
<date id="FFAC1068FFF6AE12FEF7918FFE33FD28" box="[302,438,684,710]" pageId="32" pageNumber="33" value="1987-07-24">24.vii.1987</date>
, Holzenthal, Morse, Clausen — 
<specimenCount id="9D14FD21FFF6AE12FCEE918FFC11FD28" box="[823,916,684,710]" pageId="32" pageNumber="33" type="male">7 males</specimenCount>
, 
<specimenCount id="9D14FD21FFF6AE12FC79918FFBA1FD28" box="[928,1060,684,710]" pageId="32" pageNumber="33" type="female">15 females</specimenCount>
; 
<emphasis id="B966EABAFFF6AE12FBE9918FFB06FD28" bold="true" box="[1072,1155,684,710]" pageId="32" pageNumber="33">
<collectingRegion id="49D6F84AFFF6AE12FBE9918FFB06FD28" box="[1072,1155,684,710]" country="Costa Rica" name="Limon" pageId="32" pageNumber="33">Limón</collectingRegion>
</emphasis>
: Río Barbilla, ca. 
<quantity id="4CEA9B4DFFF6AE12FEEA91F7FEF5FD00" box="[307,368,724,750]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="32" pageNumber="33" unit="km" value="8.0">8 km</quantity>
W B­Line, 
<geoCoordinate id="EE26506FFFF6AE12FDDB91F7FD11FD00" box="[514,660,724,750]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="10.066944">10°04’01”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FD7B91F7FCC3FD00" box="[674,838,724,750]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.36889">083°22’08”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FCA391F7FC3FFD00" box="[890,954,724,750]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="m" value="30.0">30 m</quantity>
, 
<date id="FFAC1068FFF6AE12FC1191F7FBBCFD00" box="[968,1081,724,750]" pageId="32" pageNumber="33" value="1986-01-31">31.i.1986</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFF6AE12FE5891DFFE5FFCF8" box="[385,474,764,790]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
, 
<specimenCount id="9D14FD21FFF6AE12FE3491DFFDECFCF8" box="[493,617,764,790]" pageId="32" pageNumber="33" type="female">5 females</specimenCount>
; 
<emphasis id="B966EABAFFF6AE12FDA291DFFC8FFCF8" bold="true" box="[635,778,764,790]" pageId="32" pageNumber="33">
<collectingRegion id="49D6F84AFFF6AE12FDA291DFFC8FFCF8" box="[635,778,764,790]" country="Costa Rica" name="Puntarenas" pageId="32" pageNumber="33">Puntarenas</collectingRegion>
</emphasis>
: Quebrada Pita, ca. 
<quantity id="4CEA9B4DFFF6AE12FBCA91DFFBD0FCF8" box="[1043,1109,764,790]" metricMagnitude="3" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="km" value="3.0">3 km</quantity>
(air) W Golfito, 
<geoCoordinate id="EE26506FFFF6AE12FED19007FE1FFCD0" box="[264,410,804,830]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="8.641944">08°38’31”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FE7F9007FDCCFCD0" box="[422,585,804,830]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.193054">083°11’35”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FDA19007FD32FCD0" box="[632,695,804,830]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="m" value="15.0">15 m</quantity>
, 
<date id="FFAC1068FFF6AE12FD1B9007FCBEFCD0" box="[706,827,804,830]" pageId="32" pageNumber="33" value="1986-02-15">15.ii.1986</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFF6AE12FB439007FA86FCD0" box="[1178,1283,804,830]" pageId="32" pageNumber="33" type="male">49 males</specimenCount>
(in alcohol); Río Bellavista, ca. 
<quantity id="4CEA9B4DFFF6AE12FD8C906FFD22FC88" box="[597,679,844,870]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="km" value="1.5">1.5 km</quantity>
NW Las Alturas, 
<geoCoordinate id="EE26506FFFF6AE12FCA4906FFB8AFC88" box="[893,1039,844,870]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="8.951111">08°57’04”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FBC2906FFB3AFC88" box="[1051,1215,844,870]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-82.846115">082°50’46”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FB29906FFEA7FC60" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="32" pageNumber="33" unit="m" value="1400.0">1400 m</quantity>
, 
<date id="FFAC1068FFF6AE12FEE99057FE2CFC60" box="[304,425,884,910]" pageId="32" pageNumber="33" value="1986-02-18">18.ii.1986</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFF6AE12FCCC9057FCF1FC60" box="[789,884,884,910]" pageId="32" pageNumber="33" type="male">2 males</specimenCount>
, 
<specimenCount id="9D14FD21FFF6AE12FC5B9057FB81FC60" box="[898,1028,884,910]" pageId="32" pageNumber="33" type="female">13 females</specimenCount>
(in alcohol); Río Ceibo, route 2 ca. 
<quantity id="4CEA9B4DFFF6AE12FE5290BFFE4DFC58" box="[395,456,924,950]" metricMagnitude="3" metricUnit="m" metricValue="6.0" pageId="32" pageNumber="33" unit="km" value="6.0">6 km</quantity>
W rd to Buenos Aires, 
<geoCoordinate id="EE26506FFFF6AE12FD0690BFFCF4FC58" box="[735,881,924,950]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="9.148889">09°08’56”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FCA790BFFBA7FC58" box="[894,1058,924,950]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.376945">083°22’37”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FB8A90BFFB1AFC58" box="[1107,1183,924,950]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="32" pageNumber="33" unit="m" value="250.0">250 m</quantity>
, 
<date id="FFAC1068FFF6AE12FB7590BFFAA3FC58" box="[1196,1318,924,950]" pageId="32" pageNumber="33" value="1986-02-20">20.ii.1986</date>
, Holzenthal, Morse, Fasth — 
<specimenCount id="9D14FD21FFF6AE12FD8490E7FD32FC30" box="[605,695,964,990]" pageId="32" pageNumber="33" type="male">4 males</specimenCount>
(in alcohol) (INBIO); Reserva Biológica Carara, Río Carara, 
<quantity id="4CEA9B4DFFF6AE12FEB490CFFE42FBE8" box="[365,455,1004,1030]" metricMagnitude="3" metricUnit="m" metricValue="4.3" pageId="32" pageNumber="33" unit="km" value="4.3">4.3 km</quantity>
(rd) E Cost. Sur, 
<geoCoordinate id="EE26506FFFF6AE12FD6790CFFCD5FBE8" box="[702,848,1004,1030]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="9.81">09°48’36”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FCBC90CFFB8CFBE8" box="[869,1033,1004,1030]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-84.571945">084°34’19”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FB9090CFFB0AFBE8" box="[1097,1167,1004,1030]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="20.0">20 m</quantity>
, 
<date id="FFAC1068FFF6AE12FB7D90CFFAA0FBE8" box="[1188,1317,1004,1030]" pageId="32" pageNumber="33" value="1991-03-12">12.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFF6AE12FD569737FC83FBC0" box="[655,774,1044,1070]" pageId="32" pageNumber="33" type="male">138 males</specimenCount>
(in alcohol), 
<specimenCount id="9D14FD21FFF6AE12FC7D9737FBB7FBC0" box="[932,1074,1044,1070]" pageId="32" pageNumber="33" type="female">355 females</specimenCount>
(in alcohol), 
<specimenCount id="9D14FD21FFF6AE12FB099737FAAFFBC0" box="[1232,1322,1044,1070]" pageId="32" pageNumber="33" type="male">3 males</specimenCount>
(pinned), 
<specimenCount id="9D14FD21FFF6AE12FEA2971FFE7EFBB8" box="[379,507,1084,1110]" pageId="32" pageNumber="33" type="female">11 females</specimenCount>
(pinned); Reserva Biológica Carara, Quebrada Bonita, 
<geoCoordinate id="EE26506FFFF6AE12FB4B971FFAA1FBB8" box="[1170,1316,1084,1110]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="9.775">09°46’30”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FED19747FE29FB90" box="[264,428,1124,1150]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-84.604996">084°36’18”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FE079747FD98FB90" box="[478,541,1124,1150]" metricMagnitude="1" metricUnit="m" metricValue="3.5" pageId="32" pageNumber="33" unit="m" value="35.0">35 m</quantity>
, 
<date id="FFAC1068FFF6AE12FDF09747FD43FB90" box="[553,710,1124,1150]" pageId="32" pageNumber="33" value="1990-05-18" valueMax="1990-05-20" valueMin="1990-05-18">18­20.v.1990</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFF6AE12FBDD9747FBEBFB90" box="[1028,1134,1124,1150]" pageId="32" pageNumber="33" type="male">89 males</specimenCount>
(in alcohol), 
<specimenCount id="9D14FD21FFF6AE12FAD79747FEC9FB48" pageId="32" pageNumber="33" type="male">11 males</specimenCount>
(pinned), 
<specimenCount id="9D14FD21FFF6AE12FE0197AFFDE4FB48" box="[472,609,1164,1190]" pageId="32" pageNumber="33" type="female">53 females</specimenCount>
(pinned); same, except 
<date id="FFAC1068FFF6AE12FC7997AFFBA5FB48" box="[928,1056,1164,1190]" pageId="32" pageNumber="33" value="1991-03-11">11.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFF6AE12FE449797FD91FB20" box="[413,532,1204,1230]" pageId="32" pageNumber="33" type="male">245 males</specimenCount>
(in alcohol), 
<specimenCount id="9D14FD21FFF6AE12FD689797FC85FB20" box="[689,768,1204,1230]" pageId="32" pageNumber="33" type="male">1 male</specimenCount>
(pinned), 
<specimenCount id="9D14FD21FFF6AE12FCA09797FC72FB20" box="[889,1015,1204,1230]" pageId="32" pageNumber="33" type="female">11 females</specimenCount>
(pinned); Río Rincón, 
<quantity id="4CEA9B4DFFF6AE12FADC9797FEA8FB18" metricMagnitude="3" metricUnit="m" metricValue="6.5" pageId="32" pageNumber="33" unit="km" value="6.5">6.5 km</quantity>
(air) S Rincón, 
<geoCoordinate id="EE26506FFFF6AE12FE3297FFFDF8FB18" box="[491,637,1244,1270]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="8.638056">08°38’17”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FD5097FFFCA8FB18" box="[649,813,1244,1270]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.48">083°28’48”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FC8797FFFC18FB18" box="[862,925,1244,1270]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="20.0">20 m</quantity>
, 
<date id="FFAC1068FFF6AE12FC7097FFFB9CFB18" box="[937,1049,1244,1270]" pageId="32" pageNumber="33" value="1987-04-07">7.iv.1987</date>
, Holzenthal, Hamilton, Heyn — 
<specimenCount id="9D14FD21FFF6AE12FEA29627FE5CFAF0" box="[379,473,1284,1310]" pageId="32" pageNumber="33" type="male">2 males</specimenCount>
(in alcohol); Parque Nacional Corcovado, Río Claro, 
<quantity id="4CEA9B4DFFF6AE12FBA99627FB43FAF0" box="[1136,1222,1284,1310]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="km" value="1.5">1.5 km</quantity>
SE Est. Sirena, 
<geoCoordinate id="EE26506FFFF6AE12FEB0960FFE7EFAA8" box="[361,507,1324,1350]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="8.471944">08°28’19”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FDD7960FFD36FAA8" box="[526,691,1324,1350]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.58806">083°35’17”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FD28960FFCB3FAA8" box="[753,822,1324,1350]" metricMagnitude="1" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="m" value="15.0">15 m</quantity>
, 
<date id="FFAC1068FFF6AE12FC90960FFC3CFAA8" box="[841,953,1324,1350]" pageId="32" pageNumber="33" value="1989-04-08">8.iv.1989</date>
, Holzenthal — 
<specimenCount id="9D14FD21FFF6AE12FB5E960FFB7FFAA8" box="[1159,1274,1324,1350]" pageId="32" pageNumber="33" type="male">33 males</specimenCount>
, 
<specimenCount id="9D14FD21FFF6AE12FAD4960FFEE6FA80" pageId="32" pageNumber="33" type="female">29 females</specimenCount>
(in alcohol); Parque Nacional Corcovado, Rio Camaronal, 
<geoCoordinate id="EE26506FFFF6AE12FB4B9677FAA1FA80" box="[1170,1316,1364,1390]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="8.481944">08°28’55”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FED1965FFE29FA78" box="[264,428,1404,1430]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.58889">083°35’20”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FE3D965FFDA2FA78" box="[484,551,1404,1430]" metricMagnitude="1" metricUnit="m" metricValue="3.0" pageId="32" pageNumber="33" unit="m" value="30.0">30 m</quantity>
, 
<date id="FFAC1068FFF6AE12FDEF965FFD31FA78" box="[566,692,1404,1430]" pageId="32" pageNumber="33" value="1989-04-13">13.iv.1989</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFF6AE12FBDD965FFBF4FA78" box="[1028,1137,1404,1430]" pageId="32" pageNumber="33" type="male">66 males</specimenCount>
(in alcohol), 
<specimenCount id="9D14FD21FFF6AE12FAC2965FFEC9FA50" pageId="32" pageNumber="33" type="male">2 males</specimenCount>
(pinned), 
<specimenCount id="9D14FD21FFF6AE12FE1F9687FDB2FA50" box="[454,567,1444,1470]" pageId="32" pageNumber="33" type="female">2 females</specimenCount>
(pinned); same, except 
<geoCoordinate id="EE26506FFFF6AE12FC8A9687FC60FA50" box="[851,997,1444,1470]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="8.481111">08°28’52”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FC289687FB13FA50" box="[1009,1174,1444,1470]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-83.593895">083°35’38”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FB1E9687FB72FA50" box="[1223,1271,1444,1470]" metricMagnitude="0" metricUnit="m" metricValue="5.0" pageId="32" pageNumber="33" unit="m" value="5.0">5 m</quantity>
, 
<date id="FFAC1068FFF6AE12FADA9687FE03FA08" pageId="32" pageNumber="33" value="1989-04-12" valueMax="1989-04-13" valueMin="1989-04-12">12­ 13.iv.1989</date>
, Holzenthal &amp; Blahnik — 
<specimenCount id="9D14FD21FFF6AE12FD1896EFFCBDFA08" box="[705,824,1484,1510]" pageId="32" pageNumber="33" type="male">162 males</specimenCount>
(in alcohol); 
<emphasis id="B966EABAFFF6AE12FC0E96EFFBC6FA08" bold="true" box="[983,1091,1484,1510]" pageId="32" pageNumber="33">
<collectingRegion id="49D6F84AFFF6AE12FC0E96EFFBC6FA08" box="[983,1091,1484,1510]" country="Costa Rica" name="San Jose" pageId="32" pageNumber="33">San José</collectingRegion>
</emphasis>
: Reserva Biológica Carara, Río del Sur, 
<quantity id="4CEA9B4DFFF6AE12FDC396D7FDF1F9E0" box="[538,628,1524,1550]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="32" pageNumber="33" unit="km" value="1.5">1.5 km</quantity>
(rd) S Carara, 
<geoCoordinate id="EE26506FFFF6AE12FC9D96D7FC53F9E0" box="[836,982,1524,1550]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="9.768888">09°46’08”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FC3296D7FB0AF9E0" box="[1003,1167,1524,1550]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FB0996D7FAA3F9E0" box="[1232,1318,1524,1550]" metricMagnitude="2" metricUnit="m" metricValue="1.6" pageId="32" pageNumber="33" unit="m" value="160.0">160 m</quantity>
, 
<date id="FFAC1068FFF6AE12FED1953FFE0FF9D8" box="[264,394,1564,1590]" pageId="32" pageNumber="33" value="1991-03-13">13.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFF6AE12FCED953FFC24F9D8" box="[820,929,1564,1590]" pageId="32" pageNumber="33" type="male">34 males</specimenCount>
(in alcohol); Reserva Biológica Carara, Río Carara in Carara, 
<geoCoordinate id="EE26506FFFF6AE12FD469567FCB4F9B0" box="[671,817,1604,1630]" direction="north" orientation="latitude" pageId="32" pageNumber="33" precision="15" value="9.778055">09°46’41”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF6AE12FC909567FC68F9B0" box="[841,1005,1604,1630]" direction="west" orientation="longitude" pageId="32" pageNumber="33" precision="15" value="-84.53111">084°31’52”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF6AE12FBED9567FB08F9B0" box="[1076,1165,1604,1630]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="32" pageNumber="33" unit="m" value="200.0">200 m</quantity>
, 
<date id="FFAC1068FFF6AE12FB7D9567FAA0F9B0" box="[1188,1317,1604,1630]" pageId="32" pageNumber="33" value="1991-03-14">14.iii.1991</date>
, Holzenthal, Muñoz, Huisman — 
<specimenCount id="9D14FD21FFF6AE12FD57954FFD72F968" box="[654,759,1644,1670]" pageId="32" pageNumber="33" type="male">91 males</specimenCount>
(in alcohol). 
<emphasis id="B966EABAFFF6AE12FC4F954FFB93F968" bold="true" box="[918,1046,1644,1670]" pageId="32" pageNumber="33">
<collectingCountry id="F3057638FFF6AE12FC4F954FFB93F968" box="[918,1046,1644,1670]" name="Panama" pageId="32" pageNumber="33">PANAMA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFF6AE12FBFC954FFB09F968" bold="true" box="[1061,1164,1644,1670]" pageId="32" pageNumber="33">
<collectingCountry id="F3057638FFF6AE12FBFC954FFB09F968" box="[1061,1164,1644,1670]" name="Panama" pageId="32" pageNumber="33">Panama</collectingCountry>
</emphasis>
: Canal Zone, Barro Colorado Island, Snyder­Molino trail, marker 3, 
<date id="FFAC1068FFF6AE12FC5395B7FBCDF940" box="[906,1096,1684,1710]" pageId="32" pageNumber="33" value="1988-06-22" valueMax="1988-09-06" valueMin="1988-06-22">22.vi.­6.ix.1988</date>
, Wolda — 
<specimenCount id="9D14FD21FFF6AE12FB1595B7FAACF940" box="[1228,1321,1684,1710]" pageId="32" pageNumber="33" type="male">2 males</specimenCount>
, 
<specimenCount id="9D14FD21FFF6AE12FED1959FFEF9F938" box="[264,380,1724,1750]" pageId="32" pageNumber="33" type="female">2 females</specimenCount>
(in alcohol); same, except 
<date id="FFAC1068FFF6AE12FD13959FFC15F938" box="[714,912,1724,1750]" pageId="32" pageNumber="33" value="1990-06-26" valueMax="1990-12-03" valueMin="1990-06-26">26.vi.­3.xii.1990</date>
, Wolda — 
<specimenCount id="9D14FD21FFF6AE12FBC7959FFBFAF938" box="[1054,1151,1724,1750]" pageId="32" pageNumber="33" type="male">2 males</specimenCount>
, 
<specimenCount id="9D14FD21FFF6AE12FB54959FFA84F938" box="[1165,1281,1724,1750]" pageId="32" pageNumber="33" type="female">1 females</specimenCount>
(in alcohol).
</paragraph>
<caption id="DF6D6620FFF7AE13FED19729FA91FBCF" box="[264,1300,1034,1058]" httpUri="https://zenodo.org/record/2646307/files/figure.svg" pageId="33" pageNumber="34" startId="33.[264,368,1034,1058]" targetBox="[94,1315,286,1008]" targetPageId="33">
<paragraph id="8BAD36A8FFF7AE13FED19729FA91FBCF" blockId="33.[264,1300,1034,1058]" box="[264,1300,1034,1058]" pageId="33" pageNumber="34">
<emphasis id="B966EABAFFF7AE13FED19729FE1DFBCC" bold="true" box="[264,408,1034,1058]" pageId="33" pageNumber="34">FIGURE 19.</emphasis>
<taxonomicName id="4C124D2BFFF7AE13FE479729FD3CFBCF" authority=", Mosely." authorityName="Mosely." box="[414,697,1034,1057]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="tojana">
<emphasis id="B966EABAFFF7AE13FE479729FDD3FBCF" box="[414,598,1034,1057]" italics="true" pageId="33" pageNumber="34">Protoptila tojana</emphasis>
, Mosely.
</taxonomicName>
Male genitalia: A—lateral; B—phallic complex, lateral.
</paragraph>
</caption>
<paragraph id="8BAD36A8FFF7AE13FED197A3FD3CFB74" blockId="33.[264,697,1152,1213]" box="[264,697,1152,1178]" pageId="33" pageNumber="34">
<emphasis id="B966EABAFFF7AE13FED197A3FD3CFB74" bold="true" box="[264,697,1152,1178]" pageId="33" pageNumber="34">
<taxonomicName id="4C124D2BFFF7AE13FED197A3FD9EFB74" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[264,539,1152,1178]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="trichoglossa" status="sp. nov.">
<emphasis id="B966EABAFFF7AE13FED197A3FD9EFB74" bold="true" box="[264,539,1152,1178]" italics="true" pageId="33" pageNumber="34">Protoptila trichoglossa</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFF7AE13FDF097A3FD3CFB74" box="[553,697,1152,1178]" pageId="33" pageNumber="34" rank="species">new species</taxonomicNameLabel>
</emphasis>
</paragraph>
<paragraph id="8BAD36A8FFF7AE13FED19780FEDFFB53" blockId="33.[264,697,1152,1213]" box="[264,346,1187,1213]" pageId="33" pageNumber="34">
<figureCitation id="13292A2DFFF7AE13FED19780FEDFFB53" box="[264,346,1187,1213]" captionStart="FIGURE 20" captionStartId="34.[264,368,1788,1812]" captionTargetBox="[295,1276,670,1767]" captionTargetId="figure@34.[295,1276,670,1768]" captionTargetPageId="34" captionText="FIGURE 20. Protoptila trichoglossa, new species. Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral (inset: apex of phallicata, dorsal)." figureDoi="http://doi.org/10.5281/zenodo.3492059" httpUri="https://zenodo.org/record/3492059/files/figure.png" pageId="33" pageNumber="34">Fig. 20</figureCitation>
</paragraph>
<paragraph id="8BAD36A8FFF7AE13FED197D0FB82FA6B" blockId="33.[264,1324,1267,1893]" pageId="33" pageNumber="34">This species is unique and easily diagnosed from all others by the structure of sternum VIII of the male, which is elongate and tapered apicoventrally and margined with enlarged, coarse setae. The posterodorsal margin of sternum VIII is also modified on either side into a short, digitate process terminating with enlarged setae.</paragraph>
<paragraph id="8BAD36A8FFF7AE13FEE196B0FBDBFA3B" blockId="33.[264,1324,1267,1893]" pageId="33" pageNumber="34">
Adult. Length of forewing: male 
<quantity id="4CEA9B4DFFF7AE13FD3396B0FC01FA43" box="[746,900,1427,1453]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="33" pageNumber="34" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6–3.8 mm</quantity>
. Color dark brown, with distinct transverse bar of whitish setae along cord and a few whitish apical spots.
</paragraph>
<paragraph id="8BAD36A8FFF7AE10FEE196C0FBEDFD98" blockId="33.[264,1324,1267,1893]" lastBlockId="34.[264,1323,284,630]" lastPageId="34" lastPageNumber="35" pageId="33" pageNumber="34">Male genitalia. Sternum VI process prominent, slightly curved, longer than wide, apex subacute. Tergum VIII posterior margin with row of very elongate setae with attenuate, curved apices; sternum VIII strongly produced ventrally, not bifurcate apically, strongly narrowed, attenuate, apex acute, dorsolateral and apical margins with thick dense setae; dorsolateral margin of sternum VIII basally with short digitate process, bearing several long thickened setae. Segment IX with anterior margin broadly rounded; sternum IX medially produced posteriorly, narrow, apically rounded, with pair of apical setae; posterolateral margin of segment IX without processes. Preanal appendages absent. Tergum X divided mesally to form sclerotized lateral parts with basal and apical sections; basal section subquadrate, longer than wide, apical section longer than basal section, nearly uniform in width, apex narrowed and forming short, acute dorsomesal process. Phallobase dorsally with large, semicircular, laterally compressed apodeme; ventrally with short, rod­like, articulated appendages with apical setae, appendages fitting into sclerotized pockets on ventral margin of phallobase; posteroventral margin of phallobase with single, cupped, upturned, apically acute basal process, but without dorsolateral processes; paramere elongate, membranous, paramere spine short, sinuously curled, acute apically; phallicata elongate, narrow, strongly arched near middle, apex expanded and bulbous; endophallus reduced, not noticeably everted; phallotremal sclerite lightly sclerotized, an incomplete basal ring, with curved, apicomesal projection.</paragraph>
<caption id="DF6D6620FFF4AE10FED195DFFBB6F8D8" ID-DOI="http://doi.org/10.5281/zenodo.3492059" ID-Zenodo-Dep="3492059" httpUri="https://zenodo.org/record/3492059/files/figure.png" pageId="34" pageNumber="35" startId="34.[264,368,1788,1812]" targetBox="[295,1276,670,1767]" targetPageId="34">
<paragraph id="8BAD36A8FFF4AE10FED195DFFBB6F8D8" blockId="34.[264,1323,1788,1846]" pageId="34" pageNumber="35">
<emphasis id="B966EABAFFF4AE10FED195DFFE1FF8FA" bold="true" box="[264,410,1788,1812]" pageId="34" pageNumber="35">FIGURE 20.</emphasis>
<taxonomicName id="4C124D2BFFF4AE10FE7D95DEFD1BF8FA" authority="Holzenthal &amp; Blahnik, 2006" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[420,670,1789,1812]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="trichoglossa" status="sp. nov.">
<emphasis id="B966EABAFFF4AE10FE7D95DEFD1BF8FA" box="[420,670,1789,1812]" italics="true" pageId="34" pageNumber="35">Protoptila trichoglossa</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel id="A25557C1FFF4AE10FD7495DFFCB6F8FD" box="[685,819,1788,1811]" pageId="34" pageNumber="35" rank="species">new species</taxonomicNameLabel>
. Male genitalia: A—lateral; B—segment VIII apex, ventral; C—phallic complex, lateral (inset: apex of phallicata, dorsal).
</paragraph>
</caption>
<paragraph id="8BAD36A8FFF5AE11FEE19238FDE4FE68" blockId="35.[264,1323,283,630]" pageId="35" pageNumber="36">
<materialsCitation id="3B7A3CF5FFF5AE11FEE19238FDE4FE68" ID-GBIF-Occurrence="2236573117" collectingDate="1986-02-18" collectionCode="UMSP" country="Costa Rica" elevation="1400" latitude="8.951111" location="Rio Bellavista" longLatPrecision="21" longitude="-82.846115" pageId="35" pageNumber="36" specimenCode="UMSP000091739" specimenCount="1" stateProvince="Puntarenas" typeStatus="holotype">
<emphasis id="B966EABAFFF5AE11FEE19238FE74FEDB" bold="true" box="[312,497,283,309]" pageId="35" pageNumber="36">
<typeStatus id="54A9880AFFF5AE11FEE19238FE2FFEDB" box="[312,426,283,309]" pageId="35" pageNumber="36" type="holotype">Holotype</typeStatus>
male
</emphasis>
: 
<emphasis id="B966EABAFFF5AE11FDDC9238FD3AFEDB" bold="true" box="[517,703,283,309]" pageId="35" pageNumber="36">
<collectingCountry id="F3057638FFF5AE11FDDC9238FD3AFEDB" box="[517,703,283,309]" name="Costa Rica" pageId="35" pageNumber="36">COSTA RICA</collectingCountry>
</emphasis>
: 
<emphasis id="B966EABAFFF5AE11FD0A9238FCE7FEDB" bold="true" box="[723,866,283,309]" pageId="35" pageNumber="36">
<collectingRegion id="49D6F84AFFF5AE11FD0A9238FCE7FEDB" box="[723,866,283,309]" country="Costa Rica" name="Puntarenas" pageId="35" pageNumber="36">Puntarenas</collectingRegion>
</emphasis>
: 
<location id="8ECD6073FFF5AE11FCAF923FFBA1FED8" LSID="urn:lsid:plazi:treatment:03BB87BEFFCEAE16FED19615FC9BFE5D:8ECD6073FFF5AE11FCAF923FFBA1FED8" box="[886,1060,284,310]" country="Costa Rica" latitude="8.951111" longLatPrecision="21" longitude="-82.846115" name="Rio Bellavista" pageId="35" pageNumber="36" stateProvince="Puntarenas">Río Bellavista</location>
, ca. 
<locationDeviation id="9A1F2F8AFFF5AE11FBBB923FFEE4FEB0" location="Las Alturas" pageId="35" pageNumber="36">
<quantity id="4CEA9B4DFFF5AE11FBBB923FFB3DFED8" box="[1122,1208,284,310]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="35" pageNumber="36" unit="km" value="1.5">1.5 km</quantity>
NW Las Alturas
</locationDeviation>
, 
<geoCoordinate id="EE26506FFFF5AE11FEAA9267FD80FEB0" box="[371,517,324,350]" direction="north" orientation="latitude" pageId="35" pageNumber="36" precision="15" value="8.951111">08°57’04”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF5AE11FDCE9267FD3EFEB0" box="[535,699,324,350]" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="15" value="-82.846115">082°50’46”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF5AE11FD2E9267FCDDFEB0" box="[759,856,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="35" pageNumber="36" unit="m" value="1400.0">
<elevation id="003FD19BFFF5AE11FD2E9267FCDDFEB0" box="[759,856,324,350]" metricMagnitude="3" metricUnit="m" metricValue="1.4" pageId="35" pageNumber="36" unit="m" value="1400.0">1400 m</elevation>
</quantity>
, 
<date id="FFAC1068FFF5AE11FCB39267FC61FEB0" box="[874,996,324,350]" pageId="35" pageNumber="36" value="1986-02-18">
<collectingDate id="EFE8E980FFF5AE11FCB39267FC61FEB0" box="[874,996,324,350]" pageId="35" pageNumber="36" value="1986-02-18">18.ii.1986</collectingDate>
</date>
, Holzenthal, Morse, Fasth (
<specimenCode id="DBB49ED3FFF5AE11FEC9924FFE62FE68" box="[272,487,364,390]" collectionCode="UMSP" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="35" pageNumber="36" type="University or college">UMSP000091739</specimenCode>
) (
<collectionCode id="ED03AE6DFFF5AE11FE26924FFDD0FE68" box="[511,597,364,390]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="35" pageNumber="36" type="University or college">UMSP</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAD36A8FFF5AE11FEE192B0FAAFFE40" blockId="35.[264,1323,283,630]" box="[312,1322,403,430]" pageId="35" pageNumber="36">
<materialsCitation id="3B7A3CF5FFF5AE11FEE192B0FBDEFE40" ID-GBIF-Occurrence="2236573144" box="[312,1115,403,430]" collectionCode="UMSP" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="B966EABAFFF5AE11FEE192B0FE30FE43" bold="true" box="[312,437,403,429]" pageId="35" pageNumber="36">
<typeStatus id="54A9880AFFF5AE11FEE192B0FE30FE43" box="[312,437,403,429]" pageId="35" pageNumber="36" type="paratype">Paratypes</typeStatus>
</emphasis>
: Same data as 
<typeStatus id="54A9880AFFF5AE11FDB792B7FD51FE40" box="[622,724,404,430]" pageId="35" pageNumber="36" type="holotype">holotype</typeStatus>
— 
<specimenCount id="9D14FD21FFF5AE11FCDC92B7FCD2FE40" box="[773,855,404,430]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFF5AE11FC2492B7FBD6FE40" box="[1021,1107,404,430]" country="USA" httpUri="http://grbio.org/cool/3avb-bd3e" name="University of Minnesota Insect Collection" pageId="35" pageNumber="36" type="University or college">UMSP</collectionCode>
)
</materialsCitation>
; 
<emphasis id="B966EABAFFF5AE11FBB292B0FAA7FE43" bold="true" box="[1131,1314,403,429]" pageId="35" pageNumber="36">
<collectingCountry id="F3057638FFF5AE11FBB292B0FAA7FE43" box="[1131,1314,403,429]" name="Costa Rica" pageId="35" pageNumber="36">COSTA RICA</collectingCountry>
</emphasis>
:
</paragraph>
</subSubSection>
<subSubSection id="C3086523FFF5AE11FED19298FD55FC15" pageId="35" pageNumber="36" type="description">
<paragraph id="8BAD36A8FFF5AE11FED19298FEF8FDC8" blockId="35.[264,1323,283,630]" pageId="35" pageNumber="36">
<materialsCitation id="3B7A3CF5FFF5AE11FED19298FEFDFDC8" ID-GBIF-Occurrence="2236573136" collectingDate="1989-04-15" collectionCode="NMNH" collectorName="Holzenthal &amp; Blahnik" county="Zona Protectora Las Tablas" elevation="1460" latitude="8.941112" location="Rio Coton" longLatPrecision="21" longitude="-82.78694" municipality="Las" pageId="35" pageNumber="36" specimenCount="1" specimenCount-male="1" stateProvince="Puntarenas">
<emphasis id="B966EABAFFF5AE11FED19298FE12FE3B" bold="true" box="[264,407,443,469]" pageId="35" pageNumber="36">
<collectingRegion id="49D6F84AFFF5AE11FED19298FE12FE3B" box="[264,407,443,469]" country="Costa Rica" name="Puntarenas" pageId="35" pageNumber="36">Puntarenas</collectingRegion>
</emphasis>
: 
<collectingCounty id="62CC4E24FFF5AE11FE6D929FFCA6FE38" box="[436,803,444,470]" pageId="35" pageNumber="36">
Zona Protectora 
<collectingMunicipality id="6BC9ACD2FFF5AE11FD4D929FFD3BFE38" box="[660,702,444,470]" pageId="35" pageNumber="36">Las</collectingMunicipality>
Tablas
</collectingCounty>
, 
<location id="8ECD6073FFF5AE11FCE5929FFC42FE38" LSID="urn:lsid:plazi:treatment:03BB87BEFFCEAE16FED19615FC9BFE5D:8ECD6073FFF5AE11FCE5929FFC42FE38" box="[828,967,444,470]" county="Zona Protectora Las Tablas" latitude="8.941112" longLatPrecision="21" longitude="-82.78694" municipality="Las" name="Rio Coton" pageId="35" pageNumber="36" stateProvince="Puntarenas">Río Cotón</location>
, 
<location id="8ECD6073FFF5AE11FC38929FFBFCFE38" LSID="urn:lsid:plazi:treatment:03BB87BEFFCEAE16FED19615FC9BFE5D:8ECD6073FFF5AE11FC38929FFBFCFE38" box="[993,1145,444,470]" county="Zona Protectora Las Tablas" latitude="8.941112" longLatPrecision="21" longitude="-82.78694" municipality="Las" name="Sitio Coton" pageId="35" pageNumber="36" stateProvince="Puntarenas">Sitio Cotón</location>
, 
<geoCoordinate id="EE26506FFFF5AE11FB4B929FFAA1FE38" box="[1170,1316,444,470]" direction="north" orientation="latitude" pageId="35" pageNumber="36" precision="15" value="8.941112">08°56’28”N</geoCoordinate>
, 
<geoCoordinate id="EE26506FFFF5AE11FED192C7FE29FE10" box="[264,428,484,510]" direction="west" orientation="longitude" pageId="35" pageNumber="36" precision="15" value="-82.78694">082°47’13”W</geoCoordinate>
, el. 
<quantity id="4CEA9B4DFFF5AE11FE3092C7FDC9FE10" box="[489,588,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="35" pageNumber="36" unit="m" value="1460.0">
<elevation id="003FD19BFFF5AE11FE3092C7FDC9FE10" box="[489,588,484,510]" metricMagnitude="3" metricUnit="m" metricValue="1.46" pageId="35" pageNumber="36" unit="m" value="1460.0">1460 m</elevation>
</quantity>
, 
<date id="FFAC1068FFF5AE11FD8792C7FD59FE10" box="[606,732,484,510]" pageId="35" pageNumber="36" value="1989-04-15">
<collectingDate id="EFE8E980FFF5AE11FD8792C7FD59FE10" box="[606,732,484,510]" pageId="35" pageNumber="36" value="1989-04-15">15.iv.1989</collectingDate>
</date>
, 
<collectorName id="26E7537EFFF5AE11FD3692C7FCEAFE10" box="[751,879,484,510]" pageId="35" pageNumber="36">Holzenthal</collectorName>
&amp; 
<collectorName id="26E7537EFFF5AE11FC7B92C7FC7AFE10" box="[930,1023,484,510]" pageId="35" pageNumber="36">Blahnik</collectorName>
— 
<specimenCount id="9D14FD21FFF5AE11FBE192C7FB0BFE10" box="[1080,1166,484,510]" pageId="35" pageNumber="36" type="male">1 male</specimenCount>
(in alcohol) (
<collectionCode id="ED03AE6DFFF5AE11FEC9912FFEF5FDC8" box="[272,368,524,550]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="35" pageNumber="36" type="Museum">NMNH</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BAD36A8FFF5AE11FEE19117FB10FD98" blockId="35.[264,1323,283,630]" pageId="35" pageNumber="36">
Etymology. The name 
<taxonomicName id="4C124D2BFFF5AE11FD919117FD5CFDA3" authorityName="Holzenthal &amp; Blahnik" authorityYear="2006" box="[584,729,564,589]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="trichoglossa">
<emphasis id="B966EABAFFF5AE11FD919117FD5CFDA3" box="[584,729,564,589]" italics="true" pageId="35" pageNumber="36">trichoglossa</emphasis>
</taxonomicName>
comes from Greek roots for “hairy tongue” and refers to the elongate, tapered, setose sternum of segment VIII of this species.
</paragraph>
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<emphasis id="B966EABAFFF5AE11FED191F7FDC8FD00" bold="true" box="[264,589,724,750]" pageId="35" pageNumber="36">ACKNOWLEDGMENTS</emphasis>
</paragraph>
<paragraph id="8BAD36A8FFF5AE11FED1903AFD55FC15" blockId="35.[264,1324,793,1019]" pageId="35" pageNumber="36">
We are grateful to Dr. Oliver S. Flint, Jr., Smithsonian Institution, for providing material for examination and for his insights into protoptiline taxonomy. We thank Desiree Robertson and Lourdes Chamorro, University of 
<collectingRegion id="49D6F84AFFF5AE11FCB3904AFC6CFC6D" box="[874,1001,873,899]" country="United States of America" name="Minnesota" pageId="35" pageNumber="36">Minnesota</collectingRegion>
, for useful discussions on morphology. Julie Martinez rendered the beautiful illustration of 
<taxonomicName id="4C124D2BFFF5AE11FBC690B2FB61FC44" box="[1055,1252,913,938]" class="Insecta" family="Glossosomatidae" genus="Protoptila" kingdom="Animalia" order="Trichoptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="ixtala">
<emphasis id="B966EABAFFF5AE11FBC690B2FB61FC44" box="[1055,1252,913,938]" italics="true" pageId="35" pageNumber="36">Protoptila ixtala</emphasis>
</taxonomicName>
. This material is based upon work supported by the National Science Foundation under Grant Nos. 9400632, 9971885, and 0 117772.
</paragraph>
</subSubSection>
<subSubSection id="C3086523FFF5AE11FED1977AFB78FB1B" pageId="35" pageNumber="36" type="reference_group">
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<author id="53AFDE4DFFF2AE16FED1925FFE1AFE7D" box="[264,415,380,403]" case="TC" fns="I" in="G.B." ins="D" ln="Wiggins" lnc="TC" npo="LnFn" pageId="36" pageNumber="37">Wiggins, G.B.</author>
(
<year id="FFBA77FBFFF2AE16FE76925FFE62FE7D" box="[431,487,380,403]" pageId="36" pageNumber="37">1996</year>
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<title id="F916DD9EFFF2AE16FE2F925FFAA1FE7D" box="[502,1316,380,403]" jopScore="0.1437139" pageId="36" pageNumber="37">
<emphasis id="B966EABAFFF2AE16FE2F925FFAA1FE7D" box="[502,1316,380,403]" italics="true" pageId="36" pageNumber="37">Larvae of the North American Caddisfly Genera (Trichoptera), 2nd edition</emphasis>
</title>
. 
<journalOrPublisher id="51D33C64FFF2AE16FEE192BFFD46FE5D" box="[312,707,412,435]" jopScore="2.1652775" pageId="36" pageNumber="37">University of Toronto Press, Toronto</journalOrPublisher>
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