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<document id="5CC38D0D37C90DE9EDC880B8FFA4319C" ID="10.11646/zootaxa.4200.3.10" ID-DOI="10.11646/zootaxa.4200.3.10" ID-GBIF-Dataset="3181840a-a6b9-476b-aaaa-350a2e2acc6f" ID-ISSN="1175-5326" ID-Zenodo-Dep="187236" ID-ZooBank="E5D3718C-87B7-42CE-A4FF-DAB9B739ED4A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1480494168585" checkinUser="plazi" docAuthor="Kóbor, P. &amp; Kondorosy, E." docDate="2016" docId="03BB87A5FF92180EF3D1F9660D29FAD4" docLanguage="en" docName="zootaxa.4200.3.10.pdf" docOrigin="Zootaxa 4200 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Germalus benyovszkyi Kobor &amp; Kondorosy, sp. nov." docType="treatment" docUuid="DCF36FFA-3368-40CE-A9E2-4EC68B7D4BBB" docUuidSource="ZooBank" docVersion="7" lastPageNumber="449" masterDocId="FF82FFDDFF96180BF346FF95096EFFE4" masterDocTitle="Germalus species of the Malagasy region (Heteroptera: Lygaeoidea: Geocoridae)" masterLastPageNumber="450" masterPageNumber="444" pageNumber="448" updateTime="1698395222755" updateUser="plazi">
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<mods:title id="DD6C971B17B809879700A31DAB8B1894">Germalus species of the Malagasy region (Heteroptera: Lygaeoidea: Geocoridae)</mods:title>
</mods:titleInfo>
<mods:name id="000271D365B5D5DEF2DBAA11675D4608" type="personal">
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<mods:namePart id="09C495680536E5560F3773233C2FDFF1">Kóbor, P.</mods:namePart>
</mods:name>
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<mods:role id="D93219B89FC003F06ED6C4B18D1B1DAA">
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<mods:namePart id="3E36CE40E94653F33B3098CBB8AD08DB">Kondorosy, E.</mods:namePart>
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<mods:typeOfResource id="76CFE77F5C637F26E9DB47B2F4BEA09F">text</mods:typeOfResource>
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<mods:title id="5E84F147C993C771FB9392453FF3E5FA">Zootaxa</mods:title>
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<mods:date id="9AEF6CA80C4D2BD88681A1B72E91E3A8">2016</mods:date>
<mods:detail id="469EE3DE9CDB39178ED0AE9ECD13EBCC" type="volume">
<mods:number id="A1E9AA4125D5E2FD324074AA3BBE49B0">4200</mods:number>
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<mods:number id="DFBBCCCC8040EFE4F30A412FD89EDB77">3</mods:number>
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<mods:start id="8E54B4C7469682DDF72D48D09204CC3C">444</mods:start>
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<mods:classification id="09278BA589B7C207071900E0F5D4976F">journal article</mods:classification>
<mods:identifier id="0E7CCB2E3949471D02AE9818A109246E" type="DOI">10.11646/zootaxa.4200.3.10</mods:identifier>
<mods:identifier id="18A0B0F7B902ED7242403917D35CCADD" type="GBIF-Dataset">3181840a-a6b9-476b-aaaa-350a2e2acc6f</mods:identifier>
<mods:identifier id="A4F5C735FBAE945E8A7E3AA42C43B078" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="422C92A4C3A70FE315CB596F7B9603CB" type="Zenodo-Dep">187236</mods:identifier>
<mods:identifier id="1EBBB0E6F7194ABCD5E88CAD0FAD1362" type="ZooBank">E5D3718C-87B7-42CE-A4FF-DAB9B739ED4A</mods:identifier>
</mods:mods>
<treatment id="03BB87A5FF92180EF3D1F9660D29FAD4" ID-DOI="http://doi.org/10.5281/zenodo.6072809" ID-GBIF-Taxon="124873617" ID-Zenodo-Dep="6072809" LSID="urn:lsid:zoobank.org:act:DCF36FFA-3368-40CE-A9E2-4EC68B7D4BBB" httpUri="http://treatment.plazi.org/id/03BB87A5FF92180EF3D1F9660D29FAD4" lastPageId="5" lastPageNumber="449" pageId="4" pageNumber="448">
<subSubSection id="C3086538FF92180FF3D1F96608E7F8CB" pageId="4" pageNumber="448" type="nomenclature">
<paragraph id="8BAD36B3FF92180FF3D1F9660A7DF8EA" blockId="4.[151,787,1779,1840]" box="[151,787,1779,1806]" pageId="4" pageNumber="448">
<heading id="D0E581DFFF92180FF3D1F9660A7DF8EA" bold="true" box="[151,787,1779,1806]" fontSize="11" level="1" pageId="4" pageNumber="448" reason="1">
<emphasis id="B966EAA1FF92180FF3D1F9660A7DF8EA" bold="true" box="[151,787,1779,1806]" pageId="4" pageNumber="448">
<taxonomicName id="4C124D30FF92180FF3D1F9660BC0F8EA" ID-CoL="8Q8DS" authority="Kobor &amp; Kondorosy" authorityName="Kobor &amp; Kondorosy" box="[151,686,1779,1806]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="448" phylum="Arthropoda" rank="species" species="benyovszkyi" status="sp. nov.">
<emphasis id="B966EAA1FF92180FF3D1F96608CAF8E9" bold="true" box="[151,420,1779,1805]" italics="true" pageId="4" pageNumber="448">Germalus benyovszkyi</emphasis>
Kóbor &amp; Kondorosy
</taxonomicName>
<taxonomicNameLabel id="A25557DAFF92180FF1F3F9610A7DF8EA" box="[693,787,1780,1806]" pageId="4" pageNumber="448" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAD36B3FF92180FF3D1F88208E7F8CB" blockId="4.[151,787,1779,1840]" box="[151,393,1815,1840]" pageId="4" pageNumber="448">
(
<figureCitation id="13292A36FF92180FF3D9F88209B0F8CB" box="[159,222,1815,1840]" captionStart="FIGURE 1" captionStartId="2.[151,250,1990,2012]" captionTargetBox="[172,1381,439,1953]" captionTargetId="figure@2.[152,1401,419,2037]" captionTargetPageId="2" captionText="FIGURE 1. Distribution records of Germalus spp." httpUri="https://zenodo.org/record/187237/files/figure.png" pageId="4" pageNumber="448">Fig. 1</figureCitation>
A–B;
<figureCitation id="13292A36FF92180FF263F8820809F8CB" box="[293,359,1815,1840]" captionStart="FIGURE 2" captionStartId="3.[151,250,1071,1093]" captionTargetBox="[151,1436,193,1050]" captionTargetId="figure@3.[151,1436,193,1050]" captionTargetPageId="3" captionText="FIGURE 2. Dorsal view of Germalus spp. A. Germalus kinbergi (Stål, 1859), Mauritius (NHMW); B. Germalus banari Kóbor &amp; Kondorosy sp. nov., Madagascar (MMBC); C. Germalus benyovszkyi Kóbor &amp; Kondorosy sp. nov., Madagascar (MNHN)." httpUri="https://zenodo.org/record/187238/files/figure.png" pageId="4" pageNumber="448">Fig. 2</figureCitation>
C.)
</paragraph>
</subSubSection>
<subSubSection id="C3086538FF92180EF3D1F8CA0B63FE6C" lastPageId="5" lastPageNumber="449" pageId="4" pageNumber="448" type="materials_examined">
<paragraph id="8BAD36B3FF92180FF3D1F8CA0844F87F" blockId="4.[151,1436,1887,2020]" pageId="4" pageNumber="448">
<materialsCitation id="3B7A3CEEFF92180FF3D1F8CA0844F87F" ID-GBIF-Occurrence="1412767734" collectingDate="1958-12" collectionCode="MNHN" collectorName="P. Girevaud" country="Madagascar" county="Matsabory" elevation="2030" location="Sambava" municipality="Marojejy" pageId="4" pageNumber="448" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
<emphasis id="B966EAA1FF92180FF3D1F8CA0828F89C" bold="true" box="[151,326,1887,1912]" pageId="4" pageNumber="448">
<typeStatus id="54A98811FF92180FF3D1F8CA09BCF89C" box="[151,210,1887,1912]" pageId="4" pageNumber="448">Type</typeStatus>
material.
</emphasis>
<typeStatus id="54A98811FF92180FF208F8CA08D5F89C" box="[334,443,1887,1912]" pageId="4" pageNumber="448" type="holotype">Holotype</typeStatus>
:
<collectingCountry id="F3057623FF92180FF280F8CA0B23F89C" box="[454,589,1887,1912]" name="Madagascar" pageId="4" pageNumber="448">Madagascar</collectingCountry>
Est / district
<location id="8ECD6068FF92180FF1AEF8CA0A20F89C" LSID="urn:lsid:plazi:treatment:03BB87A5FF92180EF3D1F9660D29FAD4:8ECD6068FF92180FF1AEF8CA0A20F89C" box="[744,846,1887,1912]" country="Madagascar" county="Matsabory" municipality="Marojejy" name="Sambava" pageId="4" pageNumber="448">Sambava</location>
/
<collectingMunicipality id="6BC9ACC9FF92180FF021F8CA0AA0F89C" box="[871,974,1887,1912]" pageId="4" pageNumber="448">Marojejy</collectingMunicipality>
/
<collectingCounty id="62CC4E3FFF92180FF0A0F8CA0D30F89C" box="[998,1118,1887,1912]" pageId="4" pageNumber="448">Matsabory</collectingCounty>
<elevation id="003FD180FF92180FF720F8CA0DDAF893" box="[1126,1204,1887,1911]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="4" pageNumber="448" unit="m" value="2030.0">
<quantity id="4CEA9B56FF92180FF720F8CA0DDAF893" box="[1126,1204,1887,1911]" metricMagnitude="3" metricUnit="m" metricValue="2.03" pageId="4" pageNumber="448" unit="m" value="2030.0">2030m</quantity>
</elevation>
/
<collectingDate id="EFE8E99BFF92180FF78AF8CA0C76F89C" box="[1228,1304,1887,1912]" pageId="4" pageNumber="448" value="1958-12">
<date id="FFAC1073FF92180FF78AF8CA0C76F89C" box="[1228,1304,1887,1912]" pageId="4" pageNumber="448" value="1958-12">XII-58</date>
</collectingDate>
<collectorName id="26E75365FF92180FF666F8CA0CF2F89C" box="[1312,1436,1887,1912]" pageId="4" pageNumber="448">P.Girevaud</collectorName>
(
<specimenCount id="9D14FD3AFF92180FF3E4F81609D8F87F" box="[162,182,1923,1947]" pageId="4" pageNumber="448" type="female">♀</specimenCount>
,
<collectionCode id="ED03AE76FF92180FF385F8160870F87F" box="[195,286,1923,1947]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="4" pageNumber="448">MNHN</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAD36B3FF92180EF381F8320B63FE6C" blockId="4.[151,1436,1887,2020]" lastBlockId="5.[151,1437,151,1328]" lastPageId="5" lastPageNumber="449" pageId="4" pageNumber="448">
<emphasis id="B966EAA1FF92180FF381F83208CCF824" bold="true" box="[199,418,1959,1984]" pageId="4" pageNumber="448">COLOURATION.</emphasis>
Body generally ochraceous. Head with thin, fuscous median line reaching to base of clypeus. Ocelli colourless. Antennal segment I fuscous except its base, segment II fuscous with base and apex black, segment III ochraceous with dark fuscous base, antennal segment IV fuscous with ochraceous base. Labial segments ochraceous, apex of segment IV black. Pronotum with pale fuscous punctuation; darker fuscous punctures in the midline. One rounded black spot at each humeral angles. Punctuation of scutellum pale brown; elevated triradiate carina ochraceous. Hemelytra pale ochraceous, nearly hyaline with pale fuscous punctuation along claval commissure and veins, costal margin narrowly black ending in oval black spot at apex of corium; corium-membrane margin fuscous; membrane hyaline. Legs ochraceous, unicolorous. Abdominal dorsum brownish, connexiva ochraceous.
</paragraph>
</subSubSection>
<subSubSection id="C3086538FF93180EF381FE0108DCFBD0" pageId="5" pageNumber="449" type="description">
<paragraph id="8BAD36B3FF93180EF381FE0108BBFC23" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="449">
<emphasis id="B966EAA1FF93180EF381FE0108AFFE49" bold="true" box="[199,449,404,429]" pageId="5" pageNumber="449">STRUCTURE. Head</emphasis>
dull, impunctate, pentagonal in dorsal view. Eyes stylate, not touching pronotum; ocelli situated near to ocular sulcus. Eye-stalks shiny. Clypeus rounded, exceeding mandibular plate. Antennal tubercles visible in dorsal view. Antennal segment I shortest, II longest,III &amp; IV subequal. Antennal segments covered with very fine, decumbent pubescence. Antennal segment IV less fusiform than
<taxonomicName id="4C124D30FF93180EF0AEFD950D3DFDFC" box="[1000,1107,512,536]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="449" phylum="Arthropoda" rank="species" species="banari">
<emphasis id="B966EAA1FF93180EF0AEFD950D3DFDFC" box="[1000,1107,512,536]" italics="true" pageId="5" pageNumber="449">G. banari</emphasis>
</taxonomicName>
. Ratio of antennal segments: 10:25:20:18. Labial segments I–III subequal, segment I exceeding well onto prosternum, segment IV shortest reaching metacoxae. Ratio of labial segments: 37:40:35:25.
<emphasis id="B966EAA1FF93180EF00FFDD20AC5FD84" bold="true" box="[841,939,583,608]" pageId="5" pageNumber="449">Thorax:</emphasis>
Pronotum carinate, gradually narrowing to anterior edge with slight impressions in the middle of lateral margin; anterior and posterior edges rounded. Disc covered with moderately dense, coarse punctuation except callosities, posterior margin and humeral angles; pronotal collar is less densely punctuate. Scutellum densely punctuate with exception of elevated triradiate carina. Hemelytra shiny. Clavus with two rows of punctures, scutellar row incomplete. Punctuation of claval commisure almost complete. Costal margin widening gradually to apical third; punctuation along corial veins complete. Membrane reaching well beyond apex of abdomen, with conspicuous venation in basal half. Thoracic pleura finely punctuate; propleuron having impunctate areas more or less under callosities. Metathoracic scent gland situated laterally at anterior margin of metapleuron; evaporative area distinct, orificia curved anteriorly. Legs shiny, impunctate, with sparse, decumbent pubescence.
<emphasis id="B966EAA1FF93180EF194FC190A32FC41" bold="true" box="[722,860,908,933]" pageId="5" pageNumber="449">Abodomen.</emphasis>
Connexiva with fine pubescence; genital capsule with sparse, semierect setae.
</paragraph>
<paragraph id="8BAD36B3FF93180EF381FC4108DCFBD0" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="449">
<emphasis id="B966EAA1FF93180EF381FC410813FC09" bold="true" box="[199,381,980,1005]" pageId="5" pageNumber="449">Measurements.</emphasis>
Body length: 7.12; head length: 0.98, width: 2.18; distance of ocelli: 0.47; antennal segments: 0.37–0.93–0.74–0.67; labial segments: 1.09–1.18–1.03–0.74; pronotum length: 1.40, width: 2.10; scutellum length: 0.86, width: 1.02.
</paragraph>
</subSubSection>
<subSubSection id="C3086538FF93180EF381FBAA0B3EFB44" pageId="5" pageNumber="449" type="discussion">
<paragraph id="8BAD36B3FF93180EF381FBAA0B3EFB44" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="449">
<emphasis id="B966EAA1FF93180EF381FBAA0817FBBC" bold="true" box="[199,377,1087,1112]" pageId="5" pageNumber="449">DISCUSSION.</emphasis>
This species is easy to distinguish from the other two species of the region due to the spots at humeral angles and corial edges, the strongly stylate eye and “striped” colouration of antennae. This is the largest known species in the genus worldwide.
</paragraph>
</subSubSection>
<subSubSection id="C3086538FF93180EF381FB390B62FB0C" pageId="5" pageNumber="449" type="etymology">
<paragraph id="8BAD36B3FF93180EF381FB390B62FB0C" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="449">
<materialsCitation id="3B7A3CEEFF93180EF381FB390B62FB0C" ID-GBIF-Occurrence="1412767726" collectingDate="1746" collectingDateMax="1786" collectingDateMin="1746" country="Madagascar" county="Moric Benyovszky" location="The" municipality="Polish-Hungarian" pageId="5" pageNumber="449" specimenCount="1">
<emphasis id="B966EAA1FF93180EF381FB3908E6FB21" bold="true" box="[199,392,1196,1221]" pageId="5" pageNumber="449">ETYMOLOGY.</emphasis>
<location id="8ECD6068FF93180EF2D4FB3908D0FB21" LSID="urn:lsid:plazi:treatment:03BB87A5FF92180EF3D1F9660D29FAD4:8ECD6068FF93180EF2D4FB3908D0FB21" box="[402,446,1196,1221]" country="Madagascar" county="Moric Benyovszky" municipality="Polish-Hungarian" name="The" pageId="5" pageNumber="449">The</location>
species is named after the
<collectingMunicipality id="6BC9ACC9FF93180EF1B9FB390AAAFB21" box="[767,964,1196,1221]" pageId="5" pageNumber="449">Polish-Hungarian</collectingMunicipality>
traveller and king of
<collectingCountry id="F3057623FF93180EF781FB390C3EFB21" box="[1223,1360,1196,1221]" name="Madagascar" pageId="5" pageNumber="449">Madagascar</collectingCountry>
, count
<collectingCounty id="62CC4E3FFF93180EF3D1FB5A0802FB0C" box="[151,364,1231,1256]" pageId="5" pageNumber="449">Móric Benyovszky</collectingCounty>
(
<collectingDate id="EFE8E99BFF93180EF23AFB5A0B6EFB0C" box="[380,512,1231,1256]" pageId="5" pageNumber="449" value="1746" valueMax="1786">
<date id="FFAC1073FF93180EF23AFB5A0B6EFB0C" box="[380,512,1231,1256]" pageId="5" pageNumber="449" value="1746" valueMax="1786">1746–1786</date>
</collectingDate>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3086538FF93180EF381FB610D29FAD4" pageId="5" pageNumber="449" type="distribution">
<paragraph id="8BAD36B3FF93180EF381FB610D29FAD4" blockId="5.[151,1437,151,1328]" pageId="5" pageNumber="449">
<emphasis id="B966EAA1FF93180EF381FB6108F3FAE9" bold="true" box="[199,413,1268,1293]" pageId="5" pageNumber="449">DISTRIBUTION.</emphasis>
<taxonomicName id="4C124D30FF93180EF2E0FB610BCAFAE8" box="[422,676,1268,1292]" class="Insecta" family="Geocoridae" genus="Germalus" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="449" phylum="Arthropoda" rank="species" species="benyovszkyi">
<emphasis id="B966EAA1FF93180EF2E0FB610BCAFAE8" box="[422,676,1268,1292]" italics="true" pageId="5" pageNumber="449">Germalus benyovszkyi</emphasis>
</taxonomicName>
has a single record from North
<collectingCountry id="F3057623FF93180EF75FFB610DCEFAE9" box="[1049,1184,1268,1293]" name="Madagascar" pageId="5" pageNumber="449">Madagascar</collectingCountry>
(
<figureCitation id="13292A36FF93180EF7F4FB610D9DFAE8" box="[1202,1267,1268,1293]" captionStart="FIGURE 1" captionStartId="2.[151,250,1990,2012]" captionTargetBox="[172,1381,439,1953]" captionTargetId="figure@2.[152,1401,419,2037]" captionTargetPageId="2" captionText="FIGURE 1. Distribution records of Germalus spp." httpUri="https://zenodo.org/record/187237/files/figure.png" pageId="5" pageNumber="449">Fig. 1</figureCitation>
B). Due to the lack of further specimens in museum collections this species can be considered rare.
</paragraph>
</subSubSection>
</treatment>
</document>
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