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<document ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E36CE21EFE45A2015E88FDC1" docLanguage="en" docName="JNATHIST.53.2526.15331584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Turritella capistrata DeVries 2019, sp. nov." docType="treatment" docVersion="19" lastPageNumber="1569" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1568" updateTime="1636522412784" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>DeVries, Thomas J.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2018-12-17</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>53</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>25</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1533</mods:start>
<mods:end>1584</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1080/00222933.2018.1524032</mods:identifier>
<mods:identifier type="GBIF-Dataset">034ebcd9-efa7-416a-822c-b6aee0e57a1c</mods:identifier>
<mods:identifier type="ISSN">1464-5262</mods:identifier>
<mods:identifier type="Zenodo-Dep">3670229</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671268" ID-GBIF-Taxon="162146476" ID-Zenodo-Dep="3671268" LSID="urn:lsid:plazi:treatment:03BB2338E36CE21EFE45A2015E88FDC1" httpUri="http://treatment.plazi.org/id/03BB2338E36CE21EFE45A2015E88FDC1" lastPageId="37" lastPageNumber="1569" pageId="36" pageNumber="1568">
<subSubSection box="[484,832,834,859]" pageId="36" pageNumber="1568" type="nomenclature">
<paragraph blockId="36.[484,832,834,893]" box="[484,832,834,859]" pageId="36" pageNumber="1568">
<heading box="[484,832,834,859]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="1568" reason="2">
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[484,729,834,859]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="1568" phylum="Mollusca" rank="species" species="capistrata" status="sp. nov.">
<emphasis bold="true" box="[484,729,834,859]" italics="true" pageId="36" pageNumber="1568">Turritella capistrata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[739,832,834,859]" pageId="36" pageNumber="1568" rank="species">sp. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection box="[558,757,868,893]" pageId="36" pageNumber="1568" type="description">
<paragraph blockId="36.[484,832,834,893]" box="[558,757,868,893]" pageId="36" pageNumber="1568">
<heading box="[558,757,868,893]" centered="true" fontSize="11" level="2" pageId="36" pageNumber="1568" reason="5">
(
<figureCitation box="[566,666,868,893]" captionStart="Figure 5" captionStartId="14.[160,225,1068,1091]" captionTargetBox="[164,1150,144,1035]" captionTargetId="figure@14.[162,1153,142,1037]" captionTargetPageId="14" captionText="Figure 5. Gastropods from the Caballas Formation. (ae) Caballasea segmentata gen. nov., sp. nov. B8769. (a) UWBM 107607, paratype, L 12.4 mm, apertural view. (b) UWBM 107607, abapertural view. (c) UWBM 107606, holotype, L 12.6 mm, abapertural view. (d) MUSM INV 262, paratype, L 8.3 mm, oblique view of parietal callus and columella. (e) UWBM 107608, paratype, L 15.3 mm, abapertural view. (f, g) Turritella capistrata sp. nov. (f) MUSM INV 259, paratype, B8772, L 16.3 mm, lateral view. (g) UWBM 107601, holotype, B8770, L 14.4 mm, apertural view. (h) Turritella saposa Olsson, 1944, UWBM 107602, B8770, L 15.5 mm. (i) Cristispira caballasensis sp. nov., UWBM 107605, holotype, B8769, L 5.1 mm, fragment. (j, k, m, n) Rhombopsis longinquus sp. nov. (j) UWBM 107630, holotype, B8772, L 18.3 mm, apertural view, outer lip missing. (k) MUSM INV 269, paratype, B8769, L 19.9 mm, abapertural view. (m) UWBM 107631, paratype, B8772, L 22.5 mm, abapertural view. (n) UWBM 107632, paratype, B8769, L 16.4 mm, lateral view of spire and columella. (l, oq) Tristeirostoma saxaquadrata sp. nov. B8771. (l) UWBM 107636, paratype, L 10.0 mm, spire. (o) UWBM 107,635, holotype, L 7.6 mm, abapertural view. (p) UWBM 107635, lateral view of columella. (q) UWBM 107635, apertural/lateral view." figureDoi="http://doi.org/10.5281/zenodo.3670237" httpUri="https://zenodo.org/record/3670237/files/figure.png" pageId="36" pageNumber="1568">Figure 5</figureCitation>
(f), 5(g))
</heading>
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="1568" type="diagnosis">
<paragraph blockId="36.[160,1156,937,1033]" box="[160,276,937,963]" pageId="36" pageNumber="1568">
<emphasis bold="true" box="[160,276,937,963]" italics="true" pageId="36" pageNumber="1568">Diagnosis</emphasis>
</paragraph>
<paragraph blockId="36.[160,1156,937,1033]" pageId="36" pageNumber="1568">Whorls imbricate, with three strong primary spiral cords, nearly evenly spaced between sutures. Lateral growth line prosocline with anterior inflexion only.</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="1568" type="description">
<paragraph blockId="36.[160,1156,1077,1415]" box="[160,294,1077,1103]" pageId="36" pageNumber="1568">
<emphasis bold="true" box="[160,294,1077,1103]" italics="true" pageId="36" pageNumber="1568">Description</emphasis>
</paragraph>
<paragraph blockId="36.[160,1156,1077,1415]" pageId="36" pageNumber="1568">
Estimated shell length to 25 mm, pleural angle about 19°. Protoconch unknown. Spire with about 10 whorls. Earliest preserved whorls straight-sided to concave, imbricate; spiral formula C1A1 and subsequently D2C1B2A1. Later whorls imbricate with strong primary spirals and insertion of secondary spirals; successive formulae D2C
<quantity box="[1035,1054,1217,1244]" metricMagnitude="3" metricUnit="kg" metricValue="1.0" pageId="36" pageNumber="1568" unit="t" value="1.0">1t</quantity>
3B2s3A1r3 and D2C
<quantity box="[255,274,1252,1278]" metricMagnitude="3" metricUnit="kg" metricValue="1.0" pageId="36" pageNumber="1568" unit="t" value="1.0">1t</quantity>
3B2s3s4A1r3r4; primary spiral
<emphasis box="[593,599,1252,1276]" italics="true" pageId="36" pageNumber="1568"></emphasis>
D
<emphasis box="[616,622,1252,1276]" italics="true" pageId="36" pageNumber="1568"></emphasis>
variably exposed, primary spiral
<emphasis box="[999,1004,1252,1276]" italics="true" pageId="36" pageNumber="1568"></emphasis>
B
<emphasis box="[1019,1025,1252,1276]" italics="true" pageId="36" pageNumber="1568"></emphasis>
sometimes weaker than primary spirals
<emphasis box="[477,483,1286,1310]" italics="true" pageId="36" pageNumber="1568"></emphasis>
A
<emphasis box="[499,505,1286,1310]" italics="true" pageId="36" pageNumber="1568"></emphasis>
and
<emphasis box="[561,567,1286,1310]" italics="true" pageId="36" pageNumber="1568"></emphasis>
C
<emphasis box="[582,588,1286,1310]" italics="true" pageId="36" pageNumber="1568"></emphasis>
. Spacing formula C27B46A77 (n = 4). Lateral growth line orthocline to slightly prosocline, inflected anteriorly only. Lateral sinus moderately deep, apex situated close to spiral
<emphasis box="[549,555,1356,1380]" italics="true" pageId="36" pageNumber="1568"></emphasis>
B
<emphasis box="[569,575,1356,1380]" italics="true" pageId="36" pageNumber="1568"></emphasis>
. Base convex with at least two spirals, basal growth line probably radial, but base mostly obscured. Allmon formula 2?-4-S-B-OR.
</paragraph>
</subSubSection>
<subSubSection pageId="36" pageNumber="1568" type="discussion">
<paragraph blockId="36.[160,1156,1459,1727]" box="[160,262,1459,1485]" pageId="36" pageNumber="1568">
<emphasis bold="true" box="[160,262,1459,1485]" italics="true" pageId="36" pageNumber="1568">Remarks</emphasis>
</paragraph>
<paragraph blockId="36.[160,1156,1459,1727]" pageId="36" pageNumber="1568">
Specimens of
<taxonomicName authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[326,542,1495,1519]" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="1568" phylum="Mollusca" rank="species" species="capistrata" status="sp. nov.">
<emphasis box="[326,542,1495,1519]" italics="true" pageId="36" pageNumber="1568">Turritella capistrata</emphasis>
</taxonomicName>
<taxonomicNameLabel box="[554,645,1495,1519]" pageId="36" pageNumber="1568" rank="species">sp. nov.</taxonomicNameLabel>
most closely resemble those of
<taxonomicName authority="Olssoni 1944" authorityName="Olssoni" authorityYear="1944" class="Gastropoda" family="Turritellidae" genus="Turritella" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="36" pageNumber="1568" phylum="Mollusca" rank="species" species="desolata">
<emphasis box="[1033,1155,1495,1519]" italics="true" pageId="36" pageNumber="1568">T. desolata</emphasis>
Olsson, 1944
</taxonomicName>
, a late Maastrichtian species from the
<collectingCountry box="[762,850,1530,1554]" name="Haiti" pageId="36" pageNumber="1568">Tortuga</collectingCountry>
Formation in the Sechura Basin of northern
<collectingCountry box="[377,428,1565,1589]" name="Peru" pageId="36" pageNumber="1568">Peru</collectingCountry>
(
<bibRefCitation author="Olsson AA" box="[446,590,1565,1589]" pageId="36" pageNumber="1568" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson 1944</bibRefCitation>
; Jaillard et al. 2005) and the Upper Cretaceous Chonta Formation of eastern
<collectingCountry box="[520,571,1599,1623]" name="Peru" pageId="36" pageNumber="1568">Peru</collectingCountry>
(
<bibRefCitation author="Singewald JT Jr." box="[592,779,1599,1623]" pageId="36" pageNumber="1568" pagination="447 - 464" refId="ref26385" refString="Singewald JT Jr. 1926. Geology of the Pichis and Pachitea Riversi Peru. Bull Geol Soc Am. 39: 447 - 464." type="journal article" year="1926">Singewald 1926</bibRefCitation>
;
<bibRefCitation author="Pilsbry HA" box="[799,942,1599,1623]" pageId="36" pageNumber="1568" pagination="137 - 153" refId="ref25981" refString="Pilsbry HA. 1944. Molluscan fossils from the Rio Pachitea and vicinity in eastern Peru. Proc Acad Nat Sci Philadelphia. 96: 137 - 153." type="journal article" year="1944">Pilsbry 1944</bibRefCitation>
; CGS Consultores Associados S.A 1997;
<bibRefCitation author="Leon Wi Cossio Ai Garcia W" box="[413,604,1634,1658]" pageId="36" pageNumber="1568" pagination="1 - 164" refId="ref24744" refString="Leon Wi Cossio Ai Garcia W. 1997. Geologia de los cuadrangulos de Obenteni y Atalaya. Inst Geologico Minero y Metalurgico Serie A Boletin. 95: 1 - 164." type="journal article" year="1997">Leon et al. 1997</bibRefCitation>
). The
<collectingCountry box="[682,770,1634,1658]" name="Haiti" pageId="36" pageNumber="1568">Tortuga</collectingCountry>
Formation species has a smaller pleural angle (12°; n = 2) and is larger (&gt;
<quantity box="[672,764,1669,1693]" metricMagnitude="-2" metricUnit="m" metricValue="5.0" pageId="36" pageNumber="1568" unit="mm" value="50.0">50 mm</quantity>
;
<bibRefCitation author="Olsson AA" box="[781,925,1669,1693]" pageId="36" pageNumber="1568" pagination="1 - 147" refId="ref25769" refString="Olsson AA. 1944. Contributions to the Tertiary paleontology of northern Peru. Part VII. The Cretaceous of the Paita region. Bulletins Am Paleontol. 19 (68): 1 - 147." type="journal article" year="1944">Olsson 1944</bibRefCitation>
) than the Caballas Formation species, but the spiral sculpture is identical.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="1569" type="etymology">
<paragraph blockId="37.[160,1155,141,236]" box="[160,285,141,167]" pageId="37" pageNumber="1569">
<emphasis bold="true" box="[160,285,141,167]" italics="true" pageId="37" pageNumber="1569">Etymology</emphasis>
</paragraph>
<paragraph blockId="37.[160,1155,141,236]" pageId="37" pageNumber="1569">
<emphasis box="[160,166,178,202]" italics="true" pageId="37" pageNumber="1569"></emphasis>
Capistrata
<emphasis box="[278,284,178,202]" italics="true" pageId="37" pageNumber="1569"></emphasis>
, Latin adjective for
<emphasis box="[519,525,178,202]" italics="true" pageId="37" pageNumber="1569"></emphasis>
bound
<emphasis box="[599,604,178,202]" italics="true" pageId="37" pageNumber="1569"></emphasis>
, referencing the seeming binding of each capstan-like whorl with coils of rope.
</paragraph>
</subSubSection>
<subSubSection pageId="37" pageNumber="1569" type="materials_examined">
<paragraph blockId="37.[160,1156,286,416]" box="[160,259,286,312]" pageId="37" pageNumber="1569">
<emphasis bold="true" box="[160,259,286,312]" italics="true" pageId="37" pageNumber="1569">Material</emphasis>
</paragraph>
<paragraph blockId="37.[160,1156,286,416]" pageId="37" pageNumber="1569">
<materialsCitation ID-GBIF-Occurrence="2572183369" box="[160,861,322,346]" collectionCode="UWBM" pageId="37" pageNumber="1569" specimenCode="UWBM 107601" specimenCount="1" typeStatus="holotype">
<specimenCode box="[160,329,322,346]" pageId="37" pageNumber="1569">UWBM 107601</specimenCode>
,
<typeStatus box="[340,443,322,346]" pageId="37" pageNumber="1569">holotype</typeStatus>
, B8770 (type locality), L (14.4), W 8.6
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572183360" collectionCode="UWBM" pageId="37" pageNumber="1569" specimenCode="UWBM 107603" specimenCount="1" typeStatus="paratype">
remainder are
<typeStatus box="[1040,1153,322,346]" pageId="37" pageNumber="1569">paratypes</typeStatus>
:
<specimenCode box="[160,329,357,381]" pageId="37" pageNumber="1569">UWBM 107603</specimenCode>
, B8770, L 24.0, W 7.1
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572183375" box="[583,1023,357,381]" collectionCode="UWBM" pageId="37" pageNumber="1569" specimenCode="UWBM 107604" specimenCount="1">
<specimenCode box="[583,752,357,381]" pageId="37" pageNumber="1569">UWBM 107604</specimenCode>
, B8770, L (18.5), W 10.4
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572183372" collectionCode="MUSM" pageId="37" pageNumber="1569" specimenCode="INV 259" specimenCount="1">
<collectionCode box="[1035,1108,357,381]" pageId="37" pageNumber="1569">MUSM</collectionCode>
<specimenCode pageId="37" pageNumber="1569">INV 259</specimenCode>
, B8772, L (16.3), W (7.5)
</materialsCitation>
;
<materialsCitation ID-GBIF-Occurrence="2572183371" box="[490,937,392,416]" collectionCode="MUSM" pageId="37" pageNumber="1569" specimenCode="INV 260" specimenCount="1">
<collectionCode box="[490,563,392,416]" pageId="37" pageNumber="1569">MUSM</collectionCode>
<specimenCode box="[572,664,392,416]" pageId="37" pageNumber="1569">INV 260</specimenCode>
, B8770, L (21.5), W (8.3)
</materialsCitation>
</paragraph>
<paragraph blockId="37.[160,1156,286,416]" box="[938,942,392,416]" pageId="37" pageNumber="1569">.</paragraph>
<paragraph blockId="37.[160,1156,465,525]" box="[160,292,465,491]" pageId="37" pageNumber="1569">
<emphasis bold="true" box="[160,292,465,491]" italics="true" pageId="37" pageNumber="1569">Occurrence</emphasis>
</paragraph>
<paragraph blockId="37.[160,1156,465,525]" box="[160,1156,501,525]" pageId="37" pageNumber="1569">
Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry box="[1099,1153,501,525]" name="Peru" pageId="37" pageNumber="1569">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>