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<document id="C86250D3DDC4FBA33061E7F8B462C9C9" ID-CLB-Dataset="24019" ID-DOI="10.1080/00222933.2018.1524032" ID-GBIF-Dataset="034ebcd9-efa7-416a-822c-b6aee0e57a1c" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670229" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941790231" checkinUser="valdenar" docAuthor="DeVries, Thomas J." docDate="2019" docId="03BB2338E36AE218FE48A4975B4BFBB0" docLanguage="en" docName="JNATHIST.53.2526.15331584.pdf.imf" docOrigin="Journal of Natural History 53 (25)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Caballasea DeVries 2019, gen. nov." docType="treatment" docVersion="22" lastPageNumber="1567" masterDocId="FF825B40E348E23BFFA1A1435A0CFFCC" masterDocTitle="Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)" masterLastPageNumber="1584" masterPageNumber="1533" pageNumber="1566" updateTime="1698783606523" updateUser="ExternalLinkService">
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<mods:title id="4F4A57863A0983E716906D43B99B740D">Early Paleogene brackish-water molluscs from the Caballas Formation of the East Pisco Basin (Southern Peru)</mods:title>
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<mods:namePart id="70CD456F82C5AA65BA485884ED009ECD">DeVries, Thomas J.</mods:namePart>
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<mods:date id="BABE277ECF11B86EE0E3FB8EBC44DDBD">2019</mods:date>
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<treatment id="03BB2338E36AE218FE48A4975B4BFBB0" ID-DOI="http://doi.org/10.5281/zenodo.3671264" ID-GBIF-Taxon="162146503" ID-Zenodo-Dep="3671264" LSID="urn:lsid:plazi:treatment:03BB2338E36AE218FE48A4975B4BFBB0" httpUri="http://treatment.plazi.org/id/03BB2338E36AE218FE48A4975B4BFBB0" lastPageId="35" lastPageNumber="1567" pageId="34" pageNumber="1566">
<subSubSection id="C308C1A5E36AE219FE48A4975937FA21" box="[489,827,1492,1517]" pageId="34" pageNumber="1566" type="nomenclature">
<paragraph id="8BAD922EE36AE219FE48A4975937FA21" blockId="34.[489,827,1492,1517]" box="[489,827,1492,1517]" pageId="34" pageNumber="1566">
<heading id="D0E52542E36AE219FE48A4975937FA21" box="[489,827,1492,1517]" centered="true" fontSize="11" level="2" pageId="34" pageNumber="1566" reason="2">
Genus
<taxonomicName id="4C12E9ADE36AE219FD9DA49758CFFA21" ID-CoL="8KRYW" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[572,707,1492,1517]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="34" pageNumber="1566" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9664E3CE36AE219FD9DA49758CFFA21" bold="true" box="[572,707,1492,1517]" italics="true" pageId="34" pageNumber="1566">Caballasea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E36AE219FD6DA4975937FA21" box="[716,827,1492,1517]" pageId="34" pageNumber="1566" rank="genus">gen. nov.</taxonomicNameLabel>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36AE219FF01A7545B8BF9B9" pageId="34" pageNumber="1566" type="materials_examined">
<paragraph id="8BAD922EE36AE219FF01A7545E1DF9E3" blockId="34.[160,1155,1559,1653]" box="[160,1041,1559,1583]" pageId="34" pageNumber="1566">
<typeStatus id="54A92C8CE36AE219FF01A7545ADAF9E3" box="[160,214,1559,1583]" pageId="34" pageNumber="1566">Type</typeStatus>
species:
<taxonomicName id="4C12E9ADE36AE219FE9EA7545832F9E3" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[319,574,1559,1583]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="segmentata" status="sp. nov.">
<emphasis id="B9664E3CE36AE219FE9EA7545832F9E3" box="[319,574,1559,1583]" italics="true" pageId="34" pageNumber="1566">Caballasea segmentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E36AE219FDE6A75458ACF9E3" box="[583,672,1559,1583]" pageId="34" pageNumber="1566" rank="species">sp. nov.</taxonomicNameLabel>
Early Paleogene, southern
<collectingCountry id="F305D2BEE36AE219FC79A7545E02F9E3" box="[984,1038,1559,1583]" name="Peru" pageId="34" pageNumber="1566">Peru</collectingCountry>
.
</paragraph>
<paragraph id="8BAD922EE36AE219FF01A7795B8BF9B9" blockId="34.[160,1155,1559,1653]" pageId="34" pageNumber="1566">
Species included:
<taxonomicName id="4C12E9ADE36AE219FED2A7795879F99E" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[371,629,1594,1618]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="segmentata" status="sp. nov.">
<emphasis id="B9664E3CE36AE219FED2A7795879F99E" box="[371,629,1594,1618]" italics="true" pageId="34" pageNumber="1566">Caballasea segmentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E36AE219FD20A77958ECF99E" box="[641,736,1594,1618]" pageId="34" pageNumber="1566" rank="species">sp. nov.</taxonomicNameLabel>
,
<taxonomicName id="4C12E9ADE36AE219FD4FA7795B8DF9B9" authority="Perrilliat et al., 2008" authorityName="Perrilliat" authorityYear="2008" class="Gastropoda" family="Pachychilidae" genus="Pachychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="34" pageNumber="1566" phylum="Mollusca" rank="species" species="lawtoni" subGenus="Pachychiloides">
<emphasis id="B9664E3CE36AE219FD4FA7795960F99E" box="[750,876,1594,1618]" italics="true" pageId="34" pageNumber="1566">Pachychilus</emphasis>
(
<emphasis id="B9664E3CE36AE219FC21A7795E12F99E" box="[896,1054,1594,1618]" italics="true" pageId="34" pageNumber="1566">Pachychiloides</emphasis>
)
<emphasis id="B9664E3CE36AE219FB93A7795E8FF99E" box="[1074,1155,1594,1618]" italics="true" pageId="34" pageNumber="1566">lawtoni</emphasis>
Perrilliat et al., 2008
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36BE218FF01A1CE5E8EFEC3" pageId="35" pageNumber="1567" type="diagnosis">
<paragraph id="8BAD922EE36BE218FF01A1CE5B18FF6B" blockId="35.[160,1155,141,271]" box="[160,276,141,167]" pageId="35" pageNumber="1567">
<emphasis id="B9664E3CE36BE218FF01A1CE5B18FF6B" bold="true" box="[160,276,141,167]" italics="true" pageId="35" pageNumber="1567">Diagnosis</emphasis>
</paragraph>
<paragraph id="8BAD922EE36BE218FF01A1F15E8EFEC3" blockId="35.[160,1155,141,271]" pageId="35" pageNumber="1567">Shell small, elongate; sutures canaliculate. Teleoconch whorls with axial ribs, rounded or erect as flanges. Aperture elongate. Outer lip not thickened; short, well-defined, very shallow anterior canal present; posterior canal absent. No varices or ventrolateral varix.</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36BE218FF01A07D59BAFC4F" pageId="35" pageNumber="1567" type="discussion">
<paragraph id="8BAD922EE36BE218FF01A07D5B0AFE94" blockId="35.[160,1156,318,899]" box="[160,262,318,344]" pageId="35" pageNumber="1567">
<emphasis id="B9664E3CE36BE218FF01A07D5B0AFE94" bold="true" box="[160,262,318,344]" italics="true" pageId="35" pageNumber="1567">Remarks</emphasis>
</paragraph>
<paragraph id="8BAD922EE36BE218FF01A0205842FD34" blockId="35.[160,1156,318,899]" pageId="35" pageNumber="1567">
In Peru,
<taxonomicName id="4C12E9ADE36BE218FF5FA0205B79FEB7" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[254,373,355,379]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="35" pageNumber="1567" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9664E3CE36BE218FF5FA0205B79FEB7" box="[254,373,355,379]" italics="true" pageId="35" pageNumber="1567">Caballasea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E36BE218FEDCA0205BEBFEB7" box="[381,487,355,379]" pageId="35" pageNumber="1567" rank="genus">gen. nov.</taxonomicNameLabel>
is known from just one species,
<taxonomicName id="4C12E9ADE36BE218FCFDA0205E57FEB7" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[860,1115,355,379]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="segmentata" status="sp. nov.">
<emphasis id="B9664E3CE36BE218FCFDA0205E57FEB7" box="[860,1115,355,379]" italics="true" pageId="35" pageNumber="1567">Caballasea segmentata</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A255F347E36BE218FBC5A0205ADDFE51" pageId="35" pageNumber="1567" rank="species">sp. nov.</taxonomicNameLabel>
In most characters, the Caballas Formation species resembles
<taxonomicName id="4C12E9ADE36BE218FBA4A0C65892FE0C" authority="Perrilliat et al., 2008" authorityName="Perrilliat" authorityYear="2008" class="Gastropoda" family="Pachychilidae" genus="Pachychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="lawtoni" subGenus="Pachychiloides">
<emphasis id="B9664E3CE36BE218FBA4A0C65E8FFE51" box="[1029,1155,389,413]" italics="true" pageId="35" pageNumber="1567">Pachychilus</emphasis>
(
<emphasis id="B9664E3CE36BE218FF06A0EB5B4AFE0C" box="[167,326,424,448]" italics="true" pageId="35" pageNumber="1567">Pachychiloides</emphasis>
)
<emphasis id="B9664E3CE36BE218FEF8A0EB5BA6FE0C" box="[345,426,424,448]" italics="true" pageId="35" pageNumber="1567">lawtoni</emphasis>
Perrilliat et al., 2008
</taxonomicName>
; a common species in Eocene fluvial red and green mudstones of the Carroza Formation of
<collectingCountry id="F305D2BEE36BE218FD5DA0885940FE2F" box="[764,844,459,483]" name="Mexico" pageId="35" pageNumber="1567">Mexico</collectingCountry>
(
<bibRefCitation id="EF83EFDFE36BE218FCFCA0885E4EFE2F" author="Perrilliat MCi Vega FJi Espinosa Bi Naranjo-Garcia E" box="[861,1090,459,483]" pageId="35" pageNumber="1567" pagination="255 - 266" refId="ref25877" refString="Perrilliat MCi Vega FJi Espinosa Bi Naranjo-Garcia E. 2008. Late Cretaceous and Paleogene freshwater gastropods from northeastern Mexico. J Paleontol. 82 (2): 255 - 266." type="journal article" year="2008">Perrilliat et al. 2008</bibRefCitation>
). The Carroza Formation specimens, for which no complete apertures are preserved, may not be properly placed in
<taxonomicName id="4C12E9ADE36BE218FE0DA3535844FDE4" box="[428,584,528,552]" class="Gastropoda" family="Pachychilidae" kingdom="Animalia" pageId="35" pageNumber="1567" phylum="Mollusca" rank="family">Pachychilidae</taxonomicName>
. All modern pachychilids, excepting
<taxonomicName id="4C12E9ADE36BE218FC5CA3535B50FD87" authority="(Linnaeusi 1758)" baseAuthorityName="Linnaeusi" baseAuthorityYear="1758" class="Gastropoda" family="Pachychilidae" genus="Faunus" kingdom="Animalia" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="ater">
<emphasis id="B9664E3CE36BE218FC5CA3535E8FFDE4" box="[1021,1155,528,552]" italics="true" pageId="35" pageNumber="1567">Faunus ater</emphasis>
(Linnaeus, 1758)
</taxonomicName>
, have apertures without anterior or posterior canals (
<bibRefCitation id="EF83EFDFE36BE218FC71A3705B67FDA1" author="Glaubrecht Mi von Rintelen T" pageId="35" pageNumber="1567" pagination="415 - 435" refId="ref23862" refString="Glaubrecht Mi von Rintelen T. 2003. Systematicsi molecular genetics and historical zoogeography of the viviparous freshwater gastropod Pseudopotamis (Cerithioideai Pachychilidae): a relic on the Torres Strait Islandsi Australia. Zool Scr. 32 (5): 415 - 435." type="journal article" year="2003">Glaubrecht and von Rintelen 2003</bibRefCitation>
). This absence of apertural canals is true for
<taxonomicName id="4C12E9ADE36BE218FCFFA3165B35FD5C" authority="(Whitei 1876)" baseAuthorityName="Whitei" baseAuthorityYear="1876" class="Gastropoda" family="Pachychilidae" genus="Pachychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="cleburni" subGenus="Pachychiloides">
<emphasis id="B9664E3CE36BE218FCFFA3165960FDA1" box="[862,876,597,621]" italics="true" pageId="35" pageNumber="1567">P</emphasis>
. (
<emphasis id="B9664E3CE36BE218FC21A3165E13FDA1" box="[896,1055,597,621]" italics="true" pageId="35" pageNumber="1567">Pachychiloides</emphasis>
)
<emphasis id="B9664E3CE36BE218FB8CA3165E8FFDA1" box="[1069,1155,597,621]" italics="true" pageId="35" pageNumber="1567">cleburni</emphasis>
(White, 1876)
</taxonomicName>
from the Cretaceous Bear River Formation of Wyoming, a species with which
<bibRefCitation id="EF83EFDFE36BE218FF4AA3D85BDFFD7F" author="Perrilliat MCi Vega FJi Espinosa Bi Naranjo-Garcia E" box="[235,467,667,691]" pageId="35" pageNumber="1567" pagination="255 - 266" refId="ref25877" refString="Perrilliat MCi Vega FJi Espinosa Bi Naranjo-Garcia E. 2008. Late Cretaceous and Paleogene freshwater gastropods from northeastern Mexico. J Paleontol. 82 (2): 255 - 266." type="journal article" year="2008">Perrilliat et al. (2008)</bibRefCitation>
compared
<taxonomicName id="4C12E9ADE36BE218FDF5A3D85979FD7F" box="[596,885,667,691]" class="Gastropoda" family="Pachychilidae" genus="Pachychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="lawtoni" subGenus="Pachychiloides">
<emphasis id="B9664E3CE36BE218FDF5A3D8586DFD7F" box="[596,609,667,691]" italics="true" pageId="35" pageNumber="1567">P</emphasis>
. (
<emphasis id="B9664E3CE36BE218FDD7A3D85918FD7F" box="[630,788,667,691]" italics="true" pageId="35" pageNumber="1567">Pachychiloides</emphasis>
)
<emphasis id="B9664E3CE36BE218FC85A3D85979FD7F" box="[804,885,667,691]" italics="true" pageId="35" pageNumber="1567">lawtoni</emphasis>
</taxonomicName>
. The Eocene
<taxonomicName id="4C12E9ADE36BE218FBADA3D85B2CFD19" authorityName="DeVries" authorityYear="2019" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="segmentata">
<emphasis id="B9664E3CE36BE218FBADA3D85B2CFD19" italics="true" pageId="35" pageNumber="1567">Caballasea segmentata</emphasis>
</taxonomicName>
, however, which so closely resembles the Carroza Formation species, has a short and very shallow anterior canal.
</paragraph>
<paragraph id="8BAD922EE36BE218FF61A24059BAFC4F" blockId="35.[160,1156,318,899]" pageId="35" pageNumber="1567">
It is proposed herein that
<taxonomicName id="4C12E9ADE36BE218FDAAA24058E6FCD7" box="[523,746,771,795]" class="Gastropoda" family="Pachychilidae" genus="Pachychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="lawtoni">
<emphasis id="B9664E3CE36BE218FDAAA24058E6FCD7" box="[523,746,771,795]" italics="true" pageId="35" pageNumber="1567">Pachychilus lawtoni</emphasis>
</taxonomicName>
from the Carroza Formation and
<taxonomicName id="4C12E9ADE36BE218FF01A2665BA1FCF1" authorityName="DeVries" authorityYear="2019" box="[160,429,805,829]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="segmentata">
<emphasis id="B9664E3CE36BE218FF01A2665BA1FCF1" box="[160,429,805,829]" italics="true" pageId="35" pageNumber="1567">Caballasea segmentata</emphasis>
</taxonomicName>
from the Caballas Formation are congeneric and that
<taxonomicName id="4C12E9ADE36BE218FF01A20B5B71FCAC" box="[160,381,840,864]" class="Gastropoda" family="Pachychilidae" genus="Pachychilus" higherTaxonomySource="CoL" kingdom="Animalia" order="Littorinimorpha" pageId="35" pageNumber="1567" phylum="Mollusca" rank="species" species="lawtoni">
<emphasis id="B9664E3CE36BE218FF01A20B5B71FCAC" box="[160,381,840,864]" italics="true" pageId="35" pageNumber="1567">Pachychilus lawtoni</emphasis>
</taxonomicName>
be transferred to
<taxonomicName id="4C12E9ADE36BE218FDC0A20B58D4FCAC" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[609,728,840,864]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="35" pageNumber="1576" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9664E3CE36BE218FDC0A20B58D4FCAC" box="[609,728,840,864]" italics="true" pageId="35" pageNumber="1567">Caballasea</emphasis>
</taxonomicName>
. The two species of
<taxonomicName id="4C12E9ADE36BE218FC7DA20B5E5FFCAC" authority="DeVries, 2019" authorityName="DeVries" authorityYear="2019" box="[988,1107,840,864]" class="Gastropoda" family="Potamididae" genus="Caballasea" higherTaxonomySource="GBIF" kingdom="Animalia" order="Neotaenioglossa" pageId="35" pageNumber="1576" phylum="Mollusca" rank="genus" status="gen. nov.">
<emphasis id="B9664E3CE36BE218FC7DA20B5E5FFCAC" box="[988,1107,840,864]" italics="true" pageId="35" pageNumber="1567">Caballasea</emphasis>
</taxonomicName>
are assigned to
<taxonomicName id="4C12E9ADE36BE218FE8AA2285BB5FC4F" box="[299,441,875,899]" class="Gastropoda" family="Potamididae" kingdom="Animalia" pageId="35" pageNumber="1567" phylum="Mollusca" rank="family">Potamididae</taxonomicName>
based on the presence of an anterior canal.
</paragraph>
</subSubSection>
<subSubSection id="C308C1A5E36BE218FF01A2EE5B4BFBB0" pageId="35" pageNumber="1567" type="etymology">
<paragraph id="8BAD922EE36BE218FF01A2EE5B17FC09" blockId="35.[160,1042,941,1005]" box="[160,283,941,965]" pageId="35" pageNumber="1567">
<emphasis id="B9664E3CE36BE218FF01A2EE5B17FC09" bold="true" box="[160,283,941,965]" italics="true" pageId="35" pageNumber="1567">Etymology</emphasis>
</paragraph>
<paragraph id="8BAD922EE36BE218FF01A2965E1EFC21" blockId="35.[160,1042,941,1005]" box="[160,1042,981,1005]" pageId="35" pageNumber="1567">
<emphasis id="B9664E3CE36BE218FF01A2965AAAFC21" box="[160,166,981,1005]" italics="true" pageId="35" pageNumber="1567"></emphasis>
Caballas
<emphasis id="B9664E3CE36BE218FEA3A2965B0BFC21" box="[258,263,981,1005]" italics="true" pageId="35" pageNumber="1567"></emphasis>
, referencing the lithostratigraphic unit in which the genus is found.
</paragraph>
<paragraph id="8BAD922EE36BE218FF01A55E5B28FBFB" blockId="35.[160,1155,1053,1148]" box="[160,292,1053,1079]" pageId="35" pageNumber="1567">
<emphasis id="B9664E3CE36BE218FF01A55E5B28FBFB" bold="true" box="[160,292,1053,1079]" italics="true" pageId="35" pageNumber="1567">Occurrence</emphasis>
</paragraph>
<paragraph id="8BAD922EE36BE218FF01A5025B4BFBB0" blockId="35.[160,1155,1053,1148]" pageId="35" pageNumber="1567">
Eocene,
<collectingCountry id="F305D2BEE36BE218FF5CA5025B43FB95" box="[253,335,1089,1113]" name="Mexico" pageId="35" pageNumber="1567">Mexico</collectingCountry>
; Lower Paleogene, Cuenca Member, Caballas Formation, East Pisco Basin, southern
<collectingCountry id="F305D2BEE36BE218FEACA5275B4FFBB0" box="[269,323,1124,1148]" name="Peru" pageId="35" pageNumber="1567">Peru</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>