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<document id="0CCA1243209D08EA751B0BAC9661C610" ID-CLB-Dataset="32165" ID-DOI="10.11646/zootaxa.4318.2.9" ID-GBIF-Dataset="1209f677-869b-4bb7-8456-997e7c214bbd" ID-ISSN="1175-5326" ID-Zenodo-Dep="886860" ID-ZooBank="86B553F5-C42D-484E-8197-Df90523Ca8C4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1504770914019" checkinUser="plazi" docAuthor="Gibon, François-Marie" docDate="2017" docId="03B9390A2717ED6C678E3FC6DA7044EE" docLanguage="en" docName="zootaxa.4318.2.9.pdf" docOrigin="Zootaxa 4318 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tinodes tafo Gibon 2017, n. sp." docType="treatment" docVersion="7" lastPageNumber="366" masterDocId="FF8041722716ED6E67193923D911430B" masterDocTitle="New species of Tinodes from Madagascar (Trichoptera, Psychomyiidae)" masterLastPageNumber="376" masterPageNumber="364" pageNumber="365" updateTime="1698468072499" updateUser="ExternalLinkService">
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<mods:title id="D166F2E6FB45DFB3B76DDDCC38601EED">New species of Tinodes from Madagascar (Trichoptera, Psychomyiidae)</mods:title>
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<mods:namePart id="8E4FAD30923614356E7FD8834FC8D448">Gibon, François-Marie</mods:namePart>
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<mods:date id="4530ECC60E58CC550DABEA9AA969A832">2017</mods:date>
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<treatment id="03B9390A2717ED6C678E3FC6DA7044EE" ID-DOI="http://doi.org/10.5281/zenodo.6023730" ID-GBIF-Taxon="133186668" ID-Zenodo-Dep="6023730" LSID="urn:lsid:plazi:treatment:03B9390A2717ED6C678E3FC6DA7044EE" httpUri="http://treatment.plazi.org/id/03B9390A2717ED6C678E3FC6DA7044EE" lastPageId="2" lastPageNumber="366" pageId="1" pageNumber="365">
<subSubSection id="C30ADB972717ED6F678E3FC6D80F442B" pageId="1" pageNumber="365" type="nomenclature">
<paragraph id="8BAF881C2717ED6F678E3FC6D86745F4" blockId="1.[151,374,1765,1824]" box="[151,374,1765,1791]" pageId="1" pageNumber="365">
<heading id="D0E73F702717ED6F678E3FC6D86745F4" bold="true" box="[151,374,1765,1791]" fontSize="11" level="1" pageId="1" pageNumber="365" reason="1">
<emphasis id="B964540E2717ED6F678E3FC6D86745F4" bold="true" box="[151,374,1765,1791]" pageId="1" pageNumber="365">
<taxonomicName id="4C10F39F2717ED6F678E3FC6D83D45F4" authority="Gibon, 2017" authorityName="Gibon" authorityYear="2017" box="[151,300,1765,1791]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="tafo" status="sp. nov.">
<emphasis id="B964540E2717ED6F678E3FC6D83D45F4" bold="true" box="[151,300,1765,1791]" italics="true" pageId="1" pageNumber="365">Tinodes tafo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257E9752717ED6F662A3FC6D86745F4" box="[307,374,1765,1791]" pageId="1" pageNumber="365" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BAF881C2717ED6F678E3E24D80F442B" blockId="1.[151,374,1765,1824]" box="[151,286,1799,1824]" pageId="1" pageNumber="365">
<figureCitation id="132B94992717ED6F678E3E24D9C9442B" box="[151,216,1799,1824]" captionStart="FIGURE 1" captionStartId="3.[151,250,1863,1885]" captionTargetBox="[166,1432,196,1839]" captionTargetId="figure@3.[151,1436,195,1841]" captionTargetPageId="3" captionText="FIGURE 1. Tinodes tafo n. sp., male genitalia: 1 A, left lateral; 1 B, dorsal, without inferior appendages; 1 C, segment IX and inferior appendages, ventral; 1 D, left lateral (specimen from Mandrare River watershed); 1 E, inferior appendages, ventral (specimen from Mandrare River watershed). Abbreviations: g. = phallic guide; g. ap. = phallic guide apodeme; inf. app. 1 = first segment of inferior appendage; inf. app. 2 = second segment of inferior appendage; int. app. = intermediate appendage; phall. = phallic apparatus; pre. app. = preanal appendage; st. IX = sternite IX; t. IX = tergite IX. Scale bar: 0.25 mm." httpUri="https://zenodo.org/record/886862/files/figure.png" pageId="1" pageNumber="365">Figs 1</figureCitation>
A1E
</paragraph>
</subSubSection>
<subSubSection id="C30ADB972717ED6C678E3E6CD8BA43F3" lastPageId="2" lastPageNumber="366" pageId="1" pageNumber="365" type="diagnosis">
<paragraph id="8BAF881C2717ED6C678E3E6CD8BA43F3" blockId="1.[151,1437,1871,2005]" lastBlockId="2.[151,1437,151,2021]" lastPageId="2" lastPageNumber="366" pageId="1" pageNumber="365">
<emphasis id="B964540E2717ED6F678E3E6CD8004463" bold="true" box="[151,273,1871,1896]" pageId="1" pageNumber="365">Diagnosis.</emphasis>
<taxonomicName id="4C10F39F2717ED6F66033E6CD8B74463" authority="Gibon, 2017" authorityName="Gibon" authorityYear="2017" box="[282,422,1871,1896]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="tafo" status="sp. nov.">
<emphasis id="B964540E2717ED6F66033E6CD8B74463" box="[282,422,1871,1896]" italics="true" pageId="1" pageNumber="365">Tinodes tafo</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A257E9752717ED6F66B63E6CD8E14463" box="[431,496,1871,1896]" pageId="1" pageNumber="365" rank="species">
<emphasis id="B964540E2717ED6F66B63E6CD8E14463" bold="true" box="[431,496,1871,1896]" pageId="1" pageNumber="365">n. sp.</emphasis>
</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C10F39F2717ED6F65DF3E72DA094463" authorityName="Melnitsky &amp; Ivanov" authorityYear="2016" box="[710,792,1871,1896]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="isalo">
<emphasis id="B964540E2717ED6F65DF3E72DA094463" box="[710,792,1871,1896]" italics="true" pageId="1" pageNumber="365">T. isalo</emphasis>
</taxonomicName>
. The two species can be distinguished by the following characters: (1) There are two sclerites in the endotheca of
<taxonomicName id="4C10F39F2717ED6F64073E56DA774487" authorityName="Gibon" authorityYear="2017" box="[798,870,1908,1932]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="tafo">
<emphasis id="B964540E2717ED6F64073E56DA774487" box="[798,870,1908,1932]" italics="true" pageId="1" pageNumber="365">T. tafo</emphasis>
</taxonomicName>
, the distal sclerite is stout, short, and spine-shaped and is also present in
<taxonomicName id="4C10F39F2717ED6F66963EBAD8F144BB" authorityName="Melnitsky &amp; Ivanov" authorityYear="2016" box="[399,480,1943,1968]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="isalo">
<emphasis id="B964540E2717ED6F66963EBAD8F144BB" box="[399,480,1943,1968]" italics="true" pageId="1" pageNumber="365">T. isalo</emphasis>
</taxonomicName>
; the proximal sclerite is long, slightly curved, and finger-shaped, but is absent in
<taxonomicName id="4C10F39F2717ED6F629E3EBAD9DC44DE" authorityName="Melnitsky &amp; Ivanov" authorityYear="2016" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="isalo">
<emphasis id="B964540E2717ED6F629E3EBAD9DC44DE" italics="true" pageId="1" pageNumber="365">T. isalo</emphasis>
</taxonomicName>
. (2) The phallic guide is long in
<taxonomicName id="4C10F39F2717ED6F65513E9EDB8044DF" authorityName="Gibon" authorityYear="2017" box="[584,657,1980,2004]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="1" pageNumber="365" phylum="Arthropoda" rank="species" species="tafo">
<emphasis id="B964540E2717ED6F65513E9EDB8044DF" box="[584,657,1980,2004]" italics="true" pageId="1" pageNumber="365">T. tafo</emphasis>
</taxonomicName>
, extending anterad into abdominal segment VIII, but is reduced to a small setose plate in
<taxonomicName id="4C10F39F2714ED6C669C39BAD8C743BB" authorityName="Melnitsky &amp; Ivanov" authorityYear="2016" box="[389,470,151,176]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="isalo">
<emphasis id="B964540E2714ED6C669C39BAD8C743BB" box="[389,470,151,176]" italics="true" pageId="2" pageNumber="366">T. isalo</emphasis>
</taxonomicName>
. (3) The harpago has a small basoventral bump when viewed laterally in
<taxonomicName id="4C10F39F2714ED6C623939BADC7843BB" authorityName="Gibon" authorityYear="2017" box="[1312,1385,152,176]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="tafo">
<emphasis id="B964540E2714ED6C623939BADC7843BB" box="[1312,1385,152,176]" italics="true" pageId="2" pageNumber="366">T. tafo</emphasis>
</taxonomicName>
, but this bump is absent in
<taxonomicName id="4C10F39F2714ED6C668E399ED8F943DE" authorityName="Melnitsky &amp; Ivanov" authorityYear="2016" box="[407,488,188,213]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="isalo">
<emphasis id="B964540E2714ED6C668E399ED8F943DE" box="[407,488,188,213]" italics="true" pageId="2" pageNumber="366">T. isalo</emphasis>
</taxonomicName>
. (4) The intermediate appendage branches of
<taxonomicName id="4C10F39F2714ED6C64EA399EDD2D43DF" authorityName="Gibon" authorityYear="2017" box="[1011,1084,188,212]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="tafo">
<emphasis id="B964540E2714ED6C64EA399EDD2D43DF" box="[1011,1084,188,212]" italics="true" pageId="2" pageNumber="366">T. tafo</emphasis>
</taxonomicName>
are stronger, more curved, and tend to be asymmetrical.
</paragraph>
</subSubSection>
<subSubSection id="C30ADB972714ED6C67DE3827DA27451A" pageId="2" pageNumber="366" type="description">
<paragraph id="8BAF881C2714ED6C67DE3827D86342BA" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="366">
<emphasis id="B964540E2714ED6C67DE3827D8484216" bold="true" box="[199,345,260,285]" pageId="2" pageNumber="366">Description.</emphasis>
General color pale brown, with dorsal sclerites slightly darker, wings brown with few small hyaline areas (in alcohol). Length of each forewing
<quantity id="4CE825F92714ED6C65C2380BDA734234" box="[731,866,296,320]" metricMagnitude="-3" metricUnit="m" metricValue="3.3499999999999996" metricValueMax="3.4" metricValueMin="3.3" pageId="2" pageNumber="366" unit="mm" value="3.3499999999999996" valueMax="3.4" valueMin="3.3">3.33.4 mm</quantity>
(n = 7) and of each hind wing
<quantity id="4CE825F92714ED6C63A23804DC534234" box="[1211,1346,295,320]" metricMagnitude="-3" metricUnit="m" metricValue="2.65" metricValueMax="2.7" metricValueMin="2.6" pageId="2" pageNumber="366" unit="mm" value="2.65" valueMax="2.7" valueMin="2.6">2.62.7 mm</quantity>
(n = 6). Forewing forks 2, 3, 4, and 5; crossveins
<emphasis id="B964540E2714ED6C657A386EDB7F426F" box="[611,622,333,356]" italics="true" pageId="2" pageNumber="366">r</emphasis>
(R1R2+3, hardly visible),
<emphasis id="B964540E2714ED6C6495386EDAA2426F" box="[908,947,333,356]" italics="true" pageId="2" pageNumber="366">r-m</emphasis>
(R4+5M1+2) and
<emphasis id="B964540E2714ED6C6375386EDD97426F" box="[1132,1158,333,356]" italics="true" pageId="2" pageNumber="366">cu</emphasis>
(Cu1Cu2); and hyaline line along stem of M present. Hind wing forks 2, 3, and 5; two anal veins; and crossveins
<emphasis id="B964540E2714ED6C63B73857DDA84280" box="[1198,1209,372,395]" italics="true" pageId="2" pageNumber="366">r</emphasis>
(R1 R2+3), and
<emphasis id="B964540E2714ED6C626D3857DC8A4280" box="[1396,1435,372,395]" italics="true" pageId="2" pageNumber="366">r-m</emphasis>
(R4+5M1+2) present.
</paragraph>
<paragraph id="8BAF881C2714ED6C67DE38E3D8FA461E" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="366">
Male genitalia (
<figureCitation id="132B94992714ED6C666638E3D8D742D2" box="[383,454,448,473]" captionStart="FIGURE 1" captionStartId="3.[151,250,1863,1885]" captionTargetBox="[166,1432,196,1839]" captionTargetId="figure@3.[151,1436,195,1841]" captionTargetPageId="3" captionText="FIGURE 1. Tinodes tafo n. sp., male genitalia: 1 A, left lateral; 1 B, dorsal, without inferior appendages; 1 C, segment IX and inferior appendages, ventral; 1 D, left lateral (specimen from Mandrare River watershed); 1 E, inferior appendages, ventral (specimen from Mandrare River watershed). Abbreviations: g. = phallic guide; g. ap. = phallic guide apodeme; inf. app. 1 = first segment of inferior appendage; inf. app. 2 = second segment of inferior appendage; int. app. = intermediate appendage; phall. = phallic apparatus; pre. app. = preanal appendage; st. IX = sternite IX; t. IX = tergite IX. Scale bar: 0.25 mm." httpUri="https://zenodo.org/record/886862/files/figure.png" pageId="2" pageNumber="366">Figs 1</figureCitation>
A1C): Sternite IX produced posteroventrad as rounded lobe, longitudinally shorter dorsal part on each side with anterodorsal tubercle, anterior and posterior margins moderately concave in lateral view; ventral part with transverse groove across middle; in ventral view lateral margins almost straight, slightly convergent posteriorly; anterior margin distinctly convex, posterior margin almost straight and with several ventrolateral setae. Tergite IX on each side composed of straight, narrow stalk, distally widening to subconical dorsal portion nearly two times longer than stalk, truncate apically in lateral view; in dorsal view almost rectangular, posterior margin very slightly concave and without setae. Tergite X indistinct. Preanal appendages longer than phallic apparatus, slender and nearly uniformly broad, only slightly sigmoid in lateral view; fine setae present along distal 2/3. Intermediate appendage arising anterolaterally at junctures of tergite and sternite IX, directed dorsad and curving posteromesad to median base, divided before mid-length into two long branches (almost reaching apices of inferior appendages), finger-shaped, branches slightly widening asymmetrically to apex; left branch arched above right one, each bearing three pointed spines: left branch with one spine inserted at 2/3 length, two others subapical and apical; right branch straight, directed posterad, with one apical and two subapical spines. Inferior appendages divided into two segments; bases of first segments fused ventrally, their resulting phallic guide apodeme directed anterad, clavate in lateral view, slender and acute apically in ventral view, posteriorly divided into pair of short processes, hook-shaped in lateral view, pointed outward in ventral view; each first segment irregularly ovoid with right-angled apicodorsal corner in lateral view; in ventral view, broad with rounded lateral margins; long, dense setae ventrally and along dorsoposterior angle; each second segment smaller than first; attached along posterior edge of first, long, finger-shaped, and with basoventral bump when viewed laterally, thin and sickle-shaped in ventral view with several thick setae along inner margin. Phallic apparatus long (longer than intermediate appendage branches but shorter than preanal appendages), curved ventrad, basal part well sclerotized, distal half greatly widened with large dorsal membranous lobe and apex with rounded excision in dorsal view; two internal phallic sclerites: proximal sclerite tubular, long, and curved slightly dorsad; distal one stout, short, and spine-shaped.
</paragraph>
<paragraph id="8BAF881C2714ED6C67DE3C02DA27451A" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="366">
Our material has three distinct places of origin, the watershed of Lake Alaotra (province of
<collectingRegion id="49D446FE2714ED6C63D23C03DC564632" box="[1227,1351,1312,1337]" country="Madagascar" name="Toamasina" pageId="2" pageNumber="366">Toamasina</collectingRegion>
, east of Ambatodrazaka), the Mandrare watershed (province of
<collectingRegion id="49D446FE2714ED6C642C3C67DA994656" box="[821,904,1348,1373]" country="Madagascar" name="Toliara" pageId="2" pageNumber="366">Toliara</collectingRegion>
), and the Betsiboka watershed (province of
<collectingRegion id="49D446FE2714ED6C678E3C4BD820468A" box="[151,305,1384,1409]" country="Madagascar" name="Antananarivo" pageId="2" pageNumber="366">Antananarivo</collectingRegion>
) (
<figureCitation id="132B94992714ED6C66533C4BD880468A" box="[330,401,1384,1409]" captionStart="FIGURE 7" captionStartId="11.[151,250,2012,2034]" captionTargetBox="[151,1436,1095,1989]" captionTargetId="figure@11.[151,1436,1095,1989]" captionTargetPageId="11" captionText="FIGURE 7. Schematic distribution maps of the Malagasy Tinodes." httpUri="https://zenodo.org/record/886876/files/figure.png" pageId="2" pageNumber="366">Fig. 7</figureCitation>
). We consider them as conspecific, but they can be distinguished by small morphological differences in the male genitalia. In the southern populations, the phallic guide is more developed and larger, the basoventral bump of the harpago is more sclerotized, the dorsal branch of the intermediate appendage is slightly longer than the ventral one (slightly shorter in the northern population) (
<figureCitation id="132B94992714ED6C63193CF7DD5846E6" box="[1024,1097,1492,1517]" captionStart="FIGURE 1" captionStartId="3.[151,250,1863,1885]" captionTargetBox="[166,1432,196,1839]" captionTargetId="figure@3.[151,1436,195,1841]" captionTargetPageId="3" captionText="FIGURE 1. Tinodes tafo n. sp., male genitalia: 1 A, left lateral; 1 B, dorsal, without inferior appendages; 1 C, segment IX and inferior appendages, ventral; 1 D, left lateral (specimen from Mandrare River watershed); 1 E, inferior appendages, ventral (specimen from Mandrare River watershed). Abbreviations: g. = phallic guide; g. ap. = phallic guide apodeme; inf. app. 1 = first segment of inferior appendage; inf. app. 2 = second segment of inferior appendage; int. app. = intermediate appendage; phall. = phallic apparatus; pre. app. = preanal appendage; st. IX = sternite IX; t. IX = tergite IX. Scale bar: 0.25 mm." httpUri="https://zenodo.org/record/886862/files/figure.png" pageId="2" pageNumber="366">Figs 1</figureCitation>
D1E). The
<typeStatus id="54AB36BE2714ED6C63FD3CF6DC0546E6" box="[1252,1300,1493,1517]" pageId="2" pageNumber="366">type</typeStatus>
material is restricted to the specimens from the province of
<collectingRegion id="49D446FE2714ED6C65AF3CDBDA22451A" box="[694,819,1528,1553]" country="Madagascar" name="Toamasina" pageId="2" pageNumber="366">Toamasina</collectingRegion>
.
</paragraph>
</subSubSection>
<subSubSection id="C30ADB972714ED6C67DE3F3FDC11445E" pageId="2" pageNumber="366" type="materials_examined">
<paragraph id="8BAF881C2714ED6C67DE3F3FD8874576" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="366">
<materialsCitation id="3B7882412714ED6C67DE3F3FD8874576" ID-GBIF-Occurrence="1612095176" collectingDate="2000-01-26" collectionCode="NMPC" collectorName="P. Chvojka" country="Madagascar" elevation="840" latitude="-17.461945" location="Amboditafonana" longLatPrecision="21" longitude="48.745556" pageId="2" pageNumber="366" specimenCount="1" stateProvince="Toamasina" typeStatus="holotype">
<typeStatus id="54AB36BE2714ED6C67DE3F3FD825453E" box="[199,308,1564,1589]" pageId="2" pageNumber="366" type="holotype">
<emphasis id="B964540E2714ED6C67DE3F3FD825453E" bold="true" box="[199,308,1564,1589]" pageId="2" pageNumber="366">Holotype</emphasis>
</typeStatus>
: Male.
<emphasis id="B964540E2714ED6C66803F3FDB61453E" bold="true" box="[409,624,1564,1589]" pageId="2" pageNumber="366">
<collectingCountry id="F307C88C2714ED6C66803F3FDB7A453E" box="[409,619,1564,1589]" name="Madagascar" pageId="2" pageNumber="366">MADAGASCAR</collectingCountry>
:
</emphasis>
<collectingRegion id="49D446FE2714ED6C65673F3FDBE6453E" box="[638,759,1564,1589]" country="Madagascar" name="Toamasina" pageId="2" pageNumber="366">Toamasina</collectingRegion>
Prov., brooklet
<quantity id="4CE825F92714ED6C64D93F3FDD07453E" box="[960,1046,1564,1589]" metricMagnitude="3" metricUnit="m" metricValue="1.5" pageId="2" pageNumber="366" unit="km" value="1.5">1.5 km</quantity>
S
<location id="8ECFDEC72714ED6C635B3F3FDC10453E" LSID="urn:lsid:plazi:treatment:03B9390A2717ED6C678E3FC6DA7044EE:8ECFDEC72714ED6C635B3F3FDC10453E" box="[1090,1281,1564,1589]" country="Madagascar" latitude="-17.461945" longLatPrecision="21" longitude="48.745556" name="Amboditafonana" pageId="2" pageNumber="366" stateProvince="Toamasina">Amboditafonana</location>
,
<geoCoordinate id="EE24EEDB2714ED6C620D3F3FDC89453E" box="[1300,1432,1564,1589]" direction="south" orientation="latitude" pageId="2" pageNumber="366" precision="15" value="-17.461945">17°2743S</geoCoordinate>
,
<geoCoordinate id="EE24EEDB2714ED6C678E3F63D80D4552" box="[151,284,1600,1625]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="15" value="48.745556">48°4444E</geoCoordinate>
, approx.
<elevation id="003D6F2F2714ED6C66953F62DB094552" box="[396,536,1600,1625]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="2" pageNumber="366" unit="m" value="840.0">
<quantity id="4CE825F92714ED6C66953F62D8C64553" box="[396,471,1601,1625]" metricMagnitude="2" metricUnit="m" metricValue="8.4" pageId="2" pageNumber="366" unit="m" value="840.0">840 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="EFEA57342714ED6C65303F63DB844552" box="[553,661,1600,1625]" pageId="2" pageNumber="366" value="2000-01-26">
<date id="FFAEAEDC2714ED6C65303F63DB844552" box="[553,661,1600,1625]" pageId="2" pageNumber="366" value="2000-01-26">26.i.2000</date>
</collectingDate>
, swept along shallow slow-flowing brooklet in deforested area,
<collectorName id="26E5EDCA2714ED6C62903F62D9E74576" pageId="2" pageNumber="366">P. Chvojka</collectorName>
leg. (
<collectionCode id="ED0110D92714ED6C662C3F46D89A4576" box="[309,395,1637,1661]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="366">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAF881C2714ED6C67DE3FABDA0B45AA" blockId="2.[151,1437,151,2021]" box="[199,794,1672,1697]" pageId="2" pageNumber="366">
<materialsCitation id="3B7882412714ED6C67DE3FABDA0B45AA" ID-GBIF-Occurrence="1612095185" box="[199,794,1672,1697]" collectingDate="2000-01-26" collectionCode="NMPC" collectorName="P. Chvojka" country="Madagascar" elevation="840" latitude="-17.461945" location="Amboditafonana" longLatPrecision="21" longitude="48.745556" pageId="2" pageNumber="366" specimenCount="3" specimenCount-male="3" stateProvince="Toamasina" typeStatus="paratype">
<emphasis id="B964540E2714ED6C67DE3FABD85945AA" bold="true" box="[199,328,1672,1697]" pageId="2" pageNumber="366">
<typeStatus id="54AB36BE2714ED6C67DE3FABD85245AA" box="[199,323,1672,1697]" pageId="2" pageNumber="366" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D1643952714ED6C66563FAAD8B945AA" box="[335,424,1672,1697]" pageId="2" pageNumber="366" type="male">3 males</specimenCount>
, same data as holotype (
<collectionCode id="ED0110D92714ED6C65A03FAADA1E45AA" box="[697,783,1673,1697]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="2" pageNumber="366">NMPC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8BAF881C2714ED6C67DE3F8FDC11445E" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="366">
<materialsCitation id="3B7882412714ED6C67DE3F8FDDDB45E2" ID-GBIF-Occurrence="1612095179" collectingDate="1995-04-26" country="Madagascar" county="Other" elevation="430" latitude="-24.043612" location="Ankazomanga" longLatPrecision="21" longitude="46.623055" municipality="Sohitay River" pageId="2" pageNumber="366" specimenCount="2" specimenCount-male="2" stateProvince="Toliara">
<emphasis id="B964540E2714ED6C67DE3F8FDB1945CE" bold="true" box="[199,520,1708,1733]" pageId="2" pageNumber="366">
<collectingCounty id="62CEF0902714ED6C67DE3F8FD81E45CE" box="[199,271,1708,1733]" pageId="2" pageNumber="366">Other</collectingCounty>
examined material.
</emphasis>
<collectingCountry id="F307C88C2714ED6C650C3F8EDBF045CE" box="[533,737,1709,1733]" name="Madagascar" pageId="2" pageNumber="366">MADAGASCAR</collectingCountry>
:
<collectingRegion id="49D446FE2714ED6C65E83F8FDA5145CE" box="[753,832,1708,1733]" country="Madagascar" name="Toliara" pageId="2" pageNumber="366">Toliara</collectingRegion>
Prov., Mandrare River watershed, tributary of the
<collectingMunicipality id="6BCB12662714ED6C678E3FF3D82545E2" box="[151,308,1744,1769]" pageId="2" pageNumber="366">Sohitay River</collectingMunicipality>
near
<location id="8ECFDEC72714ED6C66613FF3DB0C45E2" LSID="urn:lsid:plazi:treatment:03B9390A2717ED6C678E3FC6DA7044EE:8ECFDEC72714ED6C66613FF3DB0C45E2" box="[376,541,1744,1769]" country="Madagascar" county="Other" latitude="-24.043612" longLatPrecision="21" longitude="46.623055" municipality="Sohitay River" name="Ankazomanga" pageId="2" pageNumber="366" stateProvince="Toliara">Ankazomanga</location>
,
<geoCoordinate id="EE24EEDB2714ED6C65333FF3DBBF45E2" box="[554,686,1744,1769]" direction="south" orientation="latitude" pageId="2" pageNumber="366" precision="15" value="-24.043612">24°0237S</geoCoordinate>
,
<geoCoordinate id="EE24EEDB2714ED6C65A53FF3DA5045E2" box="[700,833,1744,1769]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="15" value="46.623055">46°3723E</geoCoordinate>
,
<elevation id="003D6F2F2714ED6C64563FF2DAC945E2" box="[847,984,1744,1769]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="366" unit="m" value="430.0">
<quantity id="4CE825F92714ED6C64563FF2DA8845E3" box="[847,921,1745,1769]" metricMagnitude="2" metricUnit="m" metricValue="4.3" pageId="2" pageNumber="366" unit="m" value="430.0">430 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="EFEA57342714ED6C64FE3FF3DD4E45E2" box="[999,1119,1744,1769]" pageId="2" pageNumber="366" value="1995-04-26">
<date id="FFAEAEDC2714ED6C64FE3FF3DD4E45E2" box="[999,1119,1744,1769]" pageId="2" pageNumber="366" value="1995-04-26">26.iv.1995</date>
</collectingDate>
,
<specimenCount id="9D1643952714ED6C63773FF2DDDB45E2" box="[1134,1226,1744,1769]" pageId="2" pageNumber="366" type="male">2 males</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B7882412714ED6C63C13FF3DA884406" ID-GBIF-Occurrence="1612095184" collectingDate="1996-02-02" country="Madagascar" elevation="750" latitude="-24.535276" location="Andohahela" longLatPrecision="21" longitude="46.682224" municipality="Sakamalio River" pageId="2" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Toliara">
<collectingMunicipality id="6BCB12662714ED6C63C13FF3DC8945E2" box="[1240,1432,1744,1769]" pageId="2" pageNumber="366">Sakamalio River</collectingMunicipality>
,
<location id="8ECFDEC72714ED6C678E3FD7D8304406" LSID="urn:lsid:plazi:treatment:03B9390A2717ED6C678E3FC6DA7044EE:8ECFDEC72714ED6C678E3FD7D8304406" box="[151,289,1780,1805]" country="Madagascar" latitude="-24.535276" longLatPrecision="21" longitude="46.682224" municipality="Sakamalio River" name="Andohahela" pageId="2" pageNumber="366" stateProvince="Toliara">Andohahela</location>
,
<geoCoordinate id="EE24EEDB2714ED6C662B3FD7D8A74406" box="[306,438,1780,1805]" direction="south" orientation="latitude" pageId="2" pageNumber="366" precision="15" value="-24.535276">24°3207S</geoCoordinate>
,
<geoCoordinate id="EE24EEDB2714ED6C66DE3FD7DB5C4406" box="[455,589,1780,1805]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="15" value="46.682224">46°4056E</geoCoordinate>
,
<elevation id="003D6F2F2714ED6C65473FD7DBBC4407" box="[606,685,1780,1805]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="366" unit="m" value="750.0">
<quantity id="4CE825F92714ED6C65473FD7DBBC4407" box="[606,685,1780,1805]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="2" pageNumber="366" unit="m" value="750.0">750 m</quantity>
</elevation>
,
<collectingDate id="EFEA57342714ED6C65A63FD7DA224406" box="[703,819,1780,1805]" pageId="2" pageNumber="366" value="1996-02-02">
<date id="FFAEAEDC2714ED6C65A63FD7DA224406" box="[703,819,1780,1805]" pageId="2" pageNumber="366" value="1996-02-02">02.ii.1996</date>
</collectingDate>
,
<specimenCount id="9D1643952714ED6C645D3FD6DA884406" box="[836,921,1780,1805]" pageId="2" pageNumber="366" type="male">1 male</specimenCount>
</materialsCitation>
;
<materialsCitation id="3B7882412714ED6C64B33FD7DA33443A" ID-GBIF-Occurrence="1612095182" collectingDate="1995-04-28" country="Madagascar" elevation="400" latitude="-24.3" location="Esira" longLatPrecision="21" longitude="46.68528" municipality="Abetolo River" pageId="2" pageNumber="366" specimenCount="1" specimenCount-male="1" stateProvince="Toliara">
tributary of the
<collectingMunicipality id="6BCB12662714ED6C63723FD7DC004406" box="[1131,1297,1780,1805]" pageId="2" pageNumber="366">Abetolo River</collectingMunicipality>
near
<location id="8ECFDEC72714ED6C62423FD7DC894406" LSID="urn:lsid:plazi:treatment:03B9390A2717ED6C678E3FC6DA7044EE:8ECFDEC72714ED6C62423FD7DC894406" box="[1371,1432,1780,1805]" country="Madagascar" latitude="-24.3" longLatPrecision="21" longitude="46.68528" municipality="Abetolo River" name="Esira" pageId="2" pageNumber="366" stateProvince="Toliara">Esira</location>
,
<geoCoordinate id="EE24EEDB2714ED6C678E3E3BD80B443A" box="[151,282,1816,1841]" direction="south" orientation="latitude" pageId="2" pageNumber="366" precision="15" value="-24.3">24°1800S</geoCoordinate>
,
<geoCoordinate id="EE24EEDB2714ED6C663E3E3BD8BD443A" box="[295,428,1816,1841]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="15" value="46.68528">46°4107E</geoCoordinate>
,
<elevation id="003D6F2F2714ED6C66A03E3ADB2E443A" box="[441,575,1816,1841]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="366" unit="m" value="400.0">
<quantity id="4CE825F92714ED6C66A03E3ADB10443B" box="[441,513,1817,1841]" metricMagnitude="2" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="366" unit="m" value="400.0">400 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="EFEA57342714ED6C65543E3BDBD4443A" box="[589,709,1816,1841]" pageId="2" pageNumber="366" value="1995-04-28">
<date id="FFAEAEDC2714ED6C65543E3BDBD4443A" box="[589,709,1816,1841]" pageId="2" pageNumber="366" value="1995-04-28">28.iv.1995</date>
</collectingDate>
,
<specimenCount id="9D1643952714ED6C65CB3E3ADA33443A" box="[722,802,1816,1841]" pageId="2" pageNumber="366" type="male">1 male</specimenCount>
</materialsCitation>
.
<materialsCitation id="3B7882412714ED6C64343E3BDC11445E" ID-GBIF-Occurrence="1612095174" collectingDate="1996-05-19" collectionCode="CBGP" collectorName="M. Gibon" country="Madagascar" elevation="820" latitude="-19.602777" location="Antananarivo" longLatPrecision="21" longitude="46.030556" pageId="2" pageNumber="366" specimenCount="2" specimenCount-male="2" stateProvince="Antananarivo">
<collectingRegion id="49D446FE2714ED6C64343E3BDAD7443A" box="[813,966,1816,1841]" country="Madagascar" name="Antananarivo" pageId="2" pageNumber="366">Antananarivo</collectingRegion>
Prov., Tsiribihina River watershed, small tributary,
<geoCoordinate id="EE24EEDB2714ED6C661D3E1FD896445E" box="[260,391,1852,1877]" direction="south" orientation="latitude" pageId="2" pageNumber="366" precision="15" value="-19.602777">19°3610S</geoCoordinate>
,
<geoCoordinate id="EE24EEDB2714ED6C668B3E1FDB09445E" box="[402,536,1852,1877]" direction="east" orientation="longitude" pageId="2" pageNumber="366" precision="15" value="46.030556">46°0150E</geoCoordinate>
,
<elevation id="003D6F2F2714ED6C653A3E1EDBB6445E" box="[547,679,1852,1877]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="366" unit="m" value="820.0">
<quantity id="4CE825F92714ED6C653A3E1EDB7B445F" box="[547,618,1853,1877]" metricMagnitude="2" metricUnit="m" metricValue="8.2" pageId="2" pageNumber="366" unit="m" value="820.0">820 m</quantity>
a.s.l.
</elevation>
,
<collectingDate id="EFEA57342714ED6C65AA3E1FDA35445E" box="[691,804,1852,1877]" pageId="2" pageNumber="366" value="1996-05-19">
<date id="FFAEAEDC2714ED6C65AA3E1FDA35445E" box="[691,804,1852,1877]" pageId="2" pageNumber="366" value="1996-05-19">19.v.1996</date>
</collectingDate>
,
<specimenCount id="9D1643952714ED6C64363E1EDA98445E" box="[815,905,1852,1877]" pageId="2" pageNumber="366" type="male">2 males</specimenCount>
; all F.-
<collectorName id="26E5EDCA2714ED6C64CC3E1EDD53445E" box="[981,1090,1852,1877]" pageId="2" pageNumber="366">M. Gibon</collectorName>
leg. (all
<collectionCode id="ED0110D92714ED6C63BF3E1EDDE5445E" box="[1190,1268,1853,1877]" pageId="2" pageNumber="366">CBGP</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C30ADB972714ED6C67DE3E43DBBD4497" pageId="2" pageNumber="366" type="etymology">
<paragraph id="8BAF881C2714ED6C67DE3E43DBBD4497" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="366">
<emphasis id="B964540E2714ED6C67DE3E43D85F4472" bold="true" box="[199,334,1888,1913]" pageId="2" pageNumber="366">Etymology.</emphasis>
Malagasy &quot;
<taxonomicName id="4C10F39F2714ED6C66CA3E42D8EE4472" authorityName="Gibon" authorityYear="2017" box="[467,511,1889,1913]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="species" species="tafo">
<emphasis id="B964540E2714ED6C66CA3E42D8EE4472" box="[467,511,1889,1913]" italics="true" pageId="2" pageNumber="366">tafo</emphasis>
</taxonomicName>
,&quot; meaning &quot;roof,&quot; adapted from the name of the village closest to the
<typeStatus id="54AB36BE2714ED6C62153E42DC2D4472" box="[1292,1340,1889,1913]" pageId="2" pageNumber="366">type</typeStatus>
locality, to be treated as a noun in apposition to
<taxonomicName id="4C10F39F2714ED6C65573EA7DBB44496" authorityName="Curtis" authorityYear="1834" box="[590,677,1924,1949]" class="Insecta" family="Psychomyiidae" genus="Tinodes" kingdom="Animalia" order="Trichoptera" pageId="2" pageNumber="366" phylum="Arthropoda" rank="genus">
<emphasis id="B964540E2714ED6C65573EA7DBB44496" box="[590,677,1924,1949]" italics="true" pageId="2" pageNumber="366">Tinodes</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C30ADB972714ED6C67DE3E8BDA7044EE" pageId="2" pageNumber="366" type="distribution">
<paragraph id="8BAF881C2714ED6C67DE3E8BDA7044EE" blockId="2.[151,1437,151,2021]" pageId="2" pageNumber="366">
<emphasis id="B964540E2714ED6C67DE3E8BD87144CA" bold="true" box="[199,352,1960,1985]" pageId="2" pageNumber="366">Distribution.</emphasis>
<collectingCountry id="F307C88C2714ED6C66743E8ADB2844CA" box="[365,569,1961,1985]" name="Madagascar" pageId="2" pageNumber="366">MADAGASCAR</collectingCountry>
: endemic. Recorded from the headwaters of the Alaotra, Mandrare, and Tsiribihina watersheds and probably widely distributed (
<figureCitation id="132B94992714ED6C64163EEFDA4244EE" box="[783,851,1996,2021]" captionStart="FIGURE 7" captionStartId="11.[151,250,2012,2034]" captionTargetBox="[151,1436,1095,1989]" captionTargetId="figure@11.[151,1436,1095,1989]" captionTargetPageId="11" captionText="FIGURE 7. Schematic distribution maps of the Malagasy Tinodes." httpUri="https://zenodo.org/record/886876/files/figure.png" pageId="2" pageNumber="366">Fig. 7</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>