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<document id="A0DD522F564DF27B2B0D6FBD53ED3367" ID-CLB-Dataset="24020" ID-DOI="10.1080/00222933.2019.1657513" ID-GBIF-Dataset="6be1c5ca-ab1b-4c69-8da2-b510a5ed3842" ID-ISSN="1464-5262" ID-Zenodo-Dep="3670187" ID-ZooBank="850FB31C-2B6A-4AB2-92D5-3413B193F196" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1581941708966" checkinUser="valdenar" docAuthor="Sutthacharoenthad, Waranid, Sartori, Michel &amp; Boonsoong, Boonsatien" docDate="2019" docId="039CD15EFF87FF999BB352B5FBBDFCDF" docLanguage="en" docName="JNATHIST.53.23–24.1491–1514.pdf.imf" docOrigin="Journal of Natural History 53 (23)" docStyle="DocumentStyle:485A7191CD41D6261F41521A3380DC73.9:JNatHist.2017-.journal_article.1cover" docStyleId="485A7191CD41D6261F41521A3380DC73" docStyleName="JNatHist.2017-.journal_article.1cover" docStyleVersion="9" docTitle="Thalerosphyrus thailandensis Sutthacharoenthad &amp; Sartori &amp; Boonsoong 2019, sp. nov." docType="treatment" docVersion="12" lastPageNumber="1502" masterDocId="FFA5A926FF8DFF959A14505FFF90FFE5" masterDocTitle="Integrative taxonomy of Thalerosphyrus Eatonı 1881 (Ephemeropteraı Heptageniidae) in Thailand" masterLastPageNumber="1514" masterPageNumber="1491" pageNumber="1500" updateTime="1698783594842" updateUser="ExternalLinkService">
<mods:mods id="6013E80238B765C2F62C405C67F4C9DC" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="D163DF9A8F6BAFAF29D7733BDFB4A96E">
<mods:title id="1F829E2812A0B9BF1A4849FC2E843FAB">Integrative taxonomy of Thalerosphyrus Eatonı 1881 (Ephemeropteraı Heptageniidae) in Thailand</mods:title>
</mods:titleInfo>
<mods:name id="3E6B82CDC3E82E910E116F797C7174B8" type="personal">
<mods:role id="B3483D5F392C6F86C27A8AA2DD669374">
<mods:roleTerm id="54DA2DEB45B84A5A423A22134FA72A45">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="B340CFA970C15C88C5522A0F539F8DF0">Sutthacharoenthad, Waranid</mods:namePart>
</mods:name>
<mods:name id="21DBDA50A0D5687D287082C350FA22B1" type="personal">
<mods:role id="1F838CC226D2831F1FA2B912B4541623">
<mods:roleTerm id="8253246F3E58AA274547B05B101842B9">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="5DB1DD042CDE064B8B8577DE92468BB7">Sartori, Michel</mods:namePart>
</mods:name>
<mods:name id="126FF42D63D0D64D63B372C2D0341F0E" type="personal">
<mods:role id="5A0E15886D64C250247825E8408FB07A">
<mods:roleTerm id="6792430FF70E211FD49272F1BB33EFC3">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6972C693FFC38B398589C448FD8A3205">Boonsoong, Boonsatien</mods:namePart>
</mods:name>
<mods:typeOfResource id="D188F91D0B66F1D9D6BEB1F781978464">text</mods:typeOfResource>
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<mods:titleInfo id="201C19997D5D080174BC07FFFC11A9B0">
<mods:title id="D90ED964D00EB3183DA8D68B1624FF9B">Journal of Natural History</mods:title>
</mods:titleInfo>
<mods:part id="29F5B86921D7A10064FBB461A3D0B1F9">
<mods:date id="6EEEF8E3A592AA1C4956A179DBCEEE84">2019</mods:date>
<mods:detail id="11CF71554DAD7F24B9AA47ABF6821122" type="pubDate">
<mods:number id="DCD4B9088C97746369474135A1808435">2019-09-11</mods:number>
</mods:detail>
<mods:detail id="870759FBBABFF3066D1970B81B063D4A" type="volume">
<mods:number id="8A9A3F50C0B9DAB779C6C0D094AB24C1">53</mods:number>
</mods:detail>
<mods:detail id="7E79A7940F79485317BF75C95BD74512" type="issue">
<mods:number id="3DB7BA6983DA686FE763581BCDC5B4CF">23</mods:number>
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<mods:start id="6CFA6C76D6B54F2F0FAD4BA63E9D603D">1491</mods:start>
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<mods:classification id="3D210F99B862F88B8405074222E288E5">journal article</mods:classification>
<mods:identifier id="3C9B622CAE3CEB11A9548DC565305A29" type="CLB-Dataset">24020</mods:identifier>
<mods:identifier id="0338ADBB91D1868DDB7A1C07E7E72E80" type="DOI">10.1080/00222933.2019.1657513</mods:identifier>
<mods:identifier id="D8417F8150CDC9B1BDCDB84827B7A902" type="GBIF-Dataset">6be1c5ca-ab1b-4c69-8da2-b510a5ed3842</mods:identifier>
<mods:identifier id="69D444A4B234C42200C30ADBC09260B7" type="ISSN">1464-5262</mods:identifier>
<mods:identifier id="044544695FD60517ADA10238927C614D" type="Zenodo-Dep">3670187</mods:identifier>
<mods:identifier id="9621CF4801654668294604825A848C52" type="ZooBank">850FB31C-2B6A-4AB2-92D5-3413B193F196</mods:identifier>
</mods:mods>
<treatment id="039CD15EFF87FF999BB352B5FBBDFCDF" ID-DOI="http://doi.org/10.5281/zenodo.3671296" ID-GBIF-Taxon="162146461" ID-Zenodo-Dep="3671296" LSID="urn:lsid:plazi:treatment:039CD15EFF87FF999BB352B5FBBDFCDF" httpUri="http://treatment.plazi.org/id/039CD15EFF87FF999BB352B5FBBDFCDF" lastPageId="12" lastPageNumber="1502" pageId="10" pageNumber="1500">
<subSubSection id="C32F33C3FF87FF9F9BB352B5FCECFCE6" box="[423,892,746,771]" pageId="10" pageNumber="1500" type="nomenclature">
<paragraph id="8B8A6048FF87FF9F9BB352B5FCECFCE6" blockId="10.[310,1006,746,806]" box="[423,892,746,771]" pageId="10" pageNumber="1500">
<heading id="D0C2D724FF87FF9F9BB352B5FCECFCE6" box="[423,892,746,771]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="1500" reason="2">
<emphasis id="B941BC5AFF87FF9F9BB352B5FCECFCE6" bold="true" box="[423,892,746,771]" pageId="10" pageNumber="1500">
<taxonomicName id="4C351BCBFF87FF9F9BB352B5FC9FFCE6" authority="Sutthacharoenthad &amp; Sartori &amp; Boonsoong, 2019" authorityName="Sutthacharoenthad &amp; Sartori &amp; Boonsoong" authorityYear="2019" box="[423,783,746,771]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="10" pageNumber="1500" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B941BC5AFF87FF9F9BB352B5FC9FFCE6" bold="true" box="[423,783,746,771]" italics="true" pageId="10" pageNumber="1500">Thalerosphyrus thailandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2720121FF87FF9F990C52B5FCECFCE6" box="[792,892,746,771]" pageId="10" pageNumber="1500" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F33C3FF87FF9F9B225352FC7EFCC3" box="[310,1006,781,806]" pageId="10" pageNumber="1500" type="description">
<paragraph id="8B8A6048FF87FF9F9B225352FC7EFCC3" blockId="10.[310,1006,746,806]" box="[310,1006,781,806]" pageId="10" pageNumber="1500">
<heading id="D0C2D724FF87FF9F9B225352FC7EFCC3" box="[310,1006,781,806]" centered="true" fontSize="11" level="2" pageId="10" pageNumber="1500" reason="5">
(
<figureCitation id="130E7CCDFF87FF9F9B295352FE3BFCC3" box="[317,427,781,806]" captionStart="Figure 4" captionStartId="11.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure@11.[163,1153,142,1528]" captionTargetPageId="11" captionText="Figure 4. Thalerosphyrus thailandensis sp. nov. male larva. (a) Habitus in dorsal view. (b) Habitus in ventral view. (c) Detail of abdominal segments VI–IX in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670197" httpUri="https://zenodo.org/record/3670197/files/figure.png" pageId="10" pageNumber="1500">Figures 4</figureCitation>
(a–c), 5(a–c), 6(d), 7(d), 8(b), 10(d), 11(d), 12(a–f))
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F33C3FF87FF9F9AB4532BFF6AFBB9" pageId="10" pageNumber="1500" type="materials_examined">
<paragraph id="8B8A6048FF87FF9F9AB4532BFE1BFC68" blockId="10.[160,1156,884,1116]" box="[160,395,884,909]" pageId="10" pageNumber="1500">
<emphasis id="B941BC5AFF87FF9F9AB4532BFE1BFC68" bold="true" box="[160,395,884,909]" italics="true" pageId="10" pageNumber="1500">Materials examined</emphasis>
</paragraph>
<paragraph id="8B8A6048FF87FF9F9AB453C8FB14FC37" blockId="10.[160,1156,884,1116]" pageId="10" pageNumber="1500">
<materialsCitation id="3B5D6A15FF87FF9F9AB453C8FB14FC37" ID-GBIF-Occurrence="2565890597" collectingDate="2017-02-03" collectionCode="ZMKU" collectorName="B. Boonsoong" elevation="267" latitude="15.459933" location="Soi Sawan waterfall" longLatPrecision="1" longitude="105.57835" pageId="10" pageNumber="1500" specimenCount="1" stateProvince="Ubon Ratchathani Prov." typeStatus="holotype">
<typeStatus id="548EDEEAFF87FF9F9AB453C8FE8DFC4A" box="[160,285,919,943]" pageId="10" pageNumber="1500" type="holotype">HOLOTYPE</typeStatus>
:
<specimenCount id="9D33ABC1FF87FF9F9B3B53C8FE4FFC4A" box="[303,479,919,943]" pageId="10" pageNumber="1500">mature nymph</specimenCount>
on slide,
<location id="8EEA3693FF87FF9F987053C8FCC1FC4A" LSID="urn:lsid:plazi:treatment:039CD15EFF87FF999BB352B5FBBDFCDF:8EEA3693FF87FF9F987053C8FCC1FC4A" box="[612,849,919,943]" latitude="15.459933" longLatPrecision="1" longitude="105.57835" name="Soi Sawan waterfall" pageId="10" pageNumber="1500" stateProvince="Ubon Ratchathani Prov.">Soi Sawan waterfall</location>
,
<collectingRegion id="49F1AEAAFF87FF9F997053C8FB11FC4A" box="[868,1153,919,943]" country="Thailand" name="Ubon Ratchathani" pageId="10" pageNumber="1500">Ubon Ratchathani Prov.</collectingRegion>
,
<geoCoordinate id="EE01068FFF87FF9F9AB453E5FEA0FC37" box="[160,304,954,978]" degrees="15.459933" direction="north" orientation="latitude" pageId="10" pageNumber="1500" precision="1" value="15.459933">15.459933°N</geoCoordinate>
,
<geoCoordinate id="EE01068FFF87FF9F9B5E53E5FE73FC37" box="[330,483,954,978]" degrees="105.578350" direction="east" orientation="longitude" pageId="10" pageNumber="1500" precision="1" value="105.57835">105.578350°E</geoCoordinate>
, alt.
<quantity id="4CCDCDADFF87FF9F982253E5FD1FFC37" box="[566,655,954,978]" metricMagnitude="2" metricUnit="m" metricValue="2.67" pageId="10" pageNumber="1500" unit="m" value="267.0">
<elevation id="0018877BFF87FF9F982253E5FD1FFC37" box="[566,655,954,978]" metricMagnitude="2" metricUnit="m" metricValue="2.67" pageId="10" pageNumber="1500" unit="m" value="267.0">267 m</elevation>
</quantity>
,
<date id="FF8B4688FF87FF9F98BE53E5FC9EFC37" box="[682,782,954,978]" pageId="10" pageNumber="1500" value="2017-02-03">
<collectingDate id="EFCFBF60FF87FF9F98BE53E5FC9EFC37" box="[682,782,954,978]" pageId="10" pageNumber="1500" value="2017-02-03">3-II-2017</collectingDate>
</date>
,
<collectorName id="26C0059EFF87FF9F993C53E5FC44FC37" box="[808,980,954,978]" pageId="10" pageNumber="1500">B. Boonsoong</collectorName>
leg. (
<collectionCode id="ED24F88DFF87FF9F9E2453E5FBEAFC37" box="[1072,1146,954,978]" collectionName="Ukraine, Kiev, Zoological Museum" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="10" pageNumber="1500">ZMKU</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="8B8A6048FF87FF9F9AB45383FF6AFBB9" blockId="10.[160,1156,884,1116]" pageId="10" pageNumber="1500">
<materialsCitation id="3B5D6A15FF87FF9F9AB45383FF69FBF2" ID-GBIF-Occurrence="3758461324" collectingDate="2017-02-03" collectionCode="ZMKU" collectorName="Sarika" elevation="267" isTailEnumeration="true" latitude="15.459933" location="Soi Sawan" longLatPrecision="1" longitude="105.57835" pageId="10" pageNumber="1500" specimenCount="1" stateProvince="Ubon Ratchathani" typeStatus="paratype">
<typeStatus id="548EDEEAFF87FF9F9AB45383FEB4FC11" box="[160,292,988,1012]" pageId="10" pageNumber="1500" type="paratype">PARATYPES</typeStatus>
: 1 nymph on slide and 3 nymph in ethanol same data as holotype (
<collectionCode id="ED24F88DFF87FF9F9AB253A0FF60FBF2" box="[166,240,1023,1047]" collectionName="Ukraine, Kiev, Zoological Museum" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="10" pageNumber="1500">ZMKU</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B5D6A15FF87FF9F9B1053A0FCC9FBF2" ID-GBIF-Occurrence="3758461332" box="[260,857,1023,1047]" collectingDate="2017-02-03" collectionCode="MZL" collectorName="Sarika" elevation="267" isTailEnumeration="true" latitude="15.459933" location="Soi Sawan" longLatPrecision="1" longitude="105.57835" pageId="10" pageNumber="1500" specimenCount="1" stateProvince="Ubon Ratchathani" typeStatus="holotype">
<specimenCount id="9D33ABC1FF87FF9F9B1053A0FEFEFBF2" box="[260,366,1023,1047]" pageId="10" pageNumber="1500">1 nymph</specimenCount>
in ethanol same data as
<typeStatus id="548EDEEAFF87FF9F98B653A0FC96FBF2" box="[674,774,1023,1047]" pageId="10" pageNumber="1500" type="holotype">holotype</typeStatus>
(
<collectionCode id="ED24F88DFF87FF9F990C53A0FCDEFBF2" box="[792,846,1023,1047]" country="Switzerland" httpUri="http://grbio.org/cool/pm5v-0p6q" name="Musee Zoologique" pageId="10" pageNumber="1500">MZL</collectionCode>
);
</materialsCitation>
<materialsCitation id="3B5D6A15FF87FF9F997753A0FF6AFBB9" ID-GBIF-Occurrence="3758461303" collectingDate="2016-01-23" collectionCode="ZMKU" collectorName="Sarika &amp; Boonsoong" elevation="20" isTailEnumeration="true" latitude="15.459933" location="Nakhon Nayok Prov" longLatPrecision="1" longitude="105.57835" pageId="10" pageNumber="1500" specimenCount="1" specimenCount-larva="1" stateProvince="Nakhon Nayok Prov" typeStatus="paratype">
<specimenCount id="9D33ABC1FF87FF9F997753A0FC23FBF2" box="[867,947,1023,1047]" pageId="10" pageNumber="1500" type="larva">1 larva</specimenCount>
(on slide),
<collectorName id="26C0059EFF87FF9F9E2F53A0FB10FBF2" box="[1083,1152,1023,1047]" pageId="10" pageNumber="1500">Sarika</collectorName>
,
<collectingRegion id="49F1AEAAFF87FF9F9AB4547DFE14FBDF" box="[160,388,1058,1082]" country="Thailand" name="Nakhon Nayok" pageId="10" pageNumber="1500">Nakhon Nayok Prov</collectingRegion>
.,
<geoCoordinate id="EE01068FFF87FF9F9B86547DFDB2FBDF" box="[402,546,1058,1082]" degrees="14.296194" direction="north" orientation="latitude" pageId="10" pageNumber="1500" precision="1" value="14.296194">14.296194°N</geoCoordinate>
,
<geoCoordinate id="EE01068FFF87FF9F9839547DFD56FBDF" box="[557,710,1058,1082]" degrees="101.269583" direction="east" orientation="longitude" pageId="10" pageNumber="1500" precision="1" value="101.269585">101.269583°E</geoCoordinate>
, alt.
<quantity id="4CCDCDADFF87FF9F98EE547DFCA5FBDF" box="[762,821,1058,1082]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="1500" unit="m" value="20.0">
<elevation id="0018877BFF87FF9F98EE547DFCA5FBDF" box="[762,821,1058,1082]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="10" pageNumber="1500" unit="m" value="20.0">20 m</elevation>
</quantity>
,
<date id="FF8B4688FF87FF9F9955547DFC3CFBDF" box="[833,940,1058,1082]" pageId="10" pageNumber="1500" value="2016-01-23">
<collectingDate id="EFCFBF60FF87FF9F9955547DFC3CFBDF" box="[833,940,1058,1082]" pageId="10" pageNumber="1500" value="2016-01-23">23-I-2016</collectingDate>
</date>
, B.
<collectorName id="26C0059EFF87FF9F99C7547DFBC3FBDF" box="[979,1107,1058,1082]" pageId="10" pageNumber="1500">Boonsoong</collectorName>
leg. (
<collectionCode id="ED24F88DFF87FF9F9AB2541BFF60FBB9" box="[166,240,1092,1116]" collectionName="Ukraine, Kiev, Zoological Museum" country="Ukraine" httpUri="http://grbio.org/cool/zbmn-bynd" name="Kiev Zoological Museum" pageId="10" pageNumber="1500">ZMKU</collectionCode>
).
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C32F33C3FF87FF999AB454C6FE06FD37" lastPageId="12" lastPageNumber="1502" pageId="10" pageNumber="1500" type="description">
<paragraph id="8B8A6048FF87FF9F9AB454C6FEA7F95F" blockId="10.[160,1156,1177,1722]" pageId="10" pageNumber="1500">
<emphasis id="B941BC5AFF87FF9F9AB454C6FF75FB57" bold="true" box="[160,229,1177,1202]" italics="true" pageId="10" pageNumber="1500">Head.</emphasis>
Labrum expanded laterally, ca. 3.8 times larger than long, with round apexes; dorsal surface and anterior margin covered with long thin setae (
<figureCitation id="130E7CCDFF87FF9F998254E3FB93FB31" box="[918,1027,1212,1236]" captionStart="Figure 12" captionStartId="18.[160,225,1221,1244]" captionTargetBox="[163,1153,142,1190]" captionTargetId="figure@18.[163,1153,142,1190]" captionTargetPageId="18" captionText="Figure 12. Mouthpart of Thalerosphyrus thailandensis sp. nov. (a) Labrum. (b) Right maxilla in ventral view. (c) Enlargement of left mandible in ventral view. (d) Enlargement of right mandible in ventral view. (e) Hypopharynx. (f) Labium in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670213" httpUri="https://zenodo.org/record/3670213/files/figure.png" pageId="10" pageNumber="1500">Figure 12</figureCitation>
(a)). Maxilla with long, simple setae on inner margin of galea-lacinia and with fimbriate setae scattered on ventral surface. Basal segment of maxillary palp with rows of thin and simple setae on inner and outer margins; apical segment pointed, with rows of simple setae on outer margin. Crown of the galea-lacinia of the maxillae with ca 16 combshaped setae (
<figureCitation id="130E7CCDFF87FF9F9B535535FE3DFA67" box="[327,429,1386,1410]" captionStart="Figure 12" captionStartId="18.[160,225,1221,1244]" captionTargetBox="[163,1153,142,1190]" captionTargetId="figure@18.[163,1153,142,1190]" captionTargetPageId="18" captionText="Figure 12. Mouthpart of Thalerosphyrus thailandensis sp. nov. (a) Labrum. (b) Right maxilla in ventral view. (c) Enlargement of left mandible in ventral view. (d) Enlargement of right mandible in ventral view. (e) Hypopharynx. (f) Labium in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670213" httpUri="https://zenodo.org/record/3670213/files/figure.png" pageId="10" pageNumber="1500">Figure12</figureCitation>
(b)). Mandibles strongly bent in their distal part; lateral margins with several rows of long and thin setae; left mandible with outer incisor serrate on inner margin, composed of ca. 6 teeth; inner incisor bifid, bearing two seta-like; with 6 fimbriate setae below the inner incisor (
<figureCitation id="130E7CCDFF87FF9F987D558DFD44FA0F" box="[617,724,1490,1514]" captionStart="Figure 12" captionStartId="18.[160,225,1221,1244]" captionTargetBox="[163,1153,142,1190]" captionTargetId="figure@18.[163,1153,142,1190]" captionTargetPageId="18" captionText="Figure 12. Mouthpart of Thalerosphyrus thailandensis sp. nov. (a) Labrum. (b) Right maxilla in ventral view. (c) Enlargement of left mandible in ventral view. (d) Enlargement of right mandible in ventral view. (e) Hypopharynx. (f) Labium in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670213" httpUri="https://zenodo.org/record/3670213/files/figure.png" pageId="10" pageNumber="1500">Figure 12</figureCitation>
(c)). Right mandible with outer incisor serrate on inner margin, composed of ca. 8 teeth; inner incisor trifid; 9 fimbriate setae below incisors (
<figureCitation id="130E7CCDFF87FF9F9B425648FE55F9CA" box="[342,453,1559,1583]" captionStart="Figure 12" captionStartId="18.[160,225,1221,1244]" captionTargetBox="[163,1153,142,1190]" captionTargetId="figure@18.[163,1153,142,1190]" captionTargetPageId="18" captionText="Figure 12. Mouthpart of Thalerosphyrus thailandensis sp. nov. (a) Labrum. (b) Right maxilla in ventral view. (c) Enlargement of left mandible in ventral view. (d) Enlargement of right mandible in ventral view. (e) Hypopharynx. (f) Labium in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670213" httpUri="https://zenodo.org/record/3670213/files/figure.png" pageId="10" pageNumber="1500">Figure 12</figureCitation>
(d)). Hypopharynx with robust lingua bearing a tuft of small setae, superlinguae densely covered with long and thin setae replaces before the apex by small setae up to near the lower part of superlinguae (
<figureCitation id="130E7CCDFF87FF9F994E5603FC2FF991" box="[858,959,1628,1652]" captionStart="Figure 12" captionStartId="18.[160,225,1221,1244]" captionTargetBox="[163,1153,142,1190]" captionTargetId="figure@18.[163,1153,142,1190]" captionTargetPageId="18" captionText="Figure 12. Mouthpart of Thalerosphyrus thailandensis sp. nov. (a) Labrum. (b) Right maxilla in ventral view. (c) Enlargement of left mandible in ventral view. (d) Enlargement of right mandible in ventral view. (e) Hypopharynx. (f) Labium in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670213" httpUri="https://zenodo.org/record/3670213/files/figure.png" pageId="10" pageNumber="1500">Figure12</figureCitation>
(e)). Labium with glossae rhomboid, almost straight on their inner and outer margins near the apex (
<figureCitation id="130E7CCDFF87FF9F9AB356FDFE84F95F" box="[167,276,1698,1722]" captionStart="Figure 12" captionStartId="18.[160,225,1221,1244]" captionTargetBox="[163,1153,142,1190]" captionTargetId="figure@18.[163,1153,142,1190]" captionTargetPageId="18" captionText="Figure 12. Mouthpart of Thalerosphyrus thailandensis sp. nov. (a) Labrum. (b) Right maxilla in ventral view. (c) Enlargement of left mandible in ventral view. (d) Enlargement of right mandible in ventral view. (e) Hypopharynx. (f) Labium in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670213" httpUri="https://zenodo.org/record/3670213/files/figure.png" pageId="10" pageNumber="1500">Figure 12</figureCitation>
(f)).
</paragraph>
<caption id="DF4A30C0FF86FF9E9AB45647FCC5F9A9" ID-DOI="http://doi.org/10.5281/zenodo.3670197" ID-Zenodo-Dep="3670197" httpUri="https://zenodo.org/record/3670197/files/figure.png" pageId="11" pageNumber="1501" startId="11.[160,225,1560,1583]" targetBox="[163,1153,142,1528]" targetPageId="11">
<paragraph id="8B8A6048FF86FF9E9AB45647FCC5F9A9" blockId="11.[160,1155,1560,1612]" pageId="11" pageNumber="1501">
<emphasis id="B941BC5AFF86FF9E9AB45647FF6CF9CA" bold="true" box="[160,252,1560,1583]" pageId="11" pageNumber="1501">Figure 4.</emphasis>
<taxonomicName id="4C351BCBFF86FF9E9B105647FD8AF9CA" authority="Sutthacharoenthad &amp; Sartori &amp; Boonsoong, 2019" authorityName="Sutthacharoenthad &amp; Sartori &amp; Boonsoong" authorityYear="2019" box="[260,538,1560,1583]" class="Insecta" family="Heptageniidae" genus="Thalerosphyrus" kingdom="Animalia" order="Ephemeroptera" pageId="11" pageNumber="1501" phylum="Arthropoda" rank="species" species="thailandensis" status="sp. nov.">
<emphasis id="B941BC5AFF86FF9E9B105647FD8AF9CA" box="[260,538,1560,1583]" italics="true" pageId="11" pageNumber="1501">Thalerosphyrus thailandensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A2720121FF86FF9E98365647FDE1F9CA" box="[546,625,1560,1583]" pageId="11" pageNumber="1501" rank="species">sp. nov.</taxonomicNameLabel>
male larva. (a) Habitus in dorsal view. (b) Habitus in ventral view. (c) Detail of abdominal segments VI–IX in ventral view.
</paragraph>
</caption>
<paragraph id="8B8A6048FF81FF999AB450D1FBD1FE92" blockId="12.[160,1156,142,375]" pageId="12" pageNumber="1502">
<emphasis id="B941BC5AFF81FF999AB450D1FF6AFF42" bold="true" box="[160,250,142,167]" italics="true" pageId="12" pageNumber="1502">Thorax.</emphasis>
Pronotum weakly expanded, not wider than the head and basally not fused to the mesonotum (
<figureCitation id="130E7CCDFF81FF999BF650EDFDC1FF2F" box="[482,593,178,202]" captionStart="Figure 4" captionStartId="11.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure@11.[163,1153,142,1528]" captionTargetPageId="11" captionText="Figure 4. Thalerosphyrus thailandensis sp. nov. male larva. (a) Habitus in dorsal view. (b) Habitus in ventral view. (c) Detail of abdominal segments VI–IX in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670197" httpUri="https://zenodo.org/record/3670197/files/figure.png" pageId="12" pageNumber="1502">Figures 4</figureCitation>
(a–b) &amp; (5a–b)). Fore femora with submarginal row(s) of stout bristles on the inner and outer margins. Hind femora with submarginal row of pointed bristles on the inner and outer margins and subparallel or slightly convergent margin, apex rounded (
<figureCitation id="130E7CCDFF81FF9998BD5145FC9CFED7" box="[681,780,282,306]" captionStart="Figure 7" captionStartId="15.[160,225,877,900]" captionTargetBox="[163,1153,142,845]" captionTargetId="figure@15.[163,1153,142,845]" captionTargetPageId="15" captionText="Figure 7. Bristles on the dorsal face of the hind femur. (a) Thalerosphyrus flowersi. (b) Thalerosphyrus sinuosus. (c) Thalerosphyrus vietnamensis. (d) Thalerosphyrus thailandensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3670203" httpUri="https://zenodo.org/record/3670203/files/figure.png" pageId="12" pageNumber="1502">Figure 7</figureCitation>
(d)). Hind tibia with one row of thin setae in marginal position (
<figureCitation id="130E7CCDFF81FF9998365163FD15FEB1" box="[546,645,316,340]" captionStart="Figure 8" captionStartId="15.[151,216,1712,1735]" captionTargetBox="[154,1144,1014,1681]" captionTargetId="figure@15.[154,1144,1014,1681]" captionTargetPageId="15" captionText="Figure 8. Outer margin of the hind tibia. (a) Thalerosphyrus sinuosus (arrow show two rows of setae). (b) Thalerosphyrus thailandensis sp. nov. (arrow show one rows of setae)." figureDoi="http://doi.org/10.5281/zenodo.3670205" httpUri="https://zenodo.org/record/3670205/files/figure.png" pageId="12" pageNumber="1502">Figure 8</figureCitation>
(b)) and a row of 10–12 pointed bristles in the marginal or submarginal positions. Tarsal claw with 3 teeth (
<figureCitation id="130E7CCDFF81FF9999A45100FB84FE92" box="[944,1044,351,375]" captionStart="Figure 9" captionStartId="16.[160,225,945,968]" captionTargetBox="[163,1152,142,913]" captionTargetId="figure@16.[163,1153,142,913]" captionTargetPageId="16" captionText="Figure 9. Tarsal claw. (a) Thalerosphyrus flowersi. (b) Thalerosphyrus sinuosus. (c) Thalerosphyrus vietnamensis. (d) Thalerosphyrus thailandensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3670207" httpUri="https://zenodo.org/record/3670207/files/figure.png" pageId="12" pageNumber="1502">Figure 9</figureCitation>
(d)).
</paragraph>
<paragraph id="8B8A6048FF81FF999AB451FBFE06FD37" blockId="12.[160,1156,420,722]" pageId="12" pageNumber="1502">
<emphasis id="B941BC5AFF81FF999AB451FBFE89FE58" bold="true" box="[160,281,420,445]" italics="true" pageId="12" pageNumber="1502">Abdomen.</emphasis>
Posterolateral expansions not developed on segment I and II, weakly developed on segment III–IV, moderately developed on segment V and reaching their maximum size on segment VIII (
<figureCitation id="130E7CCDFF81FF99987951B5FD4DFDE7" box="[621,733,490,514]" captionStart="Figure 4" captionStartId="11.[160,225,1560,1583]" captionTargetBox="[163,1153,142,1528]" captionTargetId="figure@11.[163,1153,142,1528]" captionTargetPageId="11" captionText="Figure 4. Thalerosphyrus thailandensis sp. nov. male larva. (a) Habitus in dorsal view. (b) Habitus in ventral view. (c) Detail of abdominal segments VI–IX in ventral view." figureDoi="http://doi.org/10.5281/zenodo.3670197" httpUri="https://zenodo.org/record/3670197/files/figure.png" pageId="12" pageNumber="1502">Figures 4</figureCitation>
(c) &amp; 5(c)). Gill I with asymmetrical elongated and rounded plate, 2 times longer than wide. Gill II–VI strongly asymmetrical, wider than long (
<figureCitation id="130E7CCDFF81FF999BAD5270FDBBFDA2" box="[441,555,559,583]" captionStart="Figure 10" captionStartId="17.[160,225,916,939]" captionTargetBox="[162,1152,142,885]" captionTargetId="figure@17.[162,1153,142,885]" captionTargetPageId="17" captionText="Figure 10. Gill I. (a) Thalerosphyrus flowersi. (b) Thalerosphyrus sinuosus. (c) Thalerosphyrus vietnamensis. (d) Thalerosphyrus thailandensis sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3670209" httpUri="https://zenodo.org/record/3670209/files/figure.png" pageId="12" pageNumber="1502">Figure 10</figureCitation>
(d)). Gill VII oval and almost symmetrical. Posterior margin of tergites with irregularly pointed teeth and a few microdenticle rows (
<figureCitation id="130E7CCDFF81FF999ABC522BFE8CFD69" box="[168,284,628,652]" captionStart="Figure 11" captionStartId="17.[159,224,1732,1755]" captionTargetBox="[161,1151,1035,1701]" captionTargetId="figure@17.[161,1151,1035,1701]" captionTargetPageId="17" captionText="Figure 11. Posterior margin of the abdominal tergite IV. (a) Thalerosphyrus flowersi. (arrow show numerous row of microdenticles). (b) Thalerosphyrus sinuosus. (c) Thalerosphyrus vietnamensis. (d) Thalerosphyrus thailandensis sp. nov. (arrow show a few row of microdenticles)." figureDoi="http://doi.org/10.5281/zenodo.3670211" httpUri="https://zenodo.org/record/3670211/files/figure.png" pageId="12" pageNumber="1502">Figure 11</figureCitation>
(d)). Cerci brownish red with medium brown bands increasing in size towards the apex. Paracercus whitish in the proximal part, with a dark brown band every 2–3 segments.
</paragraph>
</subSubSection>
<subSubSection id="C32F33C3FF81FF999AB452A1FBBDFCDF" box="[160,1069,766,826]" pageId="12" pageNumber="1502" type="etymology">
<paragraph id="8B8A6048FF81FF999AB452A1FE8DFCF2" blockId="12.[160,1069,766,826]" box="[160,285,766,791]" pageId="12" pageNumber="1502">
<emphasis id="B941BC5AFF81FF999AB452A1FE8DFCF2" bold="true" box="[160,285,766,791]" italics="true" pageId="12" pageNumber="1502">Etymology</emphasis>
</paragraph>
<paragraph id="8B8A6048FF81FF999AB4537DFBBDFCDF" blockId="12.[160,1069,766,826]" box="[160,1069,802,826]" pageId="12" pageNumber="1502">
The epithet is derived from the name of
<collectingCountry id="F32220D8FF81FF999862537DFD49FCDF" box="[630,729,802,826]" name="Thailand" pageId="12" pageNumber="1502">Thailand</collectingCountry>
, referring to the
<typeStatus id="548EDEEAFF81FF999988537DFC5EFCDF" box="[924,974,802,826]" pageId="12" pageNumber="1502">type</typeStatus>
locality.
</paragraph>
</subSubSection>
</treatment>
</document>
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