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<document id="10F3BC94FFB9EFF6C6BB9635856DED32" ID-DOI="10.5281/zenodo.5172539" ID-ISSN="1942-1354" ID-Zenodo-Dep="5172539" ID-ZooBank="D20BD560-605E-4DD6-A7C4-F55CAE8DE88B" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1628270129008" checkinUser="felipe" docAuthor="Sierra, José Monzón" docDate="2017" docId="039C87F48A7AFFDFA4D3DBF5FB5AC816" docLanguage="en" docName="InsectaMundi.0543.1-12.pdf" docOrigin="Insecta Mundi 2017 (543)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Chrysina antonkozlovi Monzon 2017, new species" docType="treatment" docVersion="3" lastPageNumber="5" masterDocId="FFA5FF8C8A7FFFD9A413D914FFDACB2C" masterDocTitle="Four new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) from Guatemala and Honduras" masterLastPageNumber="12" masterPageNumber="1" pageNumber="4" updateTime="1699038457668" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="2CCB940B8D3FFD2577062030EF7F724B" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="B4D45B24336610B9067E4CC0535DAFFB">
<mods:title id="0BE1B2095072558E00A2386D2C5C6A05">Four new species of Chrysina Kirby (Coleoptera: Scarabaeidae: Rutelinae) from Guatemala and Honduras</mods:title>
</mods:titleInfo>
<mods:name id="FAA24E565B0F88A2E2DB86D17ABD9FE8" type="personal">
<mods:role id="FDECA8FD0BAB7A155C3FCDDF3664E2AE">
<mods:roleTerm id="F092B6933F7B1247AA46BBE2B96EB03B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0E85351B90E0F2F6E2FEF133C21E5713">Sierra, José Monzón</mods:namePart>
</mods:name>
<mods:typeOfResource id="7E54B870826C3464E041C92660DF91DA">text</mods:typeOfResource>
<mods:relatedItem id="6547C4EAE2BFEEDE8B0197B54451FDE0" type="host">
<mods:titleInfo id="09AAF5FEC0EF392978A433E95A9CCC8E">
<mods:title id="AE1464C3E24704BB280E0102AC839387">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="9FA640C52B7DC8C0AED30ADCABF1EBC9">
<mods:date id="6899D37FD8B83D4652A13421ED00A42A">2017</mods:date>
<mods:detail id="37A5F55DB0FB80235DEFE73965A1A7F1" type="pubDate">
<mods:number id="53FD513D9AD8DD5B7A733503DD7E5E7F">2017-04-10</mods:number>
</mods:detail>
<mods:detail id="E4FEB1A8197F1D2EFEF5CED82CAC352A" type="volume">
<mods:number id="6BC7F9584285AA08E976D35F454B38CA">2017</mods:number>
</mods:detail>
<mods:detail id="0C5B56B9ADF818F7B1D3F820C92F20AA" type="issue">
<mods:number id="2AFAB878BBA49F6EBC4433FC1A1DF0C7">543</mods:number>
</mods:detail>
<mods:extent id="F1D78756499BC96DF672B0DF68566590" unit="page">
<mods:start id="0A622FCAA15D535D1501755A4064AFA7">1</mods:start>
<mods:end id="34BD1427F2356336FAB6E9027E62A23A">12</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="6F18165D28B17CB70DB510B95A425B09">journal article</mods:classification>
<mods:identifier id="0E0B067E544074D4A269223875C19726" type="DOI">10.5281/zenodo.5172539</mods:identifier>
<mods:identifier id="7E35E9A1155C58962F15D890720B5BFD" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="66A03ACEBFD6B411AE26C5729840584F" type="Zenodo-Dep">5172539</mods:identifier>
<mods:identifier id="8AC1FE0CF616E63770C4B5638D5E6504" type="ZooBank">D20BD560-605E-4DD6-A7C4-F55CAE8DE88B</mods:identifier>
</mods:mods>
<treatment id="039C87F48A7AFFDFA4D3DBF5FB5AC816" LSID="urn:lsid:plazi:treatment:039C87F48A7AFFDFA4D3DBF5FB5AC816" httpUri="http://treatment.plazi.org/id/039C87F48A7AFFDFA4D3DBF5FB5AC816" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4">
<subSubSection id="C32F65698A7AFFDCA4D3DBF5FCFAC9D5" box="[192,800,737,762]" pageId="5" pageNumber="4" type="nomenclature">
<paragraph id="8B8A36E28A7AFFDCA4D3DBF5FCFAC9D5" blockId="5.[192,800,737,793]" box="[192,800,737,762]" pageId="5" pageNumber="4">
<heading id="D0C2818E8A7AFFDCA4D3DBF5FCFAC9D5" bold="true" box="[192,800,737,762]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
<emphasis id="B941EAF08A7AFFDCA4D3DBF5FCFAC9D5" bold="true" box="[192,800,737,762]" pageId="5" pageNumber="4">
<taxonomicName id="4C354D618A7AFFDCA4D3DBF5FDB4C9D5" authority="Monzon" authorityName="Monzon" authorityYear="2017" box="[192,622,737,761]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antonkozlovi" status="sp. nov.">
<emphasis id="B941EAF08A7AFFDCA4D3DBF5FE2DC9D5" bold="true" box="[192,503,737,761]" italics="true" pageId="5" pageNumber="4">Chrysina antonkozlovi</emphasis>
Monzón
</taxonomicName>
,
<taxonomicNameLabel id="A272578B8A7AFFDCA668DBF6FCFAC9D5" box="[635,800,737,762]" pageId="5" pageNumber="4" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F65698A7AFFDCA4D3DA15FD6EC835" box="[192,692,769,793]" pageId="5" pageNumber="4" type="description">
<paragraph id="8B8A36E28A7AFFDCA4D3DA15FD6EC835" blockId="5.[192,800,737,793]" box="[192,692,769,793]" pageId="5" pageNumber="4">
<heading id="D0C2818E8A7AFFDCA4D3DA15FD6EC835" bold="true" box="[192,692,769,793]" fontSize="10" level="2" pageId="5" pageNumber="4" reason="3">
<emphasis id="B941EAF08A7AFFDCA4D3DA15FD6EC835" bold="true" box="[192,692,769,793]" pageId="5" pageNumber="4">
(
<figureCitation id="130E2A678A7AFFDCA4DADA15FEB1C835" box="[201,363,769,793]" captionStart="Figures 1–8" captionStartId="10.[192,288,733,755]" captionTargetBox="[197,1436,211,693]" captionTargetId="figure-67@10.[192,1440,192,720]" captionTargetPageId="10" captionText="Figures 1–8. Dorsal and ventral habitus of adult Chrysina specimens (1x). 1–2) C. alexae holotype from Purulhá, Baja Verapáz, Guatemala. 3–4) C. antonkozlovi male paratype from Cerro Minas, Lempira, Honduras. 5–6) C. maishei holotype from El Güisayote, Ocotepeque, Honduras.7–8) C. woodruffi holotype from Bulej, Huehuetenango." figureDoi="http://doi.org/10.5281/zenodo.5172541" httpUri="https://zenodo.org/record/5172541/files/figure.png" pageId="5" pageNumber="4">Figures 3, 4</figureCitation>
,
<figureCitation id="130E2A678A7AFFDCA56ADA15FE1DC835" box="[377,455,769,793]" captionStart="Figures 9–16" captionStartId="10.[192,288,1603,1625]" captionTargetBox="[192,1440,977,1589]" captionTargetId="figure-69@10.[192,1440,977,1589]" captionTargetPageId="10" captionText="Figures 9–16. Chrysina spp. structures. 9–12) Clypeus of males. 9) C. alexae. 10) C. antonkozlovi. 11) C. maishei. 12) C. woodruffi. 13–16) Mentum of males. 13) C. alexae. 14) C. antonkozlovi. 15) C. maishei. 16) C. woodruffi." figureDoi="http://doi.org/10.5281/zenodo.5172543" httpUri="https://zenodo.org/record/5172543/files/figure.png" pageId="5" pageNumber="4">10, 14</figureCitation>
,
<figureCitation id="130E2A678A7AFFDCA5C7DA15FDF8C835" box="[468,546,769,793]" captionStart="Figures 17–31" captionStartId="11.[192,288,1455,1477]" captionTargetBox="[192,1440,192,1440]" captionTargetId="figure-90@11.[192,1440,192,1440]" captionTargetPageId="11" captionText="Figures 17–31. Chrysina spp. structures. 17–20) Pygidium of males. 17) C. alexae. 18) C. antonkozlovi. 19) C. maishei. 20) C. woodruffi. 21–24) Mesometasternal protrusions of males. 21) C. alexae. 22) C. antonkozlovi. 23) C. maishei. 24) C. woodruffi. 25–27) Male metatrochanter. 25) C. alexae. 26) C. woodruffi. 27) C. baileyana. 28–31) Males metafemoral apical spine. 28) C. alexae. 29) C. karschi. 30) C. woodruffi. 31) C. baileyana." figureDoi="http://doi.org/10.5281/zenodo.5172545" httpUri="https://zenodo.org/record/5172545/files/figure.png" pageId="5" pageNumber="4">18, 22</figureCitation>
,
<figureCitation id="130E2A678A7AFFDCA623DA15FD71C835" box="[560,683,769,793]" captionStart="Figures 32–43" captionStartId="12.[192,288,1032,1054]" captionTargetBox="[195,1437,195,1005]" captionTargetId="figure-92@12.[192,1440,192,1018]" captionTargetPageId="12" captionText="Figures 32–43. Chrysina spp. genital structures, scale in mm. 32–39) Male genital capsule dorsal (d) and ventral (v) habitus. 32) C. alexae (d). 33) C. alexae (v). 34) C. antonkozlovi (d). 35) C. antonkozlovi (v). 36) C. maishei (d). 37) C. maishei (v). 38) C. woodruffi (d). 39) C. woodruffi (v). 40–42) Female inferior genital plates. 40) C. alexae. 41) C. antonkozlovi. 42) C. maishei. 43) C. woodruffi." figureDoi="http://doi.org/10.5281/zenodo.5172549" httpUri="https://zenodo.org/record/5172549/files/figure.png" pageId="5" pageNumber="4">34, 35, 41</figureCitation>
)
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32F65698A7AFFDCA4D3DA56FC25C8F6" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="8B8A36E28A7AFFDCA4D3DA56FC25C8F6" blockId="5.[190,1441,833,986]" pageId="5" pageNumber="4">
<emphasis id="B941EAF08A7AFFDCA4D3DA56FE53C875" bold="true" box="[192,393,833,858]" pageId="5" pageNumber="4">Type material.</emphasis>
<materialsCitation id="3B5D3CBF8A7AFFDCA59DDA56FD4AC8B6" collectingDate="2014-07-01" collectingDateMax="2014-07-04" collectingDateMin="2014-07-01" collectionCode="UVGC, V, HOLO-, TYPE" collectorName="P. N. Celaque &amp; M. Marquez" country="Honduras" elevation="2600" latitude="14.547223" location="Lempira" longLatPrecision="21" longitude="-88.669716" pageId="5" pageNumber="4" specimenCount="1" stateProvince="Lempira" typeStatus="holotype">
<emphasis id="B941EAF08A7AFFDCA59DDA56FDD6C876" bold="true" box="[398,524,834,858]" pageId="5" pageNumber="4">
<typeStatus id="548E88408A7AFFDCA59DDA56FDD6C876" box="[398,524,834,858]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
</emphasis>
male (
<collectionCode id="ED24AE278A7AFFDCA64FDA56FD75C876" box="[604,687,834,858]" country="Guatemala" httpUri="http://grbio.org/cool/azap-ym2d" name="Collecion de Artropodos" pageId="5" pageNumber="4">UVGC</collectionCode>
) labeled “
<collectingCountry id="F32276728A7AFFDCA735DA56FC16C876" box="[806,972,834,858]" name="Honduras" pageId="5" pageNumber="4">HONDURAS</collectingCountry>
,
<collectingRegion id="49F1F8008A7AFFDCA7C5DA56FB98C876" box="[982,1090,834,858]" country="Honduras" name="Lempira" pageId="5" pageNumber="4">Lempira</collectingRegion>
,
<collectorName id="26C053348A7AFFDCA05FDA56FB2FC876" box="[1100,1269,834,858]" pageId="5" pageNumber="4">P. N. Celaque</collectorName>
, Cerro Minas.
<quantity id="4CCD9B078A7AFFDCA4D3DA76FEC4C855" box="[192,286,866,890]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="4" unit="m" value="2600.0">
<elevation id="0018D1D18A7AFFDCA4D3DA76FEC4C855" box="[192,286,866,890]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="5" pageNumber="4" unit="m" value="2600.0">2600 m</elevation>
</quantity>
.
<date id="FF8B10228A7AFFDCA539DA76FE29C856" box="[298,499,866,890]" pageId="5" pageNumber="4" value="2014-07-01" valueMax="2014-07-04" valueMin="2014-07-01">
<collectingDate id="EFCFE9CA8A7AFFDCA539DA76FE29C856" box="[298,499,866,890]" pageId="5" pageNumber="4" value="2014-07-01" valueMax="2014-07-04" valueMin="2014-07-01">01-04 julio 2014</collectingDate>
</date>
.
<geoCoordinate id="EE0150258A7AFFDCA5ECDA76FD5CC855" box="[511,646,865,890]" degrees="14" direction="north" minutes="32" orientation="latitude" pageId="5" pageNumber="4" precision="15" seconds="50" value="14.547223">14° 32´50 N</geoCoordinate>
-
<geoCoordinate id="EE0150258A7AFFDCA685DA76FCFEC855" box="[662,804,866,890]" degrees="88" direction="west" minutes="40" orientation="longitude" pageId="5" pageNumber="4" precision="15" seconds="11" value="-88.669716">88° 40´11 W</geoCoordinate>
,
<collectionCode id="ED24AE278A7AFFDCA73CDA76FC9FC855" box="[815,837,866,889]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="5" pageNumber="4" type="Museum">V</collectionCode>
. Sinjaev &amp;
<collectorName id="26C053348A7AFFDCA7C2DA76FBB0C856" box="[977,1130,866,890]" pageId="5" pageNumber="4">M. Márquez</collectorName>
”;; on red paper, “
<collectionCode id="ED24AE278A7AFFDCA155DA76FED3C8B5" pageId="5" pageNumber="4">HOLO- TYPE</collectionCode>
<taxonomicName id="4C354D618A7AFFDCA51CDA96FD5CC8B6" authority="Monzon" authorityName="Monzon" authorityYear="2017" box="[271,646,898,922]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antonkozlovi">
<emphasis id="B941EAF08A7AFFDCA51CDA96FDFAC8B6" box="[271,544,898,922]" italics="true" pageId="5" pageNumber="4">Chrysina antonkozlovi</emphasis>
Monzón
</taxonomicName>
”
</materialsCitation>
.
<emphasis id="B941EAF08A7AFFDCA68EDA96FCF3C8B6" bold="true" box="[669,809,898,922]" pageId="5" pageNumber="4">
<typeStatus id="548E88408A7AFFDCA68EDA96FCF3C8B6" box="[669,809,898,922]" pageId="5" pageNumber="4" type="paratype">Paratypes</typeStatus>
</emphasis>
(
<specimenCount id="9D33FD6B8A7AFFDCA72BDA96FC79C8B6" box="[824,931,898,922]" pageId="5" pageNumber="4" type="male">16 males</specimenCount>
and
<specimenCount id="9D33FD6B8A7AFFDCA7CFDA96FB9EC8B6" box="[988,1092,898,922]" pageId="5" pageNumber="4" type="female">1 female</specimenCount>
) with same data as
<typeStatus id="548E88408A7AFFDCA122DA96FA47C8B6" box="[1329,1437,898,922]" pageId="5" pageNumber="4" type="holotype">holotype</typeStatus>
. All
<typeStatus id="548E88408A7AFFDCA4FFDAB6FEBCC896" box="[236,358,930,954]" pageId="5" pageNumber="4" type="paratype">paratypes</typeStatus>
with label, on yellow paper “
<typeStatus id="548E88408A7AFFDCA6F5DAB6FCA6C896" box="[742,892,930,954]" pageId="5" pageNumber="4" type="paratype">PARATYPE</typeStatus>
<taxonomicName id="4C354D618A7AFFDCA796DAB6FAD1C896" authority="Monzon" authorityName="Monzon" authorityYear="2017" box="[901,1291,930,954]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="antonkozlovi">
<emphasis id="B941EAF08A7AFFDCA796DAB6FB46C896" box="[901,1180,930,954]" italics="true" pageId="5" pageNumber="4">Chrysina antonkozlovi</emphasis>
Monzón
</taxonomicName>
”.
<typeStatus id="548E88408A7AFFDCA130DAB6FA7AC896" box="[1315,1440,930,954]" pageId="5" pageNumber="4" type="paratype">Paratypes</typeStatus>
deposited in the
<collectionCode id="ED24AE278A7AFFDCA59FDAD6FE16C8F6" box="[396,460,962,986]" pageId="5" pageNumber="4">AOK</collectionCode>
,
<collectionCode id="ED24AE278A7AFFDCA5CBDAD6FDFEC8F6" box="[472,548,962,986]" country="Honduras" httpUri="http://biocol.org/urn:lsid:biocol.org:col:33268" lsid="urn:lsid:biocol.org:col:33268" name="Escuela Agricola Panamericana" pageId="5" pageNumber="4">EAPZ</collectionCode>
,
<collectionCode id="ED24AE278A7AFFDCA63CDAD6FDA1C8F6" box="[559,635,962,986]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
,
<collectionCode id="ED24AE278A7AFFDCA694DAD6FD18C8F6" box="[647,706,962,986]" pageId="5" pageNumber="4">JMS</collectionCode>
,
<collectionCode id="ED24AE278A7AFFDCA6DCDAD6FD2CC8F5" box="[719,758,962,985]" pageId="5" pageNumber="4">KP</collectionCode>
and
<collectionCode id="ED24AE278A7AFFDCA721DAD6FCAAC8F6" box="[818,880,962,986]" country="Russia" httpUri="http://grbio.org/cool/vmfx-g997" name="Zoological Institute, Russian Academy of Sciences" pageId="5" pageNumber="4">ZISP</collectionCode>
collections.
</paragraph>
</subSubSection>
<subSubSection id="C32F65698A7AFFDFA4D3DD15FE08CA36" lastPageId="6" lastPageNumber="5" pageId="5" pageNumber="4" type="description">
<paragraph id="8B8A36E28A7AFFDCA4D3DD15FC62CD95" blockId="5.[192,1441,1025,1722]" pageId="5" pageNumber="4">
<emphasis id="B941EAF08A7AFFDCA4D3DD15FD9ECF36" bold="true" box="[192,580,1025,1050]" pageId="5" pageNumber="4">
Description.
<typeStatus id="548E88408A7AFFDCA560DD16FE2BCF36" box="[371,497,1026,1050]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
male.
</emphasis>
Length 25.0 mm; width at elytral humeri
<quantity id="4CCD9B078A7AFFDCA04BDD15FB1CCF35" box="[1112,1222,1025,1049]" metricMagnitude="-2" metricUnit="m" metricValue="1.05" pageId="5" pageNumber="4" unit="mm" value="10.5">10.5 mm</quantity>
; maximum width (middle of elytra) 12.0 mm. Color of dorsum slightly dark yellowish green (
<figureCitation id="130E2A678A7AFFDCA05EDD36FB49CF16" box="[1101,1171,1058,1082]" captionStart="Figures 1–8" captionStartId="10.[192,288,733,755]" captionTargetBox="[197,1436,211,693]" captionTargetId="figure-67@10.[192,1440,192,720]" captionTargetPageId="10" captionText="Figures 1–8. Dorsal and ventral habitus of adult Chrysina specimens (1x). 1–2) C. alexae holotype from Purulhá, Baja Verapáz, Guatemala. 3–4) C. antonkozlovi male paratype from Cerro Minas, Lempira, Honduras. 5–6) C. maishei holotype from El Güisayote, Ocotepeque, Honduras.7–8) C. woodruffi holotype from Bulej, Huehuetenango." figureDoi="http://doi.org/10.5281/zenodo.5172541" httpUri="https://zenodo.org/record/5172541/files/figure.png" pageId="5" pageNumber="4">Fig. 3</figureCitation>
); frons lighter yellowish green, clypeus coppery brown; anterior margin of clypeus and fronto-clypeal suture black; ocular canthi reddish golden; antennae reddish brown, scape dorsally greenish gold; pronotal margins greenish gold with reddish reflections; scutellum lateral margins greenish gold; elytral margins reddish gold, humeri and apical umbone yellowish green; pygidium iridescent golden green. Venter shiny yellowish green; sternites with posterior margin reddish gold; tibiae reddish brown; tarsi reddish golden green (
<figureCitation id="130E2A678A7AFFDCA4DADDF6FED5CFD6" box="[201,271,1250,1274]" captionStart="Figures 1–8" captionStartId="10.[192,288,733,755]" captionTargetBox="[197,1436,211,693]" captionTargetId="figure-67@10.[192,1440,192,720]" captionTargetPageId="10" captionText="Figures 1–8. Dorsal and ventral habitus of adult Chrysina specimens (1x). 1–2) C. alexae holotype from Purulhá, Baja Verapáz, Guatemala. 3–4) C. antonkozlovi male paratype from Cerro Minas, Lempira, Honduras. 5–6) C. maishei holotype from El Güisayote, Ocotepeque, Honduras.7–8) C. woodruffi holotype from Bulej, Huehuetenango." figureDoi="http://doi.org/10.5281/zenodo.5172541" httpUri="https://zenodo.org/record/5172541/files/figure.png" pageId="5" pageNumber="4">Fig. 4</figureCitation>
). Clypeus (
<figureCitation id="130E2A678A7AFFDCA580DDF6FE3DCFD6" box="[403,487,1250,1274]" captionStart="Figures 9–16" captionStartId="10.[192,288,1603,1625]" captionTargetBox="[192,1440,977,1589]" captionTargetId="figure-69@10.[192,1440,977,1589]" captionTargetPageId="10" captionText="Figures 9–16. Chrysina spp. structures. 9–12) Clypeus of males. 9) C. alexae. 10) C. antonkozlovi. 11) C. maishei. 12) C. woodruffi. 13–16) Mentum of males. 13) C. alexae. 14) C. antonkozlovi. 15) C. maishei. 16) C. woodruffi." figureDoi="http://doi.org/10.5281/zenodo.5172543" httpUri="https://zenodo.org/record/5172543/files/figure.png" pageId="5" pageNumber="4">Fig. 10</figureCitation>
) free margins subtrapezoidal in dorsal view, margined and raised; surface with coarse and dense punctures; frons densely punctuate, punctures dense; fronto-clypeal suture complete; interocular distance 0.8 times wider than antennal club length. Mentum (
<figureCitation id="130E2A678A7AFFDCA056DC36FB40CE16" box="[1093,1178,1314,1338]" captionStart="Figures 9–16" captionStartId="10.[192,288,1603,1625]" captionTargetBox="[192,1440,977,1589]" captionTargetId="figure-69@10.[192,1440,977,1589]" captionTargetPageId="10" captionText="Figures 9–16. Chrysina spp. structures. 9–12) Clypeus of males. 9) C. alexae. 10) C. antonkozlovi. 11) C. maishei. 12) C. woodruffi. 13–16) Mentum of males. 13) C. alexae. 14) C. antonkozlovi. 15) C. maishei. 16) C. woodruffi." figureDoi="http://doi.org/10.5281/zenodo.5172543" httpUri="https://zenodo.org/record/5172543/files/figure.png" pageId="5" pageNumber="4">Fig. 14</figureCitation>
) small, 1.3 times longer than wide; anterior margin depressed and irregular; lateral sides bottle shaped being wider and rounded distally and narrower close to anterior margin; surface with few large setigerous punctures, large narrow horizontal depression close to apex. Pronotum at base 2.7 times as wide as interocular distance; surface moderately punctate, punctures small. Lateral margin completely beaded. Elytra punctate striate; striae well defined and with punctures moderate in size and depth; intervals convex. Elytron
<quantity id="4CCD9B078A7AFFDCA537DCF5FE57CED5" box="[292,397,1505,1529]" metricMagnitude="-2" metricUnit="m" metricValue="1.65" pageId="5" pageNumber="4" unit="mm" value="16.5">16.5 mm</quantity>
long and 2.8 times as long as pronotum; lateral margin with bead complete. Pygidium (
<figureCitation id="130E2A678A7AFFDCA4DADF16FEC5CD36" box="[201,287,1538,1562]" captionStart="Figures 17–31" captionStartId="11.[192,288,1455,1477]" captionTargetBox="[192,1440,192,1440]" captionTargetId="figure-90@11.[192,1440,192,1440]" captionTargetPageId="11" captionText="Figures 17–31. Chrysina spp. structures. 17–20) Pygidium of males. 17) C. alexae. 18) C. antonkozlovi. 19) C. maishei. 20) C. woodruffi. 21–24) Mesometasternal protrusions of males. 21) C. alexae. 22) C. antonkozlovi. 23) C. maishei. 24) C. woodruffi. 25–27) Male metatrochanter. 25) C. alexae. 26) C. woodruffi. 27) C. baileyana. 28–31) Males metafemoral apical spine. 28) C. alexae. 29) C. karschi. 30) C. woodruffi. 31) C. baileyana." figureDoi="http://doi.org/10.5281/zenodo.5172545" httpUri="https://zenodo.org/record/5172545/files/figure.png" pageId="5" pageNumber="4">Fig. 18</figureCitation>
) moderately punctate, apical margin with long sparse setae; surface moderately convex and slightly prominent towards the apex. Venter with mesometasternal protrusion very short and depressed (
<figureCitation id="130E2A678A7AFFDCA4DADF56FEF9CD76" box="[201,291,1602,1626]" captionStart="Figures 17–31" captionStartId="11.[192,288,1455,1477]" captionTargetBox="[192,1440,192,1440]" captionTargetId="figure-90@11.[192,1440,192,1440]" captionTargetPageId="11" captionText="Figures 17–31. Chrysina spp. structures. 17–20) Pygidium of males. 17) C. alexae. 18) C. antonkozlovi. 19) C. maishei. 20) C. woodruffi. 21–24) Mesometasternal protrusions of males. 21) C. alexae. 22) C. antonkozlovi. 23) C. maishei. 24) C. woodruffi. 25–27) Male metatrochanter. 25) C. alexae. 26) C. woodruffi. 27) C. baileyana. 28–31) Males metafemoral apical spine. 28) C. alexae. 29) C. karschi. 30) C. woodruffi. 31) C. baileyana." figureDoi="http://doi.org/10.5281/zenodo.5172545" httpUri="https://zenodo.org/record/5172545/files/figure.png" pageId="5" pageNumber="4">Fig. 22</figureCitation>
). Metasternal sides setigerously punctate, setae long and dense. Legs with protibiae clearly tridentate, apical and medial teeth well developed, third smaller. Genitalia with parameres subsymmmetrical, apex slightly slanted to the left, apically constricted, fused except for very broad “v” shaped, apex bidentate; length of genital capsule 8.0 mm (
<figureCitation id="130E2A678A7AFFDCA735DFB6FC71CD95" box="[806,939,1697,1722]" captionStart="Figures 32–43" captionStartId="12.[192,288,1032,1054]" captionTargetBox="[195,1437,195,1005]" captionTargetId="figure-92@12.[192,1440,192,1018]" captionTargetPageId="12" captionText="Figures 32–43. Chrysina spp. genital structures, scale in mm. 32–39) Male genital capsule dorsal (d) and ventral (v) habitus. 32) C. alexae (d). 33) C. alexae (v). 34) C. antonkozlovi (d). 35) C. antonkozlovi (v). 36) C. maishei (d). 37) C. maishei (v). 38) C. woodruffi (d). 39) C. woodruffi (v). 40–42) Female inferior genital plates. 40) C. alexae. 41) C. antonkozlovi. 42) C. maishei. 43) C. woodruffi." figureDoi="http://doi.org/10.5281/zenodo.5172549" httpUri="https://zenodo.org/record/5172549/files/figure.png" pageId="5" pageNumber="4">Fig. 34, 35</figureCitation>
).
</paragraph>
<paragraph id="8B8A36E28A7AFFDCA4D3DFF6FDF8CC56" blockId="5.[192,1440,1762,1914]" pageId="5" pageNumber="4">
<emphasis id="B941EAF08A7AFFDCA4D3DFF6FEF7CDD6" bold="true" box="[192,301,1762,1786]" pageId="5" pageNumber="4">Female.</emphasis>
Similar to male except body stout; tarsi less robust and apical sternite not depressed; antennal club short (interocular distance 1.2 times wider than antennal club length). Length
<quantity id="4CCD9B078A7AFFDCA0D2DE15FAF7CC35" box="[1217,1325,1793,1817]" metricMagnitude="-2" metricUnit="m" metricValue="2.75" pageId="5" pageNumber="4" unit="mm" value="27.5">27.5 mm</quantity>
; width at elytral humeri 13.0 mm; maximum width 15.0 mm. Inferior genital plates subsymmetrical; external margin with side pointing spine process and with long fine sparse setae (
<figureCitation id="130E2A678A7AFFDCA020DE56FB5DCC76" box="[1075,1159,1858,1882]" captionStart="Figures 32–43" captionStartId="12.[192,288,1032,1054]" captionTargetBox="[195,1437,195,1005]" captionTargetId="figure-92@12.[192,1440,192,1018]" captionTargetPageId="12" captionText="Figures 32–43. Chrysina spp. genital structures, scale in mm. 32–39) Male genital capsule dorsal (d) and ventral (v) habitus. 32) C. alexae (d). 33) C. alexae (v). 34) C. antonkozlovi (d). 35) C. antonkozlovi (v). 36) C. maishei (d). 37) C. maishei (v). 38) C. woodruffi (d). 39) C. woodruffi (v). 40–42) Female inferior genital plates. 40) C. alexae. 41) C. antonkozlovi. 42) C. maishei. 43) C. woodruffi." figureDoi="http://doi.org/10.5281/zenodo.5172549" httpUri="https://zenodo.org/record/5172549/files/figure.png" pageId="5" pageNumber="4">Fig. 41</figureCitation>
) (based on female with database label “JMS10798”).
</paragraph>
<paragraph id="8B8A36E28A79FFDFA4D3D9D5FE08CA36" blockId="6.[192,1440,193,282]" pageId="6" pageNumber="5">
<emphasis id="B941EAF08A79FFDFA4D3D9D5FE96CBF5" bold="true" box="[192,332,193,217]" pageId="6" pageNumber="5">Variation.</emphasis>
Males: Length 20.5–25.0 mm; width at elytral humeri
<quantity id="4CCD9B078A79FFDFA7E0D9D5FBA4CBF5" box="[1011,1150,193,217]" metricMagnitude="-2" metricUnit="m" metricValue="1.025" metricValueMax="1.1" metricValueMin="0.95" pageId="6" pageNumber="5" unit="mm" value="10.25" valueMax="11.0" valueMin="9.5">9.5–11 mm</quantity>
; maximum width 10.0– 13.00 mm.
<typeStatus id="548E88408A79FFDFA55ED9F6FE50CBD6" box="[333,394,226,250]" pageId="6" pageNumber="5">Type</typeStatus>
series very homogeneous yellowish green,
<specimenCount id="9D33FD6B8A79FFDFA7B4D9F6FB86CBD6" box="[935,1116,226,250]" pageId="6" pageNumber="5" type="generic">two specimens</specimenCount>
reddish brown with most margins golden green.
</paragraph>
</subSubSection>
<subSubSection id="C32F65698A79FFDFA4D3D856FBD8CA56" pageId="6" pageNumber="5" type="etymology">
<paragraph id="8B8A36E28A79FFDFA4D3D856FBD8CA56" blockId="6.[192,1440,322,378]" pageId="6" pageNumber="5">
<emphasis id="B941EAF08A79FFDFA4D3D856FE86CA76" bold="true" box="[192,348,322,346]" pageId="6" pageNumber="5">Etymology.</emphasis>
Named in honor of my friend Anton Olegovich Kozlov from
<collectingRegion id="49F1F8008A79FFDFA02AD856FB40CA75" box="[1081,1178,322,345]" country="Russia" name="Moscow" pageId="6" pageNumber="5">Moscow</collectingRegion>
(
<collectingCountry id="F32276728A79FFDFA0BAD856FADACA76" box="[1193,1280,322,346]" name="Russia" pageId="6" pageNumber="5">Russia</collectingCountry>
), a great and enthusiastic entomologist that brought this species to my attention.
</paragraph>
</subSubSection>
<subSubSection id="C32F65698A79FFDFA4D3D8B5FDAFC9B6" pageId="6" pageNumber="5" type="diagnosis">
<paragraph id="8B8A36E28A79FFDFA4D3D8B5FDAFC9B6" blockId="6.[192,1440,417,666]" pageId="6" pageNumber="5">
<emphasis id="B941EAF08A79FFDFA4D3D8B5FE8ACA95" bold="true" box="[192,336,417,441]" pageId="6" pageNumber="5">Diagnosis.</emphasis>
This is one of the five green species in the adelaida group in northern Central America (including the two described in this publication). This species seems to be a sister species of
<taxonomicName id="4C354D618A79FFDFA13AD8D6FE9DCAD6" authority="(Moron)" baseAuthorityName="Moron" baseAuthorityYear="1990" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="B941EAF08A79FFDFA13AD8D6FF04CAD6" italics="true" pageId="6" pageNumber="5">C. centralis</emphasis>
(Morón)
</taxonomicName>
with which it shares many morphological characters. The two can be easily differentiated from the other three green species of the group in the region by having the elytral humeri and umbone yellowish green, very similar to color of disc (
<taxonomicName id="4C354D618A79FFDFA6E8DB36FC39C916" authority="(Ohaus)" baseAuthorityName="Ohaus" baseAuthorityYear="1930" box="[763,995,546,570]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="pehlkei">
<emphasis id="B941EAF08A79FFDFA6E8DB36FCADC916" box="[763,887,546,570]" italics="true" pageId="6" pageNumber="5">C. pehlkei</emphasis>
(Ohaus)
</taxonomicName>
,
<taxonomicName id="4C354D618A79FFDFA7E0DB36FB00C916" authority="Monzon" authorityName="Monzon" authorityYear="2010" box="[1011,1242,546,570]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="hawksi">
<emphasis id="B941EAF08A79FFDFA7E0DB36FBB5C916" box="[1011,1135,546,570]" italics="true" pageId="6" pageNumber="5">C. hawksi</emphasis>
Monzón
</taxonomicName>
and
<taxonomicName id="4C354D618A79FFDFA108DB36FA7AC916" authorityName="Sierra" authorityYear="2017" box="[1307,1440,546,570]" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="maishei">
<emphasis id="B941EAF08A79FFDFA108DB36FA7AC916" box="[1307,1440,546,570]" italics="true" pageId="6" pageNumber="5">C. maishei</emphasis>
</taxonomicName>
have golden green elytral humeri and umbone, different from disc). It can be easily separated from
<taxonomicName id="4C354D618A79FFDFA196DB56FEF0C956" baseAuthorityName="Moron" baseAuthorityYear="1990" class="Insecta" family="Rutelidae" genus="Chrysina" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="5" phylum="Arthropoda" rank="species" species="centralis">
<emphasis id="B941EAF08A79FFDFA196DB56FEF0C956" italics="true" pageId="6" pageNumber="5">C. centralis</emphasis>
</taxonomicName>
by the male antennal club, which is longer than the interocular distance, and the different shape of male and female genitalia.
</paragraph>
</subSubSection>
<subSubSection id="C32F65698A79FFDFA4D3DBD5FB5AC816" pageId="6" pageNumber="5" type="distribution">
<paragraph id="8B8A36E28A79FFDFA4D3DBD5FB5AC816" blockId="6.[192,1440,705,826]" pageId="6" pageNumber="5">
<emphasis id="B941EAF08A79FFDFA4D3DBD5FDE8C9F6" bold="true" box="[192,562,705,730]" pageId="6" pageNumber="5">Distribution and remarks.</emphasis>
This species is currently known only from the
<typeStatus id="548E88408A79FFDFA083DBD6FB1FC9F6" box="[1168,1221,706,730]" pageId="6" pageNumber="5">type</typeStatus>
locality, Celaque National Park, an isolated mountain located south of
<collectingRegion id="49F1F8008A79FFDFA74BDBF6FC3FC9D5" box="[856,997,738,762]" country="Guatemala" name="Santa Rosa" pageId="6" pageNumber="5">Santa Rosa</collectingRegion>
de
<collectingRegion id="49F1F8008A79FFDFA002DBF6FBBAC9D6" box="[1041,1120,738,762]" country="Honduras" name="Copan" pageId="6" pageNumber="5">Copán</collectingRegion>
(
<figureCitation id="130E2A678A79FFDFA062DBF6FB10C9D6" box="[1137,1226,738,762]" captionStart="Figure 44" captionStartId="13.[192,276,1172,1194]" captionTargetBox="[192,1440,192,1151]" captionTargetId="figure-42@13.[192,1440,192,1151]" captionTargetPageId="13" captionText="Figure 44. Distribution map for Chrysina alexae n. sp., C. antonkozlovi n. sp., C. maishei n. sp. and C. woodruffi n. sp. in Guatemala and Honduras." figureDoi="http://doi.org/10.5281/zenodo.5172553" httpUri="https://zenodo.org/record/5172553/files/figure.png" pageId="6" pageNumber="5">Fig. 44</figureCitation>
). It is interesting that even though some collecting has been conducted in the park by several researchers, this species was never collected. It seems to be isolated to the higher altitudes of the park.
</paragraph>
</subSubSection>
</treatment>
</document>
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