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<mods:title id="C15CC7BA61784AAB4E20A263DE0030F9">Revision of the horseface loaches (Cobitidae, Acantopsis), with descriptions of three new species from Southeast Asia</mods:title>
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<mods:namePart id="9378282D3B346701B4DAC482D14AC758">Nithirojpakdee, Patchara</mods:namePart>
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<mods:namePart id="B54806FDF3BFF906F606ACDB4BF2D7B5">Tangjitjaroen, Weerapongse</mods:namePart>
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<mods:namePart id="28BD49C9D944C8B4EE519752EE456941">Pfeiffer, John M.</mods:namePart>
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<treatment id="537A87CE835DFFAEFF04FA74FCC8FEE4" ID-DOI="http://doi.org/10.5281/zenodo.6010732" ID-GBIF-Taxon="135489600" ID-Zenodo-Dep="6010732" LSID="urn:lsid:plazi:treatment:537A87CE835DFFAEFF04FA74FCC8FEE4" httpUri="http://treatment.plazi.org/id/537A87CE835DFFAEFF04FA74FCC8FEE4" lastPageId="18" lastPageNumber="169" pageId="16" pageNumber="167">
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<paragraph id="DB6C36D8835DFFACFF04FA74FDA6FA27" blockId="16.[151,583,1414,1508]" box="[151,583,1414,1440]" pageId="16" pageNumber="167">
<heading id="802481B4835DFFACFF04FA74FDA6FA27" bold="true" box="[151,583,1414,1440]" fontSize="11" level="1" pageId="16" pageNumber="167" reason="1">
<taxonomicName id="1CD34D5B835DFFACFF04FA74FDA6FA27" ID-CoL="64G3J" authority="(Blyth, 1860)" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[151,583,1414,1440]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="spectabilis">
<emphasis id="E9A7EACA835DFFACFF04FA74FDA6FA27" bold="true" box="[151,583,1414,1440]" pageId="16" pageNumber="167">
<emphasis id="E9A7EACA835DFFACFF04FA74FE7EFA27" bold="true" box="[151,415,1414,1440]" italics="true" pageId="16" pageNumber="167">Acantopsis spectabilis</emphasis>
(
<bibRefCitation id="BF424B29835DFFACFE3DFA74FDDEFA27" author="Blyth" box="[430,575,1414,1440]" pageId="16" pageNumber="167" refString="Blyth, E. (1860) Report on some fishes receiVed chiefly from the Sitang riVer and its tributary streams, Tenasserim ProVinces. Journal of the Asiatic Society of Bengal, 29, 138 - 174." type="journal article" year="1860">Blyth, 1860</bibRefCitation>
)
</emphasis>
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</paragraph>
<paragraph id="DB6C36D8835DFFACFF04FA5AFF0AFA63" blockId="16.[151,583,1414,1508]" pageId="16" pageNumber="167">
Spectacular Horseface Loach (
<figureCitation id="43E82A5D835DFFACFF0CFA39FF02FA63" box="[159,227,1483,1508]" captionStart="FIGURE 9" captionStartId="17.[151,250,1739,1761]" captionTargetBox="[151,1436,541,1717]" captionTargetId="figure@17.[151,1436,541,1717]" captionTargetPageId="17" captionText="FIGURE 9. Acantopsis spectabilis. (A) Lateral and dorsal VieWs, CAS 238298, 163.0 mm SL. (B) Lateral VieWs, CAS 88783, 112.9 mm SL and 56.2 mm SL." httpUri="https://zenodo.org/record/1039429/files/figure.png" pageId="16" pageNumber="167">Fig. 9</figureCitation>
)
</paragraph>
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<subSubSection id="93C96553835DFFACFF04F9E1FCAAF923" pageId="16" pageNumber="167" type="reference_group">
<paragraph id="DB6C36D8835DFFACFF04F9E1FCDFF9CF" blockId="16.[151,1436,1555,1700]" pageId="16" pageNumber="167">
<materialsCitation id="6BBB3C85835DFFACFF04F9E1FCDFF9CF" ID-GBIF-Occurrence="1676480050" collectingDate="2017-02-06" collectorName="L. Kosygin" country="Myanmar" location="Burma" pageId="16" pageNumber="167" specimenCount="2" typeStatus="holotype">
<treatmentCitationGroup id="FBC311F6835DFFACFF04F9E1FCDFF9CF" pageId="16" pageNumber="167">
<taxonomicName id="1CD34D5B835DFFACFF04F9E1FD82F9AE" ID-CoL="77Z9F" authority="Blyth, 1860: 167" authorityName="Blyth" authorityPageNumber="167" authorityYear="1860" box="[151,611,1555,1577]" class="Actinopterygii" family="Cobitidae" genus="Prostheacanthus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="spectabilis">
<emphasis id="E9A7EACA835DFFACFF04F9E1FE52F9AE" box="[151,435,1555,1577]" italics="true" pageId="16" pageNumber="167">Prostheacanthus spectabilis</emphasis>
<treatmentCitation id="5A7210C9835DFFACFE2BF9E1FD82F9AE" author="Blyth" box="[440,611,1555,1577]" page="167" pageId="16" pageNumber="167" year="1860">
<bibRefCitation id="BF424B29835DFFACFE2BF9E1FDD2F9AE" author="Blyth" box="[440,563,1555,1577]" pageId="16" pageNumber="167" refString="Blyth, E. (1860) Report on some fishes receiVed chiefly from the Sitang riVer and its tributary streams, Tenasserim ProVinces. Journal of the Asiatic Society of Bengal, 29, 138 - 174." type="journal article" year="1860">Blyth, 1860</bibRefCitation>
: 167
</treatmentCitation>
</taxonomicName>
(
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locality:
<collectingCountry id="A3C47648835DFFACFD6FF9E6FCA4F9AE" box="[764,837,1556,1577]" name="Myanmar" pageId="16" pageNumber="167">Burma</collectingCountry>
: Tenasserim proVinces;
<typeStatus id="0468887A835DFFACFBA4F9E6FB91F9AE" box="[1079,1136,1556,1577]" pageId="16" pageNumber="167">types</typeStatus>
: none knoWn, possibly at ZSI but cannot be located [
<collectorName id="7626530E835DFFACFE3FF9C1FDC3F9CF" box="[428,546,1586,1608]" pageId="16" pageNumber="167">L. Kosygin</collectorName>
, pers. comm.,
<collectingDate id="BF29E9F0835DFFACFD26F9C0FCD1F9CF" box="[693,816,1586,1608]" pageId="16" pageNumber="167" value="2017-02-06">
<date id="AF6D1018835DFFACFD26F9C0FCD1F9CF" box="[693,816,1586,1608]" pageId="16" pageNumber="167" value="2017-02-06">6 Feb. 2017</date>
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</paragraph>
<paragraph id="DB6C36D8835DFFACFF04F9A3FDA2F9E0" blockId="16.[151,1436,1555,1700]" box="[151,579,1617,1639]" pageId="16" pageNumber="167">
<treatmentCitationGroup id="FBC311F6835DFFACFF04F9A3FDA2F9E0" box="[151,579,1617,1639]" pageId="16" pageNumber="167">
<taxonomicName id="1CD34D5B835DFFACFF04F9A3FE99F9E0" ID-CoL="64G3J" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[151,376,1617,1639]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="spectabilis">
<emphasis id="E9A7EACA835DFFACFF04F9A3FE99F9E0" box="[151,376,1617,1639]" italics="true" pageId="16" pageNumber="167">Acantopsis spectabilis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="5A7210C9835DFFACFE05F9A3FDA2F9E0" author="Kottelat" box="[406,579,1617,1639]" page="22" pageId="16" pageNumber="167" year="2012">
<bibRefCitation id="BF424B29835DFFACFE05F9A3FDC7F9E0" author="Kottelat" box="[406,550,1617,1639]" pageId="16" pageNumber="167" refString="Kottelat, M. (2012) Conspectus cobitidum: an inVentory of the loaches of the World (Teleostei: Cypriniformes: Cobitoidei). Raffles Bulletin of Zoology Supplement, 26, 1 - 199." type="journal article" year="2012">Kottelat 2012</bibRefCitation>
:22
</treatmentCitation>
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</paragraph>
<paragraph id="DB6C36D8835DFFACFF04F982FCAAF923" blockId="16.[151,1436,1555,1700]" pageId="16" pageNumber="167">
<materialsCitation id="6BBB3C85835DFFACFF04F982FCAAF923" ID-GBIF-Occurrence="1676480042" collectionCode="MUMF" country="India" county="ChindWin RiVer" location="Yu RiVer" municipality="Lokchao" pageId="16" pageNumber="167" specimenCode="MUMF 3044" specimenCount="2" stateProvince="Manipur" typeStatus="holotype">
<treatmentCitationGroup id="FBC311F6835DFFACFF04F982FCAAF923" pageId="16" pageNumber="167">
<taxonomicName id="1CD34D5B835DFFACFF04F982FCC5F902" ID-CoL="64FRK" authority="VishWanath and Laisram, 2005: 433" authorityName="Vishwanath and Laisram" authorityPageNumber="433" authorityYear="2005" box="[151,804,1647,1669]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="multistigmatus">
<emphasis id="E9A7EACA835DFFACFF04F982FE42F902" box="[151,419,1647,1669]" italics="true" pageId="16" pageNumber="167">Acantopsis multistigmatus</emphasis>
<treatmentCitation id="5A7210C9835DFFACFE3FF99DFCC5F902" author="VishWanath" box="[428,804,1647,1669]" page="433" pageId="16" pageNumber="167" year="2005">
<bibRefCitation id="BF424B29835DFFACFE3FF99DFD10F902" author="VishWanath" box="[428,753,1647,1669]" pageId="16" pageNumber="167" refString="VishWanath, W. &amp; Laisram, J. (2005) A neW fish of the genus Acantopsis Van Hasselt (Cypriniformes: Cobitidae) from Manipur, Indin [India]. Journal of the Bombay Natural History Society, 101, 433 - 436." type="journal article" year="2005">VishWanath and Laisram, 2005</bibRefCitation>
: 433
</treatmentCitation>
</taxonomicName>
, fig. 1 (
<typeStatus id="0468887A835DFFACFCEBF99DFC44F902" box="[888,933,1647,1669]" pageId="16" pageNumber="167">type</typeStatus>
locality:
<collectingCountry id="A3C47648835DFFACFB9AF99DFBA0F902" box="[1033,1089,1647,1669]" name="India" pageId="16" pageNumber="167">India</collectingCountry>
:
<collectingRegion id="1917F83A835DFFACFBDFF99DFB46F902" box="[1100,1191,1647,1669]" country="India" name="Manipur" pageId="16" pageNumber="167">Manipur</collectingRegion>
:
<collectingCounty id="320D4E54835DFFACFB20F99DFAB6F902" box="[1203,1367,1647,1669]" pageId="16" pageNumber="167">ChindWin RiVer</collectingCounty>
basin,
<collectingMunicipality id="3B08ACA2835DFFACFF54F97CFEC1F923" box="[199,288,1678,1700]" pageId="16" pageNumber="167">Lokchao</collectingMunicipality>
stream, tributary of
<location id="DE0C6003835DFFACFE7CF97DFDACF923" LSID="urn:lsid:plazi:treatment:537A87CE835DFFAEFF04FA74FCC8FEE4:DE0C6003835DFFACFE7CF97DFDACF923" box="[495,589,1678,1700]" country="India" county="ChindWin RiVer" municipality="Lokchao" name="Yu RiVer" pageId="16" pageNumber="167" stateProvince="Manipur">Yu RiVer</location>
;
<typeStatus id="0468887A835DFFACFDC4F97CFD55F923" box="[599,692,1678,1700]" pageId="16" pageNumber="167" type="holotype">holotype</typeStatus>
:
<specimenCode id="8B759EA3835DFFACFD2EF97DFCA5F923" box="[701,836,1678,1700]" collectionCode="MUMF" country="India" httpUri="http://grbio.org/cool/x8gv-47he" name="Department of Life Sciences" pageId="16" pageNumber="167">MUMF 3044</specimenCode>
)
</treatmentCitationGroup>
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="93C96553835DFFADFF04F922FD98FE72" lastPageId="17" lastPageNumber="168" pageId="16" pageNumber="167" type="diagnosis">
<paragraph id="DB6C36D8835DFFACFF04F922FE7BF81A" blockId="16.[151,1437,1744,2022]" pageId="16" pageNumber="167">
<emphasis id="E9A7EACA835DFFACFF04F922FEF0F96E" bold="true" box="[151,273,1744,1769]" pageId="16" pageNumber="167">Diagnosis.</emphasis>
<taxonomicName id="1CD34D5B835DFFACFE88F923FDF9F96E" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[283,536,1745,1769]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="spectabilis">
<emphasis id="E9A7EACA835DFFACFE88F923FDF9F96E" box="[283,536,1745,1769]" italics="true" pageId="16" pageNumber="167">Acantopsis spectabilis</emphasis>
</taxonomicName>
differs from all other species of
<taxonomicName id="1CD34D5B835DFFACFC0FF923FBF6F96F" authorityName="Dialuzona Van Hasselt" authorityYear="1823" box="[924,1047,1745,1768]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="genus">
<emphasis id="E9A7EACA835DFFACFC0FF923FBF6F96F" box="[924,1047,1745,1768]" italics="true" pageId="16" pageNumber="167">Acantopsis</emphasis>
</taxonomicName>
(
<tableCitation id="96510363835DFFACFBBAF922FB63F96E" box="[1065,1154,1744,1769]" captionStart="TABLE 4" captionStartId="15.[120,195,534,553]" captionText="TABLE 4. Diagnostic features of species of Acantopsis. * Data from Siebert (1991 b), Sontirat (1999), or Vishwanath and Laisram (2004). Methods from various sources may not be consistent." pageId="16" pageNumber="167">Table 4</tableCitation>
) by combination of 34 black spots along upper lobe, 23 larger, often darker, black spots on lower lobe of caudal fin (smaller black spots in middle of fin); 12 bands on anal and paired fins; 23 pairs of labial barbels, 9½ branched dorsal rays; 10 pectoral rays; 912 dark saddles along dorsal midline; 915 black spots along side of body; 23 rows of large dark spots on side of head, 4344 total vertebrae, body depth 9.011.6% SL, body width 6.89.4% SL, caudal-peduncle length 12.916.6% SL.
</paragraph>
<paragraph id="DB6C36D8835DFFADFF54F85AFD98FE72" blockId="16.[151,1437,1744,2022]" lastBlockId="17.[151,1436,151,501]" lastPageId="17" lastPageNumber="168" pageId="16" pageNumber="167">
<emphasis id="E9A7EACA835DFFACFF54F85AFE8CF846" bold="true" box="[199,365,1960,1985]" pageId="16" pageNumber="167">Comparisons.</emphasis>
The unique color pattern on caudal fin distinguishes
<taxonomicName id="1CD34D5B835DFFACFC4FF85BFB97F846" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[988,1142,1961,1985]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="spectabilis">
<emphasis id="E9A7EACA835DFFACFC4FF85BFB97F846" box="[988,1142,1961,1985]" italics="true" pageId="16" pageNumber="167">A. spectabilis</emphasis>
</taxonomicName>
from all other species of
<taxonomicName id="1CD34D5B835DFFACFF04F83FFEF3F863" authorityName="Dialuzona Van Hasselt" authorityYear="1823" box="[151,274,1997,2020]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="genus">
<emphasis id="E9A7EACA835DFFACFF04F83FFEF3F863" box="[151,274,1997,2020]" italics="true" pageId="16" pageNumber="167">Acantopsis</emphasis>
</taxonomicName>
. It is most similar to
<taxonomicName id="1CD34D5B835DFFACFD92F83FFD42F862" authorityName="Sontirat" authorityYear="1999" box="[513,675,1997,2021]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="16" pageNumber="167" phylum="Chordata" rank="species" species="thiemmedhi">
<emphasis id="E9A7EACA835DFFACFD92F83FFD42F862" box="[513,675,1997,2021]" italics="true" pageId="16" pageNumber="167">A. thiemmedhi</emphasis>
</taxonomicName>
, which differs in having 1 large black blotch on each lobe of caudal fin. All other species of
<taxonomicName id="1CD34D5B835CFFADFE34FF6AFDC3FF28" authorityName="Dialuzona Van Hasselt" authorityYear="1823" box="[423,546,152,175]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="genus">
<emphasis id="E9A7EACA835CFFADFE34FF6AFDC3FF28" box="[423,546,152,175]" italics="true" pageId="17" pageNumber="168">Acantopsis</emphasis>
</taxonomicName>
have no or dusky bands on caudal fin. It further differs from
<taxonomicName id="1CD34D5B835CFFADFB45FF6AFA98FF37" authorityName="Sontirat" authorityYear="1999" box="[1238,1401,152,176]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="thiemmedhi">
<emphasis id="E9A7EACA835CFFADFB45FF6AFA98FF37" box="[1238,1401,152,176]" italics="true" pageId="17" pageNumber="168">A. thiemmedhi</emphasis>
</taxonomicName>
by having 23 (vs. 1) pairs of labial barbels and in having usually 9½ (vs. 10½) branched dorsal rays and in lacking large black blotch on tip of anterior rays of dorsal fin. It further differs from
<taxonomicName id="1CD34D5B835CFFADFC6BFF12FB8BFF7F" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[1016,1130,224,248]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="dinema">
<emphasis id="E9A7EACA835CFFADFC6BFF12FB8BFF7F" box="[1016,1130,224,248]" italics="true" pageId="17" pageNumber="168">A. dinema</emphasis>
</taxonomicName>
by having dark large spots on side of head, no row of black specks beneath midlateral row of black spots, usually 9½ (vs. 10½) branched dorsal rays, smaller body depth (9.011.6 vs. 12.014.5% SL). It further differs from
<taxonomicName id="1CD34D5B835CFFADFBF3FEDAFB1AFEC7" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[1120,1275,296,320]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rungthipae">
<emphasis id="E9A7EACA835CFFADFBF3FEDAFB1AFEC7" box="[1120,1275,296,320]" italics="true" pageId="17" pageNumber="168">A. rungthipae</emphasis>
</taxonomicName>
and
<taxonomicName id="1CD34D5B835CFFADFAA7FEDAFA99FEB8" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[1332,1400,296,319]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="ioa">
<emphasis id="E9A7EACA835CFFADFAA7FEDAFA99FEB8" box="[1332,1400,296,319]" italics="true" pageId="17" pageNumber="168">A. ioa</emphasis>
</taxonomicName>
by having 23 (vs. 0) pairs of labial barbels, from
<taxonomicName id="1CD34D5B835CFFADFD3DFEBEFCA8FEE3" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[686,841,332,356]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="rungthipae">
<emphasis id="E9A7EACA835CFFADFD3DFEBEFCA8FEE3" box="[686,841,332,356]" italics="true" pageId="17" pageNumber="168">A. rungthipae</emphasis>
</taxonomicName>
in lacking distinct small black spot on upper margin of caudal-fin base, and from
<taxonomicName id="1CD34D5B835CFFADFE49FE82FDFCFE00" authorityName="Boyd &amp; Nithirojpakdee &amp; Deein &amp; Vidthayanon &amp; Grudpan &amp; Tangjitjaroen &amp; Pfeiffer &amp; Randall &amp; Srisombat &amp; Page" authorityYear="2017" box="[474,541,368,391]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="ioa">
<emphasis id="E9A7EACA835CFFADFE49FE82FDFCFE00" box="[474,541,368,391]" italics="true" pageId="17" pageNumber="168">A. ioa</emphasis>
</taxonomicName>
in having 912 (vs. 1523) dark saddles along dorsal midline, 915 (vs. 1321) black spots along side of body, large dark spots on side of head, 4344 (vs. 4648) total vertebrae. It further differs from
<taxonomicName id="1CD34D5B835CFFADFF4BFE4AFE89FE57" authorityName="van Hasselt" authorityYear="1823" box="[216,360,440,464]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="dialuzona">
<emphasis id="E9A7EACA835CFFADFF4BFE4AFE89FE57" box="[216,360,440,464]" italics="true" pageId="17" pageNumber="168">A. dialuzona</emphasis>
</taxonomicName>
by lacking a well developed ocellus on the upper margin of the caudal-fin base and having usually 9½ (vs. 10½) branched dorsal rays.
</paragraph>
</subSubSection>
<caption id="8FAC6650835CFFADFF04F939FE37F978" httpUri="https://zenodo.org/record/1039429/files/figure.png" pageId="17" pageNumber="168" targetBox="[151,1436,541,1717]" targetPageId="17">
<paragraph id="DB6C36D8835CFFADFF04F939FE37F978" blockId="17.[151,1436,1739,1791]" pageId="17" pageNumber="168">
<emphasis id="E9A7EACA835CFFADFF04F939FEF2F966" bold="true" box="[151,275,1739,1761]" pageId="17" pageNumber="168">FIGURE 9.</emphasis>
<emphasis id="E9A7EACA835CFFADFE8AF939FDE2F966" box="[281,515,1739,1761]" italics="true" pageId="17" pageNumber="168">
<taxonomicName id="1CD34D5B835CFFADFE8AF939FE1FF966" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[281,510,1739,1761]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="17" pageNumber="168" phylum="Chordata" rank="species" species="spectabilis">Acantopsis spectabilis</taxonomicName>
.
</emphasis>
(A) Lateral and dorsal VieWs, CAS 238298, 163.0 mm SL. (B) Lateral VieWs, CAS 88783, 112.9 mm SL and 56.2 mm SL.
</paragraph>
</caption>
<subSubSection id="93C96553835CFFAEFF54F8DEFB92FF70" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168" type="description">
<paragraph id="DB6C36D8835CFFADFF54F8DEFAE0F80B" blockId="17.[151,1436,1836,2038]" pageId="17" pageNumber="168">
<emphasis id="E9A7EACA835CFFADFF54F8DEFEB8F8C2" bold="true" box="[199,345,1836,1861]" pageId="17" pageNumber="168">Description.</emphasis>
As in description of genus;
<tableCitation id="96510363835CFFADFD0BF8DEFD16F8C2" box="[664,759,1836,1861]" captionStart="TABLE 1" captionStartId="4.[151,239,152,175]" captionText="TABLE 1. Frequency distributions of labial barbels and meristic characteristics in species of Acantopsis." pageId="17" pageNumber="168">Tables 1</tableCitation>
,
<tableCitation id="96510363835CFFADFC95F8DFFCF7F8C2" box="[774,790,1837,1861]" captionStart="TABLE 2" captionStartId="6.[120,202,556,577]" captionText="TABLE 2. Morphometric data for recognized species of Acantopsis." pageId="17" pageNumber="168">2</tableCitation>
,
<tableCitation id="96510363835CFFADFCB1F8DFFCD2F8C2" box="[802,819,1837,1861]" captionStart="TABLE 4" captionStartId="15.[120,195,534,553]" captionText="TABLE 4. Diagnostic features of species of Acantopsis. * Data from Siebert (1991 b), Sontirat (1999), or Vishwanath and Laisram (2004). Methods from various sources may not be consistent." pageId="17" pageNumber="168">4</tableCitation>
. Head long (21.825.7% SL); snout long (60.170.3% HL); 23 pairs of labial barbels; 9½, rarely 8½, branched dorsal rays; 10, rarely 11, pectoral rays; 3132 abdominal + 12 caudal = 4344 total vertebrae (N=3). Maximum SL =
<quantity id="1C2B9B3D835CFFADFCAAF881FC4CF80C" box="[825,941,1907,1932]" metricMagnitude="-1" metricUnit="m" metricValue="2.125" pageId="17" pageNumber="168" unit="mm" value="212.5">212.5 mm</quantity>
, TL =
<quantity id="1C2B9B3D835CFFADFC6AF881FB8AF80C" box="[1017,1131,1907,1932]" metricMagnitude="-1" metricUnit="m" metricValue="2.425" pageId="17" pageNumber="168" unit="mm" value="242.5">242.5 mm</quantity>
(NIFI 2456).
</paragraph>
<paragraph id="DB6C36D8835CFFAEFF54F864FB92FF70" blockId="17.[151,1436,1836,2038]" lastBlockId="18.[151,1436,151,355]" lastPageId="18" lastPageNumber="169" pageId="17" pageNumber="168">
<emphasis id="E9A7EACA835CFFADFF54F864FEF1F828" bold="true" box="[199,272,1942,1967]" pageId="17" pageNumber="168">Color.</emphasis>
Dark spots in 1 row on top of head, scattered or in 23 rows on side of head. Black line from eye to tip of snout. Nine12 dark saddles along dorsal midline; 915 dark spots or blotches along to slightly below lateral line from head to caudal fin, varying from large and rounded to flattened and elongate; dark spots and blotches on upper side of body, none on lower side. Two3 bold, irregular bands of blotches on caudal fin, darkest at extremity of lower lobe and along margins. One2 dark bands on anal and paired fins. Small black spot near tip of anterior rays of dorsal fin. No small black spot or ocellus near upper margin of caudal-fin origin.
</paragraph>
</subSubSection>
<subSubSection id="93C96553835FFFAEFF54FEF0FCC8FEE4" pageId="18" pageNumber="169" type="distribution">
<paragraph id="DB6C36D8835FFFAEFF54FEF0FCC8FEE4" blockId="18.[151,1436,151,355]" pageId="18" pageNumber="169">
<emphasis id="E9A7EACA835FFFAEFF54FEF0FE81FE9C" bold="true" box="[199,352,258,283]" pageId="18" pageNumber="169">Distribution.</emphasis>
<taxonomicName id="1CD34D5B835FFFAEFEF8FEF1FD89FE9D" baseAuthorityName="Blyth" baseAuthorityYear="1860" box="[363,616,258,282]" class="Actinopterygii" family="Cobitidae" genus="Acantopsis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cypriniformes" pageId="18" pageNumber="169" phylum="Chordata" rank="species" species="spectabilis">
<emphasis id="E9A7EACA835FFFAEFEF8FEF1FD89FE9D" box="[363,616,258,282]" italics="true" pageId="18" pageNumber="169">Acantopsis spectabilis</emphasis>
</taxonomicName>
is known from the
<collectingRegion id="1917F83A835FFFAEFCC6FEF0FC29FE9C" box="[853,968,258,283]" country="Myanmar" name="Ayeyawady" pageId="18" pageNumber="169">Irrawaddy</collectingRegion>
River basin of
<collectingCountry id="A3C47648835FFFAEFB17FEF1FB10FE9C" box="[1156,1265,259,283]" name="Myanmar" pageId="18" pageNumber="169">Myanmar</collectingCountry>
and
<collectingCountry id="A3C47648835FFFAEFABDFEF0FA8BFE9C" box="[1326,1386,258,283]" name="India" pageId="18" pageNumber="169">India</collectingCountry>
, the Salween River basin of
<collectingCountry id="A3C47648835FFFAEFE56FED4FDC8FEB8" box="[453,553,294,319]" name="Thailand" pageId="18" pageNumber="169">Thailand</collectingCountry>
and
<collectingCountry id="A3C47648835FFFAEFDE3FED5FD3EFEB8" box="[624,735,295,319]" name="Myanmar" pageId="18" pageNumber="169">Myanmar</collectingCountry>
, and the Tenasserim provinces region (Dawei River, and presumably the Ataran River basin) of
<collectingCountry id="A3C47648835FFFAEFDDFFEB9FD58FEE4" box="[588,697,331,355]" name="Myanmar" pageId="18" pageNumber="169">Myanmar</collectingCountry>
(
<figureCitation id="43E82A5D835FFFAEFD5BFEB8FCFAFEE4" box="[712,795,330,355]" captionStart="FIGURE 10" captionStartId="18.[151,250,1996,2018]" captionTargetBox="[184,1387,413,1968]" captionTargetId="figure@18.[173,1409,398,1997]" captionTargetPageId="18" captionText="FIGURE 10. Localities of specimens examined of Acantopsis spectabilis, including the type locality of A. multistigmatus." httpUri="https://zenodo.org/record/1039431/files/figure.png" pageId="18" pageNumber="169">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>