treatments-xml/data/1C/11/87/1C1187FCEB62ED4BFF9EFC0C62DC82CC.xml

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<document id="BEF7124E27C80527133FBDDD50A1B9DA" ID-CLB-Dataset="299696" ID-DOI="10.5281/zenodo.11512952" ID-GBIF-Dataset="2b56e3b0-2274-4963-b243-9c991b07072c" ID-ISSN="0068-547X" ID-Zenodo-Dep="11512952" IM.bibliography_approvedBy="julia" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="julia" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="julia" IM.treatmentCitations_approvedBy="julia" IM.treatments_approvedBy="julia" checkinTime="1717721037103" checkinUser="felipe" docAuthor="Kavanaugh, David H. &amp; Cueva-Dabkoski, Mollie" docDate="2023" docId="1C1187FCEB62ED4BFF9EFC0C62DC82CC" docLanguage="en" docName="ProcCalifAcadSci.67.21.493-537.pdf" docOrigin="Proceedings of the California Academy of Sciences 67 (21)" docStyle="DocumentStyle:E411DA009AF5E7F9A7E023431E2B7D5B.2:ProcCalifAcadSci.2015-.journal_article" docStyleId="E411DA009AF5E7F9A7E023431E2B7D5B" docStyleName="ProcCalifAcadSci.2015-.journal_article" docStyleVersion="2" docTitle="Cyclosomus flexuosus" docType="treatment" docVersion="5" lastPageNumber="507" masterDocId="E028FF84EB68ED45FFE5FFAF6327867B" masterDocTitle="A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia." masterLastPageNumber="537" masterPageNumber="493" pageNumber="503" updateTime="1720721490288" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AC1B0DBC7BE5106EA94B45F234F31E53">A Review of Genus Cyclosomus Latreille (Coleoptera: Carabidae: Lebiinae: Cyclosomini) in Asia.</mods:title>
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<mods:namePart id="1703AEA3A38CA0D83565D2F50D529E60">Kavanaugh, David H.</mods:namePart>
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<mods:namePart id="88D4350AAC59E447F8E1FC3E0E1F34FE">Cueva-Dabkoski, Mollie</mods:namePart>
<mods:affiliation id="32F105C3C2ED1DC978C377DE373382AE">Student Science Fellow, California Academy of Sciences, San Francisco, and &amp; 2355 Pearl Street, Santa Monica, CA 90405.</mods:affiliation>
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<mods:date id="DFA15140F3E12DBCDB45CC66304964E4">2023</mods:date>
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<treatment id="1C1187FCEB62ED4BFF9EFC0C62DC82CC" ID-DOI="http://doi.org/10.5281/zenodo.12726721" ID-GBIF-Taxon="235039987" ID-Zenodo-Dep="12726721" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC" httpUri="http://treatment.plazi.org/id/1C1187FCEB62ED4BFF9EFC0C62DC82CC" lastPageId="14" lastPageNumber="507" pageId="10" pageNumber="503">
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<paragraph id="940736EAEB62ED4FFF9EFC0C62D385C1" blockId="10.[123,1175,931,1725]" box="[123,500,931,955]" pageId="10" pageNumber="503">
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<taxonomicName id="53B84D69EB62ED4FFF9EFC0C62D385C1" ID-CoL="32X56" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[123,500,931,955]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFF9EFC0C62D385C1" bold="true" box="[123,500,931,955]" pageId="10" pageNumber="503">
<emphasis id="A6CCEAF8EB62ED4FFF9EFC0C624885C0" bold="true" box="[123,367,931,955]" italics="true" pageId="10" pageNumber="503">Cyclosomus flexuosus</emphasis>
(Fabricius)
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<subSubSection id="DCA26561EB62ED4FFF9EFC6C62C285A2" box="[123,485,963,985]" pageId="10" pageNumber="503" type="description">
<paragraph id="940736EAEB62ED4FFF9EFC6C62C285A2" blockId="10.[123,1175,931,1725]" box="[123,485,963,985]" pageId="10" pageNumber="503">
<figureCitation id="0C832A6FEB62ED4FFF9EFC6C63F185A2" box="[123,214,963,985]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="10" pageNumber="503">Figures 5</figureCitation>
,
<figureCitation id="0C832A6FEB62ED4FFF07FC6C63D785A2" box="[226,240,963,985]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="10" pageNumber="503">6</figureCitation>
,
<figureCitation id="0C832A6FEB62ED4FFF1FFC6C620285A2" box="[250,293,963,985]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="10" pageNumber="503">14B</figureCitation>
,
<figureCitation id="0C832A6FEB62ED4FFECAFC6C627E85A2" box="[303,345,963,985]" captionStart="FIGURE" captionStartId="34.[171,183,869,888]" captionTargetBox="[140,1176,216,846]" captionTargetId="figure-9@34.[120,1176,216,846]" captionTargetPageId="34" captionText="FIGURE 15. Prosternal intercoxal process, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Cinkona, Anamalia Hills, Tamil Nadu, India); C. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); D. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512990" httpUri="https://zenodo.org/record/11512990/files/figure.png" pageId="10" pageNumber="503">15B</figureCitation>
,
<figureCitation id="0C832A6FEB62ED4FFE81FC6C62A985A2" box="[356,398,963,985]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="10" pageNumber="503">16B</figureCitation>
,
<figureCitation id="0C832A6FEB62ED4FFE7DFC6C62E585A2" box="[408,450,963,985]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="10" pageNumber="503">17B</figureCitation>
,
<figureCitation id="0C832A6FEB62ED4FFE28FC6C62C285A2" box="[461,485,963,985]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="10" pageNumber="503">20</figureCitation>
</paragraph>
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<taxonomicName id="53B84D69EB62ED4FFF9EFC4762D28585" authority="Fabricius, 1775: 246" authorityName="Fabricius" authorityPageNumber="246" authorityYear="1775" box="[123,501,1000,1022]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFF9EFC47620B8585" box="[123,300,1000,1022]" italics="true" pageId="10" pageNumber="503">Carabus flexuosus</emphasis>
<bibRefCitation id="F0294B1BEB62ED4FFED7FC4762D28585" author="FABRICIUS, J. C." box="[306,501,1000,1022]" pageId="10" pageNumber="503" refId="ref23315" refString="FABRICIUS, J. C. 1775. Systema entomologica, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Libraria Kortii, Flensburgi et Lipsiae, xxxii + 832 pp." type="book" year="1775">Fabricius, 1775:246</bibRefCitation>
</taxonomicName>
.
<typeStatus id="4B038848EB62ED4FFE18FC4761AF8585" box="[509,648,1000,1022]" pageId="10" pageNumber="503" type="lectotype">LECTOTYPE</typeStatus>
, here designated, a male, deposited in
<collectionCode id="F2A9AE2FEB62ED4FFC12FC4667678585" box="[1015,1088,1001,1022]" pageId="10" pageNumber="503">ZMUK</collectionCode>
, labeled: “
<taxonomicName id="53B84D69EB62ED4FFF51FBAA624A8260" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[180,365,1029,1051]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">Scolytus flexuosus</taxonomicName>
“ [handwritten]/ “
<typeStatus id="4B038848EB62ED4FFDF2FBAA61858260" box="[535,674,1029,1051]" pageId="10" pageNumber="503" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="53B84D69EB62ED4FFD4CFBAA60D38260" ID-CoL="QWMS" authority="Fabricius 1775" authorityName="Fabricius" authorityYear="1775" box="[681,1012,1029,1051]" class="Insecta" family="Carabidae" genus="Carabus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFD4CFBAA607B8260" box="[681,860,1029,1051]" italics="true" pageId="10" pageNumber="503">Carabus flexuosus</emphasis>
<bibRefCitation id="F0294B1BEB62ED4FFC86FBAA60D38260" author="FABRICIUS, J. C." box="[867,1012,1029,1051]" pageId="10" pageNumber="503" refId="ref23315" refString="FABRICIUS, J. C. 1775. Systema entomologica, sistens insectorum classes, ordines, genera, species, adiectis synonymis, locis, descriptionibus, observationibus. Libraria Kortii, Flensburgi et Lipsiae, xxxii + 832 pp." type="book" year="1775">Fabricius 1775</bibRefCitation>
</taxonomicName>
design. by D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2022” [red label].
<typeStatus id="4B038848EB62ED4FFD56FB8C61C48243" box="[691,739,1059,1080]" pageId="10" pageNumber="503">Type</typeStatus>
locality: eastern
<collectingCountry id="ECAF767AEB62ED4FFC6CFB8D609A8243" box="[905,957,1058,1080]" name="India" pageId="10" pageNumber="503">India</collectingCountry>
.
</paragraph>
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<taxonomicName id="53B84D69EB62ED4FFF9EFBEF62BA822D" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[123,413,1088,1110]" class="Insecta" family="Curculionidae" genus="Scolytus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFF9EFBEF620D822D" box="[123,298,1088,1110]" italics="true" pageId="10" pageNumber="503">Scolytus flexuosus</emphasis>
(Fabricius)
</taxonomicName>
,
<bibRefCitation id="F0294B1BEB62ED4FFE42FBEF614B822D" author="FABRICIUS, J. C." box="[423,620,1088,1110]" pageId="10" pageNumber="503" pagination="213 - 228" refId="ref23357" refString="FABRICIUS, J. C. 1790. Nova insectorum genera. Skrivter af Naturhistorie-Selskabet 1: 213 - 228." type="journal article" year="1790">Fabricius (1790:221</bibRefCitation>
;
<bibRefCitation id="F0294B1BEB62ED4FFD92FBEF61F1822D" author="FABRICIUS, J. C." box="[631,726,1088,1110]" pageId="10" pageNumber="503" refId="ref23378" refString="FABRICIUS, J. C. 1792. Entomologia systematica emendate et aucta, secundum classes, ordines, genera, species adiectis synonymis, locis, observationibus, descriptionibus. Tome I. Pars I. Proft, Hafniae, xx + 330 pp." type="book" year="1792">1792:180</bibRefCitation>
).
</paragraph>
<paragraph id="940736EAEB62ED4FFF9EFBF2607A82EB" blockId="10.[123,1175,931,1725]" pageId="10" pageNumber="503">
<taxonomicName id="53B84D69EB62ED4FFF9EFBF262E08208" ID-CoL="32X56" authority="(Fabricius)" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[123,455,1117,1139]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFF9EFBF262768208" box="[123,337,1117,1139]" italics="true" pageId="10" pageNumber="503">Cyclosomus flexuosus</emphasis>
(Fabricius)
</taxonomicName>
,
<bibRefCitation id="F0294B1BEB62ED4FFE37FBF261878208" author="ANDREWES, H. E." box="[466,672,1117,1139]" pageId="10" pageNumber="503" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:153</bibRefCitation>
and 166, 1924a:464, 1927:105, 1930:365);
<bibRefCitation id="F0294B1BEB62ED4FFBBCFBF267B28208" author="HELLER, K. M." box="[1113,1173,1117,1139]" pageId="10" pageNumber="503" pagination="295 - 307" refId="ref23510" refString="HELLER, K. M. 1923. Some new Malayan Carabidae, especially Philippine. The Philippine Journal of Science 23: 295 - 307 + 1 pl." type="journal article" year="1923">Heller</bibRefCitation>
(1923:302);
<bibRefCitation id="F0294B1BEB62ED4FFEC6FBD562F282EB" author="CSIKI, E." box="[291,469,1146,1168]" pageId="10" pageNumber="503" pagination="1279 - 1598" refId="ref23258" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1295)</bibRefCitation>
;
<bibRefCitation id="F0294B1BEB62ED4FFE05FBD461BF82EB" author="LORENZ, W." box="[480,664,1146,1168]" pageId="10" pageNumber="503" refId="ref23840" refString="LORENZ, W. 2005. Systematic List of Extant Ground Beetles of the World (Insecta Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae). Second edition. W. Lorenz, Tutzing, 530 pp." type="book" year="2005">Lorenz (2005:452)</bibRefCitation>
;
<bibRefCitation id="F0294B1BEB62ED4FFD47FBD5607E82EB" author="KAVANAUGH, D. H." box="[674,857,1146,1168]" pageId="10" pageNumber="503" pagination="267 - 298" refId="ref23574" refString="KAVANAUGH, D. H. 2015. A Review of the Genus Cyclosomus Latreille (Coleoptera: Carabidae: Cyclosomini) in the Afrotropical Region. Proceedings of the California Academy of Sciences (Series 4), 62: 267 - 298." type="journal article" year="2015">Kavanaugh (2015)</bibRefCitation>
.
</paragraph>
<paragraph id="940736EAEB62ED4FFF9EFB3862DE83CF" blockId="10.[123,1175,931,1725]" pageId="10" pageNumber="503">
<taxonomicName id="53B84D69EB62ED4FFF9EFB38613982D6" authority="Nietner, 1857: 312" authorityName="Nietner" authorityPageNumber="312" authorityYear="1857" box="[123,542,1175,1197]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="A6CCEAF8EB62ED4FFF9EFB38624582D6" box="[123,354,1175,1197]" italics="true" pageId="10" pageNumber="503">Cyclosomus dytiscoides</emphasis>
<bibRefCitation id="F0294B1BEB62ED4FFE8EFB38613982D6" author="NIETNER, J." box="[363,542,1175,1197]" pageId="10" pageNumber="503" pagination="132 - 153" refId="ref23924" refString="NIETNER, J. 1857. Entomological papers, being chiefly descriptions of new Ceylon Coleoptera with such observations on their habits etc., as appear in any way interesting. Journal of the Asiatic Society of Bengal 26: 132 - 153." type="journal article" year="1857">Nietner, 1857:312</bibRefCitation>
</taxonomicName>
.
<typeStatus id="4B038848EB62ED4FFDCFFB38619282D6" box="[554,693,1175,1197]" pageId="10" pageNumber="503" type="lectotype">LECTOTYPE</typeStatus>
, here designated, a female, deposited in
<collectionCode id="F2A9AE2FEB62ED4FFBA9FB3767B582D6" box="[1100,1170,1176,1197]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="503">MFNB</collectionCode>
, labeled: “
<collectingCountry id="ECAF767AEB62ED4FFEEFFB1B627482B1" box="[266,339,1204,1226]" name="Sri Lanka" pageId="10" pageNumber="503">Ceylon</collectingCountry>
,
<location id="91676031EB62ED4FFE84FB1B628C82B1" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB62ED4FFE84FB1B628C82B1" box="[353,427,1204,1226]" country="Sri Lanka" municipality="Type" name="Nietner" pageId="10" pageNumber="503">Nietner</location>
” [handwritten]/ “
<collectingMunicipality id="7463AC90EB62ED4FFDBFFB1B61AA82B1" box="[602,653,1204,1226]" pageId="10" pageNumber="503">Type</collectingMunicipality>
” [red label]/ “
<typeStatus id="4B038848EB62ED4FFCC4FB1B60B882B1" box="[801,927,1204,1226]" pageId="10" pageNumber="503" type="syntype">SYNTYPUS</typeStatus>
<taxonomicName id="53B84D69EB62ED4FFC4CFB1B67B282B1" authority="Nietner, 1857: 312" authorityName="Nietner" authorityPageNumber="312" authorityYear="1857" box="[937,1173,1204,1226]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="dytiscoides">Cyclosomus dytiscoides</taxonomicName>
<bibRefCitation id="F0294B1BEB62ED4FFF4EFB7E6216829C" author="NIETNER, J." box="[171,305,1233,1255]" pageId="10" pageNumber="503" pagination="132 - 153" refId="ref23924" refString="NIETNER, J. 1857. Entomological papers, being chiefly descriptions of new Ceylon Coleoptera with such observations on their habits etc., as appear in any way interesting. Journal of the Asiatic Society of Bengal 26: 132 - 153." type="journal article" year="1857">Nietner 1857</bibRefCitation>
” [red label]/ “
<typeStatus id="4B038848EB62ED4FFE2BFB7E617E829C" box="[462,601,1233,1255]" pageId="10" pageNumber="503" type="lectotype">LECTOTYPE</typeStatus>
<taxonomicName id="53B84D69EB62ED4FFD83FB7E60C4829C" authority="Nietner 1857" authorityName="Nietner" authorityYear="1857" box="[614,995,1233,1255]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="A6CCEAF8EB62ED4FFD83FB7E6075829C" box="[614,850,1233,1255]" italics="true" pageId="10" pageNumber="503">Cyclosomus dytiscoides</emphasis>
<bibRefCitation id="F0294B1BEB62ED4FFCBBFB7E60C4829C" author="NIETNER, J." box="[862,995,1233,1255]" pageId="10" pageNumber="503" pagination="132 - 153" refId="ref23924" refString="NIETNER, J. 1857. Entomological papers, being chiefly descriptions of new Ceylon Coleoptera with such observations on their habits etc., as appear in any way interesting. Journal of the Asiatic Society of Bengal 26: 132 - 153." type="journal article" year="1857">Nietner 1857</bibRefCitation>
</taxonomicName>
design. by D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2023” [red label].
<typeStatus id="4B038848EB62ED4FFD2BFB40607D837E" box="[718,858,1263,1285]" pageId="10" pageNumber="503" type="paralectotype">Paralectotypes</typeStatus>
examined:
<specimenCount id="82BEFD63EB62ED4FFC36FB406763837E" box="[979,1092,1263,1285]" count="1" pageId="10" pageNumber="503" type="female">one female</specimenCount>
, also in
<collectionCode id="F2A9AE2FEB62ED4FFF4EFAA263D68359" box="[171,241,1293,1314]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="10" pageNumber="503">MFNB</collectionCode>
, labeled: “
<taxonomicName id="53B84D69EB62ED4FFEBBFAA361868359" authority="Nietner" authorityName="Nietner" box="[350,673,1292,1314]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="dytiscoides">Cyclosomus dytiscoides Nietner</taxonomicName>
” [handwritten]/ “Type” [red label]/ “
<typeStatus id="4B038848EB62ED4FFBFDFAA367B18359" box="[1048,1174,1292,1314]" pageId="10" pageNumber="503" type="syntype">SYNTYPUS</typeStatus>
<taxonomicName id="53B84D69EB62ED4FFF4EFA8661028344" ID-CoL="8LCTN" authority="Nietner 1857" authorityName="Nietner" authorityYear="1857" box="[171,549,1321,1343]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="dytiscoides">
Cyclosomus dytiscoides
<bibRefCitation id="F0294B1BEB62ED4FFE47FA8661028344" author="NIETNER, J." box="[418,549,1321,1343]" pageId="10" pageNumber="503" pagination="132 - 153" refId="ref23924" refString="NIETNER, J. 1857. Entomological papers, being chiefly descriptions of new Ceylon Coleoptera with such observations on their habits etc., as appear in any way interesting. Journal of the Asiatic Society of Bengal 26: 132 - 153." type="journal article" year="1857">Nietner 1857</bibRefCitation>
</taxonomicName>
labeled by MNHUB 2013” [red label]/ “
<typeStatus id="4B038848EB62ED4FFC28FA8667B18344" box="[973,1174,1321,1343]" pageId="10" pageNumber="503" type="paralectotype">PARALECTOTYPE</typeStatus>
<taxonomicName id="53B84D69EB62ED4FFF4EFAE961348327" ID-CoL="8LCTN" authority="Nietner 1857" authorityName="Nietner" authorityYear="1857" box="[171,531,1350,1372]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="A6CCEAF8EB62ED4FFF4EFAE962B78327" box="[171,400,1350,1372]" italics="true" pageId="10" pageNumber="503">Cyclosomus dytiscoides</emphasis>
<bibRefCitation id="F0294B1BEB62ED4FFE70FAE961348327" author="NIETNER, J." box="[405,531,1350,1372]" pageId="10" pageNumber="503" pagination="132 - 153" refId="ref23924" refString="NIETNER, J. 1857. Entomological papers, being chiefly descriptions of new Ceylon Coleoptera with such observations on their habits etc., as appear in any way interesting. Journal of the Asiatic Society of Bengal 26: 132 - 153." type="journal article" year="1857">Nietner 1857</bibRefCitation>
</taxonomicName>
design. by D.H. Kavanaugh &amp; M. Cueva-Dabkoski 2023” [yellow label].
<typeStatus id="4B038848EB62ED4FFF1DFACB620E8302" box="[248,297,1380,1401]" pageId="10" pageNumber="503">Type</typeStatus>
locality:
<collectingCountry id="ECAF767AEB62ED4FFE72FACC61248302" box="[407,515,1379,1401]" name="Sri Lanka" pageId="10" pageNumber="503">Sri Lanka</collectingCountry>
,
<collectingRegion id="567CF808EB62ED4FFDF0FACB61EA8302" box="[533,717,1379,1401]" country="Sri Lanka" name="Western" pageId="10" pageNumber="503">Western Province</collectingRegion>
, Negombo.
<bibRefCitation id="F0294B1BEB62ED4FFCB0FACC67038302" author="CHAUDOIR, M." box="[853,1060,1379,1401]" pageId="10" pageNumber="503" pagination="1 - 84" refId="ref23225" refString="CHAUDOIR, M. 1876. Etude monographique des Masoreides, des Tetragonoderides et du genre Nemotarsus. Bulletin de la Societe Imperiale des Naturalistes de Moscou 51: 1 - 84." type="journal article" year="1876">Chaudoir (1876:31)</bibRefCitation>
;
<bibRefCitation id="F0294B1BEB62ED4FFBD3FACC67B28302" author="BOUCHARD, J." box="[1078,1173,1379,1401]" pageId="10" pageNumber="503" pagination="169 - 176" refId="ref23110" refString="BOUCHARD, J. 1903. Insectes recueillis par M. le Professeur Dr Forster a Borneo, Java et Sumatra (Palembang). Annales de la Societe Entomologique de France 72: 169 - 176." type="journal article" year="1903">Bouchard</bibRefCitation>
(1903:174);
<bibRefCitation id="F0294B1BEB62ED4FFEC0FA2F62D283ED" author="ANDREWES, H. E." box="[293,501,1408,1430]" pageId="10" pageNumber="503" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:154</bibRefCitation>
, 1927:105);
<bibRefCitation id="F0294B1BEB62ED4FFD92FA2F600A83ED" author="CSIKI, E." box="[631,813,1408,1430]" pageId="10" pageNumber="503" pagination="1279 - 1598" refId="ref23258" refString="CSIKI, E. 1932. Coleopterorum catalogus. Pars 124. Carabidae: Harpalinae VII. W. Junk, Berlin, pp. 1279 - 1598." type="book chapter" year="1932">Csiki (1932:1295)</bibRefCitation>
;
<bibRefCitation id="F0294B1BEB62ED4FFCDEFA2F670783ED" author="KAVANAUGH, D. H." box="[827,1056,1408,1430]" pageId="10" pageNumber="503" pagination="267 - 298" refId="ref23574" refString="KAVANAUGH, D. H. 2015. A Review of the Genus Cyclosomus Latreille (Coleoptera: Carabidae: Cyclosomini) in the Afrotropical Region. Proceedings of the California Academy of Sciences (Series 4), 62: 267 - 298." type="journal article" year="2015">Kavanaugh (2015:281)</bibRefCitation>
. Synonymy proposed by
<bibRefCitation id="F0294B1BEB62ED4FFEC2FA3162DE83CF" author="ANDREWES, H. E." box="[295,505,1438,1460]" pageId="10" pageNumber="503" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:154)</bibRefCitation>
</paragraph>
<paragraph id="940736EAEB62ED4FFF4EFA6C606F8060" blockId="10.[123,1175,931,1725]" pageId="10" pageNumber="503">
<emphasis id="A6CCEAF8EB62ED4FFF4EFA6C610A83A1" bold="true" box="[171,557,1475,1498]" pageId="10" pageNumber="503">Notes on types and nomenclature.</emphasis>
We follow
<bibRefCitation id="F0294B1BEB62ED4FFD4DFA6D60A383A1" author="ANDREWES, H. E." box="[680,900,1474,1498]" pageId="10" pageNumber="503" pagination="145 - 195" refId="ref22984" refString="ANDREWES, H. E. 1921. Notes on synonymy and some types of Oriental Carabidae in various foreign collections. Transactions of the Entomological Society of London 1921: 145 - 195." type="journal article" year="1921">Andrewes (1921:31)</bibRefCitation>
in recognizing the Fabrician specimen at
<collectionCode id="F2A9AE2FEB62ED4FFEDCFA4B62AF8380" box="[313,392,1508,1531]" pageId="10" pageNumber="503">ZMUK</collectionCode>
as the type of
<taxonomicName id="53B84D69EB62ED4FFDCAFA4B61948380" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[559,691,1507,1531]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFDCAFA4B61948380" box="[559,691,1507,1531]" italics="true" pageId="10" pageNumber="503">C. flexuosus</emphasis>
</taxonomicName>
and provide
<typeStatus id="4B038848EB62ED4FFCA0FA4C60808380" box="[837,935,1507,1531]" pageId="10" pageNumber="503" type="lectotype">lectotype</typeStatus>
designations for both
<taxonomicName id="53B84D69EB62ED4FFF9EF9AB63DA8060" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[123,253,1539,1563]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFF9EF9AB63DA8060" box="[123,253,1539,1563]" italics="true" pageId="10" pageNumber="503">C. flexuosus</emphasis>
</taxonomicName>
and
<taxonomicName id="53B84D69EB62ED4FFED6F9AB62ED8060" authority="Nietner, 1857: 312" authorityName="Nietner" authorityPageNumber="312" authorityYear="1857" box="[307,458,1539,1563]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="A6CCEAF8EB62ED4FFED6F9AB62ED8060" box="[307,458,1539,1563]" italics="true" pageId="10" pageNumber="503">C. dytiscoides</emphasis>
</taxonomicName>
to promote nomenclatural stability.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB62ED48FF4EF98B6103854C" lastPageId="13" lastPageNumber="506" pageId="10" pageNumber="503" type="diagnosis">
<paragraph id="940736EAEB62ED49FF4EF98B639083BD" blockId="10.[123,1175,931,1725]" lastBlockId="12.[123,1174,1230,1702]" lastPageId="12" lastPageNumber="505" pageId="10" pageNumber="503">
<emphasis id="A6CCEAF8EB62ED4FFF4EF98B62388040" bold="true" box="[171,287,1572,1595]" pageId="10" pageNumber="503">Diagnosis.</emphasis>
Adults of
<taxonomicName id="53B84D69EB62ED4FFE69F98B612A8047" baseAuthorityName="Fabricius" baseAuthorityYear="1775" box="[396,525,1572,1596]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="503" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB62ED4FFE69F98B612A8047" box="[396,525,1572,1596]" italics="true" pageId="10" pageNumber="503">C. flexuosus</emphasis>
</taxonomicName>
can be distinguished from those of other
<taxonomicName id="53B84D69EB62ED4FFC27F98B67B28047" box="[962,1173,1572,1596]" pageId="10" pageNumber="503">
<emphasis id="A6CCEAF8EB62ED4FFC27F98B67648047" box="[962,1091,1572,1596]" italics="true" pageId="10" pageNumber="503">Cyclosomus</emphasis>
species
</taxonomicName>
in Asia by the following combination of character states: Body size larger for genus, BL males =
<quantity id="53409B0FEB62ED4FFF9EF9CB62368007" box="[123,273,1636,1660]" metricMagnitude="-3" metricUnit="m" metricValue="8.8" metricValueMax="9.4" metricValueMin="8.2" pageId="10" pageNumber="503" unit="mm" value="8.8" valueMax="9.4" valueMin="8.2">8.2 to 9.4 mm</quantity>
, females =
<quantity id="53409B0FEB62ED4FFE6FF9CB61068007" box="[394,545,1636,1660]" metricMagnitude="-3" metricUnit="m" metricValue="9.05" metricValueMax="9.8" metricValueMin="8.3" pageId="10" pageNumber="503" unit="mm" value="9.05" valueMax="9.8" valueMin="8.3">8.3 to 9.8 mm</quantity>
; body form (
<figureCitation id="0C832A6FEB62ED4FFD4CF9CB60138007" box="[681,820,1636,1660]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="10" pageNumber="503">Figs. 5A, 5C</figureCitation>
,
<figureCitation id="0C832A6FEB62ED4FFCDAF9CB60AB8007" box="[831,908,1636,1660]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="10" pageNumber="503">6A, 6B</figureCitation>
) slightly elongate ovoid; pronotum (
<figureCitation id="0C832A6FEB62ED4FFF14F92A627780E6" box="[241,336,1669,1693]" captionStart="FIGURE" captionStartId="33.[171,183,1573,1592]" captionTargetBox="[139,1158,216,1560]" captionTargetId="figure-14@33.[135,1162,216,1560]" captionTargetPageId="33" captionText="FIGURE 14. Pronota, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Bhalukpong, Arunachal Pradesh, India); B. C. flexuosus (Fabricius) (Hetauda, Bagmati Province, Nepal); C. C. inustus Andrewes, holotype (Hong Kong, China); D. C. marginatus Motschulsky (India); E. C. pallidus Kavanaugh and Cueva- Dabkoski sp. nov. (Karachi, Sindh Province, Pakistan); F. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); G. C. sumatrensis Bouchard (Taman Negra National Park, Pahang, Malaysia); H. C. suturalis (Wiedemann), lectotype (“Bengala”, India); I. C. vespertilio Cueva-Dabkoski and Kavanaugh sp. nov. Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512988" httpUri="https://zenodo.org/record/11512988/files/figure.png" pageId="10" pageNumber="503">Fig. 14B</figureCitation>
) relatively broader (ratio PWM/PL = 2.45 to 2.69), disc light reddish brown to rufopiceous, with lateral pale bands moderately broad but not sharply defined, anterior angles broad and broadly rounded apically, lateral margins smoothly arcuate, not sinuate near anterior angles; elytra with pattern of dark maculae more extensively developed than average in most specimens (
<figureCitation id="0C832A6FEB64ED49FF2FFAA1620D835D" box="[202,298,1294,1318]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="12" pageNumber="505">Figs. 5C</figureCitation>
,
<figureCitation id="0C832A6FEB64ED49FED3FAA1627F835D" box="[310,344,1294,1318]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="12" pageNumber="505">6A</figureCitation>
) but highly varied and less than average (
<figureCitation id="0C832A6FEB64ED49FCC4FAA160A5835D" box="[801,898,1294,1318]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="12" pageNumber="505">Figs. 5A</figureCitation>
,
<figureCitation id="0C832A6FEB64ED49FC6AFAA16089835D" box="[911,942,1294,1318]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="12" pageNumber="505">6B</figureCitation>
) in a few specimens, preapical dark spot present; elytral epipleuron with long setae over entire length (although many of these are broken off in many specimens); male median lobe with apical lamella slightly recurved dorsally in relation to ventral curvature of shaft in lateral aspect (
<figureCitation id="0C832A6FEB64ED49FCD6FAC160B383FD" box="[819,916,1390,1414]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="12" pageNumber="505">Fig. 16B</figureCitation>
), apical lamella broadly triangular with a distinct, more narrowly-rounded projection apicomedially in dorsal aspect (
<figureCitation id="0C832A6FEB64ED49FB8FFA21638D83BD" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="12" pageNumber="505">Fig. 17B</figureCitation>
).
</paragraph>
<caption id="C0C76662EB63ED4EFF4EF986625F8014" ID-DOI="http://doi.org/10.5281/zenodo.11512964" ID-Zenodo-Dep="11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="11" pageNumber="504" startId="11.[171,183,1577,1596]" targetBox="[203,1083,216,1560]" targetPageId="11" targetType="figure">
<paragraph id="940736EAEB63ED4EFF4EF986625F8014" blockId="11.[123,1173,1577,1647]" pageId="11" pageNumber="504">
<smallCapsWord id="92E1A036EB63ED4EFF4EF98663CF8040" baselines="1591,1591" box="[171,232,1577,1596]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="11" pageNumber="504">FIGURE</smallCapsWord>
5.
<taxonomicName id="53B84D69EB63ED4EFEE1F98661118047" authority="Fabricius. A." authorityName="Fabricius. A." box="[260,566,1577,1596]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="504" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB63ED4EFEE1F98662E78047" box="[260,448,1577,1596]" italics="true" pageId="11" pageNumber="504">Cyclosomus flexuosus</emphasis>
Fabricius. A.
</taxonomicName>
Holotype male, dorsal habitus; B. Labels associated with
<taxonomicName id="53B84D69EB63ED4EFBC9F98667B28047" box="[1068,1173,1577,1596]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="504" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB63ED4EFBC9F98667B28047" box="[1068,1173,1577,1596]" italics="true" pageId="11" pageNumber="504">C. flexuosus</emphasis>
</taxonomicName>
holotype; C.
<taxonomicName id="53B84D69EB63ED4EFF0CF9ED62DE802E" authority="Nietner" authorityName="Nietner" box="[233,505,1602,1621]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="504" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="A6CCEAF8EB63ED4EFF0CF9ED6293802E" box="[233,436,1602,1621]" italics="true" pageId="11" pageNumber="504">Cyclosomus dytiscoides</emphasis>
Nietner
</taxonomicName>
, lectotype female, dorsal habitus; D. Labels associated with
<taxonomicName id="53B84D69EB63ED4EFC1CF9ED6756802E" box="[1017,1137,1602,1621]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="504" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="A6CCEAF8EB63ED4EFC1CF9ED6756802E" box="[1017,1137,1602,1621]" italics="true" pageId="11" pageNumber="504">C. dytiscoides</emphasis>
</taxonomicName>
lectotype. Scale lines = 1.0 mm.
</paragraph>
</caption>
<caption id="C0C76662EB64ED49FF4EFBD8601D82DF" ID-DOI="http://doi.org/10.5281/zenodo.11512968" ID-Zenodo-Dep="11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="12" pageNumber="505" startId="12.[171,183,1143,1162]" targetBox="[120,1176,216,1117]" targetPageId="12" targetType="figure">
<paragraph id="940736EAEB64ED49FF4EFBD8601D82DF" blockId="12.[123,1174,1143,1188]" pageId="12" pageNumber="505">
<smallCapsWord id="92E1A036EB64ED49FF4EFBD863CF82F2" baselines="1157,1157" box="[171,232,1143,1162]" lowerCaseFontSize="6" mainFontSize="8" normCase="title" normString="Figure" pageId="12" pageNumber="505">FIGURE</smallCapsWord>
6.
<taxonomicName id="53B84D69EB64ED49FEEDFBD8610682F1" authority="Fabricius" authorityName="Fabricius" box="[264,545,1143,1162]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="505" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB64ED49FEEDFBD862E182F1" box="[264,454,1143,1162]" italics="true" pageId="12" pageNumber="505">Cyclosomus flexuosus</emphasis>
Fabricius
</taxonomicName>
, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm.
</paragraph>
</caption>
<paragraph id="940736EAEB64ED49FF4EFA60671D80DD" blockId="12.[123,1174,1230,1702]" pageId="12" pageNumber="505">
In describing
<taxonomicName id="53B84D69EB64ED49FED9FA6061B6839D" authority=", Kavanaugh (2015)" authorityName=", Kavanaugh" authorityYear="2015" box="[316,657,1486,1510]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="505" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB64ED49FED9FA60629A839D" box="[316,445,1486,1510]" italics="true" pageId="12" pageNumber="505">C. flexuosus</emphasis>
,
<bibRefCitation id="F0294B1BEB64ED49FE2FFA6161B6839D" author="KAVANAUGH, D. H." box="[458,657,1486,1510]" pageId="12" pageNumber="505" pagination="267 - 298" refId="ref23574" refString="KAVANAUGH, D. H. 2015. A Review of the Genus Cyclosomus Latreille (Coleoptera: Carabidae: Cyclosomini) in the Afrotropical Region. Proceedings of the California Academy of Sciences (Series 4), 62: 267 - 298." type="journal article" year="2015">Kavanaugh (2015)</bibRefCitation>
</taxonomicName>
incorrectly stated “Male with middle tarsomeres 1 to 3 not laterally expanded and without ventral pads of adhesive setae.” In fact,
<taxonomicName id="53B84D69EB64ED49FBF5FA4067B2807D" authorityName=", Kavanaugh" authorityYear="2015" box="[1040,1173,1518,1542]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="505" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB64ED49FBF5FA4067B2807D" box="[1040,1173,1518,1542]" italics="true" pageId="12" pageNumber="505">C. flexuosus</emphasis>
</taxonomicName>
males, like those of all the other
<taxonomicName id="53B84D69EB64ED49FE16F9A161EB805D" box="[499,716,1550,1574]" pageId="12" pageNumber="505">
<emphasis id="A6CCEAF8EB64ED49FE16F9A16153805D" box="[499,628,1550,1574]" italics="true" pageId="12" pageNumber="505">Cyclosomus</emphasis>
species
</taxonomicName>
represented in Asia for which males are known, have middle tarsomeres 1 to 3 distinctly expanded in comparison with those in females and have two longitudinal rows of squamatous adhesive setae ventrally on tarsomeres 1 to 3. In contrast, most (four of six) species in the Afrotropical Region have the male middle tarsi similar to those of females in form (i.e., more slender) and in the absence of ventral adhesive setae.
</paragraph>
<paragraph id="940736EAEB65ED48FF4EFF74676487AF" blockId="13.[123,1174,219,1725]" pageId="13" pageNumber="506">
This species demonstrates the greatest range of variation in elytral dark color pattern seen among all species in the Asian fauna. The extremes are represented in
<figureCitation id="0C832A6FEB65ED48FC8EFF54608C8768" box="[875,939,251,275]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="13" pageNumber="506">Fig. 6</figureCitation>
, with the most broadly developed dark pattern (
<figureCitation id="0C832A6FEB65ED48FE44FEB462D18748" box="[417,502,283,307]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="13" pageNumber="506">Fig. 6A</figureCitation>
) covering more than 50% of the elytral surface, and the least developed pattern (
<figureCitation id="0C832A6FEB65ED48FEA9FE9462B88728" box="[332,415,315,339]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="13" pageNumber="506">Fig. 6B</figureCitation>
) restricted to the basal dark band, a typical longitudinal dark band, a relatively narrow middle dark band extended only to interval 4, and a small preapical spot. The patterns seen in the
<typeStatus id="4B038848EB65ED48FEA9FED2625D87EE" box="[332,378,381,405]" pageId="13" pageNumber="506">type</typeStatus>
specimens of
<taxonomicName id="53B84D69EB65ED48FDF4FED361B587EF" authorityName=", Kavanaugh" authorityYear="2015" box="[529,658,380,404]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="506" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB65ED48FDF4FED361B587EF" box="[529,658,380,404]" italics="true" pageId="13" pageNumber="506">C. flexuosus</emphasis>
</taxonomicName>
(
<figureCitation id="0C832A6FEB65ED48FD45FED361D487EF" box="[672,755,380,404]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="13" pageNumber="506">Fig. 5A</figureCitation>
) and its junior synonym,
<taxonomicName id="53B84D69EB65ED48FBE5FED367B187EF" box="[1024,1174,380,404]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="506" phylum="Arthropoda" rank="species" species="dytiscoides">
<emphasis id="A6CCEAF8EB65ED48FBE5FED367B187EF" box="[1024,1174,380,404]" italics="true" pageId="13" pageNumber="506">C. dytiscoides</emphasis>
</taxonomicName>
(
<figureCitation id="0C832A6FEB65ED48FF66FE3363F087CF" box="[131,215,412,436]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="13" pageNumber="506">Fig. 5B</figureCitation>
), are close to the opposite extremes for the species, so it is certainly understandable that they would have been described as distinct species on the basis of different color patterns.
</paragraph>
<paragraph id="940736EAEB65ED48FF4EFE736103854C" blockId="13.[123,1174,219,1725]" pageId="13" pageNumber="506">
Members of this species share epipleura having relatively long setae throughout their length only with those of
<taxonomicName id="53B84D69EB65ED48FEA4FE5262FF846E" box="[321,472,509,533]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="506" phylum="Arthropoda" rank="species" species="philippinus">
<emphasis id="A6CCEAF8EB65ED48FEA4FE5262FF846E" box="[321,472,509,533]" italics="true" pageId="13" pageNumber="506">C. philippinus</emphasis>
</taxonomicName>
; but they differ from the latter in having a prepical pale spot present and pronota with the pale lateral bands less well-defined medially. Males of the two species differ markedly in the curvature of the male median lobe in lateral aspect (compare
<figureCitation id="0C832A6FEB65ED48FBE4FD926778842E" box="[1025,1119,573,597]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="13" pageNumber="506">Fig. 16B</figureCitation>
with
<figureCitation id="0C832A6FEB65ED48FF9EFDF263FE840E" box="[123,217,605,629]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="13" pageNumber="506">Fig. 16E</figureCitation>
) and shape of the apical lamella in dorsal aspect (compare
<figureCitation id="0C832A6FEB65ED48FCBEFDF2609E840E" box="[859,953,605,629]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="13" pageNumber="506">Fig. 17B</figureCitation>
with
<figureCitation id="0C832A6FEB65ED48FC1DFDF2677E840E" box="[1016,1113,605,629]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="13" pageNumber="506">Fig. 17E</figureCitation>
). The elytra color pattern of darker specimens of
<taxonomicName id="53B84D69EB65ED48FDB3FDD161FC84ED" authorityName=", Kavanaugh" authorityYear="2015" box="[598,731,638,662]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="506" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB65ED48FDB3FDD161FC84ED" box="[598,731,638,662]" italics="true" pageId="13" pageNumber="506">C. flexuosus</emphasis>
</taxonomicName>
is most similar to that seen in
<taxonomicName id="53B84D69EB65ED48FBD6FDD163E684CD" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="506" phylum="Arthropoda" rank="species" species="sumatrensis">
<emphasis id="A6CCEAF8EB65ED48FBD6FDD163E684CD" italics="true" pageId="13" pageNumber="506">C. sumatrensis</emphasis>
</taxonomicName>
(
<figureCitation id="0C832A6FEB65ED48FF2AFD31623C84CD" box="[207,283,670,694]" captionStart="FIGURE" captionStartId="25.[171,183,1138,1157]" captionTargetBox="[120,1176,216,1118]" captionTargetId="figure-226@25.[120,1176,216,1118]" captionTargetPageId="25" captionText="FIGURE 11. Cyclosomus sumatrensis Bouchard, dorsal habitus. A. Dark male (Kras, Kediri, East Java, Java, Indonesia); B. Pale male (Taman Negara, Pahang, Malaysia). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512980" httpUri="https://zenodo.org/record/11512980/files/figure.png" pageId="13" pageNumber="506">Fig. 11</figureCitation>
) and some
<taxonomicName id="53B84D69EB65ED48FE73FD31613584CD" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[406,530,670,694]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="13" pageNumber="506" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A6CCEAF8EB65ED48FE73FD31613584CD" box="[406,530,670,694]" italics="true" pageId="13" pageNumber="506">C. suturalis</emphasis>
</taxonomicName>
(
<figureCitation id="0C832A6FEB65ED48FDC5FD3161A784CD" box="[544,640,670,694]" captionStart="FIGURE" captionStartId="28.[171,183,1524,1543]" captionTargetBox="[120,1173,216,1504]" captionTargetId="figure-9@28.[120,1173,216,1504]" captionTargetPageId="28" captionText="FIGURE 12. Cyclosomus suturalis (Wiedemann). A. Scolytus suturalis Wiedemann, lectotype male, dorsal habitus; B. Labels associated with lectotype; C. Dark male ((Longchuan Jiang at Nongkan village, Zhefang Township, Mangshi, Yunnan, China). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512982" httpUri="https://zenodo.org/record/11512982/files/figure.png" pageId="13" pageNumber="506">Fig. 12C</figureCitation>
) specimens; but members of both of these species have long epipleural setae only on the humeral and subhumeral regions and progressively and distinctly shorter setae to toward the apex. Also, males of these species have the male median lobe less curved in lateral aspect (
<figureCitation id="0C832A6FEB65ED48FE49FD516117856D" box="[428,560,766,790]" captionStart="FIGURE" captionStartId="35.[171,183,1494,1513]" captionTargetBox="[120,1163,216,1464]" captionTargetId="figure-14@35.[120,1176,216,1479]" captionTargetPageId="35" captionText="FIGURE 16. Male genitalia, Cyclosomus spp., left lateral aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512992" httpUri="https://zenodo.org/record/11512992/files/figure.png" pageId="13" pageNumber="506">Figs. 16F, G</figureCitation>
) and the apical lamella of distinctly different shape (compare
<figureCitation id="0C832A6FEB65ED48FF55FCB06229854C" box="[176,270,799,823]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="13" pageNumber="506">Fig. 17B</figureCitation>
with
<figureCitation id="0C832A6FEB65ED48FEA9FCB0613F854C" box="[332,536,799,823]" captionStart="FIGURE" captionStartId="36.[171,183,1563,1582]" captionTargetBox="[120,1164,216,1526]" captionTargetId="figure-9@36.[120,1176,216,1548]" captionTargetPageId="36" captionText="FIGURE 17. Male genitalia, Cyclosomus spp., dorsal aspect. A. C. acutangulus Kavanaugh and Cueva-Dabkoski (Longchuan Jiang at Longjiang Bridge, Wuhe Township, Tengchong County, Yunnan, China); B. C. flexuosus Dejean (Karaikal, Puducherry, India); C. C. inustus Andrewes (“China”); D. C. marginatus Motschulsky (“Bengal”, India); E. C. philippinus Heller (Pili, Zambales Mountains, Luzon, Philippines); F. C. sumatrensis Bouchard (Taman Negara, Pahang, Malaysia); G. C. suturalis (Wiedemann), lectotype (“Bengala”, India); H. C. verspertilio Cueva-Dabkoski and Kavanaugh sp. nov. (New Delhi, Delhi, India). Scale lines = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.11512994" httpUri="https://zenodo.org/record/11512994/files/figure.png" pageId="13" pageNumber="506">Figs. 17F and 17G</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB65ED48FF4EFCEF67788583" pageId="13" pageNumber="506" type="biology_ecology">
<paragraph id="940736EAEB65ED48FF4EFCEF67788583" blockId="13.[123,1174,219,1725]" pageId="13" pageNumber="506">
<emphasis id="A6CCEAF8EB65ED48FF4EFCEF62BE852C" bold="true" box="[171,409,832,855]" pageId="13" pageNumber="506">Habitat distribution.</emphasis>
Not yet reported, but presumed to be restricted to sandy substrates in the vicinity of streams or other water bodies and any associated dune systems, and likely also to occur in such sandy areas along ocean shores. Several localities from coastal areas both on the Indian Subcontinent and in
<collectingCountry id="ECAF767AEB65ED48FE4CFC30613485CC" box="[425,531,927,951]" name="Sri Lanka" pageId="13" pageNumber="506">Sri Lanka</collectingCountry>
likely represent occurrences in coastal sand beach or dune habitats. The apparent elevational range of this species extends from at or near sea level (
<quantity id="53409B0FEB65ED48FBADFC6F675485AC" box="[1096,1139,960,984]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="13" pageNumber="506" unit="m" value="3.0">3 m</quantity>
on Mannar Island,
<collectingCountry id="ECAF767AEB65ED48FEC1FC4F62A88583" box="[292,399,992,1016]" name="Sri Lanka" pageId="13" pageNumber="506">Sri Lanka</collectingCountry>
) to at least
<quantity id="53409B0FEB65ED48FDE9FC4F61768583" box="[524,593,992,1016]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="13" pageNumber="506" unit="m" value="500.0">500 m</quantity>
(near Kurseong in
<collectingRegion id="567CF808EB65ED48FCC5FC4E608D8583" box="[800,938,992,1016]" country="India" name="West Bengal" pageId="13" pageNumber="506">West Bengal</collectingRegion>
, northern
<collectingCountry id="ECAF767AEB65ED48FBFDFC4F67748583" box="[1048,1107,992,1016]" name="India" pageId="13" pageNumber="506">India</collectingCountry>
).
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB65ED4BFF4EFBAE62DE842F" lastPageId="14" lastPageNumber="507" pageId="13" pageNumber="506" type="materials_examined">
<paragraph id="940736EAEB65ED4BFF4EFBAE62DE842F" blockId="13.[123,1174,219,1725]" lastBlockId="14.[123,1174,219,1207]" lastPageId="14" lastPageNumber="507" pageId="13" pageNumber="506">
<emphasis id="A6CCEAF8EB65ED48FF4EFBAE62FB8263" bold="true" box="[171,476,1025,1048]" pageId="13" pageNumber="506">Geographical distribution.</emphasis>
We have examined a total of
<specimenCount id="82BEFD63EB65ED48FCC9FBAF60EB8263" box="[812,972,1024,1048]" count="176" pageId="13" pageNumber="506" type="generic">176 specimens</specimenCount>
(
<specimenCount id="82BEFD63EB65ED48FC3BFBAF67668263" box="[990,1089,1024,1048]" count="90" pageId="13" pageNumber="506" type="male">90 males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FB9EFBAF63F78243" count="86" pageId="13" pageNumber="506" type="female">86 females</specimenCount>
) from the following localities:
<emphasis id="A6CCEAF8EB65ED48FDADFB8E604F8243" bold="true" box="[584,872,1057,1080]" pageId="13" pageNumber="506">
<collectingCountry id="ECAF767AEB65ED48FDADFB8E60268243" box="[584,769,1057,1080]" name="Bangladesh" pageId="13" pageNumber="506">BANGLADESH</collectingCountry>
/
<collectingCountry id="ECAF767AEB65ED48FCFCFB8E604F8243" box="[793,872,1057,1080]" name="India" pageId="13" pageNumber="506">INDIA</collectingCountry>
</emphasis>
: “Bengala” ([
<specimenCount id="82BEFD63EB65ED48FBE9FB8F67B68243" box="[1036,1169,1056,1080]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFBEF63E98223" box="[123,206,1088,1112]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="506">MFNB</collectionCode>
]).
<materialsCitation id="24D03CB7EB65ED48FF0AFBEE603D8223" ID-GBIF-Occurrence="4905152355" box="[239,794,1088,1112]" collectingDate="2006-05-26" collectingDateMax="2006-06-03" collectingDateMin="2006-05-26" collectionCode="IRSNB" collectorName="P. Pacholatko" country="India" elevation="150" location="INDIA" pageId="13" pageNumber="506" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<emphasis id="A6CCEAF8EB65ED48FF0AFBEE62198223" bold="true" box="[239,318,1089,1112]" pageId="13" pageNumber="506">
<collectingCountry id="ECAF767AEB65ED48FF0AFBEE62198223" box="[239,318,1089,1112]" name="India" pageId="13" pageNumber="506">INDIA</collectingCountry>
</emphasis>
([
<specimenCount id="82BEFD63EB65ED48FEBEFBEF62F28223" box="[347,469,1088,1112]" count="2" pageId="13" pageNumber="506" type="male">two males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FDFEFBEE61B98223" box="[539,670,1088,1112]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FD57FBEF602F8223" box="[690,776,1088,1112]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
]):
</materialsCitation>
<materialsCitation id="24D03CB7EB65ED48FCCDFBEF63D282E2" ID-GBIF-Occurrence="4905152334" collectingDate="2006-05-26" collectingDateMax="2006-06-03" collectingDateMin="2006-05-26" collectionCode="IRSNB" collectorName="P. Pacholatko" country="India" elevation="150" location="INDIA" pageId="13" pageNumber="506" specimenCount="3" specimenCount-female="1" specimenCount-male="2" stateProvince="Arunachal Pradesh">
<collectingRegion id="567CF808EB65ED48FCCDFBEF67278223" box="[808,1024,1088,1112]" country="India" name="Arunachal Pradesh" pageId="13" pageNumber="506">Arunachal Pradesh</collectingRegion>
:
<location id="91676031EB65ED48FBF6FBEF67B28223" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FBF6FBEF67B28223" box="[1043,1173,1088,1112]" country="India" name="Bhalukpong" pageId="13" pageNumber="506" stateProvince="Arunachal Pradesh">Bhalukpong</location>
(27.0333°/92.5833°,
<quantity id="53409B0FEB65ED48FE83FBCE62918202" box="[358,438,1121,1145]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="506" unit="m" value="150.0">
<elevation id="1F95D1D9EB65ED48FE83FBCE62918202" box="[358,438,1121,1145]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="13" pageNumber="506" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="E006102AEB65ED48FE29FBCE61EE8202" box="[460,713,1121,1145]" pageId="13" pageNumber="506" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">
<collectingDate id="F042E9C2EB65ED48FE29FBCE61EE8202" box="[460,713,1121,1145]" pageId="13" pageNumber="506" value="2006-05-26" valueMax="2006-06-03" valueMin="2006-05-26">26 May-3 June 2006</collectingDate>
</date>
,
<collectorName id="394D533CEB65ED48FD3BFBCD605E8202" box="[734,889,1121,1145]" pageId="13" pageNumber="506">P. Pacholátko</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FC1AFBCD67B68202" box="[1023,1169,1121,1145]" count="2" pageId="13" pageNumber="506" type="female">two females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFB2E63CF82E2" box="[123,232,1153,1177]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FEEFFB2E61D382E2" ID-GBIF-Occurrence="4905152324" box="[266,756,1153,1177]" collectionCode="NHMUK" country="India" location="Kinara" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="Gujarat">
<collectingRegion id="567CF808EB65ED48FEEFFB2E627882E2" box="[266,351,1153,1177]" country="India" name="Gujarat" pageId="13" pageNumber="506">Gujarat</collectingRegion>
:
<location id="91676031EB65ED48FE97FB2E629C82E2" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FE97FB2E629C82E2" box="[370,443,1153,1177]" country="India" name="Kinara" pageId="13" pageNumber="506" stateProvince="Gujarat">Kinara</location>
([
<specimenCount id="82BEFD63EB65ED48FE3FFB2E614682E2" box="[474,609,1153,1177]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FD93FB2E61C482E2" box="[630,739,1153,1177]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
]);
</materialsCitation>
<materialsCitation id="24D03CB7EB65ED48FCE0FB2E63D382C2" ID-GBIF-Occurrence="4905152358" collectionCode="NHMUK" country="India" location="Kinara" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="Gujarat">
<collectingRegion id="567CF808EB65ED48FCE0FB2E605082E2" box="[773,887,1153,1177]" country="India" name="Karnataka" pageId="13" pageNumber="506">Karnataka</collectingRegion>
:
<location id="91676031EB65ED48FC6EFB2E60CD82E2" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FC6EFB2E60CD82E2" box="[907,1002,1153,1177]" country="India" name="Belagavi" pageId="13" pageNumber="506" stateProvince="Karnataka">Belagavi</location>
([
<specimenCount id="82BEFD63EB65ED48FBEFFB2E67B682E2" box="[1034,1169,1153,1177]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFB0E63C082C2" box="[123,231,1185,1209]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FEE6FB0E61BC82C2" ID-GBIF-Occurrence="4905152321" box="[259,667,1185,1209]" collectionCode="NHMUK" country="India" location="Shivamogga" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="19" specimenCount-male="1" stateProvince="Karnataka">
<location id="91676031EB65ED48FEE6FB0E62AF82C2" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FEE6FB0E62AF82C2" box="[259,392,1185,1209]" country="India" name="Shivamogga" pageId="13" pageNumber="506" stateProvince="Karnataka">Shivamogga</location>
([
<specimenCount id="82BEFD63EB65ED48FE47FB0E612982C2" box="[418,526,1185,1209]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FDF8FB0E61AE82C2" box="[541,649,1185,1209]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
]);
</materialsCitation>
<materialsCitation id="24D03CB7EB65ED48FD43FB0E612582A2" ID-GBIF-Occurrence="4905152319" collectionCode="NHMUK" country="India" location="Shivamogga" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="19" specimenCount-male="1" stateProvince="Karnataka">
<collectingRegion id="567CF808EB65ED48FD43FB0E61D682C2" box="[678,753,1185,1209]" country="India" name="Kerala" pageId="13" pageNumber="506">Kerala</collectingRegion>
:
<location id="91676031EB65ED48FCE5FB0E607E82C2" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FCE5FB0E607E82C2" box="[768,857,1185,1209]" country="India" name="Malabar" pageId="13" pageNumber="506" stateProvince="Kerala">Malabar</location>
([
<specimenCount id="82BEFD63EB65ED48FC96FB0E60FE82C2" box="[883,985,1185,1209]" count="16" pageId="13" pageNumber="506" type="male">16 males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FBF0FB0E67B682C2" box="[1045,1169,1185,1209]" count="17" pageId="13" pageNumber="506" type="female">17 females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFB6E63E882A2" box="[123,207,1217,1241]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
], [
<specimenCount id="82BEFD63EB65ED48FF0EFB6D625F82A2" box="[235,376,1217,1241]" count="2" pageId="13" pageNumber="506" type="female">two females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FE6CFB6E62D282A2" box="[393,501,1217,1241]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FDF6FB6E63F98281" ID-GBIF-Occurrence="4905152359" collectionCode="IRSNB" country="India" location="Barwai" pageId="13" pageNumber="506" specimenCount="4" specimenCount-female="3" specimenCount-male="1" stateProvince="Madhya Pradesh">
<collectingRegion id="567CF808EB65ED48FDF6FB6E61E882A2" box="[531,719,1217,1241]" country="India" name="Madhya Pradesh" pageId="13" pageNumber="506">Madhya Pradesh</collectingRegion>
:
<location id="91676031EB65ED48FD3AFB6E600B82A2" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FD3AFB6E600B82A2" box="[735,812,1217,1241]" country="India" name="Barwai" pageId="13" pageNumber="506" stateProvince="Madhya Pradesh">Barwai</location>
([
<specimenCount id="82BEFD63EB65ED48FCA3FB6E609D82A2" box="[838,954,1217,1241]" count="1" pageId="13" pageNumber="506" type="male">one males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FC1CFB6E67B682A2" box="[1017,1169,1217,1241]" count="3" pageId="13" pageNumber="506" type="female">three females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFB4D63F78281" box="[123,208,1250,1274]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FF0BFB4D61CC8281" ID-GBIF-Occurrence="4905152370" box="[238,747,1250,1274]" collectionCode="NHMUK" country="India" location="Brahmapur" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="Odisha">
<collectingRegion id="567CF808EB65ED48FF0BFB4D62198281" box="[238,318,1250,1274]" country="India" name="Odisha" pageId="13" pageNumber="506">Odisha</collectingRegion>
:
<location id="91676031EB65ED48FEA8FB4D62E28281" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FEA8FB4D62E28281" box="[333,453,1250,1274]" country="India" name="Brahmapur" pageId="13" pageNumber="506" stateProvince="Odisha">Brahmapur</location>
([
<specimenCount id="82BEFD63EB65ED48FE05FB4D61458281" box="[480,610,1250,1274]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FD97FB4D61F98281" box="[626,734,1250,1274]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FD1EFB4D63C78341" ID-GBIF-Occurrence="4905152327" collectingDate="1955-08" collectingDateMax="1962-07" collectingDateMin="1955-08" collectionCode="EMEC, NMNH" collectorName="P. S. Nathan" country="India" location="Karaikal" municipality="Karaikal" pageId="13" pageNumber="506" specimenCount="13" specimenCount-female="4" specimenCount-male="9" stateProvince="Puducherry">
<collectingRegion id="567CF808EB65ED48FD1EFB4D605D8281" box="[763,890,1250,1274]" country="India" name="Puducherry" pageId="13" pageNumber="506">Puducherry</collectingRegion>
:
<collectingMunicipality id="7463AC90EB65ED48FC6FFB4D60C18281" box="[906,998,1250,1274]" pageId="13" pageNumber="506">Karaikal</collectingMunicipality>
(
<location id="91676031EB65ED48FC1CFB4D677E8281" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FC1CFB4D677E8281" box="[1017,1113,1250,1274]" country="India" municipality="Karaikal" name="Karaikal" pageId="13" pageNumber="506" stateProvince="Puducherry">Karaikal</location>
,
<date id="E006102AEB65ED48FB8CFB4D63948361" pageId="13" pageNumber="506" value="1962-07">
<collectingDate id="F042E9C2EB65ED48FB8CFB4D63948361" pageId="13" pageNumber="506" value="1962-07">July 1962</collectingDate>
</date>
,
<collectorName id="394D533CEB65ED48FF21FAAD62628361" box="[196,325,1282,1306]" pageId="13" pageNumber="506">P.S. Nathan</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FE26FAAD611B8361" box="[451,572,1282,1306]" count="8" pageId="13" pageNumber="506" type="male">eight male</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FD99FAAD602C8361" box="[636,779,1282,1306]" count="4" pageId="13" pageNumber="506" type="female">four females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FCF9FAAD604C8361" box="[796,875,1282,1306]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:1013" lsid="urn:lsid:biocol.org:col:1013" name="Essig Museum of Entomology" pageId="13" pageNumber="506" type="University or college">EMEC</collectionCode>
],
<date id="E006102AEB65ED48FC67FAAD67378361" box="[898,1040,1282,1306]" pageId="13" pageNumber="506" value="1955-08">
<collectingDate id="F042E9C2EB65ED48FC67FAAD67378361" box="[898,1040,1282,1306]" pageId="13" pageNumber="506" value="1955-08">August 1955</collectingDate>
</date>
[
<specimenCount id="82BEFD63EB65ED48FBC1FAAC67B68361" box="[1060,1169,1282,1306]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFA8D63F48341" box="[123,211,1314,1338]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="13" pageNumber="506" type="Museum">NMNH</collectionCode>
])
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FF08FA8D61CB8341" ID-GBIF-Occurrence="4905152314" box="[237,748,1314,1338]" collectingDate="1956-07" collectingDateMax="1962-07" collectingDateMin="1955-08" collectionCode="CAS" collectorName="P. S. Nathan" country="India" isTailEnumeration="true" location="Karaikal" municipality="Karaikal" pageId="13" pageNumber="506" specimenCount="23" specimenCount-female="7" specimenCount-male="16" stateProvince="Puducherry">
<date id="E006102AEB65ED48FF08FA8D627F8341" box="[237,344,1314,1338]" pageId="13" pageNumber="506" value="1956-07">
<collectingDate id="F042E9C2EB65ED48FF08FA8D627F8341" box="[237,344,1314,1338]" pageId="13" pageNumber="506" value="1956-07">July 1956</collectingDate>
</date>
[
<specimenCount id="82BEFD63EB65ED48FE8DFA8D62EC8341" box="[360,459,1314,1338]" count="16" pageId="13" pageNumber="506" type="male">16 males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FDE1FA8C61B88341" box="[516,671,1314,1338]" count="7" pageId="13" pageNumber="506" type="female">seven females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FD48FA8D61C18341" box="[685,742,1314,1338]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="13" pageNumber="506" type="Museum">CAS</collectionCode>
]
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FD1CFA8C63D48301" ID-GBIF-Occurrence="4905152311" collectingDate="1931-06" collectingDateMax="1932-05-10" collectingDateMin="1931-06" collectionCode="MFNB, NHMUK" country="India" location="Nedungadu" pageId="13" pageNumber="506" specimenCount="6" specimenCount-female="4" specimenCount-male="2" stateProvince="Puducherry">
[
<specimenCount id="82BEFD63EB65ED48FD1AFA8C60428341" box="[767,869,1314,1338]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FC78FA8C67038341" box="[925,1060,1314,1338]" count="2" pageId="13" pageNumber="506" type="female">two females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FBD4FA8D67A48341" box="[1073,1155,1314,1338]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="506">MFNB</collectionCode>
]), (
<location id="91676031EB65ED48FF67FAED62268321" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FF67FAED62268321" box="[130,257,1346,1370]" country="India" name="Nedungadu" pageId="13" pageNumber="506" stateProvince="Puducherry">Nedungadu</location>
,
<date id="E006102AEB65ED48FEF6FAEC62A08321" box="[275,391,1346,1370]" pageId="13" pageNumber="506" value="1931-06">
<collectingDate id="F042E9C2EB65ED48FEF6FAEC62A08321" box="[275,391,1346,1370]" pageId="13" pageNumber="506" value="1931-06">June 1931</collectingDate>
</date>
[
<specimenCount id="82BEFD63EB65ED48FE7EFAEC61388321" box="[411,543,1346,1370]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FDD5FAEC61BD8321" box="[560,666,1347,1370]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
],
<date id="E006102AEB65ED48FD56FAED60688321" box="[691,847,1346,1371]" pageId="13" pageNumber="506" value="1932-05-10">
<collectingDate id="F042E9C2EB65ED48FD56FAED60688321" box="[691,847,1346,1371]" pageId="13" pageNumber="506" value="1932-05-10">10 May 1932</collectingDate>
</date>
[
<specimenCount id="82BEFD63EB65ED48FC81FAEC60E88321" box="[868,975,1346,1370]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FBF4FAEC67B68321" box="[1041,1169,1346,1370]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFACD63C08301" box="[123,231,1378,1402]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FF1BFACD63F983E0" ID-GBIF-Occurrence="4905152346" collectingDate="1951-07" collectionCode="IRSNB" collectorName="M. Maindron" country="India" location="Malabar Coast" municipality="Mahe" pageId="13" pageNumber="506" specimenCount="15" specimenCount-female="3" specimenCount-male="12" stateProvince="Puducherry">
<collectingMunicipality id="7463AC90EB65ED48FF1BFACD621C8301" box="[254,315,1378,1402]" pageId="13" pageNumber="506">Mahé</collectingMunicipality>
(
<location id="91676031EB65ED48FEACFACD62CE8301" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FEACFACD62CE8301" box="[329,489,1378,1402]" country="India" municipality="Mahe" name="Malabar Coast" pageId="13" pageNumber="506" stateProvince="Puducherry">Malabar Coast</location>
,
<date id="E006102AEB65ED48FE11FACD61788301" box="[500,607,1378,1402]" pageId="13" pageNumber="506" value="1951-07">
<collectingDate id="F042E9C2EB65ED48FE11FACD61788301" box="[500,607,1378,1402]" pageId="13" pageNumber="506" value="1951-07">July 1951</collectingDate>
</date>
,
<collectorName id="394D533CEB65ED48FD8FFACC61DF8301" box="[618,760,1378,1402]" pageId="13" pageNumber="506">M. Maindron</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FC8FFACD60EC8301" box="[874,971,1378,1402]" count="12" pageId="13" pageNumber="506" type="male">12 males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FC1AFACD67B68301" box="[1023,1169,1378,1402]" count="3" pageId="13" pageNumber="506" type="female">three females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EFA2C63F783E0" box="[123,208,1411,1435]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FF0FFA2C62C483C0" ID-GBIF-Occurrence="4905152304" collectingDate="1966-05" collectionCode="MFNB" collectorName="P. S. Nathan" country="India" elevation="1070" location="Cinkona" municipality="Anaimalai Hills" pageId="13" pageNumber="506" specimenCount="5" specimenCount-male="5" stateProvince="Tamil Nadu">
<collectingRegion id="567CF808EB65ED48FF0FFA2C624A83E0" box="[234,365,1411,1435]" country="India" name="Tamil Nadu" pageId="13" pageNumber="506">Tamil Nadu</collectingRegion>
:
<collectingMunicipality id="7463AC90EB65ED48FE92FA2C610583E0" box="[375,546,1411,1435]" pageId="13" pageNumber="506">Anaimalai Hills</collectingMunicipality>
(
<location id="91676031EB65ED48FDD4FA2C61B783E0" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FDD4FA2C61B783E0" box="[561,656,1411,1435]" country="India" municipality="Anaimalai Hills" name="Cinkona" pageId="13" pageNumber="506" stateProvince="Tamil Nadu">Cinkona</location>
,
<quantity id="53409B0FEB65ED48FD79FA2C61D783E1" box="[668,752,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="13" pageNumber="506" unit="m" value="1070.0">
<elevation id="1F95D1D9EB65ED48FD79FA2C61D783E1" box="[668,752,1411,1435]" metricMagnitude="3" metricUnit="m" metricValue="1.0699999999999998" pageId="13" pageNumber="506" unit="m" value="1070.0">1070 m</elevation>
</quantity>
,
<date id="E006102AEB65ED48FD19FA2C604A83E0" box="[764,877,1411,1435]" pageId="13" pageNumber="506" value="1966-05">
<collectingDate id="F042E9C2EB65ED48FD19FA2C604A83E0" box="[764,877,1411,1435]" pageId="13" pageNumber="506" value="1966-05">May 1966</collectingDate>
</date>
,
<collectorName id="394D533CEB65ED48FC9CFA2C60D183E0" box="[889,1014,1411,1435]" pageId="13" pageNumber="506">P.S. Nathan</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FB8CFA2C639E83C0" count="5" pageId="13" pageNumber="506" type="male">five males</specimenCount>
and ten females;
<collectionCode id="F2A9AE2FEB65ED48FE66FA0C62F283C0" box="[387,469,1443,1467]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="506">MFNB</collectionCode>
])
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FE14FA0C672E83C0" ID-GBIF-Occurrence="4905152329" box="[497,1033,1443,1467]" collectionCode="NHMUK" country="India" location="Chennai" pageId="13" pageNumber="506" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Tamil Nadu">
<location id="91676031EB65ED48FE14FA0C616D83C0" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FE14FA0C616D83C0" box="[497,586,1443,1467]" country="India" name="Chennai" pageId="13" pageNumber="506" stateProvince="Tamil Nadu">Chennai</location>
([
<specimenCount id="82BEFD63EB65ED48FD87FA0C61EC83C0" box="[610,715,1443,1467]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FCE0FA0B60A583C0" box="[773,898,1443,1467]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FC75FA0C60DB83C0" box="[912,1020,1443,1467]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FBF2FA0C61C683A0" ID-GBIF-Occurrence="4905152351" collectingDate="1958-10" collectionCode="NHMUK" country="India" location="Coimbatore" pageId="13" pageNumber="506" specimenCount="5" specimenCount-female="2" specimenCount-male="3" stateProvince="Tamil Nadu">
<location id="91676031EB65ED48FBF2FA0C67B283C0" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FBF2FA0C67B283C0" box="[1047,1173,1443,1467]" country="India" name="Coimbatore" pageId="13" pageNumber="506" stateProvince="Tamil Nadu">Coimbatore</location>
(
<date id="E006102AEB65ED48FF67FA6C623283A0" box="[130,277,1475,1499]" pageId="13" pageNumber="506" value="1958-10">
<collectingDate id="F042E9C2EB65ED48FF67FA6C623283A0" box="[130,277,1475,1499]" pageId="13" pageNumber="506" value="1958-10">October 1958</collectingDate>
</date>
[
<specimenCount id="82BEFD63EB65ED48FEC4FA6C62B983A0" box="[289,414,1475,1499]" count="3" pageId="13" pageNumber="506" type="male">three males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FE34FA6B617F83A0" box="[465,600,1475,1499]" count="2" pageId="13" pageNumber="506" type="female">two females</specimenCount>
];
<collectionCode id="F2A9AE2FEB65ED48FD8CFA6C61F283A0" box="[617,725,1475,1499]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FD09FA6C67B683A0" ID-GBIF-Occurrence="4905152325" box="[748,1169,1475,1499]" collectionCode="IRSNB" country="India" location="Ramanathapuram" pageId="13" pageNumber="506" specimenCount="1" specimenCount-male="1" stateProvince="Tamil Nadu">
<location id="91676031EB65ED48FD09FA6C608F83A0" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FD09FA6C608F83A0" box="[748,936,1475,1499]" country="India" name="Ramanathapuram" pageId="13" pageNumber="506" stateProvince="Tamil Nadu">Ramanathapuram</location>
([
<specimenCount id="82BEFD63EB65ED48FC58FA6C670383A0" box="[957,1060,1475,1499]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FBCBFA6C67A483A0" box="[1070,1155,1475,1499]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
])
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FF9EFA4C60FB8380" ID-GBIF-Occurrence="4905152313" box="[123,988,1507,1531]" collectionCode="IRSNB" collectorName="P. du Breuil" country="India" location="Tiruchirappalli" pageId="13" pageNumber="506" specimenCount="5" specimenCount-female="4" specimenCount-male="1" stateProvince="Tamil Nadu">
<location id="91676031EB65ED48FF9EFA4C623D8380" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FF9EFA4C623D8380" box="[123,282,1507,1531]" country="India" name="Tiruchirappalli" pageId="13" pageNumber="506" stateProvince="Tamil Nadu">Tiruchirappalli</location>
(1905,
<collectorName id="394D533CEB65ED48FE8EFA4B62CF8380" box="[363,488,1507,1531]" pageId="13" pageNumber="506">P. du Breuil</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FDBDFA4B619B8380" box="[600,700,1507,1531]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FD0AFA4C605F8380" box="[751,888,1507,1531]" count="4" pageId="13" pageNumber="506" type="female">four females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FC66FA4C60F08380" box="[899,983,1507,1531]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
]
</materialsCitation>
,
<materialsCitation id="24D03CB7EB65ED48FC02FA4B63FC8047" ID-GBIF-Occurrence="4905152366" collectionCode="IRSNB, MFNB" collectorName="R. P. Castets" country="India" pageId="13" pageNumber="506" specimenCount="8" specimenCount-female="5" specimenCount-male="3" stateProvince="Tamil Nadu">
<collectorName id="394D533CEB65ED48FC02FA4B67418380" box="[999,1126,1507,1531]" pageId="13" pageNumber="506">R.P. Castets</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FF22F9AB620A8060" box="[199,301,1539,1563]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FE80F9AB62CB8060" box="[357,492,1539,1563]" count="2" pageId="13" pageNumber="506" type="female">two females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FE1FF9AC61688060" box="[506,591,1539,1563]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
],([
<specimenCount id="82BEFD63EB65ED48FD82F9AC61FE8060" box="[615,729,1539,1563]" count="2" pageId="13" pageNumber="506" type="male">two males</specimenCount>
and
<specimenCount id="82BEFD63EB65ED48FCF4F9AB60BF8060" box="[785,920,1539,1563]" count="2" pageId="13" pageNumber="506" type="female">two females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FC43F9AC60DD8060" box="[934,1018,1539,1563]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
], [
<specimenCount id="82BEFD63EB65ED48FBF6F9AB67B68060" box="[1043,1169,1539,1563]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FF9EF98B63E98047" box="[123,206,1572,1596]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="506">MFNB</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FF03F98B62EF8027" ID-GBIF-Occurrence="4905152322" collectingDate="1918-11" collectionCode="NHMUK" collectorName="H. G. Champion" country="India" location="West Almora" municipality="Kumaon Division" pageId="13" pageNumber="506" specimenCount="2" specimenCount-female="2" stateProvince="Uttarakhand">
<collectingRegion id="567CF808EB65ED48FF03F98B62498047" box="[230,366,1572,1596]" country="India" name="Uttarakhand" pageId="13" pageNumber="506">Uttarakhand</collectingRegion>
:
<collectingMunicipality id="7463AC90EB65ED48FE9CF98A611F8047" box="[377,568,1572,1596]" pageId="13" pageNumber="506">Kumaon Division</collectingMunicipality>
(
<location id="91676031EB65ED48FDA0F98B61F18047" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FDA0F98B61F18047" box="[581,726,1572,1596]" country="India" municipality="Kumaon Division" name="West Almora" pageId="13" pageNumber="506" stateProvince="Uttarakhand">West Almora</location>
,
<date id="E006102AEB65ED48FD04F98B60B68047" box="[737,913,1572,1596]" pageId="13" pageNumber="506" value="1918-11">
<collectingDate id="F042E9C2EB65ED48FD04F98B60B68047" box="[737,913,1572,1596]" pageId="13" pageNumber="506" value="1918-11">November 1918</collectingDate>
</date>
,
<collectorName id="394D533CEB65ED48FC79F98B67618047" box="[924,1094,1572,1596]" pageId="13" pageNumber="506">H.G. Champion</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FF48F9EA621B8027" box="[173,316,1604,1628]" count="2" pageId="13" pageNumber="506" type="female">two females</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FEAAF9EB629C8027" box="[335,443,1604,1628]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FE3EF9EA63F98007" ID-GBIF-Occurrence="4905152373" collectionCode="IRSNB" country="India" location="Kurseong" pageId="13" pageNumber="506" specimenCount="4" specimenCount-female="1" specimenCount-male="3" stateProvince="West Bengal">
<collectingRegion id="567CF808EB65ED48FE3EF9EA61498027" box="[475,622,1604,1628]" country="India" name="West Bengal" pageId="13" pageNumber="506">West Bengal</collectingRegion>
:
<location id="91676031EB65ED48FD9AF9EA61C08026" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FD9AF9EA61C08026" box="[639,743,1605,1629]" country="India" name="Kurseong" pageId="13" pageNumber="506" stateProvince="West Bengal">Kurseong</location>
(1904 [
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and
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;
<collectionCode id="F2A9AE2FEB65ED48FF9EF9CB63F78007" box="[123,208,1636,1660]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FF0EF9CB617D8007" ID-GBIF-Occurrence="4905152374" box="[235,602,1636,1660]" collectionCode="MFNB" country="India" isTailEnumeration="true" location="Ind. Or." pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="West Bengal">
<location id="91676031EB65ED48FF13F9CB626A8007" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FF13F9CB626A8007" box="[246,333,1636,1660]" name="Ind. Or." pageId="13" pageNumber="506">Ind. Or.</location>
” ([
<specimenCount id="82BEFD63EB65ED48FE8BF9CB62CA8007" box="[366,493,1636,1660]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FE1CF9CB616B8007" box="[505,588,1636,1660]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="13" pageNumber="506">MFNB</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FD82F9CB67188007" ID-GBIF-Occurrence="4905152301" box="[615,1087,1636,1660]" collectionCode="IRSNB" country="India" location="Inde Meridionale" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="West Bengal">
<location id="91676031EB65ED48FD95F9CB60088007" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FD95F9CB60088007" box="[624,815,1636,1660]" country="India" name="Inde Meridionale" pageId="13" pageNumber="506" stateProvince="West Bengal">Inde Meridionale</location>
” ([
<specimenCount id="82BEFD63EB65ED48FCB5F9CB60E98007" box="[848,974,1636,1660]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FC3EF9CB67178007" box="[987,1072,1636,1660]" country="Belgium" httpUri="http://grbio.org/cool/c7r2-61q8" name="Institut Royal des Sciences Naturelles de Belgique" pageId="13" pageNumber="506">IRSNB</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB65ED48FBAEF9CB62F280E7" ID-GBIF-Occurrence="4905152363" collectionCode="NHMUK" country="India" location="South India" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="1" stateProvince="West Bengal">
<location id="91676031EB65ED48FBB0F9CB639380E7" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FBB0F9CB639380E7" country="India" name="South India" pageId="13" pageNumber="506" stateProvince="West Bengal">South India</location>
” ([
<specimenCount id="82BEFD63EB65ED48FF36F92B627780E7" box="[211,336,1668,1692]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FEBEF92B62E080E7" box="[347,455,1668,1692]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
.
<materialsCitation id="24D03CB7EB65ED48FE05F92A607780E7" ID-GBIF-Occurrence="4905152309" box="[480,848,1668,1692]" collectionCode="NHMUK" country="Nepal" location="NEPAL" pageId="13" pageNumber="506" specimenCount="1" specimenCount-female="1">
<emphasis id="A6CCEAF8EB65ED48FE05F92A611F80E7" bold="true" box="[480,568,1669,1692]" pageId="13" pageNumber="506">
<collectingCountry id="ECAF767AEB65ED48FE05F92A611F80E7" box="[480,568,1669,1692]" name="Nepal" pageId="13" pageNumber="506">NEPAL</collectingCountry>
</emphasis>
([
<specimenCount id="82BEFD63EB65ED48FDABF92B61EC80E7" box="[590,715,1668,1692]" count="1" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB65ED48FD33F92B606480E7" box="[726,835,1668,1692]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="13" pageNumber="506" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
:
<materialsCitation id="24D03CB7EB65ED4BFCB9F92B623D8688" ID-GBIF-Occurrence="4905152347" collectingDate="1960-09-26" collectionCode="CAS" collectorName="H. Brydon" country="Nepal" elevation="540" lastPageId="14" lastPageNumber="507" location="Hetauda" municipality="Chisapani Garhi District" pageId="13" pageNumber="506" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Bagmati">
<collectingRegion id="567CF808EB65ED48FCB9F92B670780E7" box="[860,1056,1668,1692]" country="Nepal" name="Bagmati" pageId="13" pageNumber="506">Bagmati Province</collectingRegion>
:
<collectingMunicipality id="7463AC90EB65ED48FBCEF92B623480C6" pageId="13" pageNumber="506">Chisapani Garhi District</collectingMunicipality>
(
<location id="91676031EB65ED48FEC3F90A62A480C6" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB65ED48FEC3F90A62A480C6" box="[294,387,1701,1725]" country="Nepal" municipality="Chisapani Garhi District" name="Hetauda" pageId="13" pageNumber="506" stateProvince="Bagmati">Hetauda</location>
,
<quantity id="53409B0FEB65ED48FE77F90A62FA80C7" box="[402,477,1701,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="13" pageNumber="506" unit="m" value="540.0">
<elevation id="1F95D1D9EB65ED48FE77F90A62FA80C7" box="[402,477,1701,1725]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="13" pageNumber="506" unit="m" value="540.0">540 m</elevation>
</quantity>
,
<date id="E006102AEB65ED48FE08F90A61EE80C6" box="[493,713,1701,1725]" pageId="13" pageNumber="506" value="1960-09-26">
<collectingDate id="F042E9C2EB65ED48FE08F90A61EE80C6" box="[493,713,1701,1725]" pageId="13" pageNumber="506" value="1960-09-26">26 September 1960</collectingDate>
</date>
,
<collectorName id="394D533CEB65ED48FD3DF90A606980C6" box="[728,846,1701,1725]" pageId="13" pageNumber="506">H. Brydon</collectorName>
collector [
<specimenCount id="82BEFD63EB65ED48FC2CF90A671680C6" box="[969,1073,1701,1725]" count="1" pageId="13" pageNumber="506" type="male">one male</specimenCount>
and
<specimenCount id="82BEFD63EB65ED4BFB8AF90A63E08688" count="1" lastPageId="14" lastPageNumber="507" pageId="13" pageNumber="506" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFF37FF74622B8688" box="[210,268,219,243]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15690" lsid="urn:lsid:biocol.org:col:34700" name="California Academy of Sciences" pageId="14" pageNumber="507" type="Museum">CAS</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB66ED4BFEC3FF7461648768" ID-GBIF-Occurrence="4905152317" collectingDate="2000-06-28" collectionCode="NHMUK" collectorName="T. W. Harman" country="Nepal" elevation="200" location="Babi River" municipality="Bardiya National Park" pageId="14" pageNumber="507" specimenCount="1" specimenCount-female="1" stateProvince="Lumbini">
<collectingRegion id="567CF808EB66ED4BFEC3FF7462CC8688" box="[294,491,219,243]" country="Nepal" name="Lumbini" pageId="14" pageNumber="507">Lumbini Province</collectingRegion>
:
<collectingMunicipality id="7463AC90EB66ED4BFE13FF7461C28688" box="[502,741,219,243]" pageId="14" pageNumber="507">Bardiya National Park</collectingMunicipality>
(
<location id="91676031EB66ED4BFD16FF74604C8688" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFD16FF74604C8688" box="[755,875,219,243]" country="Nepal" municipality="Bardiya National Park" name="Babi River" pageId="14" pageNumber="507" stateProvince="Lumbini">Babi River</location>
,
<quantity id="53409B0FEB66ED4BFC93FF74609B8689" box="[886,956,219,243]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="507" unit="m" value="200.0">
<elevation id="1F95D1D9EB66ED4BFC93FF74609B8689" box="[886,956,219,243]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="14" pageNumber="507" unit="m" value="200.0">200 m</elevation>
</quantity>
,
<date id="E006102AEB66ED4BFC22FF74677F8688" box="[967,1112,219,243]" pageId="14" pageNumber="507" value="2000-06-28">
<collectingDate id="F042E9C2EB66ED4BFC22FF74677F8688" box="[967,1112,219,243]" pageId="14" pageNumber="507" value="2000-06-28">28 June 2000</collectingDate>
</date>
,
<collectorName id="394D533CEB66ED4BFB86FF7463F68768" pageId="14" pageNumber="507">T.W. Harman</collectorName>
collector [
<specimenCount id="82BEFD63EB66ED4BFEA7FF5362998768" box="[322,446,251,275]" count="1" pageId="14" pageNumber="507" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFE2CFF5461128768" box="[457,565,251,275]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="507" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
.
<materialsCitation id="24D03CB7EB66ED4BFDA8FF53625C8748" ID-GBIF-Occurrence="4905152375" collectionCode="MFNB, NHMUK" country="Sri Lanka" location="SRI LANKA" pageId="14" pageNumber="507" specimenCount="4" specimenCount-female="2" specimenCount-male="2">
<emphasis id="A6CCEAF8EB66ED4BFDA8FF5361C68768" bold="true" box="[589,737,252,275]" pageId="14" pageNumber="507">
<collectingCountry id="ECAF767AEB66ED4BFDA8FF5361C68768" box="[589,737,252,275]" name="Sri Lanka" pageId="14" pageNumber="507">SRI LANKA</collectingCountry>
</emphasis>
([
<specimenCount id="82BEFD63EB66ED4BFD13FF5460548768" box="[758,883,251,275]" count="1" pageId="14" pageNumber="507" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFC9BFF5460F78768" box="[894,976,251,275]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="507">MFNB</collectionCode>
]), ([
<specimenCount id="82BEFD63EB66ED4BFC12FF54674F8768" box="[1015,1128,251,275]" count="2" pageId="14" pageNumber="507" type="male">two males</specimenCount>
and
<specimenCount id="82BEFD63EB66ED4BFF9EFEB363D28748" box="[123,245,283,307]" count="1" pageId="14" pageNumber="507" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFEE4FEB462498748" box="[257,366,283,307]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="507" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
:
<materialsCitation id="24D03CB7EB66ED4BFE62FEB463C587E8" ID-GBIF-Occurrence="4905152365" collectingDate="1970-10-10" collectingDateMax="1977-10-26" collectingDateMin="1970-10-10" collectionCode="A, NMNH" collectorName=" O. S. Flint Jr." country="Sri Lanka" elevation="150" location="Kitulgala" pageId="14" pageNumber="507" specimenCount="3" specimenCount-male="3" stateProvince="Sabaragamuva Province">
<collectingRegion id="567CF808EB66ED4BFE62FEB461A88748" box="[391,655,283,307]" inferredBySuffix="true" pageId="14" pageNumber="507">Sabaragamuva Province</collectingRegion>
:
<location id="91676031EB66ED4BFD7FFEB461DA8748" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFD7FFEB461DA8748" box="[666,765,283,307]" country="Sri Lanka" name="Kitulgala" pageId="14" pageNumber="507" stateProvince="Sabaragamuva Province">Kitulgala</location>
(
<collectorName id="394D533CEB66ED4BFCE8FEB460DF8748" box="[781,1016,283,307]" pageId="14" pageNumber="507">Kitulgala Rest House</collectorName>
,
<quantity id="53409B0FEB66ED4BFBE1FEB4676C8748" box="[1028,1099,283,307]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="507" unit="m" value="150.0">
<elevation id="1F95D1D9EB66ED4BFBE1FEB4676C8748" box="[1028,1099,283,307]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="507" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="E006102AEB66ED4BFBB2FEB462358728" pageId="14" pageNumber="507" value="1977-10-24" valueMax="1977-10-26" valueMin="1977-10-24">
<collectingDate id="F042E9C2EB66ED4BFBB2FEB462358728" pageId="14" pageNumber="507" value="1977-10-24" valueMax="1977-10-26" valueMin="1977-10-24">24-26 October 1977</collectingDate>
</date>
,
<collectorName id="394D533CEB66ED4BFEC5FE9362EE8728" box="[288,457,315,339]" pageId="14" pageNumber="507">K.V. Krombein</collectorName>
,
<collectorName id="394D533CEB66ED4BFE33FE9361458728" box="[470,610,315,339]" pageId="14" pageNumber="507">T. Wijesinhe</collectorName>
,
<collectorName id="394D533CEB66ED4BFD95FE93602D872F" box="[624,778,316,340]" pageId="14" pageNumber="507">M. Jayaweera</collectorName>
, and P.
<collectionCode id="F2A9AE2FEB66ED4BFCBEFE9460578728" box="[859,880,315,339]" pageId="14" pageNumber="507">A</collectionCode>
.
<collectorName id="394D533CEB66ED4BFC98FE9360CC8728" box="[893,1003,316,339]" pageId="14" pageNumber="507">Panawatta</collectorName>
collectors [
<specimenCount id="82BEFD63EB66ED4BFB89FE9363958708" count="1" pageId="14" pageNumber="507" type="male">one male</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFF5AFEF462308708" box="[191,279,347,371]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="507" type="Museum">NMNH</collectionCode>
]),
<collectorName id="394D533CEB66ED4BFED6FEF4629A8708" box="[307,445,347,371]" pageId="14" pageNumber="507">Uggalkaltota</collectorName>
(
<quantity id="53409B0FEB66ED4BFE2BFEF461308708" box="[462,535,347,371]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="507" unit="m" value="150.0">
<elevation id="1F95D1D9EB66ED4BFE2BFEF461308708" box="[462,535,347,371]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="14" pageNumber="507" unit="m" value="150.0">150 m</elevation>
</quantity>
,
<date id="E006102AEB66ED4BFDC0FEF460238708" box="[549,772,347,371]" pageId="14" pageNumber="507" value="1970-10-10" valueMax="1970-10-14" valueMin="1970-10-10">
<collectingDate id="F042E9C2EB66ED4BFDC0FEF460238708" box="[549,772,347,371]" pageId="14" pageNumber="507" value="1970-10-10" valueMax="1970-10-14" valueMin="1970-10-10">10-14 October 1970</collectingDate>
</date>
,
<collectorName id="394D533CEB66ED4BFCF4FEF460588708" box="[785,895,347,371]" pageId="14" pageNumber="507">O.S. Flint</collectorName>
,
<collectorName id="394D533CEB66ED4BFC68FEF360818708" box="[909,934,348,371]" pageId="14" pageNumber="507">Jr.</collectorName>
collector [
<specimenCount id="82BEFD63EB66ED4BFBF9FEF367B58708" box="[1052,1170,347,371]" count="2" pageId="14" pageNumber="507" type="male">two males</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFF9EFED463F387E8" box="[123,212,379,403]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="507" type="Museum">NMNH</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB66ED4BFF09FED4615A87C8" ID-GBIF-Occurrence="4905152302" collectionCode="NMNH" collectorName="W. N. Mathis &amp; T. Wijesinhe &amp; L. Jayawickrema" country="Sri Lanka" location="May" municipality="Polonnaruwa" pageId="14" pageNumber="507" specimenCount="1" specimenCount-female="1" stateProvince="North Central">
<collectingRegion id="567CF808EB66ED4BFF09FED462CE87E8" box="[236,489,379,403]" country="Sri Lanka" name="North Central" pageId="14" pageNumber="507">North Central Province</collectingRegion>
:
<collectingMunicipality id="7463AC90EB66ED4BFE17FED4615887E8" box="[498,639,379,403]" pageId="14" pageNumber="507">Polonnaruwa</collectingMunicipality>
(4
<location id="91676031EB66ED4BFD46FED361F387EF" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFD46FED361F387EF" box="[675,724,380,404]" country="Sri Lanka" municipality="Polonnaruwa" name="May" pageId="14" pageNumber="507" stateProvince="North Central">May</location>
19080,
<collectorName id="394D533CEB66ED4BFCCCFED3609387E8" box="[809,948,379,403]" pageId="14" pageNumber="507">W.N. Mathis</collectorName>
,
<collectorName id="394D533CEB66ED4BFC5BFED3676087E8" box="[958,1095,379,403]" pageId="14" pageNumber="507">T. Wijesinhe</collectorName>
, and
<collectorName id="394D533CEB66ED4BFB9AFED3623387C8" pageId="14" pageNumber="507">L. Jayawickrema</collectorName>
collectors [
<specimenCount id="82BEFD63EB66ED4BFE6AFE33612C87C8" box="[399,523,411,435]" count="1" pageId="14" pageNumber="507" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFDF3FE34614887C8" box="[534,623,411,435]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="507" type="Museum">NMNH</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB66ED4BFD62FE3467B687A8" ID-GBIF-Occurrence="4905152306" collectingDate="1981-04-11" collectingDateMax="1981-04-12" collectingDateMin="1981-04-11" collectionCode="NMNH" collectorName="Krombein &amp; Weeratunga &amp; P. Leanage" country="Sri Lanka" location="Ma Villu" municipality="Kondachchi" pageId="14" pageNumber="507" specimenCount="1" specimenCount-female="1" stateProvince="Northern">
<collectingRegion id="567CF808EB66ED4BFD62FE34607687C8" box="[647,849,411,435]" country="Sri Lanka" name="Northern" pageId="14" pageNumber="507">Northern Province</collectingRegion>
:
<collectingMunicipality id="7463AC90EB66ED4BFCBEFE3460FB87C8" box="[859,988,411,435]" pageId="14" pageNumber="507">Kondachchi</collectingMunicipality>
(
<location id="91676031EB66ED4BFC0FFE34676987C8" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFC0FFE34676987C8" box="[1002,1102,411,435]" country="Sri Lanka" municipality="Kondachchi" name="Ma Villu" pageId="14" pageNumber="507" stateProvince="Northern">Ma Villu</location>
,
<date id="E006102AEB66ED4BFBBDFE3463D087A8" pageId="14" pageNumber="507" value="1981-04-11" valueMax="1981-04-12" valueMin="1981-04-11">
<collectingDate id="F042E9C2EB66ED4BFBBDFE3463D087A8" pageId="14" pageNumber="507" value="1981-04-11" valueMax="1981-04-12" valueMin="1981-04-11">11-12 April 1981</collectingDate>
</date>
, K.V.
<collectorName id="394D533CEB66ED4BFEA7FE14629687A8" box="[322,433,443,467]" pageId="14" pageNumber="507">Krombein</collectorName>
, L.
<collectorName id="394D533CEB66ED4BFE06FE13614287AF" box="[483,613,444,468]" pageId="14" pageNumber="507">Weeratunga</collectorName>
, and
<collectorName id="394D533CEB66ED4BFD43FE13603987AF" box="[678,798,444,468]" pageId="14" pageNumber="507">P. Leanage</collectorName>
collectors [
<specimenCount id="82BEFD63EB66ED4BFC41FE13670287A8" box="[932,1061,443,467]" count="1" pageId="14" pageNumber="507" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFBD1FE1367AC87A8" box="[1076,1163,444,467]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="507" type="Museum">NMNH</collectionCode>
]
</materialsCitation>
,
<materialsCitation id="24D03CB7EB66ED4BFF9EFE7362B4846F" ID-GBIF-Occurrence="4905152348" collectingDate="1970-03-24" collectionCode="NMNH" collectorName="Davis &amp; Rowe" country="Sri Lanka" elevation="3" location="Mannar Island" pageId="14" pageNumber="507" specimenCount="7" specimenCount-female="1" specimenCount-male="6" stateProvince="Northern">
<location id="91676031EB66ED4BFF9EFE736231878F" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFF9EFE736231878F" box="[123,278,476,500]" country="Sri Lanka" name="Mannar Island" pageId="14" pageNumber="507" stateProvince="Northern">Mannar Island</location>
(
<locationDeviation id="85B52FC8EB66ED4BFEC3FE736121878F" box="[294,518,476,500]" location="Pesalai" pageId="14" pageNumber="507">
<quantity id="53409B0FEB66ED4BFEC3FE736255878F" box="[294,370,476,500]" metricMagnitude="3" metricUnit="m" metricValue="3.2" pageId="14" pageNumber="507" unit="km" value="3.2">3.2 km</quantity>
W of Pesalai
</locationDeviation>
,
<quantity id="53409B0FEB66ED4BFDF7FE7361198788" box="[530,574,476,500]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="507" unit="m" value="3.0">
<elevation id="1F95D1D9EB66ED4BFDF7FE7361198788" box="[530,574,476,500]" metricMagnitude="0" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="507" unit="m" value="3.0">3 m</elevation>
</quantity>
,
<date id="E006102AEB66ED4BFDAFFE7361D5878F" box="[586,754,476,500]" pageId="14" pageNumber="507" value="1970-03-24">
<collectingDate id="F042E9C2EB66ED4BFDAFFE7361D5878F" box="[586,754,476,500]" pageId="14" pageNumber="507" value="1970-03-24">24 March 1970</collectingDate>
</date>
,
<collectorName id="394D533CEB66ED4BFD1BFE73601B878F" box="[766,828,476,500]" pageId="14" pageNumber="507">Davis</collectorName>
and
<collectorName id="394D533CEB66ED4BFC97FE7360978788" box="[882,944,476,499]" pageId="14" pageNumber="507">Rowe</collectorName>
collectors [
<specimenCount id="82BEFD63EB66ED4BFBCBFE7367B2878F" box="[1070,1173,476,500]" count="6" pageId="14" pageNumber="507" type="male">six males</specimenCount>
and
<specimenCount id="82BEFD63EB66ED4BFF4DFE536206846F" box="[168,289,508,532]" count="1" pageId="14" pageNumber="507" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFEC9FE5362A2846F" box="[300,389,508,532]" country="USA" httpUri="http://grbio.org/cool/142r-0w94" name="Smithsonian Institution, National Museum of Natural History" pageId="14" pageNumber="507" type="Museum">NMNH</collectionCode>
])
</materialsCitation>
;
<materialsCitation id="24D03CB7EB66ED4BFE78FE5360C7846F" ID-GBIF-Occurrence="4905152328" box="[413,992,508,532]" collectionCode="NHMUK" country="Sri Lanka" location="Colombo" pageId="14" pageNumber="507" specimenCount="1" specimenCount-female="1" stateProvince="Western">
<collectingRegion id="567CF808EB66ED4BFE78FE536179846F" box="[413,606,508,532]" country="Sri Lanka" name="Western" pageId="14" pageNumber="507">Western Province</collectingRegion>
:
<location id="91676031EB66ED4BFD8DFE5361EC846F" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFD8DFE5361EC846F" box="[616,715,508,532]" country="Sri Lanka" name="Colombo" pageId="14" pageNumber="507" stateProvince="Western">Colombo</location>
([
<specimenCount id="82BEFD63EB66ED4BFD05FE53607A846F" box="[736,861,508,532]" count="1" pageId="14" pageNumber="507" type="female">one female</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFC8DFE5360F3846F" box="[872,980,508,532]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="507" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
,
<materialsCitation id="24D03CB7EB66ED4BFC09FE536266844F" ID-GBIF-Occurrence="4905152335" collectionCode="MFNB" country="Sri Lanka" location="Negombo" pageId="14" pageNumber="507" specimenCount="2" specimenCount-female="2" stateProvince="Western">
<location id="91676031EB66ED4BFC09FE536772846F" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFC09FE536772846F" box="[1004,1109,508,532]" country="Sri Lanka" name="Negombo" pageId="14" pageNumber="507" stateProvince="Western">Negombo</location>
([
<specimenCount id="82BEFD63EB66ED4BFB8FFE5363F6844F" count="2" pageId="14" pageNumber="507" type="female">two females</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFF05FDB36214844F" box="[224,307,540,564]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="14" pageNumber="507">MFNB</collectionCode>
])
</materialsCitation>
.
<materialsCitation id="24D03CB7EB66ED4BFEB5FDB363D6842F" ID-GBIF-Occurrence="4905152357" collectingDate="1909-07-05" collectionCode="NHMUK" country="Sri Lanka" location="Specimens" pageId="14" pageNumber="507" specimenCount="1" specimenCount-male="1" stateProvince="Western">
<location id="91676031EB66ED4BFEB5FDB362E3844F" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFEB5FDB362E3844F" box="[336,452,540,564]" country="Sri Lanka" name="Specimens" pageId="14" pageNumber="507" stateProvince="Western">Specimens</location>
without or with illegible locality data: (
<date id="E006102AEB66ED4BFC6BFDB36733844F" box="[910,1044,540,564]" pageId="14" pageNumber="507" value="1909-07-05">
<collectingDate id="F042E9C2EB66ED4BFC6BFDB36733844F" box="[910,1044,540,564]" pageId="14" pageNumber="507" value="1909-07-05">5 July 1909</collectingDate>
</date>
[
<specimenCount id="82BEFD63EB66ED4BFBC3FDB267B6844F" box="[1062,1169,540,564]" count="1" pageId="14" pageNumber="507" type="male">one male</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFF9EFD9363C3842F" box="[123,228,572,596]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="507" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
<materialsCitation id="24D03CB7EB66ED4BFF1DFD9362D3842F" ID-GBIF-Occurrence="4905152369" box="[248,500,572,596]" collectingDate="1909-07-05" collectionCode="NHMUK" country="Sri Lanka" isTailEnumeration="true" location="Specimens" pageId="14" pageNumber="507" specimenCount="1" specimenCount-male="1" stateProvince="Western">
([
<specimenCount id="82BEFD63EB66ED4BFEE2FD936248842F" box="[263,367,572,596]" count="1" pageId="14" pageNumber="507" type="male">one male</specimenCount>
;
<collectionCode id="F2A9AE2FEB66ED4BFE9EFD9362C0842F" box="[379,487,572,596]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="14" pageNumber="507" type="Museum">NHMUK</collectionCode>
])
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB66ED4BFF4EFDF361E4854F" pageId="14" pageNumber="507" type="distribution">
<paragraph id="940736EAEB66ED4BFF4EFDF361E4854F" blockId="14.[123,1174,219,1207]" pageId="14" pageNumber="507">
The known geographical range of
<taxonomicName id="53B84D69EB66ED4BFDC5FDF26184840F" authorityName=", Kavanaugh" authorityYear="2015" box="[544,675,604,628]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB66ED4BFDC5FDF26184840F" box="[544,675,604,628]" italics="true" pageId="14" pageNumber="507">C. flexuosus</emphasis>
</taxonomicName>
(
<figureCitation id="0C832A6FEB66ED4BFD56FDF36025840F" box="[691,770,604,628]" captionStart="FIGURE" captionStartId="39.[171,183,837,856]" captionTargetBox="[168,1128,216,825]" captionTargetId="figure-14@39.[168,1128,216,825]" captionTargetPageId="39" captionText="FIGURE 20. Map showing distribution of locality records for Cyclosomus specimens examined. Yellow circles = C. acutangulus Kavanaugh and Cueva-Dabkoski; white triangles = C. flexuosus Dejean; yellow triangle = locality where C. acutangulus and C. flexuosus have been found together; white squares = C. inustus Andrewes. Modified from Wikimedia Commons, World Atlas of the World, at URL: &lt;http://upload.wikimedia.org/wikipedia/commons/8/8f/ Whole_world_land_and_oceans_12000.jpg&gt;. Scale line = 500 km." figureDoi="http://doi.org/10.5281/zenodo.11513000" httpUri="https://zenodo.org/record/11513000/files/figure.png" pageId="14" pageNumber="507">Fig. 20</figureCitation>
) covers most of the Indian Subcontinent, from the southern base of the
<collectingMunicipality id="7463AC90EB66ED4BFDE4FDD361D784EF" box="[513,752,636,660]" pageId="14" pageNumber="507">Himalayan Mountains</collectingMunicipality>
in the north to
<collectingCountry id="ECAF767AEB66ED4BFC79FDD3672084EF" box="[924,1031,636,660]" name="Sri Lanka" pageId="14" pageNumber="507">Sri Lanka</collectingCountry>
in the south. We have examined specimens from
<collectingRegion id="567CF808EB66ED4BFDFFFD33620B84AF" isEnumeration="true" pageId="14" pageNumber="507">Arunachal Pradesh, Gujarat, Karnataka, Kerala, Madhya Pradesh, Odisha</collectingRegion>
, Puducherry,
<collectingRegion id="567CF808EB66ED4BFE24FD1361F084AF" box="[449,727,700,724]" isEnumeration="true" pageId="14" pageNumber="507">Tamil Nadu, Uttarakhand</collectingRegion>
, and
<collectingRegion id="567CF808EB66ED4BFCF4FD1260BD84AF" box="[785,922,700,724]" country="India" name="West Bengal" pageId="14" pageNumber="507">West Bengal</collectingRegion>
states in
<collectingCountry id="ECAF767AEB66ED4BFBE6FD13671B84AF" box="[1027,1084,700,724]" name="India" pageId="14" pageNumber="507">India</collectingCountry>
, as well as
<collectingCountry id="ECAF767AEB66ED4BFF7EFD7363FA848F" box="[155,221,732,756]" name="Nepal" pageId="14" pageNumber="507">Nepal</collectingCountry>
, and
<collectingCountry id="ECAF767AEB66ED4BFEF9FD7362AB848F" box="[284,396,732,756]" name="Sri Lanka" pageId="14" pageNumber="507">Sri Lanka</collectingCountry>
.
<bibRefCitation id="F0294B1BEB66ED4BFE7CFD736143848F" author="KAVANAUGH, D. H." box="[409,612,732,756]" pageId="14" pageNumber="507" pagination="267 - 298" refId="ref23574" refString="KAVANAUGH, D. H. 2015. A Review of the Genus Cyclosomus Latreille (Coleoptera: Carabidae: Cyclosomini) in the Afrotropical Region. Proceedings of the California Academy of Sciences (Series 4), 62: 267 - 298." type="journal article" year="2015">Kavanaugh (2015)</bibRefCitation>
reported on a disjunct record of this species from “Nubia” (historical name for the area including southern
<collectingCountry id="ECAF767AEB66ED4BFD3DFD52603F856E" box="[728,792,765,789]" name="Egypt" pageId="14" pageNumber="507">Egypt</collectingCountry>
and northern
<collectingCountry id="ECAF767AEB66ED4BFC4EFD5360C9856F" box="[939,1006,764,788]" name="Sudan" pageId="14" pageNumber="507">Sudan</collectingCountry>
centered on the
<location id="91676031EB66ED4BFF9EFCB26211854E" LSID="urn:lsid:plazi:treatment:1C1187FCEB62ED4BFF9EFC0C62DC82CC:91676031EB66ED4BFF9EFCB26211854E" box="[123,310,797,821]" country="Sri Lanka" municipality="Himalayan Mountains" name="Nile River valley" pageId="14" pageNumber="507" stateProvince="Arunachal Pradesh, Gujarat, Karnataka, Kerala, Madhya Pradesh, Odisha">Nile River valley</location>
) and commented on that occurrence.
</paragraph>
</subSubSection>
<subSubSection id="DCA26561EB66ED4BFF4EFC9162DC82CC" pageId="14" pageNumber="507" type="description">
<paragraph id="940736EAEB66ED4BFF4EFC91606685EE" blockId="14.[123,1174,219,1207]" pageId="14" pageNumber="507">
<emphasis id="A6CCEAF8EB66ED4BFF4EFC91629B852E" bold="true" box="[171,444,830,853]" pageId="14" pageNumber="507">Geographical variation.</emphasis>
Although there is evident individual variation (
<figureCitation id="0C832A6FEB66ED4BFC20FC926772852E" box="[965,1109,829,853]" captionStart="FIGURE" captionStartId="11.[171,183,1577,1596]" captionTargetBox="[203,1083,216,1560]" captionTargetId="figure-14@11.[203,1093,216,1560]" captionTargetPageId="11" captionText="FIGURE 5. Cyclosomus flexuosus Fabricius. A. Holotype male, dorsal habitus; B. Labels associated with C. flexuosus holotype; C. Cyclosomus dytiscoides Nietner, lectotype female, dorsal habitus; D. Labels associated with C. dytiscoides lectotype. Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512964" httpUri="https://zenodo.org/record/11512964/files/figure.png" pageId="14" pageNumber="507">Figs. 5A, 5C</figureCitation>
,
<figureCitation id="0C832A6FEB66ED4BFB87FC926757852E" box="[1122,1136,829,853]" captionStart="FIGURE" captionStartId="12.[171,183,1143,1162]" captionTargetBox="[120,1176,216,1117]" captionTargetId="figure-224@12.[120,1176,216,1127]" captionTargetPageId="12" captionText="FIGURE 6. Cyclosomus flexuosus Fabricius, dorsal habitus. A. Dark male (Cinkona, Anaimalai Hills, Tamil Nadu, India); B. Pale female (Tiruchirappalli, Tamil Nadu, India). Scale lines = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.11512968" httpUri="https://zenodo.org/record/11512968/files/figure.png" pageId="14" pageNumber="507">6</figureCitation>
) in development of the dark color pattern of the elytra throughout the range of this species, we could not discern any particular geographic component to that variation.
</paragraph>
<paragraph id="940736EAEB66ED4BFF4EFC3162DC82CC" blockId="14.[123,1174,219,1207]" pageId="14" pageNumber="507">
<emphasis id="A6CCEAF8EB66ED4BFF4EFC31607585CE" bold="true" box="[171,850,925,949]" pageId="14" pageNumber="507">
Geographical relationships with other
<taxonomicName id="53B84D69EB66ED4BFD8BFC32607585CE" box="[622,850,925,949]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="A6CCEAF8EB66ED4BFD8BFC3261D585CE" bold="true" box="[622,754,925,949]" italics="true" pageId="14" pageNumber="507">Cyclosomus</emphasis>
species.
</taxonomicName>
</emphasis>
This species has been found synotopic only with
<taxonomicName id="53B84D69EB66ED4BFEB2FC1162DE85AE" box="[343,505,957,981]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="acutangulus">
<emphasis id="A6CCEAF8EB66ED4BFEB2FC1162DE85AE" box="[343,505,957,981]" italics="true" pageId="14" pageNumber="507">C. acutangulus</emphasis>
</taxonomicName>
(in
<collectingRegion id="567CF808EB66ED4BFDC1FC1261D685AE" box="[548,753,957,981]" country="India" name="Arunachal Pradesh" pageId="14" pageNumber="507">Arunachal Pradesh</collectingRegion>
,
<collectingCountry id="ECAF767AEB66ED4BFD18FC12601285AE" box="[765,821,957,981]" name="India" pageId="14" pageNumber="507">India</collectingCountry>
) at the northwesternmost known locality for the latter species. The range of
<taxonomicName id="53B84D69EB66ED4BFDB5FC7161F4858E" authorityName=", Kavanaugh" authorityYear="2015" box="[592,723,989,1013]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB66ED4BFDB5FC7161F4858E" box="[592,723,989,1013]" italics="true" pageId="14" pageNumber="507">C. flexuosus</emphasis>
</taxonomicName>
overlaps that of
<taxonomicName id="53B84D69EB66ED4BFC69FC716739858E" authority="Kavanaugh &amp; Cueva-Dabkoski, 2023" authorityName="Kavanaugh &amp; Cueva-Dabkoski" authorityYear="2023" box="[908,1054,989,1013]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="vespertilio" status="sp. nov.">
<emphasis id="A6CCEAF8EB66ED4BFC69FC716739858E" box="[908,1054,989,1013]" italics="true" pageId="14" pageNumber="507">C. vespertilio</emphasis>
</taxonomicName>
<taxonomicNameLabel id="BDFF5783EB66ED4BFBC0FC71675E858E" box="[1061,1145,990,1013]" pageId="14" pageNumber="507" rank="species">sp. nov.</taxonomicNameLabel>
in northern
<collectingCountry id="ECAF767AEB66ED4BFF38FC526233826E" box="[221,276,1021,1045]" name="India" pageId="14" pageNumber="507">India</collectingCountry>
and
<collectingCountry id="ECAF767AEB66ED4BFEAEFC5262AA826E" box="[331,397,1021,1045]" name="Nepal" pageId="14" pageNumber="507">Nepal</collectingCountry>
, but these two species have not yet been found together. Known localities for
<taxonomicName id="53B84D69EB66ED4BFF29FBB16269824E" authorityName=", Kavanaugh" authorityYear="2015" box="[204,334,1053,1077]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB66ED4BFF29FBB16269824E" box="[204,334,1053,1077]" italics="true" pageId="14" pageNumber="507">C. flexuosus</emphasis>
</taxonomicName>
in northern West Bengal and
<taxonomicName id="53B84D69EB66ED4BFD6FFBB16021824E" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[650,774,1053,1077]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A6CCEAF8EB66ED4BFD6FFBB16021824E" box="[650,774,1053,1077]" italics="true" pageId="14" pageNumber="507">C. suturalis</emphasis>
</taxonomicName>
in southernmost Sikkim are less than
<quantity id="53409B0FEB66ED4BFF9EFB916398822D" box="[123,191,1086,1110]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="14" pageNumber="507" unit="km" value="30.0">30 km</quantity>
apart, so at least some overlap in their ranges is likely. Because of uncertainty with respect to the respective geographical ranges of
<taxonomicName id="53B84D69EB66ED4BFDC2FBF161AC820D" authorityName="Andrewes" authorityYear="1924" box="[551,651,1118,1142]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="inustis">
<emphasis id="A6CCEAF8EB66ED4BFDC2FBF161AC820D" box="[551,651,1118,1142]" italics="true" pageId="14" pageNumber="507">C. inustis</emphasis>
</taxonomicName>
,
<emphasis id="A6CCEAF8EB66ED4BFD7DFBF16008820D" box="[664,815,1118,1142]" italics="true" pageId="14" pageNumber="507">
C,
<taxonomicName id="53B84D69EB66ED4BFD53FBF06008820D" box="[694,815,1119,1142]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="marginatus">marginatus</taxonomicName>
</emphasis>
, and
<taxonomicName id="53B84D69EB66ED4BFC8CFBF160C3820D" baseAuthorityName="Wiedemann" baseAuthorityYear="1819" box="[873,996,1118,1142]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="suturalis">
<emphasis id="A6CCEAF8EB66ED4BFC8CFBF160C3820D" box="[873,996,1118,1142]" italics="true" pageId="14" pageNumber="507">C. suturalis</emphasis>
</taxonomicName>
(see discussions below), it is possible but not yet confirmed that one or more of these overlap with
<taxonomicName id="53B84D69EB66ED4BFC12FBD0675D82ED" authorityName=", Kavanaugh" authorityYear="2015" box="[1015,1146,1150,1174]" class="Insecta" family="Carabidae" genus="Cyclosomus" kingdom="Animalia" order="Coleoptera" pageId="14" pageNumber="507" phylum="Arthropoda" rank="species" species="flexuosus">
<emphasis id="A6CCEAF8EB66ED4BFC12FBD0675D82ED" box="[1015,1146,1150,1174]" italics="true" pageId="14" pageNumber="507">C. flexuosus</emphasis>
</taxonomicName>
at least in the eastern part of its range.
</paragraph>
</subSubSection>
</treatment>
</document>