treatments-xml/data/03/C3/D1/03C3D14BFFCA507CFC72350F97315664.xml

<document ID-DOI="10.1590/S1679-62252005000400006" ID-ISSN="1982-0224" ID-Zenodo-Dep="5417869" LSID="urn:lsid:zoobank.org:act:CDBCD8DB-5A2E-4027-B802-982E712A4BBC" approvalRequired="2" approvalRequired_for_textStreams="2" checkinTime="1630367863910" checkinUser="carolina" docAuthor="Ribeiro, Alexandre C., Carvalho, Murilo &amp; Melo, Alex L. A." docDate="2005" docId="03C3D14BFFCA507CFC72350F97315664" docLanguage="en" docName="NeotropIchthyol.2005.35.pdf" docOrigin="Neotropical Ichthyology 3 (4)" docSource="http://www.scielo.br/scielo.php?script=sci_arttext&amp;pid=S1679-62252005000400006&amp;lng=en&amp;tlng=en" docStyle="DocumentStyle:85A40B78B715C4497F837D9D78407CE5.4:NeotropIchthyol.2003-2013.journal_article" docStyleId="85A40B78B715C4497F837D9D78407CE5" docStyleName="NeotropIchthyol.2003-2013.journal_article" docStyleVersion="4" docTitle="Otothyropsis marapoama Ribeiro &amp; Carvalho &amp; Melo 2005, new species" docType="treatment" docUuid="CDBCD8DB-5A2E-4027-B802-982E712A4BBC" docUuidSource="ZooBank" docVersion="3" lastPageId="5" lastPageNumber="494" masterDocId="FFFAA933FFC85079FFC6372F95425306" masterDocTitle="Description and relationships of Otothyropsis marapoama, a new genus and species of Hypoptopomatine catfish (Siluriformes: Loricariidae) from rio Tietê basin, southeastern Brazil" masterLastPageNumber="498" masterPageNumber="489" pageId="2" pageNumber="491" updateTime="1630680565616" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title>Description and relationships of Otothyropsis marapoama, a new genus and species of Hypoptopomatine catfish (Siluriformes: Loricariidae) from rio Tietê basin, southeastern Brazil</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Ribeiro, Alexandre C.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Carvalho, Murilo</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Melo, Alex L. A.</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
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<mods:titleInfo>
<mods:title>Neotropical Ichthyology</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2005</mods:date>
<mods:detail type="pubDate">
<mods:number>2005-12-31</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>3</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
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<mods:identifier type="DOI">10.1590/S1679-62252005000400006</mods:identifier>
<mods:identifier type="ISSN">1982-0224</mods:identifier>
<mods:identifier type="Zenodo-Dep">5417869</mods:identifier>
<mods:identifier type="ZooBank">6A56E88A-127D-48CD-89EC-33BDE38A037C</mods:identifier>
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<treatment LSID="urn:lsid:zoobank.org:act:CDBCD8DB-5A2E-4027-B802-982E712A4BBC" httpUri="http://treatment.plazi.org/id/03C3D14BFFCA507CFC72350F97315664" lastPageId="5" lastPageNumber="494" pageId="2" pageNumber="491">
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<paragraph blockId="2.[948,1372,544,600]" pageId="2" pageNumber="491">
<emphasis bold="true" box="[948,1372,544,568]" pageId="2" pageNumber="491">
<taxonomicName authority="Ribeiro &amp; Carvalho &amp; Melo, 2005" authorityName="Ribeiro &amp; Carvalho &amp; Melo" authorityYear="2005" box="[948,1228,544,568]" class="Actinopterygii" family="Loricariidae" genus="Otothyropsis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="491" phylum="Chordata" rank="species" species="marapoama" status="new species">
<emphasis bold="true" box="[948,1228,544,568]" italics="true" pageId="2" pageNumber="491">Otothyropsis marapoama</emphasis>
</taxonomicName>
, 
<taxonomicNameLabel box="[1241,1372,545,568]" pageId="2" pageNumber="491" rank="species">new species</taxonomicNameLabel>
</emphasis>
<figureCitation box="[1134,1186,577,600]" captionStart="Fig" captionStartId="1.[151,189,981,1004]" captionTargetBox="[359,1266,192,964]" captionTargetId="figure-506@1.[359,1266,191,964]" captionTargetPageId="1" captionText="Fig. 1. Otothyropsis marapoama, holotype, LIRP 5640, 38.5 mm SL. Brazil, São Paulo, rio Tietê basin, Município de Marapoama, ribeirãoCubatão,off theroad between Marapoaraand Elisiário(21º11’35&quot;S 49º07’22&quot;W).Dorsal(a), lateral (b), andventral (c)views." figureDoi="http://doi.org/10.5281/zenodo.5417871" httpUri="https://zenodo.org/record/5417871/files/figure.png" pageId="2" pageNumber="491">Fig.1</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="2" pageNumber="491" type="materials_examined">
<paragraph blockId="2.[839,1482,641,1079]" pageId="2" pageNumber="491">
<materialsCitation collectingDate="2003-02-10" collectionCode="LIRP" collectorName="de Marapoama &amp; A. L. A. Melo." country="Brazil" county="rio Tiete" latitude="-21.193054" location="Cubatao" longLatPrecision="21" longitude="-49.122776" municipality="Municipio de Marapoama" pageId="2" pageNumber="491" specimenCode="LIRP 5640" specimenCount="1" stateProvince="Sao Paulo" typeStatus="holotype">
<emphasis bold="true" box="[839,948,641,664]" pageId="2" pageNumber="491">
<typeStatus box="[839,943,641,664]" pageId="2" pageNumber="491">Holotype</typeStatus>
:
</emphasis>
<specimenCode box="[952,1066,641,664]" collectionCode="LIRP" country="Brazil" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="2" pageNumber="491">LIRP 5640</specimenCode>
, 
<quantity box="[1075,1165,641,664]" metricMagnitude="-2" metricUnit="m" metricValue="3.85" pageId="2" pageNumber="491" unit="mm" value="38.5">38.5 mm</quantity>
SL, 
<collectingCountry box="[1210,1275,641,664]" name="Brazil" pageId="2" pageNumber="491">Brazil</collectingCountry>
, 
<collectingRegion box="[1283,1387,641,664]" country="Brazil" name="Sao Paulo" pageId="2" pageNumber="491">São Paulo</collectingRegion>
, 
<collectingCounty box="[1396,1481,641,664]" pageId="2" pageNumber="491">rio Tietê</collectingCounty>
basin, 
<collectingMunicipality box="[908,1189,673,696]" pageId="2" pageNumber="491">
Município 
<collectorName box="[1027,1189,673,696]" pageId="2" pageNumber="491">de Marapoama</collectorName>
</collectingMunicipality>
, ribeirão 
<location LSID="urn:lsid:plazi:treatment:03C3D14BFFCA507CFC72350F97315664:8EB53686FFCA507BFACC358E902551BE" box="[1290,1383,673,696]" country="Brazil" county="rio Tiete" latitude="-21.193054" longLatPrecision="21" longitude="-49.122776" municipality="Municipio de Marapoama" name="Cubatao" pageId="2" pageNumber="491" stateProvince="Sao Paulo">Cubatão</location>
, road 
<locationDeviation firstLocation="Marapoama" pageId="2" pageNumber="491" secondLocation="Elisiario">between Marapoama and Elisiário</locationDeviation>
(
<geoCoordinate box="[1205,1329,705,728]" degrees="21" direction="south" minutes="11" orientation="latitude" pageId="2" pageNumber="491" precision="15" seconds="35" value="-21.193054">21º11’35&quot;S</geoCoordinate>
<geoCoordinate box="[1338,1473,705,728]" degrees="49" direction="west" minutes="07" orientation="longitude" pageId="2" pageNumber="491" precision="15" seconds="22" value="-49.122776">49º07’22&quot;W</geoCoordinate>
) (
<figureCitation box="[847,905,737,760]" captionStart="Fig" captionStartId="2.[839,881,1858,1881]" captionTargetBox="[839,1480,1372,1837]" captionTargetId="figure-730@2.[839,1480,1372,1837]" captionTargetPageId="2" captionText="Fig. 2. Southeastern Brazil showing type-locality of Otothyropsis marapoama in the middle stretch of the rio Tietê basin (1: rio Tietê; 2: rio Grande; 3: rio Paraná; 4: rio Paranapanema; 5: rio Ribeira de Iguape; 6: rio Paraíba do Sul)." figureDoi="http://doi.org/10.5281/zenodo.5417873" httpUri="https://zenodo.org/record/5417873/files/figure.png" pageId="2" pageNumber="491">Fig.2</figureCitation>
), 
<date box="[922,1053,737,760]" pageId="2" pageNumber="491" value="2003-02-10">
<collectingDate box="[922,1053,737,760]" pageId="2" pageNumber="491" value="2003-02-10">10 Feb 2003</collectingDate>
</date>
; 
<collectorName box="[1060,1209,737,760]" pageId="2" pageNumber="491">A. L. A. Melo.</collectorName>
</materialsCitation>
</paragraph>
<paragraph blockId="2.[839,1482,641,1079]" pageId="2" pageNumber="491">
<materialsCitation collectingDate="2003-02-10" collectionCode="LIRP" collectorName="de Marapoama &amp; A. L. A. Melo." country="Brazil" county="rio Tiete" latitude="-21.193054" location="Cubatao" longLatPrecision="21" longitude="-49.122776" municipality="Municipio de Marapoama" pageId="2" pageNumber="491" specimenCode="LIRP 4621, 3" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<emphasis bold="true" box="[839,953,769,792]" pageId="2" pageNumber="491">
<typeStatus box="[839,950,769,792]" pageId="2" pageNumber="491">Paratypes</typeStatus>
:
</emphasis>
<specimenCode box="[957,1067,769,792]" collectionCode="LIRP" country="Brazil" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="2" pageNumber="491">LIRP 4621</specimenCode>
, 
<specimenCode box="[1075,1089,769,792]" collectionCode="LIRP" country="Brazil" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="2" pageNumber="491">3</specimenCode>
, 
<quantity box="[1097,1236,769,792]" metricMagnitude="-2" metricUnit="m" metricValue="2.995" metricValueMax="3.2299999999999995" metricValueMin="2.76" pageId="2" pageNumber="491" unit="mm" value="29.95" valueMax="32.3" valueMin="27.6">27.6-32.3 mm</quantity>
SL (1, 
<quantity box="[1302,1390,769,792]" metricMagnitude="-2" metricUnit="m" metricValue="2.76" pageId="2" pageNumber="491" unit="mm" value="27.6">27.6 mm</quantity>
SL c&amp;s), collected with holotype
</materialsCitation>
; 
<materialsCitation collectingDate="2004-07-14" collectionCode="MZUSP" collectorName="Melo &amp; A. C. Ribeiro &amp; Carvalho &amp; A. K. Takako" country="Brazil" pageId="2" pageNumber="491" specimenCode="MZUSP 87892, 1" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCode box="[1100,1264,801,824]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="491">MZUSP 87892</specimenCode>
, 
<specimenCode box="[1273,1286,801,824]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="491">1</specimenCode>
, 
<quantity box="[1298,1392,801,824]" metricMagnitude="-2" metricUnit="m" metricValue="2.86" pageId="2" pageNumber="491" unit="mm" value="28.6">28.6 mm</quantity>
SL, collected at type locality, 
<date box="[1063,1183,833,856]" pageId="2" pageNumber="491" value="2004-07-14">
<collectingDate box="[1063,1183,833,856]" pageId="2" pageNumber="491" value="2004-07-14">14 Jul 2004</collectingDate>
</date>
, A. L.A. 
<collectorName box="[1275,1333,833,856]" pageId="2" pageNumber="491">Melo</collectorName>
, 
<collectorName box="[1339,1477,833,856]" pageId="2" pageNumber="491">A. C. Ribeiro</collectorName>
, M. 
<collectorName box="[873,968,865,888]" pageId="2" pageNumber="491">Carvalho</collectorName>
&amp; 
<collectorName box="[996,1134,865,888]" pageId="2" pageNumber="491">A. K. Takako</collectorName>
</materialsCitation>
; 
<materialsCitation collectionCode="LIRP" country="Brazil" pageId="2" pageNumber="491" specimenCode="LIRP 5641, 20" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCode box="[1142,1256,865,888]" collectionCode="LIRP" country="Brazil" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="2" pageNumber="491">LIRP 5641</specimenCode>
, 
<specimenCode box="[1265,1293,865,888]" collectionCode="LIRP" country="Brazil" httpUri="http://grbio.org/cool/7bc6-1gdv" name="Laboratorio de Ictiologia, Faculdade de Filosofia" pageId="2" pageNumber="491">20</specimenCode>
, 18.0-29.0 mm SL (4, 19.8-29.0 mm SL c&amp;s; 4 stained with alizarin red solution)
</materialsCitation>
; 
<materialsCitation box="[839,1211,929,952]" collectionCode="MZUSP" country="Brazil" pageId="2" pageNumber="491" specimenCode="MZUSP 87893, 10, 18.7" specimenCount="1" stateProvince="Sao Paulo" typeStatus="paratype">
<specimenCode box="[839,994,929,952]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="491">MZUSP 87893</specimenCode>
, 
<specimenCode box="[1001,1028,929,952]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="491">10</specimenCode>
, 
<specimenCode box="[1036,1079,929,952]" collectionCode="MZUSP" country="Brazil" httpUri="http://grbio.org/cool/6yx0-3nm7" name="Museu de Zoologia da Universidade de Sao Paulo" pageId="2" pageNumber="491">18.7</specimenCode>
-23.0 mm SL
</materialsCitation>
, 
<materialsCitation accessionNumber="MCP 38303" collectingDate="2004-07-14" collectionCode="MCP" collectorName="A. L. A. Melo &amp; A. C. Ribeiro &amp; Carvalho &amp; A. K. Takako" country="Brazil" county="Cubatao" latitude="-21.248167" location="SaoPaulo" longLatPrecision="1" longitude="-49.02189" municipality="Sitio Cubatao" pageId="2" pageNumber="491" specimenCode="10, 18.2, 22.8" specimenCount="1" typeStatus="paratype">
<accessionNumber box="[1219,1343,929,952]" httpUri="http://www.ncbi.nlm.nih.gov/protein/MCP38303" pageId="2" pageNumber="491">
<collectionCode box="[1219,1274,929,952]" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="491">MCP</collectionCode>
38303
</accessionNumber>
, 
<specimenCode box="[1350,1377,929,952]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="491">10</specimenCode>
, 
<specimenCode box="[1385,1429,929,952]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="491">18.2</specimenCode>
- 
<specimenCode box="[1436,1481,929,952]" collectionCode="MCP" country="Brazil" httpUri="http://grbio.org/cool/bqgs-f9q8" name="Pontificia Universidade Catolica do Rio Grande do Sul" pageId="2" pageNumber="491">22.8</specimenCode>
mmSLandDZSJRP 7887, 8, 19.3-21.9 mmSL, 
<collectingCountry box="[1308,1370,961,984]" name="Brazil" pageId="2" pageNumber="491">Brazil</collectingCountry>
, 
<location LSID="urn:lsid:plazi:treatment:03C3D14BFFCA507CFC72350F97315664:8EB53686FFCA507BFAA734EE908750DE" box="[1377,1477,961,984]" country="Brazil" county="Cubatao" latitude="-21.248167" longLatPrecision="1" longitude="-49.02189" municipality="Sitio Cubatao" name="SaoPaulo" pageId="2" pageNumber="491">SãoPaulo</location>
, 
<location LSID="urn:lsid:plazi:treatment:03C3D14BFFCA507CFC72350F97315664:8EB53686FFCA507BFC8134CE96EA50FE" box="[839,936,993,1016]" country="Brazil" county="Cubatao" latitude="-21.248167" longLatPrecision="1" longitude="-49.02189" municipality="Sitio Cubatao" name="rio Tiete" pageId="2" pageNumber="491">rio Tietê</location>
basin, 
<location LSID="urn:lsid:plazi:treatment:03C3D14BFFCA507CFC72350F97315664:8EB53686FFCA507BFC3934CE905B50FE" box="[1023,1305,993,1016]" country="Brazil" county="Cubatao" latitude="-21.248167" longLatPrecision="1" longitude="-49.02189" municipality="Sitio Cubatao" name="Municipio de Catanduva" pageId="2" pageNumber="491">Município de Catanduva</location>
, 
<collectingMunicipality box="[1320,1477,993,1016]" pageId="2" pageNumber="491">Sítio Cubatão</collectingMunicipality>
, córrego 
<collectingCounty box="[922,1009,1025,1048]" pageId="2" pageNumber="491">Cubatão</collectingCounty>
(
<geoCoordinate box="[1020,1159,1025,1048]" degrees="21" direction="south" minutes="14" orientation="latitude" pageId="2" pageNumber="491" precision="1" seconds="53.4" value="-21.248167">21º14’53.4&quot;S</geoCoordinate>
<geoCoordinate box="[1163,1314,1025,1048]" degrees="49" direction="west" minutes="01" orientation="longitude" pageId="2" pageNumber="491" precision="1" seconds="18.8" value="-49.02189">49º01’18.8&quot;W</geoCoordinate>
), 
<date box="[1330,1449,1025,1048]" pageId="2" pageNumber="491" value="2004-07-14">
<collectingDate box="[1330,1449,1025,1048]" pageId="2" pageNumber="491" value="2004-07-14">14 Jul 2004</collectingDate>
</date>
, 
<collectorName pageId="2" pageNumber="491">A. L. A. Melo</collectorName>
, 
<collectorName box="[962,1104,1056,1079]" pageId="2" pageNumber="491">A. C. Ribeiro</collectorName>
, M. 
<collectorName box="[1148,1245,1056,1079]" pageId="2" pageNumber="491">Carvalho</collectorName>
&amp; 
<collectorName box="[1275,1417,1056,1079]" pageId="2" pageNumber="491">A. K. Takako</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection box="[839,1109,1120,1143]" pageId="2" pageNumber="491" type="diagnosis">
<paragraph blockId="2.[839,1109,1120,1143]" box="[839,1109,1120,1143]" pageId="2" pageNumber="491">
<emphasis bold="true" box="[839,957,1120,1143]" pageId="2" pageNumber="491">Diagnosis.</emphasis>
As for genus.
</paragraph>
</subSubSection>
<subSubSection lastPageId="5" lastPageNumber="494" pageId="2" pageNumber="491" type="description">
<paragraph blockId="2.[839,1481,1184,1303]" lastBlockId="3.[831,1474,194,665]" lastPageId="3" lastPageNumber="492" pageId="2" pageNumber="491">
<emphasis bold="true" box="[839,972,1184,1207]" pageId="2" pageNumber="491">Description</emphasis>
. Morphometrics and meristics given in 
<tableCitation box="[1393,1481,1184,1207]" captionStart="Table 1" captionStartId="4.[161,224,194,217]" captionText="Table 1. Descriptive morphometrics of Otothyropsis marapoama. Values are given for the holotype (separately) and 14 paratypes. Hol = Holotype; SD = Standard Deviation." pageId="2" pageNumber="491">Tables 1</tableCitation>
and 
<tableCitation box="[882,895,1216,1239]" captionStart="Table 2" captionStartId="6.[159,222,194,217]" captionText="Table 2. Frequency distribution and summary of meristics for Otothyropsis marapoama. Plate counts were made in 8 specimens stained with alizarin red solution. Holotype values are marked with an asterisk. Meristics marked with double asterisks were counted in c&amp;s specimens only." pageId="2" pageNumber="491">2</tableCitation>
respectively. Dorsal profile of body gently arched from snout tip to dorsal-fin origin; slightly concave to straight at dorsal-fin base; straight from end of dorsal-fin base to cau- dal-fin origin. Ventral profile of body straight to slightly concave from snout tip to base of pectoral fin; straight from that point to anal-fin origin, slightly concave at anal-fin base, and then straight from end of anal-fin base to caudal-fin base. Greatest body depth at dorsal-fin origin. Least body depth at caudal peduncle. Greatest body width at opercular or cleithral regions. Trunk gradually tapering to caudal-fin base. Trunk and caudal peduncle rounded (smaller individuals) to slightly square (in the 
<typeStatus box="[295,390,450,473]" pageId="3" pageNumber="492">holotype</typeStatus>
, the larger specimen with 
<quantity box="[657,749,450,473]" metricMagnitude="-2" metricUnit="m" metricValue="3.5799999999999996" pageId="3" pageNumber="492" unit="mm" value="35.8">35.8 mm</quantity>
SL) in cross section, trunk slightly flattened at base of dorsal and pectoral fins and laterally compressed caudally. Body progressively narrowing caudally from cleithrum. Head wide and rounded anteriorly. Interorbital region flat to slightly convex, with latter margin slightly elevated. Snout straight at mid-line and concave in the region anterior to nares in both sides of head. Eye laterally placed. Superior margin of orbit conspicuously elevated, forming a ridge around orbital margin. Region of pterotic-supracleithrum posterior to orbital margin slightly depressed. Lateral wall of pterotic pierced by enlarged fenestrae, which are visible externally (
<figureCitation box="[1184,1247,322,345]" captionStart="Fig" captionStartId="3.[151,190,1954,1977]" captionTargetBox="[331,1285,723,1919]" captionTargetId="figure-300@3.[321,1304,723,1929]" captionTargetPageId="3" captionText="Fig. 3. Skull of Otothyropsis marapoama, paratype, LIRP 5641, 29.0 mm SL, a) dorsal view; b) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5417875" httpUri="https://zenodo.org/record/5417875/files/figure.png" pageId="3" pageNumber="492">Fig. 3</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5417873" ID-Zenodo-Dep="5417873" httpUri="https://zenodo.org/record/5417873/files/figure.png" pageId="2" pageNumber="491" startId="2.[839,881,1858,1881]" targetBox="[839,1480,1372,1837]" targetPageId="2">
<paragraph blockId="2.[839,1482,1858,1977]" pageId="2" pageNumber="491">
<emphasis bold="true" box="[839,920,1858,1881]" pageId="2" pageNumber="491">Fig. 2.</emphasis>
Southeastern Brazil showing type-locality of 
<taxonomicName authorityName="Ribeiro &amp; Carvalho &amp; Melo" authorityYear="2005" box="[839,1108,1889,1913]" class="Actinopterygii" family="Loricariidae" genus="Otothyropsis" kingdom="Animalia" order="Siluriformes" pageId="2" pageNumber="491" phylum="Chordata" rank="species" species="marapoama">
<emphasis box="[839,1108,1889,1913]" italics="true" pageId="2" pageNumber="491">Otothyropsis marapoama</emphasis>
</taxonomicName>
in the middle stretch of the rio Tietê basin (1: rio Tietê; 2: rio Grande; 3: rio Paraná; 4: rio Paranapanema; 5: rio Ribeira de Iguape; 6: rio Paraíba do Sul).
</paragraph>
</caption>
<paragraph blockId="3.[831,1474,194,665]" lastBlockId="4.[158,801,1057,1208]" lastPageId="4" lastPageNumber="493" pageId="3" pageNumber="492">
Body surface without well-developed crests. Body and abdomen entirely covered by plates, except for small ventral surface in front of anus, region overlying opening of swim bladder capsule, and around pectoral, pelvic, and anal-fin insertions. Rostral plates well-developed and projecting ventrally over the ventral margin of snout (
<figureCitation box="[1204,1285,514,537]" captionStart="Fig" captionStartId="3.[151,190,1954,1977]" captionTargetBox="[331,1285,723,1919]" captionTargetId="figure-300@3.[321,1304,723,1929]" captionTargetPageId="3" captionText="Fig. 3. Skull of Otothyropsis marapoama, paratype, LIRP 5641, 29.0 mm SL, a) dorsal view; b) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5417875" httpUri="https://zenodo.org/record/5417875/files/figure.png" pageId="3" pageNumber="492">Fig. 3b</figureCitation>
). Head and body plates covered with pointed odontodes, mostly uniform in size and distribution, not arranged in conspicuous rows. Odontodes from dorsal and ventral surface of snout slightly larger than remaining ones. Odontodes at tip of supraoccipital and compound petrotic-supracleithrum-posttemporal slightly larger than remaining odontodes of head, but not forming conspicuous crests (
<figureCitation box="[272,335,1121,1144]" captionStart="Fig" captionStartId="3.[151,190,1954,1977]" captionTargetBox="[331,1285,723,1919]" captionTargetId="figure-300@3.[321,1304,723,1929]" captionTargetPageId="3" captionText="Fig. 3. Skull of Otothyropsis marapoama, paratype, LIRP 5641, 29.0 mm SL, a) dorsal view; b) lateral view." figureDoi="http://doi.org/10.5281/zenodo.5417875" httpUri="https://zenodo.org/record/5417875/files/figure.png" pageId="4" pageNumber="493">Fig. 3</figureCitation>
). Pectoral girdle almost completely exposed, except for small median area covered by skin (
<figureCitation box="[641,717,1153,1176]" captionStart="Fig" captionStartId="4.[158,197,1923,1946]" captionTargetBox="[312,1315,1260,1894]" captionTargetId="figure-350@4.[312,1321,1251,1902]" captionTargetPageId="4" captionText="Fig. 4. Otothyropsis marapoama, paratype, LIRP 5641, 29.0 mm SL, a) pectoral girdle and abdominal platelets in ventral view, anterior to top; b) basipterygium in dorsal view, anterior to top." figureDoi="http://doi.org/10.5281/zenodo.5417877" httpUri="https://zenodo.org/record/5417877/files/figure.png" pageId="4" pageNumber="493">Fig. 4a</figureCitation>
). Pelvic girdle with solid basipterygium anteriorly (
<figureCitation box="[600,676,1185,1208]" captionStart="Fig" captionStartId="4.[158,197,1923,1946]" captionTargetBox="[312,1315,1260,1894]" captionTargetId="figure-350@4.[312,1321,1251,1902]" captionTargetPageId="4" captionText="Fig. 4. Otothyropsis marapoama, paratype, LIRP 5641, 29.0 mm SL, a) pectoral girdle and abdominal platelets in ventral view, anterior to top; b) basipterygium in dorsal view, anterior to top." figureDoi="http://doi.org/10.5281/zenodo.5417877" httpUri="https://zenodo.org/record/5417877/files/figure.png" pageId="4" pageNumber="493">Fig. 4b</figureCitation>
).
</paragraph>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5417875" ID-Zenodo-Dep="5417875" box="[151,1280,1953,1977]" httpUri="https://zenodo.org/record/5417875/files/figure.png" pageId="3" pageNumber="492" startId="3.[151,190,1954,1977]" targetBox="[331,1285,723,1919]" targetPageId="3">
<paragraph blockId="3.[151,1280,1953,1977]" box="[151,1280,1953,1977]" pageId="3" pageNumber="492">
<emphasis bold="true" box="[151,219,1954,1977]" pageId="3" pageNumber="492">Fig. 3.</emphasis>
Skull of 
<emphasis box="[313,589,1953,1977]" italics="true" pageId="3" pageNumber="492">
<taxonomicName authorityName="Ribeiro &amp; Carvalho &amp; Melo" authorityYear="2005" box="[313,585,1953,1977]" class="Actinopterygii" family="Loricariidae" genus="Otothyropsis" kingdom="Animalia" order="Siluriformes" pageId="3" pageNumber="492" phylum="Chordata" rank="species" species="marapoama">Otothyropsis marapoama</taxonomicName>
,
</emphasis>
paratype, LIRP 5641, 29.0 mm SL, a) dorsal view; b) lateral view.
</paragraph>
</caption>
<caption pageId="4" pageNumber="493" startId="4.[161,224,194,217]">
<paragraph blockId="4.[161,804,193,281]" pageId="4" pageNumber="493">
<emphasis bold="true" box="[161,245,194,217]" pageId="4" pageNumber="493">Table 1</emphasis>
. Descriptive morphometrics of 
<taxonomicName authorityName="Ribeiro &amp; Carvalho &amp; Melo" authorityYear="2005" class="Actinopterygii" family="Loricariidae" genus="Otothyropsis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="493" phylum="Chordata" rank="species" species="marapoama">
<emphasis italics="true" pageId="4" pageNumber="493">Otothyropsis marapoama</emphasis>
</taxonomicName>
. Values are given for the holotype (separately) and 14 paratypes. Hol = Holotype; SD = Standard Deviation.
</paragraph>
</caption>
<paragraph blockId="4.[839,1482,194,569]" pageId="4" pageNumber="493">Tip of adpressed pectoral fin reaching to middle of pelvic fin, with posterior margin slightly rounded. Well-developed slit present posterodorsally to pectoral insertion. Tip of adpressed pelvic fin reaching anus. Pelvic-fin spine smaller than branched rays and supporting medially projected odontodes. Dorsal-fin origin at vertical behind end of pelvicfin base. Dorsal-fin spinelet small, with dorsal-fin locking mechanism non-functional. Adipose fin absent. Ventral caudal-fin lobe slightly longer than dorsal lobe.</paragraph>
<paragraph pageId="4" pageNumber="493">
<table box="[166,793,310,1021]" gridcols="6" gridrows="27" pageId="4" pageNumber="493">
<tr box="[166,793,310,329]" gridrow="0" pageId="4" pageNumber="493">
<td box="[166,419,310,329]" gridcol="0" gridrow="0" pageId="4" pageNumber="493">
<emphasis bold="true" box="[166,261,310,329]" pageId="4" pageNumber="493">Character</emphasis>
</td>
<td box="[447,746,310,329]" colspan="4" colspanRight="3" gridcol="1" gridrow="0" pageId="4" pageNumber="493">
<emphasis bold="true" box="[449,746,310,329]" pageId="4" pageNumber="493">Hol Minimum Maximum Mean</emphasis>
</td>
<td box="[765,793,310,329]" gridcol="5" gridrow="0" pageId="4" pageNumber="493">
<emphasis bold="true" box="[765,793,310,329]" pageId="4" pageNumber="493">SD</emphasis>
</td>
</tr>
<tr box="[166,793,338,357]" gridrow="1" pageId="4" pageNumber="493">
<td box="[166,419,338,357]" gridcol="0" gridrow="1" pageId="4" pageNumber="493">Standard length (mm)</td>
<td box="[447,485,338,357]" gridcol="1" gridrow="1" pageId="4" pageNumber="493">38.5</td>
<td box="[523,561,338,357]" gridcol="2" gridrow="1" pageId="4" pageNumber="493">19.6</td>
<td box="[621,659,338,357]" gridcol="3" gridrow="1" pageId="4" pageNumber="493">38.5</td>
<td box="[700,746,338,357]" gridcol="4" gridrow="1" pageId="4" pageNumber="493">25.6</td>
<td box="[765,793,338,357]" gridcol="5" gridrow="1" pageId="4" pageNumber="493">5.2</td>
</tr>
<tr box="[166,793,365,384]" gridrow="2" pageId="4" pageNumber="493" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td box="[166,419,365,384]" gridcol="0" gridrow="2" pageId="4" pageNumber="493">
<emphasis bold="true" box="[166,419,365,384]" pageId="4" pageNumber="493">Percents of standard length</emphasis>
</td>
</tr>
<tr box="[166,793,391,410]" gridrow="3" pageId="4" pageNumber="493">
<td box="[166,419,391,410]" gridcol="0" gridrow="3" pageId="4" pageNumber="493">Predorsal length</td>
<td box="[447,485,391,410]" gridcol="1" gridrow="3" pageId="4" pageNumber="493">46.0</td>
<td box="[523,561,391,410]" gridcol="2" gridrow="3" pageId="4" pageNumber="493">44.4</td>
<td box="[621,659,391,410]" gridcol="3" gridrow="3" pageId="4" pageNumber="493">50.3</td>
<td box="[700,746,391,410]" gridcol="4" gridrow="3" pageId="4" pageNumber="493">46.8</td>
<td box="[765,793,391,410]" gridcol="5" gridrow="3" pageId="4" pageNumber="493">1.9</td>
</tr>
<tr box="[166,793,417,436]" gridrow="4" pageId="4" pageNumber="493">
<td box="[166,419,417,436]" gridcol="0" gridrow="4" pageId="4" pageNumber="493">Preanal length</td>
<td box="[447,485,417,436]" gridcol="1" gridrow="4" pageId="4" pageNumber="493">61.8</td>
<td box="[523,561,417,436]" gridcol="2" gridrow="4" pageId="4" pageNumber="493">58.4</td>
<td box="[621,659,417,436]" gridcol="3" gridrow="4" pageId="4" pageNumber="493">64.1</td>
<td box="[700,746,417,436]" gridcol="4" gridrow="4" pageId="4" pageNumber="493">61.6</td>
<td box="[765,793,417,436]" gridcol="5" gridrow="4" pageId="4" pageNumber="493">1.8</td>
</tr>
<tr box="[166,793,444,463]" gridrow="5" pageId="4" pageNumber="493">
<td box="[166,419,444,463]" gridcol="0" gridrow="5" pageId="4" pageNumber="493">Head length</td>
<td box="[447,485,444,463]" gridcol="1" gridrow="5" pageId="4" pageNumber="493">32.4</td>
<td box="[523,561,444,463]" gridcol="2" gridrow="5" pageId="4" pageNumber="493">30.2</td>
<td box="[621,659,444,463]" gridcol="3" gridrow="5" pageId="4" pageNumber="493">38.2</td>
<td box="[700,746,444,463]" gridcol="4" gridrow="5" pageId="4" pageNumber="493">35.5</td>
<td box="[765,793,444,463]" gridcol="5" gridrow="5" pageId="4" pageNumber="493">2.3</td>
</tr>
<tr box="[166,793,471,490]" gridrow="6" pageId="4" pageNumber="493">
<td box="[166,419,471,490]" gridcol="0" gridrow="6" pageId="4" pageNumber="493">Cleithral width</td>
<td box="[447,485,471,490]" gridcol="1" gridrow="6" pageId="4" pageNumber="493">23.9</td>
<td box="[523,561,471,490]" gridcol="2" gridrow="6" pageId="4" pageNumber="493">21.8</td>
<td box="[621,659,471,490]" gridcol="3" gridrow="6" pageId="4" pageNumber="493">25.3</td>
<td box="[700,746,471,490]" gridcol="4" gridrow="6" pageId="4" pageNumber="493">23.7</td>
<td box="[765,793,471,490]" gridcol="5" gridrow="6" pageId="4" pageNumber="493">1.0</td>
</tr>
<tr box="[166,793,497,516]" gridrow="7" pageId="4" pageNumber="493">
<td box="[166,419,497,516]" gridcol="0" gridrow="7" pageId="4" pageNumber="493">Dorsal-fin spine length</td>
<td box="[447,485,497,516]" gridcol="1" gridrow="7" pageId="4" pageNumber="493">25.7</td>
<td box="[523,561,497,516]" gridcol="2" gridrow="7" pageId="4" pageNumber="493">22.7</td>
<td box="[621,659,497,516]" gridcol="3" gridrow="7" pageId="4" pageNumber="493">33.3</td>
<td box="[700,746,497,516]" gridcol="4" gridrow="7" pageId="4" pageNumber="493">26.3</td>
<td box="[765,793,497,516]" gridcol="5" gridrow="7" pageId="4" pageNumber="493">2.5</td>
</tr>
<tr box="[166,793,524,543]" gridrow="8" pageId="4" pageNumber="493">
<td box="[166,419,524,543]" gridcol="0" gridrow="8" pageId="4" pageNumber="493">Folded dorsal-fin length</td>
<td box="[447,485,524,543]" gridcol="1" gridrow="8" pageId="4" pageNumber="493">25.7</td>
<td box="[523,561,524,543]" gridcol="2" gridrow="8" pageId="4" pageNumber="493">25.4</td>
<td box="[621,659,524,543]" gridcol="3" gridrow="8" pageId="4" pageNumber="493">33.3</td>
<td box="[700,746,524,543]" gridcol="4" gridrow="8" pageId="4" pageNumber="493">27.7</td>
<td box="[765,793,524,543]" gridcol="5" gridrow="8" pageId="4" pageNumber="493">2.1</td>
</tr>
<tr box="[166,793,551,570]" gridrow="9" pageId="4" pageNumber="493">
<td box="[166,419,551,570]" gridcol="0" gridrow="9" pageId="4" pageNumber="493">Base of dorsal-fin length</td>
<td box="[447,485,551,570]" gridcol="1" gridrow="9" pageId="4" pageNumber="493">15.3</td>
<td box="[523,561,551,570]" gridcol="2" gridrow="9" pageId="4" pageNumber="493">11.2</td>
<td box="[621,659,551,570]" gridcol="3" gridrow="9" pageId="4" pageNumber="493">17.6</td>
<td box="[700,746,551,570]" gridcol="4" gridrow="9" pageId="4" pageNumber="493">14.6</td>
<td box="[765,793,551,570]" gridcol="5" gridrow="9" pageId="4" pageNumber="493">1.8</td>
</tr>
<tr box="[166,793,577,596]" gridrow="10" pageId="4" pageNumber="493">
<td box="[166,419,577,596]" gridcol="0" gridrow="10" pageId="4" pageNumber="493">Thorax length</td>
<td box="[447,485,577,596]" gridcol="1" gridrow="10" pageId="4" pageNumber="493">18.1</td>
<td box="[523,561,577,596]" gridcol="2" gridrow="10" pageId="4" pageNumber="493">14.5</td>
<td box="[621,659,577,596]" gridcol="3" gridrow="10" pageId="4" pageNumber="493">20.7</td>
<td box="[700,746,577,596]" gridcol="4" gridrow="10" pageId="4" pageNumber="493">17.9</td>
<td box="[765,793,577,596]" gridcol="5" gridrow="10" pageId="4" pageNumber="493">1.7</td>
</tr>
<tr box="[166,793,604,623]" gridrow="11" pageId="4" pageNumber="493">
<td box="[166,419,604,623]" gridcol="0" gridrow="11" pageId="4" pageNumber="493">Pectoral-fin spine length</td>
<td box="[447,485,604,623]" gridcol="1" gridrow="11" pageId="4" pageNumber="493">23.5</td>
<td box="[523,561,604,623]" gridcol="2" gridrow="11" pageId="4" pageNumber="493">23.5</td>
<td box="[621,659,604,623]" gridcol="3" gridrow="11" pageId="4" pageNumber="493">31.1</td>
<td box="[700,746,604,623]" gridcol="4" gridrow="11" pageId="4" pageNumber="493">27.6</td>
<td box="[765,793,604,623]" gridcol="5" gridrow="11" pageId="4" pageNumber="493">1.9</td>
</tr>
<tr box="[166,793,630,649]" gridrow="12" pageId="4" pageNumber="493">
<td box="[166,419,630,649]" gridcol="0" gridrow="12" pageId="4" pageNumber="493">Abdomen length</td>
<td box="[447,485,630,649]" gridcol="1" gridrow="12" pageId="4" pageNumber="493">14.1</td>
<td box="[523,561,630,649]" gridcol="2" gridrow="12" pageId="4" pageNumber="493">10.4</td>
<td box="[621,659,630,649]" gridcol="3" gridrow="12" pageId="4" pageNumber="493">14.9</td>
<td box="[700,746,630,649]" gridcol="4" gridrow="12" pageId="4" pageNumber="493">13.4</td>
<td box="[765,793,630,649]" gridcol="5" gridrow="12" pageId="4" pageNumber="493">1.2</td>
</tr>
<tr box="[166,793,657,676]" gridrow="13" pageId="4" pageNumber="493">
<td box="[166,419,657,676]" gridcol="0" gridrow="13" pageId="4" pageNumber="493">Pelvic-fin spine length</td>
<td box="[447,485,657,676]" gridcol="1" gridrow="13" pageId="4" pageNumber="493">12.8</td>
<td box="[523,561,657,676]" gridcol="2" gridrow="13" pageId="4" pageNumber="493">12.8</td>
<td box="[621,659,657,676]" gridcol="3" gridrow="13" pageId="4" pageNumber="493">19.0</td>
<td box="[700,746,657,676]" gridcol="4" gridrow="13" pageId="4" pageNumber="493">16.2</td>
<td box="[765,793,657,676]" gridcol="5" gridrow="13" pageId="4" pageNumber="493">1.8</td>
</tr>
<tr box="[166,793,683,702]" gridrow="14" pageId="4" pageNumber="493">
<td box="[166,419,683,702]" gridcol="0" gridrow="14" pageId="4" pageNumber="493">Postanal length</td>
<td box="[447,485,683,702]" gridcol="1" gridrow="14" pageId="4" pageNumber="493">34.5</td>
<td box="[523,561,683,702]" gridcol="2" gridrow="14" pageId="4" pageNumber="493">32.3</td>
<td box="[621,659,683,702]" gridcol="3" gridrow="14" pageId="4" pageNumber="493">40.3</td>
<td box="[700,746,683,702]" gridcol="4" gridrow="14" pageId="4" pageNumber="493">35.8</td>
<td box="[765,793,683,702]" gridcol="5" gridrow="14" pageId="4" pageNumber="493">2.3</td>
</tr>
<tr box="[166,793,710,729]" gridrow="15" pageId="4" pageNumber="493">
<td box="[166,419,710,729]" gridcol="0" gridrow="15" pageId="4" pageNumber="493">Caudal-peduncle depth</td>
<td box="[447,485,710,729]" gridcol="1" gridrow="15" pageId="4" pageNumber="493">9.2</td>
<td box="[523,561,710,729]" gridcol="2" gridrow="15" pageId="4" pageNumber="493">9.0</td>
<td box="[621,659,710,729]" gridcol="3" gridrow="15" pageId="4" pageNumber="493">11.0</td>
<td box="[700,746,710,729]" gridcol="4" gridrow="15" pageId="4" pageNumber="493">9.8</td>
<td box="[765,793,710,729]" gridcol="5" gridrow="15" pageId="4" pageNumber="493">0.5</td>
</tr>
<tr box="[166,793,737,756]" gridrow="16" pageId="4" pageNumber="493">
<td box="[166,419,737,756]" gridcol="0" gridrow="16" pageId="4" pageNumber="493">Anal width</td>
<td box="[447,485,737,756]" gridcol="1" gridrow="16" pageId="4" pageNumber="493">13.3</td>
<td box="[523,561,737,756]" gridcol="2" gridrow="16" pageId="4" pageNumber="493">9.6</td>
<td box="[621,659,737,756]" gridcol="3" gridrow="16" pageId="4" pageNumber="493">14.0</td>
<td box="[700,746,737,756]" gridcol="4" gridrow="16" pageId="4" pageNumber="493">11.8</td>
<td box="[765,793,737,756]" gridcol="5" gridrow="16" pageId="4" pageNumber="493">1.3</td>
</tr>
<tr box="[166,793,763,782]" gridrow="17" pageId="4" pageNumber="493">
<td box="[166,419,763,782]" gridcol="0" gridrow="17" pageId="4" pageNumber="493">Snout-opercle length</td>
<td box="[447,485,763,782]" gridcol="1" gridrow="17" pageId="4" pageNumber="493">23.5</td>
<td box="[523,561,763,782]" gridcol="2" gridrow="17" pageId="4" pageNumber="493">21.9</td>
<td box="[621,659,763,782]" gridcol="3" gridrow="17" pageId="4" pageNumber="493">26.6</td>
<td box="[700,746,763,782]" gridcol="4" gridrow="17" pageId="4" pageNumber="493">24.1</td>
<td box="[765,793,763,782]" gridcol="5" gridrow="17" pageId="4" pageNumber="493">1.2</td>
</tr>
<tr box="[166,793,790,809]" gridrow="18" pageId="4" pageNumber="493" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td box="[166,419,790,809]" gridcol="0" gridrow="18" pageId="4" pageNumber="493">
<emphasis bold="true" box="[166,381,790,809]" pageId="4" pageNumber="493">Percents of head length</emphasis>
</td>
</tr>
<tr box="[166,793,816,835]" gridrow="19" pageId="4" pageNumber="493">
<td box="[166,419,816,835]" gridcol="0" gridrow="19" pageId="4" pageNumber="493">Head width</td>
<td box="[447,485,816,835]" gridcol="1" gridrow="19" pageId="4" pageNumber="493">72.8</td>
<td box="[523,561,816,835]" gridcol="2" gridrow="19" pageId="4" pageNumber="493">6.7</td>
<td box="[621,659,816,835]" gridcol="3" gridrow="19" pageId="4" pageNumber="493">12.5</td>
<td box="[700,746,816,835]" gridcol="4" gridrow="19" pageId="4" pageNumber="493">9.1</td>
<td box="[765,793,816,835]" gridcol="5" gridrow="19" pageId="4" pageNumber="493">1.6</td>
</tr>
<tr box="[166,793,843,862]" gridrow="20" pageId="4" pageNumber="493">
<td box="[166,419,843,862]" gridcol="0" gridrow="20" pageId="4" pageNumber="493">Head depth</td>
<td box="[447,485,843,862]" gridcol="1" gridrow="20" pageId="4" pageNumber="493">44.2</td>
<td box="[523,561,843,862]" gridcol="2" gridrow="20" pageId="4" pageNumber="493">51.2</td>
<td box="[621,659,843,862]" gridcol="3" gridrow="20" pageId="4" pageNumber="493">79.3</td>
<td box="[700,746,843,862]" gridcol="4" gridrow="20" pageId="4" pageNumber="493">59.7</td>
<td box="[765,793,843,862]" gridcol="5" gridrow="20" pageId="4" pageNumber="493">7.6</td>
</tr>
<tr box="[166,793,869,888]" gridrow="21" pageId="4" pageNumber="493">
<td box="[166,419,869,888]" gridcol="0" gridrow="21" pageId="4" pageNumber="493">Snout length</td>
<td box="[447,485,869,888]" gridcol="1" gridrow="21" pageId="4" pageNumber="493">50.3</td>
<td box="[523,561,869,888]" gridcol="2" gridrow="21" pageId="4" pageNumber="493">40.0</td>
<td box="[621,659,869,888]" gridcol="3" gridrow="21" pageId="4" pageNumber="493">54.8</td>
<td box="[700,746,869,888]" gridcol="4" gridrow="21" pageId="4" pageNumber="493">44.5</td>
<td box="[765,793,869,888]" gridcol="5" gridrow="21" pageId="4" pageNumber="493">3.5</td>
</tr>
<tr box="[166,793,896,915]" gridrow="22" pageId="4" pageNumber="493">
<td box="[166,419,896,915]" gridcol="0" gridrow="22" pageId="4" pageNumber="493">Interorbital length</td>
<td box="[447,485,896,915]" gridcol="1" gridrow="22" pageId="4" pageNumber="493">46.2</td>
<td box="[523,561,896,915]" gridcol="2" gridrow="22" pageId="4" pageNumber="493">46.5</td>
<td box="[621,659,896,915]" gridcol="3" gridrow="22" pageId="4" pageNumber="493">57.2</td>
<td box="[700,746,896,915]" gridcol="4" gridrow="22" pageId="4" pageNumber="493">50.4</td>
<td box="[765,793,896,915]" gridcol="5" gridrow="22" pageId="4" pageNumber="493">3.0</td>
</tr>
<tr box="[166,793,923,942]" gridrow="23" pageId="4" pageNumber="493">
<td box="[166,419,923,942]" gridcol="0" gridrow="23" pageId="4" pageNumber="493">Orbital diameter</td>
<td box="[447,485,923,942]" gridcol="1" gridrow="23" pageId="4" pageNumber="493">15.5</td>
<td box="[523,561,923,942]" gridcol="2" gridrow="23" pageId="4" pageNumber="493">34.6</td>
<td box="[621,659,923,942]" gridcol="3" gridrow="23" pageId="4" pageNumber="493">47.9</td>
<td box="[700,746,923,942]" gridcol="4" gridrow="23" pageId="4" pageNumber="493">39.5</td>
<td box="[765,793,923,942]" gridcol="5" gridrow="23" pageId="4" pageNumber="493">4.0</td>
</tr>
<tr box="[166,793,949,968]" gridrow="24" pageId="4" pageNumber="493">
<td box="[166,419,949,968]" gridcol="0" gridrow="24" pageId="4" pageNumber="493">Barbel length</td>
<td box="[447,485,949,968]" gridcol="1" gridrow="24" pageId="4" pageNumber="493">8.6</td>
<td box="[523,561,949,968]" gridcol="2" gridrow="24" pageId="4" pageNumber="493">13.6</td>
<td box="[621,659,949,968]" gridcol="3" gridrow="24" pageId="4" pageNumber="493">19.9</td>
<td box="[700,746,949,968]" gridcol="4" gridrow="24" pageId="4" pageNumber="493">15.4</td>
<td box="[765,793,949,968]" gridcol="5" gridrow="24" pageId="4" pageNumber="493">1.7</td>
</tr>
<tr box="[166,793,976,995]" gridrow="25" pageId="4" pageNumber="493" rowspan-1="1" rowspan-2="1" rowspan-3="1" rowspan-4="1" rowspan-5="1">
<td box="[166,419,976,995]" gridcol="0" gridrow="25" pageId="4" pageNumber="493">
<emphasis bold="true" box="[166,408,976,995]" pageId="4" pageNumber="493">Percents of cleithral width</emphasis>
</td>
</tr>
<tr box="[166,793,1002,1021]" gridrow="26" pageId="4" pageNumber="493">
<td box="[166,419,1002,1021]" gridcol="0" gridrow="26" pageId="4" pageNumber="493">Anal width</td>
<td box="[447,485,1002,1021]" gridcol="1" gridrow="26" pageId="4" pageNumber="493">55.4</td>
<td box="[523,561,1002,1021]" gridcol="2" gridrow="26" pageId="4" pageNumber="493">43.2</td>
<td box="[621,659,1002,1021]" gridcol="3" gridrow="26" pageId="4" pageNumber="493">57.0</td>
<td box="[700,746,1002,1021]" gridcol="4" gridrow="26" pageId="4" pageNumber="493">49.5</td>
<td box="[765,793,1002,1021]" gridcol="5" gridrow="26" pageId="4" pageNumber="493">4.4</td>
</tr>
</table>
</paragraph>
<paragraph blockId="4.[839,1482,194,569]" pageId="4" pageNumber="493">Lips roundish, papillose. Maxillary barbel present. Teeth slender, bifid, with larger medial cusp bladelike and slightly rounded, and smaller lateral cusp minute and pointed.</paragraph>
<paragraph blockId="4.[839,1482,610,1209]" pageId="4" pageNumber="493">
<emphasis bold="true" box="[839,1017,610,633]" pageId="4" pageNumber="493">Color in alcohol</emphasis>
. Background color of dorsal surface of head and body dark gray to brown. Body, in larger specimens, completely covered by white spots, ranging in shape from vermiculated to ovoid and often coalescing with each other. Smaller specimens more regular gray to brown overall color pattern. On head, dark pigmentation more concentrated between nares, and along lateral head margin. On body, dark pigmentation concentrated at lateral regions. Ventral surface of body mostly unpigmented, with small scattered spots, which are more concentrated around oral disk and pectoralfin insertion.
</paragraph>
<paragraph blockId="4.[839,1482,610,1209]" lastBlockId="5.[831,1474,202,289]" lastPageId="5" lastPageNumber="494" pageId="4" pageNumber="493">
Unbranched rays of dorsal, pectoral, pelvic and anal fins with five to seven, five to eight, two to five, and four to six dark transverse stripes, respectively. Branched rays of each of these fins with series of spots aligned in continuous transverse stripes. Caudal-fin coloration variable, with pattern ranging from striped along whole fin, to single large spot on fin base and scattered chromatophores distally. Ventral lobe of caudal fin usually more heavily pigmented than dorsal lobe. 
<emphasis bold="true" box="[831,958,202,225]" pageId="5" pageNumber="494">Etymology.</emphasis>
The specific epithet 
<taxonomicName authorityName="Ribeiro &amp; Carvalho &amp; Melo" authorityYear="2005" box="[1171,1299,202,225]" class="Actinopterygii" family="Loricariidae" genus="Otothyropsis" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="494" phylum="Chordata" rank="species" species="marapoama">
<emphasis box="[1171,1299,202,225]" italics="true" pageId="5" pageNumber="494">marapoama</emphasis>
</taxonomicName>
is in reference to the county of the city in which the new species was collected. It is treated as a noun in apposition.
</paragraph>
</subSubSection>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5417877" ID-Zenodo-Dep="5417877" httpUri="https://zenodo.org/record/5417877/files/figure.png" pageId="4" pageNumber="493" startId="4.[158,197,1923,1946]" targetBox="[312,1315,1260,1894]" targetPageId="4">
<paragraph blockId="4.[158,1482,1922,1978]" pageId="4" pageNumber="493">
<emphasis bold="true" box="[158,226,1923,1946]" pageId="4" pageNumber="493">Fig. 4.</emphasis>
<emphasis box="[232,508,1922,1946]" italics="true" pageId="4" pageNumber="493">
<taxonomicName authorityName="Ribeiro &amp; Carvalho &amp; Melo" authorityYear="2005" box="[232,504,1922,1946]" class="Actinopterygii" family="Loricariidae" genus="Otothyropsis" kingdom="Animalia" order="Siluriformes" pageId="4" pageNumber="493" phylum="Chordata" rank="species" species="marapoama">Otothyropsis marapoama</taxonomicName>
,
</emphasis>
paratype, LIRP 5641, 29.0 mm SL, a) pectoral girdle and abdominal platelets in ventral view, anterior to top; b) basipterygium in dorsal view, anterior to top.
</paragraph>
</caption>
<caption ID-DOI="http://doi.org/10.5281/zenodo.5417879" ID-Zenodo-Dep="5417879" httpUri="https://zenodo.org/record/5417879/files/figure.png" pageId="5" pageNumber="494" startId="5.[151,190,1008,1031]" targetBox="[159,780,204,970]" targetPageId="5">
<paragraph blockId="5.[151,794,1007,1223]" pageId="5" pageNumber="494">
<emphasis bold="true" box="[151,218,1008,1031]" pageId="5" pageNumber="494">Fig. 5.</emphasis>
<emphasis box="[223,499,1007,1031]" italics="true" pageId="5" pageNumber="494">
<taxonomicName authorityName="Ribeiro &amp; Carvalho &amp; Melo" authorityYear="2005" box="[223,494,1007,1031]" class="Actinopterygii" family="Loricariidae" genus="Otothyropsis" kingdom="Animalia" order="Siluriformes" pageId="5" pageNumber="494" phylum="Chordata" rank="species" species="marapoama">Otothyropsis marapoama</taxonomicName>
,
</emphasis>
a) branchial skeleton in dorsal view, except for upper elements of right side in ventral view, anterior to top, paratype, LIRP 5641, 24.2.mm SL; b) Upper pharyngeal toothplate in lateral view, showing its reduced condition relative to fourth infra-pharyngobranchial, Anterior to left paratype LIRP 4621, 27.6 mm SL (see 
<bibRefCitation author="Schaefer, S" pageId="5" pageNumber="494" pagination="1 - 41" refId="ref6508" refString="Schaefer, S. A. 1991. Phylogenetic analysis of the loricariid subfamily Hypoptopomatinae (Pisces: Siluroidei: Loricariidae), with comments on generic diagnoses and geographic distribution. Zoological Journal of the Linnean Society, 102: 1 - 41." type="journal article" year="1991">Schaefer, 1991</bibRefCitation>
, fig. 5 a, b and c for comparisons).
</paragraph>
</caption>
<subSubSection pageId="5" pageNumber="494" type="distribution">
<paragraph blockId="5.[151,793,1291,1378]" pageId="5" pageNumber="494">
<emphasis bold="true" box="[151,299,1291,1314]" pageId="5" pageNumber="494">Distribution.</emphasis>
Known from two nearby localities in the same stream along the middle stretch of the rio Tietê, a tributary of the rio 
<collectingRegion box="[224,297,1355,1378]" country="Brazil" name="Parana" pageId="5" pageNumber="494">Paraná</collectingRegion>
of southeastern 
<collectingCountry box="[470,536,1355,1378]" name="Brazil" pageId="5" pageNumber="494">Brazil</collectingCountry>
(
<figureCitation box="[549,614,1355,1378]" captionStart="Fig" captionStartId="2.[839,881,1858,1881]" captionTargetBox="[839,1480,1372,1837]" captionTargetId="figure-730@2.[839,1480,1372,1837]" captionTargetPageId="2" captionText="Fig. 2. Southeastern Brazil showing type-locality of Otothyropsis marapoama in the middle stretch of the rio Tietê basin (1: rio Tietê; 2: rio Grande; 3: rio Paraná; 4: rio Paranapanema; 5: rio Ribeira de Iguape; 6: rio Paraíba do Sul)." figureDoi="http://doi.org/10.5281/zenodo.5417873" httpUri="https://zenodo.org/record/5417873/files/figure.png" pageId="5" pageNumber="494">Fig. 2</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>