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<document id="41078035E8A1D5811C095F81126252CD" ID-DOI="10.25674/SO94iSS3iD303" ID-ISSN="2509-9523" ID-Zenodo-Dep="13193962" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722631614709" checkinUser="felipe" docAuthor="Frisch, Johannes &amp; Mainda, Tobias" docDate="2022" docId="039B5575FF9ADF793AB7FD50FBE7FDA3" docLanguage="en" docName="SoilOrg.94.3.139-147.pdf" docOrigin="Soil Organisms 94 (3)" docSource="http://dx.doi.org/10.25674/so94iss3id303" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Scopaeus balkei Frisch &amp; Mainda, 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="146" masterDocId="FFA22D0DFF9EDF7E3A19FFECFFA6FFF1" masterDocTitle="The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species" masterLastPageNumber="147" masterPageNumber="139" pageNumber="143" updateTime="1722679412379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
<mods:mods id="594BA6E97FC48F7D9E2100CA2128E585" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="9E91D37711AB5F43B1FFB8199418F861">
<mods:title id="48FA55FDB99E8A96F120F9C40AD2E9FA">The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species</mods:title>
</mods:titleInfo>
<mods:name id="98D40E5384BFA48AF86A1C4C8E55848C" type="personal">
<mods:role id="A77956843052A910F2109678DAF27CB7">
<mods:roleTerm id="72B2B57F33BF0EC387190EB5786D6E59">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="27C18489D4C3113080A0F3A4D6C4068C">Frisch, Johannes</mods:namePart>
<mods:affiliation id="ED398E1CD84A9346F6ED25288620BC23">MUSeUM füR NatURKUnDe BeRLin, Leibniz InStitUte fOR EvOLUtiOn anD BiODiveRSity Science,</mods:affiliation>
</mods:name>
<mods:name id="7AE018248C8B76E9C0ED14752EB68943" type="personal">
<mods:role id="A7425EB51B4C2BE2F2B048F0039411EF">
<mods:roleTerm id="B2B53FB64D6585AC9DDD66E5099D849E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="7FF1BB162AFD9CD0B356AC65B2394B3B">Mainda, Tobias</mods:namePart>
<mods:affiliation id="1A7AF68F2DFF9DCED97B57C9F77D5426">ZOOLOgiScheS MUSeUM, UniveRSität GReifSWaLD, LOitzeR StRasse 26, 17489 GReifSWaLD, GeRMany</mods:affiliation>
</mods:name>
<mods:typeOfResource id="E382A7090ECAECC946BB92B65DC65413">text</mods:typeOfResource>
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<mods:titleInfo id="1B56B517FA143C1E49EADEEAFA6ABE83">
<mods:title id="18FB1744B0875548751E90DFA8BBD184">Soil Organisms</mods:title>
</mods:titleInfo>
<mods:part id="FA3E86311EE1E7F9FFE89F11DD10D159">
<mods:date id="A66176D57B603C4E589A2174E2876D47">2022</mods:date>
<mods:detail id="7E18CF53C168269DEDFF08A8DED30D3C" type="pubDate">
<mods:number id="4A6D759F1F2CCFE0B4904752674F127F">2022-12-01</mods:number>
</mods:detail>
<mods:detail id="3F9E4A198D8789C866919B377C936514" type="volume">
<mods:number id="4ECB458EE6420CD154FB7A2C20DAD9D9">94</mods:number>
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<mods:detail id="C98C35476F141428AB6D5E1C43A9A6CE" type="issue">
<mods:number id="EAB97B0A7A0748535D900AB8CB88AC10">3</mods:number>
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<mods:start id="8FED16C94579A263383A70AB9A991991">139</mods:start>
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<mods:location id="753CF420D52333D159985069CC2592E5">
<mods:url id="E7A7984AFEA8C39944B0C7C40CBF7C78">http://dx.doi.org/10.25674/so94iss3id303</mods:url>
</mods:location>
<mods:classification id="5805A0BC556E04D364808D685DADDEF9">journal article</mods:classification>
<mods:identifier id="9A0C5422F5E2B6EB4BE8FD2399667594" type="DOI">10.25674/SO94iSS3iD303</mods:identifier>
<mods:identifier id="AD5DE2DB905F1936758BFE4B5258EC37" type="ISSN">2509-9523</mods:identifier>
<mods:identifier id="EBD42D1F39CB48B23DA876C05C66C25F" type="Zenodo-Dep">13193962</mods:identifier>
</mods:mods>
<treatment id="039B5575FF9ADF793AB7FD50FBE7FDA3" LSID="urn:lsid:plazi:treatment:039B5575FF9ADF793AB7FD50FBE7FDA3" httpUri="http://treatment.plazi.org/id/039B5575FF9ADF793AB7FD50FBE7FDA3" lastPageId="7" lastPageNumber="146" pageId="4" pageNumber="143">
<subSubSection id="C328B7E8FF9ADF7A3AB7FD50FE56FD27" box="[174,496,699,726]" pageId="4" pageNumber="143" type="nomenclature">
<paragraph id="8B8DE463FF9ADF7A3AB7FD50FE56FD27" blockId="4.[174,496,699,759]" box="[174,496,699,726]" pageId="4" pageNumber="143">
<heading id="D0C5530FFF9ADF7A3AB7FD50FE56FD27" box="[174,496,699,726]" fontSize="11" level="2" pageId="4" pageNumber="143" reason="5">
<emphasis id="B9463871FF9ADF7A3AB7FD50FE56FD27" bold="true" box="[174,496,699,726]" pageId="4" pageNumber="143">
<taxonomicName id="4C329FE0FF9ADF7A3AB7FD50FED6FD27" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[174,368,699,726]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="balkei" status="sp. nov.">
<emphasis id="B9463871FF9ADF7A3AB7FD50FED6FD27" bold="true" box="[174,368,699,726]" italics="true" pageId="4" pageNumber="143">Scopaeus balkei</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A275850AFF9ADF7A3B61FD50FE56FD27" box="[376,496,700,726]" pageId="4" pageNumber="143" rank="species">spec. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF9ADF7A3AB7FD33FE7CFD06" box="[174,474,735,759]" pageId="4" pageNumber="143" type="description">
<paragraph id="8B8DE463FF9ADF7A3AB7FD33FE7CFD06" blockId="4.[174,496,699,759]" box="[174,474,735,759]" pageId="4" pageNumber="143">
(
<figureCitation id="1309F8E6FF9ADF7A3AACFD33FF5FFD06" box="[181,249,735,759]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="4" pageNumber="143">Figs 3</figureCitation>
,
<figureCitation id="1309F8E6FF9ADF7A3B1CFD33FEB2FD06" box="[261,276,735,759]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="4" pageNumber="143">6</figureCitation>
,
<figureCitation id="1309F8E6FF9ADF7A3B39FD33FEC6FD06" box="[288,352,735,759]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="4" pageNumber="143">11–13</figureCitation>
,
<figureCitation id="1309F8E6FF9ADF7A3B72FD33FE75FD06" box="[363,467,735,759]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="4" pageNumber="143">19, 20, 24</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF9ADF7A3AB7FCCDFCDEFEBA" pageId="4" pageNumber="143" type="materials_examined">
<paragraph id="8B8DE463FF9ADF7A3AB7FCCDFCDEFEBA" blockId="4.[151,764,800,956]" lastBlockId="4.[786,1399,241,956]" pageId="4" pageNumber="143">
<emphasis id="B9463871FF9ADF7A3AB7FCCDFEC8FCC9" bold="true" box="[174,366,800,825]" pageId="4" pageNumber="143">Type specimens</emphasis>
:
<materialsCitation id="3B5AEE3EFF9ADF7A3B93FCCCFE3CFC6A" collectingDate="2019-11-06" collectionCode="ZMBC" collectorName="Bretschneider" country="Indonesia" elevation="300" location="Cyclops Mts" pageId="4" pageNumber="143" specimenCount="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="holotype">
<typeStatus id="54895AC1FF9ADF7A3B93FCCCFE48FCC9" box="[394,494,800,824]" pageId="4" pageNumber="143" type="holotype">Holotype</typeStatus>
<specimenCount id="9D342FEAFF9ADF7A381BFCCDFDB0FCC9" box="[514,534,801,824]" count="1" pageId="4" pageNumber="143" type="male">♂</specimenCount>
,
<collectingCountry id="F325A4F3FF9ADF7A3836FCCCFD3DFCC9" box="[559,667,800,824]" name="Indonesia" pageId="4" pageNumber="143">Indonesia</collectingCountry>
,
<collectingRegion id="49F62A81FF9ADF7A38AAFCCDFD5EFCC9" box="[691,760,801,824]" country="Indonesia" name="Papua Barat" pageId="4" pageNumber="143">Papua</collectingRegion>
,
<location id="8EEDB2B8FF9ADF7A3A8EFCADFE88FCA8" LSID="urn:lsid:plazi:treatment:039B5575FF9ADF793AB7FD50FBE7FDA3:8EEDB2B8FF9ADF7A3A8EFCADFE88FCA8" box="[151,302,833,857]" country="Indonesia" name="Cyclops Mts" pageId="4" pageNumber="143" stateProvince="Papua Barat">Cyclops Mts</location>
(02°33’13’’S, 140°34’34’’E), vegetated sandbank of a small stream,
<quantity id="4CCA4986FF9ADF7A3BE9FC8EFD9BFC8B" box="[496,573,866,890]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="143" unit="m" value="300.0">
<elevation id="001F0350FF9ADF7A3BE9FC8EFD9BFC8B" box="[496,573,866,890]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="143" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FF8CC2A3FF9ADF7A3849FC8EFD65FC8B" box="[592,707,866,890]" pageId="4" pageNumber="143" value="2019-11-06">
<collectingDate id="EFC83B4BFF9ADF7A3849FC8EFD65FC8B" box="[592,707,866,890]" pageId="4" pageNumber="143" value="2019-11-06">11.06.2019</collectingDate>
</date>
, leg.
<collectorName id="26C781B5FF9ADF7A3A8EFC6FFE8BFC6A" box="[151,301,899,923]" pageId="4" pageNumber="143">Bretschneider</collectorName>
(
<collectionCode id="ED237CA6FF9ADF7A3B59FC6FFE34FC6A" box="[320,402,899,923]" pageId="4" pageNumber="143">ZMBC</collectionCode>
)
</materialsCitation>
.
<typeStatus id="54895AC1FF9ADF7A3BB1FC68FDB5FC6D" box="[424,531,900,924]" pageId="4" pageNumber="143" type="paratype">Paratypes</typeStatus>
(
<specimenCount id="9D342FEAFF9ADF7A383EFC6FFD12FC6A" box="[551,692,899,923]" count="5" pageId="4" pageNumber="143" type="generic">5 specimens</specimenCount>
):
<materialsCitation id="3B5AEE3EFF9ADF7A38D3FC6FFE50FC4D" collectingDate="2019-11-06" collectionCode="MFNB" collectorName="Bretschneider" country="Indonesia" elevation="300" location="Cyclops Mts" pageId="4" pageNumber="143" specimenCount="4" specimenCount-male="4" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="9D342FEAFF9ADF7A38D3FC6FFD51FC6A" box="[714,759,899,923]" count="4" pageId="4" pageNumber="143" type="male">4 ♂</specimenCount>
, same data as holotype (
<collectionCode id="ED237CA6FF9ADF7A3B85FC48FE48FC4D" box="[412,494,932,956]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="4" pageNumber="143">MFNB</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B5AEE3EFF9ADF7A381AFC48FCD2FEBA" collectingDate="2019-05-28" collectionCode="MFNB" collectorName="Bretschneider" country="Indonesia" elevation="300" location="Foja Mts" pageId="4" pageNumber="143" specimenCount="1" specimenCount-male="1" stateProvince="Papua Barat" typeStatus="paratype">
<specimenCount id="9D342FEAFF9ADF7A381AFC48FD8BFC4A" box="[515,557,932,956]" count="1" pageId="4" pageNumber="143" type="male">1 ♂</specimenCount>
,
<collectingCountry id="F325A4F3FF9ADF7A3823FC48FD00FC4D" box="[570,678,932,956]" name="Indonesia" pageId="4" pageNumber="143">Indonesia</collectingCountry>
,
<collectingRegion id="49F62A81FF9ADF7A38AAFC48FD5EFC4A" box="[691,760,932,955]" country="Indonesia" name="Papua Barat" pageId="4" pageNumber="143">Papua</collectingRegion>
,
<location id="8EEDB2B8FF9ADF7A390BFF1DFCD3FEF8" LSID="urn:lsid:plazi:treatment:039B5575FF9ADF793AB7FD50FBE7FDA3:8EEDB2B8FF9ADF7A390BFF1DFCD3FEF8" box="[786,885,241,265]" country="Indonesia" name="Foja Mts" pageId="4" pageNumber="143" stateProvince="Papua Barat">Foja Mts</location>
(02°27’32’’S, 138°46’30’’E), gravel bank of a small stream,
<quantity id="4CCA4986FF9ADF7A39D9FEFEFBA0FEDB" box="[960,1030,274,298]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="143" unit="m" value="300.0">
<elevation id="001F0350FF9ADF7A39D9FEFEFBA0FEDB" box="[960,1030,274,298]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="4" pageNumber="143" unit="m" value="300.0">300 m</elevation>
</quantity>
,
<date id="FF8CC2A3FF9ADF7A3E04FEFEFB34FEDB" box="[1053,1170,274,298]" pageId="4" pageNumber="143" value="2019-05-28">
<collectingDate id="EFC83B4BFF9ADF7A3E04FEFEFB34FEDB" box="[1053,1170,274,298]" pageId="4" pageNumber="143" value="2019-05-28">28.05.2019</collectingDate>
</date>
, leg.
<collectorName id="26C781B5FF9ADF7A3EF8FEFEFAD1FEDB" box="[1249,1399,274,298]" pageId="4" pageNumber="143">Bretschneider</collectorName>
(
<collectionCode id="ED237CA6FF9ADF7A3903FEDFFCCAFEBA" box="[794,876,307,331]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="4" pageNumber="143">MFNB</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF9ADF793930FEB8FDE3F9FD" lastPageId="7" lastPageNumber="146" pageId="4" pageNumber="143" type="diagnosis">
<paragraph id="8B8DE463FF9ADF7A3930FEB8FBDCFC8B" blockId="4.[786,1399,241,956]" pageId="4" pageNumber="143">
<emphasis id="B9463871FF9ADF7A3930FEB8FC3FFE9D" bold="true" box="[809,921,340,364]" pageId="4" pageNumber="143">Diagnosis</emphasis>
: Body shape and coloring as in
<figureCitation id="1309F8E6FF9ADF7A3EECFEB8FA92FE9D" box="[1269,1332,340,364]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="4" pageNumber="143">Fig. 3</figureCitation>
. Total body length
<quantity id="4CCA4986FF9ADF7A3985FE99FB80FE7C" box="[924,1062,373,397]" metricMagnitude="-3" metricUnit="m" metricValue="3.75" metricValueMax="3.9" metricValueMin="3.6" pageId="4" pageNumber="143" unit="mm" value="3.75" valueMax="3.9" valueMin="3.6">3.6 – 3.9 mm</quantity>
; forebody length
<quantity id="4CCA4986FF9ADF7A3EFEFE99FAD5FE7C" box="[1255,1395,373,397]" metricMagnitude="-3" metricUnit="m" metricValue="2.2" metricValueMax="2.3" metricValueMin="2.1" pageId="4" pageNumber="143" unit="mm" value="2.2" valueMax="2.3" valueMin="2.1">2.1 – 2.3 mm</quantity>
. Head slightly elongate, non-pyriform but subcircular, 1.08 – 1.16 (
<geoCoordinate id="EE0682A4FF9ADF7A3989FE59FC7FFE3E" box="[912,985,437,463]" degrees="1.12" direction="west" orientation="longitude" pageId="4" pageNumber="143" precision="555" value="-1.12">Ø 1.12</geoCoordinate>
) times as long as wide, with convex tempora evenly narrowed towards round posterior margin, across eyes 1.27 – 1.33 (
<geoCoordinate id="EE0682A4FF9ADF7A3E74FE1BFB10FDE1" box="[1133,1206,503,528]" degrees="1.36" direction="west" orientation="longitude" pageId="4" pageNumber="143" precision="555" value="-1.36">Ø 1.36</geoCoordinate>
) times as wide as across tempora at half of their length from eyes to neck constriction. Eyes 0.6 – 0.63 (
<geoCoordinate id="EE0682A4FF9ADF7A3E45FDD5FB0EFDA3" box="[1116,1192,569,594]" degrees="0.61" direction="west" orientation="longitude" pageId="4" pageNumber="143" precision="555" value="-0.61">Ø 0.61</geoCoordinate>
) times as long as tempora. Elytra about as long as pronotum. Penultimate segment of antenna about 1.2 times as long as wide. Tibia of mesothoracic leg 6.7 – 8.2 (
<geoCoordinate id="EE0682A4FF9ADF7A3E81FD70FB73FD44" box="[1176,1237,668,693]" degrees="7.6" direction="west" orientation="longitude" pageId="4" pageNumber="143" precision="5555" value="-7.6">Ø 7.6</geoCoordinate>
) times as long as wide. Body black-brown except for yellow distal fifth of elytra, medium brown maxillary palpi, yellow-brown legs with medium brown femora, and antennae with medium brown segments 1 and 2 followed by blackened segments 3 – 6, which are gradually lighter towards yellowish brown segments 7 – 11.
</paragraph>
<paragraph id="8B8DE463FF9ADF793930FC6FFDE3F9FD" blockId="4.[786,1399,241,956]" lastBlockId="7.[189,803,1162,1975]" lastPageId="7" lastPageNumber="146" pageId="4" pageNumber="143">
<emphasis id="B9463871FF9ADF7A3930FC6FFCC5FC6A" bold="true" box="[809,867,899,923]" pageId="4" pageNumber="143">Male</emphasis>
: Aedeagus (
<figureCitation id="1309F8E6FF9ADF7A39EDFC6FFB9EFC6A" box="[1012,1080,899,923]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="4" pageNumber="143">Fig. 6</figureCitation>
) about
<quantity id="4CCA4986FF9ADF7A3E8DFC6FFB4FFC6A" box="[1172,1257,899,923]" metricMagnitude="-4" metricUnit="m" metricValue="7.0" pageId="4" pageNumber="143" unit="mm" value="0.7">0.7 mm</quantity>
long; apical lobes in lateral view with ventrally curved dorsal margin with longitudinally bent apex and almost right angled extended ventrally (
<figureCitation id="1309F8E6FF99DF793B8EFB47FE44FB32" box="[407,482,1195,1219]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Fig. 11</figureCitation>
), in dorsal view (
<figureCitation id="1309F8E6FF99DF7938BAFB47FD49FB32" box="[675,751,1195,1219]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Fig. 13</figureCitation>
) not widened laterally, but convex towards round apex, their distoventral margins circular in ventral view (
<figureCitation id="1309F8E6FF99DF7938DDFB01FCB3FAF4" box="[708,789,1261,1285]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Fig. 12</figureCitation>
); lateral lobes about as wide as long (
<figureCitation id="1309F8E6FF99DF793855FAE2FD6FFAD7" box="[588,713,1294,1318]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Figs 12, 13</figureCitation>
); dorsal lobe stout with round apex in dorsal view (
<figureCitation id="1309F8E6FF99DF7938DBFAC2FCB3FAB6" box="[706,789,1326,1351]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Fig. 13</figureCitation>
). Asetose median depression of abdominal sternite VII (
<figureCitation id="1309F8E6FF99DF793ADCFA9CFEA8FA79" box="[197,270,1392,1416]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="7" pageNumber="146">Fig. 19</figureCitation>
) with parallel lateral margins. Abdominal sternite VIII (
<figureCitation id="1309F8E6FF99DF793AE7FA7DFEE8FA58" box="[254,334,1425,1449]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="7" pageNumber="146">Fig. 20</figureCitation>
) with median lobe of posterior margin as long as lateroposterior angles; end of median lobe notably emarginate; four distinct, pigmented macrosetae situated close to posterior margin of sternite.
</paragraph>
</subSubSection>
<caption id="DF4DB4EBFF9ADF7A3A8EF89FFE89F847" ID-DOI="http://doi.org/10.5281/zenodo.13193968" ID-Zenodo-Dep="13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="4" pageNumber="143" startId="4.[151,224,1907,1929]" targetBox="[227,1323,999,1870]" targetPageId="4" targetType="figure">
<paragraph id="8B8DE463FF9ADF7A3A8EF89FFE89F847" blockId="4.[151,1399,1907,1974]" pageId="4" pageNumber="143">
<emphasis id="B9463871FF9ADF7A3A8EF89FFEABF879" bold="true" box="[151,269,1907,1929]" pageId="4" pageNumber="143">Figures 5–7.</emphasis>
Aedeagus (in lateral view) of
<taxonomicName id="4C329FE0FF9ADF7A383AF89FFD5BF879" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[547,765,1907,1928]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="arfakmontium">
<emphasis id="B9463871FF9ADF7A383AF89FFD5BF879" box="[547,765,1907,1928]" italics="true" pageId="4" pageNumber="143">Scopaeus arfakmontium</emphasis>
</taxonomicName>
, holotype, Indonesia, West Papua, Arfak Mts (
<emphasis id="B9463871FF9ADF7A3EB1F89FFB15F879" bold="true" box="[1192,1203,1907,1928]" pageId="4" pageNumber="143">5</emphasis>
),
<taxonomicName id="4C329FE0FF9ADF7A3EDEF89FFAB3F879" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1223,1301,1907,1928]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF9ADF7A3EDEF89FFAB3F879" box="[1223,1301,1907,1928]" italics="true" pageId="4" pageNumber="143">S. balkei</emphasis>
</taxonomicName>
, holotype, Indonesia, Papua, Cyclops Mts (
<emphasis id="B9463871FF9ADF7A3BD7F866FE7FF86E" bold="true" box="[462,473,1930,1951]" pageId="4" pageNumber="143">6</emphasis>
),
<taxonomicName id="4C329FE0FF9ADF7A3BE9F866FDCEF86E" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[496,616,1930,1951]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="143" phylum="Arthropoda" rank="species" species="potamides">
<emphasis id="B9463871FF9ADF7A3BE9F866FDCEF86E" box="[496,616,1930,1951]" italics="true" pageId="4" pageNumber="143">S. potamides</emphasis>
</taxonomicName>
, holotype, Solomon Islands, Kolombangara (
<emphasis id="B9463871FF9ADF7A3E00F866FB82F86E" bold="true" box="[1049,1060,1930,1951]" pageId="4" pageNumber="143">7</emphasis>
).
<emphasis id="B9463871FF9ADF7A3E25F865FB63F86E" bold="true" box="[1084,1221,1929,1951]" pageId="4" pageNumber="143">Abbreviations</emphasis>
:
<emphasis id="B9463871FF9ADF7A3ECCF866FB4EF86E" bold="true" box="[1237,1256,1930,1951]" pageId="4" pageNumber="143">dl</emphasis>
– distal lobes,
<emphasis id="B9463871FF9ADF7A3A8EF84CFF16F844" bold="true" box="[151,176,1952,1973]" pageId="4" pageNumber="143">pb</emphasis>
– phallobase.
</paragraph>
</caption>
<caption id="DF4DB4EBFF9BDF7B3AA4F898FABDF846" ID-DOI="http://doi.org/10.5281/zenodo.13193970" ID-Zenodo-Dep="13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="5" pageNumber="144" startId="5.[189,262,1908,1930]" targetBox="[236,1378,299,1853]" targetPageId="5" targetType="figure">
<paragraph id="8B8DE463FF9BDF7B3AA4F898FABDF846" blockId="5.[189,1437,1908,1975]" pageId="5" pageNumber="144">
<emphasis id="B9463871FF9BDF7B3AA4F898FE99F878" bold="true" box="[189,319,1908,1930]" pageId="5" pageNumber="144">Figures 8–16.</emphasis>
Apical portion of aedeagus in lateral (
<emphasis id="B9463871FF9BDF7B38BDF898FD09F878" bold="true" box="[676,687,1908,1929]" pageId="5" pageNumber="144">8</emphasis>
,
<emphasis id="B9463871FF9BDF7B38A5F898FD77F878" bold="true" box="[700,721,1908,1929]" pageId="5" pageNumber="144">11</emphasis>
,
<emphasis id="B9463871FF9BDF7B38C7F898FD53F878" bold="true" box="[734,757,1908,1929]" pageId="5" pageNumber="144">14</emphasis>
), ventral (
<emphasis id="B9463871FF9BDF7B394FF898FCC7F878" bold="true" box="[854,865,1908,1929]" pageId="5" pageNumber="144">9</emphasis>
,
<emphasis id="B9463871FF9BDF7B3977F898FC23F878" bold="true" box="[878,901,1908,1929]" pageId="5" pageNumber="144">12</emphasis>
,
<emphasis id="B9463871FF9BDF7B398BF898FC0FF878" bold="true" box="[914,937,1908,1929]" pageId="5" pageNumber="144">15</emphasis>
), dorsal view (
<emphasis id="B9463871FF9BDF7B3E2CF898FBEAF878" bold="true" box="[1077,1100,1908,1929]" pageId="5" pageNumber="144">10</emphasis>
,
<emphasis id="B9463871FF9BDF7B3E40F898FBD6F878" bold="true" box="[1113,1136,1908,1929]" pageId="5" pageNumber="144">13</emphasis>
,
<emphasis id="B9463871FF9BDF7B3E65F898FB35F878" bold="true" box="[1148,1171,1908,1929]" pageId="5" pageNumber="144">16</emphasis>
) of
<taxonomicName id="4C329FE0FF9BDF7B3EA2F898FA30F878" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1211,1430,1908,1929]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="144" phylum="Arthropoda" rank="species" species="arfakmontium">
<emphasis id="B9463871FF9BDF7B3EA2F898FA30F878" box="[1211,1430,1908,1929]" italics="true" pageId="5" pageNumber="144">Scopaeus arfakmontium</emphasis>
</taxonomicName>
, holotype, Indonesia, West Papua, Arfak Mts (
<emphasis id="B9463871FF9BDF7B3844F866FD2CF86E" bold="true" box="[605,650,1930,1952]" pageId="5" pageNumber="144">8–10</emphasis>
),
<taxonomicName id="4C329FE0FF9BDF7B3887F867FD4AF851" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[670,748,1931,1952]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="144" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF9BDF7B3887F867FD4AF851" box="[670,748,1931,1952]" italics="true" pageId="5" pageNumber="144">S. balkei</emphasis>
</taxonomicName>
, holotype, Indonesia, Papua, Cyclops Mts (
<emphasis id="B9463871FF9BDF7B3E65F866FB12F86E" bold="true" box="[1148,1204,1930,1952]" pageId="5" pageNumber="144">11–13</emphasis>
),
<taxonomicName id="4C329FE0FF9BDF7B3EDEF867FA9DF851" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1223,1339,1931,1952]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="144" phylum="Arthropoda" rank="species" species="potamides">
<emphasis id="B9463871FF9BDF7B3EDEF867FA9DF851" box="[1223,1339,1931,1952]" italics="true" pageId="5" pageNumber="144">S. potamides</emphasis>
</taxonomicName>
, holotype, Solomon Islands, Kolombangara (
<emphasis id="B9463871FF9BDF7B3BEDF84DFD8BF847" bold="true" box="[500,557,1953,1975]" pageId="5" pageNumber="144">14–16</emphasis>
).
<emphasis id="B9463871FF9BDF7B3826F84DFD6EF846" bold="true" box="[575,712,1953,1975]" pageId="5" pageNumber="144">Abbreviations</emphasis>
:
<emphasis id="B9463871FF9BDF7B38CDF84DFD40F847" bold="true" box="[724,742,1953,1974]" pageId="5" pageNumber="144">al</emphasis>
– apical lobes,
<emphasis id="B9463871FF9BDF7B396DF84DFC21F847" bold="true" box="[884,903,1953,1974]" pageId="5" pageNumber="144">dl</emphasis>
– dorsal lobe,
<emphasis id="B9463871FF9BDF7B3E14F84DFBB3F846" bold="true" box="[1037,1045,1953,1975]" pageId="5" pageNumber="144">f</emphasis>
– flagellum,
<emphasis id="B9463871FF9BDF7B3E95F84DFB3FF847" bold="true" box="[1164,1177,1953,1974]" pageId="5" pageNumber="144">ll</emphasis>
– lateral lobe.
</paragraph>
</caption>
<caption id="DF4DB4EBFF98DF783A8EF8B1FD30F846" ID-DOI="http://doi.org/10.5281/zenodo.13193972" ID-Zenodo-Dep="13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="6" pageNumber="145" startId="6.[151,224,1885,1907]" targetBox="[222,1362,228,1856]" targetPageId="6" targetType="figure">
<paragraph id="8B8DE463FF98DF783A8EF8B1FD30F846" blockId="6.[151,1399,1885,1975]" pageId="6" pageNumber="145">
<emphasis id="B9463871FF98DF783A8EF8B1FE81F883" bold="true" box="[151,295,1885,1907]" pageId="6" pageNumber="145">Figures 17–24.</emphasis>
Abdominal sternite VII (
<emphasis id="B9463871FF98DF783802F8B1FD94F883" bold="true" box="[539,562,1885,1906]" pageId="6" pageNumber="145">17</emphasis>
,
<emphasis id="B9463871FF98DF783858F8B1FDFEF883" bold="true" box="[577,600,1885,1906]" pageId="6" pageNumber="145">19</emphasis>
,
<emphasis id="B9463871FF98DF78387EF8B1FDD8F883" bold="true" box="[615,638,1885,1906]" pageId="6" pageNumber="145">21</emphasis>
) and abdominal sternite VIII (
<emphasis id="B9463871FF98DF7839BCF8B1FC1AF883" bold="true" box="[933,956,1885,1906]" pageId="6" pageNumber="145">18</emphasis>
,
<emphasis id="B9463871FF98DF7839D2F8B1FC44F883" bold="true" box="[971,994,1885,1906]" pageId="6" pageNumber="145">20</emphasis>
,
<emphasis id="B9463871FF98DF7839E9F8B1FBA1F883" bold="true" box="[1008,1031,1885,1906]" pageId="6" pageNumber="145">22</emphasis>
) of
<taxonomicName id="4C329FE0FF98DF783E2DF8B1FAB4F883" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1076,1298,1885,1906]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="arfakmontium">
<emphasis id="B9463871FF98DF783E2DF8B1FAB4F883" box="[1076,1298,1885,1906]" italics="true" pageId="6" pageNumber="145">Scopaeus arfakmontium</emphasis>
</taxonomicName>
, holotype, Indonesia, West Papua, Arfak Mts (
<emphasis id="B9463871FF98DF783BC0F898FE56F878" bold="true" box="[473,496,1908,1929]" pageId="6" pageNumber="145">17</emphasis>
,
<emphasis id="B9463871FF98DF783BE5F898FDB5F878" bold="true" box="[508,531,1908,1929]" pageId="6" pageNumber="145">18</emphasis>
),
<taxonomicName id="4C329FE0FF98DF78383CF898FDD5F878" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[549,627,1908,1929]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF98DF78383CF898FDD5F878" box="[549,627,1908,1929]" italics="true" pageId="6" pageNumber="145">S. balkei</emphasis>
</taxonomicName>
, holotype, Indonesia, Papua, Cyclops Mts (
<emphasis id="B9463871FF98DF783E19F898FBB1F878" bold="true" box="[1024,1047,1908,1929]" pageId="6" pageNumber="145">19</emphasis>
,
<emphasis id="B9463871FF98DF783E3BF898FB9FF878" bold="true" box="[1058,1081,1908,1929]" pageId="6" pageNumber="145">20</emphasis>
),
<taxonomicName id="4C329FE0FF98DF783E55F898FB66F878" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1100,1216,1908,1929]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="potamides">
<emphasis id="B9463871FF98DF783E55F898FB66F878" box="[1100,1216,1908,1929]" italics="true" pageId="6" pageNumber="145">S. potamides</emphasis>
</taxonomicName>
, paratype, Solomon Islands, Kolombangara (
<emphasis id="B9463871FF98DF783B67F866FE33F86E" bold="true" box="[382,405,1930,1951]" pageId="6" pageNumber="145">21</emphasis>
,
<emphasis id="B9463871FF98DF783BBDF866FE1DF86E" bold="true" box="[420,443,1930,1951]" pageId="6" pageNumber="145">22</emphasis>
),
<taxonomicName id="4C329FE0FF98DF783BC8F867FDEAF86E" authorityName="Cameron" authorityYear="1938" box="[465,588,1930,1952]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="kokodanus">
<emphasis id="B9463871FF98DF783BC8F867FDEAF86E" box="[465,588,1930,1952]" italics="true" pageId="6" pageNumber="145">S. kokodanus</emphasis>
</taxonomicName>
(
<emphasis id="B9463871FF98DF783844F866FDD2F86E" bold="true" box="[605,628,1930,1951]" pageId="6" pageNumber="145">23</emphasis>
). Figs
<emphasis id="B9463871FF98DF7838A2F866FD74F86E" bold="true" box="[699,722,1930,1951]" pageId="6" pageNumber="145">23</emphasis>
,
<emphasis id="B9463871FF98DF7838F8F866FD5EF86E" bold="true" box="[737,760,1930,1951]" pageId="6" pageNumber="145">24</emphasis>
: sperm pumps of (
<emphasis id="B9463871FF98DF7839ABF866FC6FF86E" bold="true" box="[946,969,1930,1951]" pageId="6" pageNumber="145">23</emphasis>
)
<taxonomicName id="4C329FE0FF98DF7839C0F867FBF3F86E" authorityName="Cameron" authorityYear="1938" box="[985,1109,1930,1952]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="kokodanus">
<emphasis id="B9463871FF98DF7839C0F867FBF3F86E" box="[985,1109,1930,1952]" italics="true" pageId="6" pageNumber="145">S. kokodanus</emphasis>
</taxonomicName>
, Papua New Guinea, Morobe, (
<emphasis id="B9463871FF98DF783A86F84DFF10F847" bold="true" box="[159,182,1953,1974]" pageId="6" pageNumber="145">24</emphasis>
)
<taxonomicName id="4C329FE0FF98DF783ADAF84DFEB6F847" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[195,272,1953,1974]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="145" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF98DF783ADAF84DFEB6F847" box="[195,272,1953,1974]" italics="true" pageId="6" pageNumber="145">S. balkei</emphasis>
</taxonomicName>
, paratype, Indonesia, Papua, Cyclops Mts.
</paragraph>
</caption>
<caption id="DF4DB4EBFF99DF793AA4FBC1FE13FBA8" ID-DOI="http://doi.org/10.5281/zenodo.13193978" ID-Zenodo-Dep="13193978" httpUri="https://zenodo.org/record/13193978/files/figure.png" pageId="7" pageNumber="146" startId="7.[189,253,1069,1091]" targetBox="[189,801,239,1046]" targetPageId="7" targetType="figure">
<paragraph id="8B8DE463FF99DF793AA4FBC1FE13FBA8" blockId="7.[189,802,1069,1113]" pageId="7" pageNumber="146">
<emphasis id="B9463871FF99DF793AA4FBC1FE83FBB3" bold="true" box="[189,293,1069,1091]" pageId="7" pageNumber="146">Figure 25.</emphasis>
Habitat of
<taxonomicName id="4C329FE0FF99DF793B84FBC1FD90FBB3" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[413,566,1069,1090]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF99DF793B84FBC1FD90FBB3" box="[413,566,1069,1090]" italics="true" pageId="7" pageNumber="146">Scopaeus balkei</emphasis>
</taxonomicName>
in the Foja Mountains, Indonesia, Papua Province.
</paragraph>
</caption>
<subSubSection id="C328B7E8FF99DF793ACDF9F8FBD8FE7C" pageId="7" pageNumber="146" type="distribution">
<paragraph id="8B8DE463FF99DF793ACDF9F8FD24F99F" blockId="7.[189,803,1162,1975]" pageId="7" pageNumber="146">
<emphasis id="B9463871FF99DF793ACDF9F8FEC2F9DD" bold="true" box="[212,356,1556,1580]" pageId="7" pageNumber="146">Distribution</emphasis>
:
<taxonomicName id="4C329FE0FF99DF793B6EF9F9FD88F9DC" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[375,558,1557,1581]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF99DF793B6EF9F9FD88F9DC" box="[375,558,1557,1581]" italics="true" pageId="7" pageNumber="146">Scopaeus balkei</emphasis>
</taxonomicName>
was collected in the Cyclops and Foja Mountains (
<figureCitation id="1309F8E6FF99DF79381FF9DAFDF2F9BC" box="[518,596,1589,1614]" captionStart="Figure 25" captionStartId="7.[189,253,1069,1091]" captionTargetBox="[189,801,239,1046]" captionTargetId="figure-632@7.[189,804,236,1046]" captionTargetPageId="7" captionText="Figure 25. Habitat of Scopaeus balkei in the Foja Mountains, Indonesia, Papua Province." figureDoi="http://doi.org/10.5281/zenodo.13193978" httpUri="https://zenodo.org/record/13193978/files/figure.png" pageId="7" pageNumber="146">Fig. 25</figureCitation>
) in the Indonesian province of Papua, western New
<collectingCountry id="F325A4F3FF99DF793833F9BAFDD8F99F" box="[554,638,1622,1646]" name="Guinea-Bissau" pageId="7" pageNumber="146">Guinea</collectingCountry>
.
</paragraph>
<paragraph id="8B8DE463FF99DF793ACDF99BFBD8FE7C" blockId="7.[189,803,1162,1975]" lastBlockId="7.[824,1437,241,594]" pageId="7" pageNumber="146">
<emphasis id="B9463871FF99DF793ACDF99BFE10F97E" bold="true" box="[212,438,1655,1679]" pageId="7" pageNumber="146">Comparative notes</emphasis>
: Among the members of the
<taxonomicName id="4C329FE0FF99DF793AA4F975FE0DF941" authorityName="Cameron" authorityYear="1938" box="[189,427,1688,1712]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="kokodanus">
<emphasis id="B9463871FF99DF793AA4F975FE0DF941" box="[189,427,1688,1712]" italics="true" pageId="7" pageNumber="146">Scopaeus kokodanus</emphasis>
</taxonomicName>
species group,
<taxonomicName id="4C329FE0FF99DF79386DF975FD7BF941" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[628,733,1688,1712]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF99DF79386DF975FD7BF941" box="[628,733,1688,1712]" italics="true" pageId="7" pageNumber="146">S. balkei</emphasis>
</taxonomicName>
(
<figureCitation id="1309F8E6FF99DF7938EEF974FF6CF920" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="7" pageNumber="146">Fig. 3</figureCitation>
) differs from both
<taxonomicName id="4C329FE0FF99DF793BD9F955FDFDF920" authorityName="Cameron" authorityYear="1938" box="[448,603,1721,1745]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="kokodanus">
<emphasis id="B9463871FF99DF793BD9F955FDFDF920" box="[448,603,1721,1745]" italics="true" pageId="7" pageNumber="146">S. kokodanus</emphasis>
</taxonomicName>
(
<figureCitation id="1309F8E6FF99DF79386AF955FD1CF920" box="[627,698,1721,1745]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="7" pageNumber="146">Fig. 1</figureCitation>
) and
<taxonomicName id="4C329FE0FF99DF793914F955FEF0F903" authorityName="Frisch &amp; Mainda" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="arfakmontium">
<emphasis id="B9463871FF99DF793914F955FEF0F903" italics="true" pageId="7" pageNumber="146">S. arfakmontium</emphasis>
</taxonomicName>
(
<figureCitation id="1309F8E6FF99DF793B73F936FE08F903" box="[362,430,1754,1778]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="7" pageNumber="146">Fig. 2</figureCitation>
) by smaller body size and non-pyriform head. The species can moreover be separated from
<taxonomicName id="4C329FE0FF99DF793AE1F8F0FE0BF8C5" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[248,429,1820,1844]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="arfakmontium">
<emphasis id="B9463871FF99DF793AE1F8F0FE0BF8C5" box="[248,429,1820,1844]" italics="true" pageId="7" pageNumber="146">S. arfakmontium</emphasis>
</taxonomicName>
by the yellow apical margin of the elytra, the smaller aedeagus with stouter lateral lobes, laterally not widened, but ventrally angled extended apical lobes and round apex of the dorsal lobe (
<figureCitation id="1309F8E6FF99DF7938DCF892FF7EF846" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Figs 11– 13</figureCitation>
), and by the posterior margin of abdominal sternite VII with shorter median lobe and the presence of four distinct, dark macrosetae (
<figureCitation id="1309F8E6FF99DF793E68FEFEFB6EFEDB" box="[1137,1224,274,298]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="7" pageNumber="146">Fig. 20</figureCitation>
).
<taxonomicName id="4C329FE0FF99DF793EFDFEFFFA3AFEDB" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1252,1436,274,298]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="balkei">
<emphasis id="B9463871FF99DF793EFDFEFFFA3AFEDB" box="[1252,1436,274,298]" italics="true" pageId="7" pageNumber="146">Scopaeus balkei</emphasis>
</taxonomicName>
differs from
<taxonomicName id="4C329FE0FF99DF7939DEFED8FBF0FEBA" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[967,1110,307,331]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="potamides">
<emphasis id="B9463871FF99DF7939DEFED8FBF0FEBA" box="[967,1110,307,331]" italics="true" pageId="7" pageNumber="146">S. potamides</emphasis>
</taxonomicName>
by the darker body coloring and characters of the aedeagus and abdominal sternites VII and VIII described below.
</paragraph>
</subSubSection>
<subSubSection id="C328B7E8FF99DF793956FE7AFBE7FDA3" pageId="7" pageNumber="146" type="etymology">
<paragraph id="8B8DE463FF99DF793956FE7AFBE7FDA3" blockId="7.[824,1437,241,594]" pageId="7" pageNumber="146">
<emphasis id="B9463871FF99DF793956FE7AFC6AFE5F" bold="true" box="[847,972,406,430]" pageId="7" pageNumber="146">Etymology</emphasis>
: With the choice of the epithet
<taxonomicName id="4C329FE0FF99DF793F53FE7AFA35FE5F" authorityName="Frisch &amp; Mainda" authorityYear="2022" box="[1354,1427,406,430]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="balkei">
‘
<emphasis id="B9463871FF99DF793F48FE7AFA35FE5F" box="[1361,1427,406,430]" italics="true" pageId="7" pageNumber="146">balkei</emphasis>
</taxonomicName>
’ (Latinized noun, derived from the surname ‘Balke’, genitive) we warmly dedicate the species to Michael Balke, coleopterist at the Zoologische Staatssammlung Munich, who actively supports the second author in his entomological activities.
</paragraph>
</subSubSection>
</treatment>
</document>
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