234 lines
36 KiB
XML
234 lines
36 KiB
XML
<document id="E6C815F31CF09C34EB2CF89D092553CC" ID-DOI="10.25674/SO94iSS3iD303" ID-ISSN="2509-9523" ID-Zenodo-Dep="13193962" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1722631614709" checkinUser="felipe" docAuthor="Frisch, Johannes & Mainda, Tobias" docDate="2022" docId="039B5575FF99DF763956FD77FD62FBEE" docLanguage="en" docName="SoilOrg.94.3.139-147.pdf" docOrigin="Soil Organisms 94 (3)" docSource="http://dx.doi.org/10.25674/so94iss3id303" docStyle="DocumentStyle:EA2ECAC87EC21C07AB805D5B9B4601F2.2:SoilOrg.2018-.journal_article" docStyleId="EA2ECAC87EC21C07AB805D5B9B4601F2" docStyleName="SoilOrg.2018-.journal_article" docStyleVersion="2" docTitle="Scopaeus potamides Frisch & Mainda, 2022, spec. nov." docType="treatment" docVersion="2" lastPageNumber="147" masterDocId="FFA22D0DFF9EDF7E3A19FFECFFA6FFF1" masterDocTitle="The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species" masterLastPageNumber="147" masterPageNumber="139" pageNumber="146" updateTime="1722679412379" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1EB0F9ABFE3CEE2660BD39275E805748">The Scopaeus kokodanus species group (Coleoptera: Staphylinidae: Paederinae) from New Guinea and the Solomon Islands, with description of three new species</mods:title>
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<mods:namePart id="4AEEE0449FC09C158AF927553838D37D">Frisch, Johannes</mods:namePart>
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<mods:affiliation id="D3355B0085567CD7154D19C305BA2020">MUSeUM füR NatURKUnDe BeRLin, Leibniz InStitUte fOR EvOLUtiOn anD BiODiveRSity Science,</mods:affiliation>
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<mods:roleTerm id="A859C651D449E1BC63D5D5D2B161F01F">Author</mods:roleTerm>
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<mods:namePart id="949E3C90721739565FBF786237C0C0F1">Mainda, Tobias</mods:namePart>
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<mods:affiliation id="94D574FE23F64B988C0745B31E1E96C6">ZOOLOgiScheS MUSeUM, UniveRSität GReifSWaLD, LOitzeR StRasse 26, 17489 GReifSWaLD, GeRMany</mods:affiliation>
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<mods:typeOfResource id="02C5C4E83F1E5CADBE5ADB22FA9AEEDD">text</mods:typeOfResource>
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<mods:title id="CD201D84326694385A72AF2E92133267">Soil Organisms</mods:title>
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<mods:part id="7F7C8C56C4E917C83C5A67BCEC8D724C">
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<mods:date id="F04A2903AAF972413F8B0BCB6CB81184">2022</mods:date>
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<mods:number id="334AB047DB2C70EE0CDA047ECF6D1BFC">2022-12-01</mods:number>
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<mods:number id="CD81E9D1662E1D199FF0C985716D7CD7">94</mods:number>
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<mods:number id="CA3FFBBB4E747A316BF7FF9BD50FB453">3</mods:number>
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<mods:start id="206F0CB6544AF41217705F9DA0A17D65">139</mods:start>
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<mods:url id="BFCA8EFCB052A0D1EDC0E1EA1625229D">http://dx.doi.org/10.25674/so94iss3id303</mods:url>
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<mods:classification id="D42729C9B354F6E4EB2D94346E0D3FAB">journal article</mods:classification>
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<mods:identifier id="6B25D1C1F4F1CD971C1D16663E26D88E" type="DOI">10.25674/SO94iSS3iD303</mods:identifier>
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<mods:identifier id="A13C678F7316FE5F734B51C1CCC4ABA9" type="ISSN">2509-9523</mods:identifier>
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<mods:identifier id="08C37BE6FDA6F133B537DF3A6292A5B6" type="Zenodo-Dep">13193962</mods:identifier>
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<treatment id="039B5575FF99DF763956FD77FD62FBEE" LSID="urn:lsid:plazi:treatment:039B5575FF99DF763956FD77FD62FBEE" httpUri="http://treatment.plazi.org/id/039B5575FF99DF763956FD77FD62FBEE" lastPageId="8" lastPageNumber="147" pageId="7" pageNumber="146">
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<subSubSection id="C328B7E8FF99DF793956FD77FB62FD44" box="[847,1220,667,694]" pageId="7" pageNumber="146" type="nomenclature">
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<paragraph id="8B8DE463FF99DF793956FD77FB62FD44" blockId="7.[847,1220,667,726]" box="[847,1220,667,694]" pageId="7" pageNumber="146">
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<heading id="D0C5530FFF99DF793956FD77FB62FD44" box="[847,1220,667,694]" fontSize="11" level="2" pageId="7" pageNumber="146" reason="5">
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<emphasis id="B9463871FF99DF793956FD77FB62FD44" bold="true" box="[847,1220,667,694]" pageId="7" pageNumber="146">
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<taxonomicName id="4C329FE0FF99DF793956FD77FBE3FD47" authorityName="Frisch & Mainda" authorityYear="2022" box="[847,1093,667,694]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="potamides" status="sp. nov.">
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<emphasis id="B9463871FF99DF793956FD77FBE3FD47" bold="true" box="[847,1093,667,694]" italics="true" pageId="7" pageNumber="146">Scopaeus potamides</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A275850AFF99DF793E55FD77FB62FD44" box="[1100,1220,667,693]" pageId="7" pageNumber="146" rank="species">spec. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C328B7E8FF99DF793956FD52FBF2FD27" box="[847,1108,702,726]" pageId="7" pageNumber="146" type="description">
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<paragraph id="8B8DE463FF99DF793956FD52FBF2FD27" blockId="7.[847,1220,667,726]" box="[847,1108,702,726]" pageId="7" pageNumber="146">
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(
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<figureCitation id="1309F8E6FF99DF79394FFD52FC3DFD27" box="[854,923,702,726]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="7" pageNumber="146">Figs 4</figureCitation>
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,
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<figureCitation id="1309F8E6FF99DF7939BFFD52FC12FD24" box="[934,948,702,725]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="7" pageNumber="146">7</figureCitation>
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,
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<figureCitation id="1309F8E6FF99DF7939A6FD52FBA4FD27" box="[959,1026,702,726]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">14–16</figureCitation>
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,
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<figureCitation id="1309F8E6FF99DF793E15FD52FBEBFD27" box="[1036,1101,702,726]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="7" pageNumber="146">21, 22</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C328B7E8FF99DF793956FCECFA8AFC6A" pageId="7" pageNumber="146" type="materials_examined">
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<paragraph id="8B8DE463FF99DF793956FCECFA8AFC6A" blockId="7.[824,1437,767,1976]" pageId="7" pageNumber="146">
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<emphasis id="B9463871FF99DF793956FCECFBB6FCE6" bold="true" box="[847,1040,767,792]" pageId="7" pageNumber="146">Type specimens</emphasis>
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:
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<materialsCitation id="3B5AEE3EFF99DF793E34FCECFA3FFC8B" collectingDate="1965-07-09" collectionCode="NHML" collectorName="Royal Society Expedition & British Museum" country="Solomon Islands" county="Kolombara River" location="Kolombangara" municipality="Camp" pageId="7" pageNumber="146" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
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<typeStatus id="54895AC1FF99DF793E34FCECFB37FCE9" box="[1069,1169,768,792]" pageId="7" pageNumber="146" type="holotype">Holotype</typeStatus>
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<specimenCount id="9D342FEAFF99DF793EBFFCECFB1CFCE6" box="[1190,1210,768,791]" count="1" pageId="7" pageNumber="146" type="male">♂</specimenCount>
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,
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<collectingCountry id="F325A4F3FF99DF793ECDFCECFA3EFCE9" box="[1236,1432,768,792]" name="Solomon Islands" pageId="7" pageNumber="146">Solomon Islands</collectingCountry>
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,
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<location id="8EEDB2B8FF99DF793921FCCCFC7CFCC9" LSID="urn:lsid:plazi:treatment:039B5575FF99DF763956FD77FD62FBEE:8EEDB2B8FF99DF793921FCCCFC7CFCC9" box="[824,986,800,824]" country="Solomon Islands" county="Kolombara River" municipality="Camp" name="Kolombangara" pageId="7" pageNumber="146">Kolombangara</location>
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,
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<location id="8EEDB2B8FF99DF7939F0FCCCFB86FCC9" LSID="urn:lsid:plazi:treatment:039B5575FF99DF763956FD77FD62FBEE:8EEDB2B8FF99DF7939F0FCCCFB86FCC9" box="[1001,1056,800,824]" country="Solomon Islands" county="Kolombara River" municipality="Camp" name="Kuzi" pageId="7" pageNumber="146">Kuzi</location>
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,
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<collectingMunicipality id="6BE97E19FF99DF793E36FCCDFBC9FCC8" box="[1071,1135,801,825]" pageId="7" pageNumber="146">Camp</collectingMunicipality>
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1,
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<date id="FF8CC2A3FF99DF793E8EFCCCFAACFCC9" box="[1175,1290,800,824]" pageId="7" pageNumber="146" value="1965-07-09">
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<collectingDate id="EFC83B4BFF99DF793E8EFCCCFAACFCC9" box="[1175,1290,800,824]" pageId="7" pageNumber="146" value="1965-07-09">07.09.1965</collectingDate>
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</date>
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, flood refuse between stones, banks of
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<collectingCounty id="62EC9CEFFF99DF793E44FCADFABBFCA8" box="[1117,1309,833,857]" pageId="7" pageNumber="146">Kolombara River</collectingCounty>
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, leg.
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<collectorName id="26C781B5FF99DF793F44FCADFBA8FC8B" pageId="7" pageNumber="146">Royal Society Expedition</collectorName>
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,
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<collectorName id="26C781B5FF99DF793E07FC8EFB74FC8B" box="[1054,1234,866,890]" pageId="7" pageNumber="146">British Museum</collectorName>
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1966-1 (
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<collectionCode id="ED237CA6FF99DF793F25FC8EFA37FC8B" box="[1340,1425,866,890]" country="Libya" httpUri="http://grbio.org/cool/49nc-28y0" name="Natural History Museum, Tripoli" pageId="7" pageNumber="146">NHML</collectionCode>
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)
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</materialsCitation>
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.
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<materialsCitation id="3B5AEE3EFF99DF793921FC68FA8FFC6A" box="[824,1321,899,924]" collectingDate="1965-07-09" collectionCode="MFNB" collectorName="Royal Society Expedition & British Museum" country="Solomon Islands" county="Kolombara River" location="Kolombangara" municipality="Camp" pageId="7" pageNumber="146" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
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<typeStatus id="54895AC1FF99DF793921FC68FC3EFC6D" box="[824,920,900,924]" pageId="7" pageNumber="146" type="paratype">Paratype</typeStatus>
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<specimenCount id="9D342FEAFF99DF793986FC6FFC61FC6A" box="[927,967,899,923]" count="1" pageId="7" pageNumber="146" type="male">1 ♂</specimenCount>
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, same data as holotype (
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<collectionCode id="ED237CA6FF99DF793ED7FC6FFA86FC6A" box="[1230,1312,899,923]" country="Italy" httpUri="http://grbio.org/cool/irpn-c0nq" name="Museo Friulano di Storia Naturale" pageId="7" pageNumber="146">MFNB</collectionCode>
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)
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</materialsCitation>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="C328B7E8FF99DF793956FC48FC43F8C5" pageId="7" pageNumber="146" type="diagnosis">
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<paragraph id="8B8DE463FF99DF793956FC48FBABFA79" blockId="7.[824,1437,767,1976]" pageId="7" pageNumber="146">
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<emphasis id="B9463871FF99DF793956FC48FC1BFC4D" bold="true" box="[847,957,932,956]" pageId="7" pageNumber="146">Diagnosis</emphasis>
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: Body shape and coloring as in
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<figureCitation id="1309F8E6FF99DF793F00FC48FAFDFC4D" box="[1305,1371,932,956]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="7" pageNumber="146">Fig. 4</figureCitation>
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. Total body length
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<quantity id="4CCA4986FF99DF7939D8FC29FBEBFC2D" box="[961,1101,965,989]" metricMagnitude="-3" metricUnit="m" metricValue="3.7" metricValueMax="3.8" metricValueMin="3.6" pageId="7" pageNumber="146" unit="mm" value="3.7" valueMax="3.8" valueMin="3.6">3.6 – 3.8 mm</quantity>
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; forebody length
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<quantity id="4CCA4986FF99DF793F14FC29FA3FFC2D" box="[1293,1433,965,989]" metricMagnitude="-3" metricUnit="m" metricValue="2.3499999999999996" metricValueMax="2.4" metricValueMin="2.3" pageId="7" pageNumber="146" unit="mm" value="2.3499999999999996" valueMax="2.4" valueMin="2.3">2.3 – 2.4 mm</quantity>
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||
. Head slightly elongate, non-pyriform but subcircular, 1.1 times as long as wide, with convex tempora evenly narrowed towards round posterior margin, across eyes 1.17 – 1.28 (
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<geoCoordinate id="EE0682A4FF99DF7939A2FBABFBAFFB91" box="[955,1033,1095,1120]" degrees="1.23" direction="west" orientation="longitude" pageId="7" pageNumber="146" precision="555" value="-1.23">Ø 1.23</geoCoordinate>
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||
) times as wide as across tempora at half of their length from eyes to neck constriction. Eyes 0.64 – 0.71 (
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<geoCoordinate id="EE0682A4FF99DF7939A6FB65FBB6FB53" box="[959,1040,1161,1186]" degrees="0.68" direction="west" orientation="longitude" pageId="7" pageNumber="146" precision="555" value="-0.68">Ø 0.68</geoCoordinate>
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||
) times as long as tempora. Elytra 1.1 times as long as pronotum. Penultimate segment of antenna about 1.3 times as long as wide. Tibia of mesothoracic leg about seven times as long as wide. Head and pronotum reddish brown, elytra dark brown except for yellow posterior fifth, abdomen dark brown, appendages (maxillary palps, antennae, legs) unicolorous light yellow-brown.
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</paragraph>
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||
<paragraph id="8B8DE463FF99DF793956FA7DFC43F8C5" blockId="7.[824,1437,767,1976]" pageId="7" pageNumber="146">
|
||
<emphasis id="B9463871FF99DF793956FA7DFC2FFA58" bold="true" box="[847,905,1425,1449]" pageId="7" pageNumber="146">Male</emphasis>
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: Aedeagus (
|
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<figureCitation id="1309F8E6FF99DF793E00FA7DFBFBFA58" box="[1049,1117,1425,1449]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="7" pageNumber="146">Fig. 7</figureCitation>
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) about
|
||
<quantity id="4CCA4986FF99DF793EA1FA7DFAA9FA58" box="[1208,1295,1425,1449]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="146" unit="mm" value="0.8">0.8 mm</quantity>
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long; apical lobes in lateral view with strongly S-shaped ventroapical margin and obtusely extended ventrally (
|
||
<figureCitation id="1309F8E6FF99DF793F01FA3FFACAFA1A" box="[1304,1388,1491,1515]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Fig. 14</figureCitation>
|
||
), in ventral and dorsal view (
|
||
<figureCitation id="1309F8E6FF99DF793E5CFA18FB11F9FD" box="[1093,1207,1524,1548]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Fig. 15, 16</figureCitation>
|
||
) stout and somewhat narrowed towards clipped apex; lateral lobes about as wide as long (
|
||
<figureCitation id="1309F8E6FF99DF7939CDF9DAFBF6F9BC" box="[980,1104,1589,1614]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Figs 15, 16</figureCitation>
|
||
); dorsal lobe stout with round apex (
|
||
<figureCitation id="1309F8E6FF99DF793966F9BAFC74F99F" box="[895,978,1622,1646]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="7" pageNumber="146">Fig. 16</figureCitation>
|
||
). Abdominal sternite VII (
|
||
<figureCitation id="1309F8E6FF99DF793F11F9BAFAFEF99E" box="[1288,1368,1622,1647]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="7" pageNumber="146">Fig. 21</figureCitation>
|
||
) with asetose median depression notably narrowed towards posterior margin. Abdominal sternite VIII (
|
||
<figureCitation id="1309F8E6FF99DF793F08F974FAF8F941" box="[1297,1374,1688,1712]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="7" pageNumber="146">Fig. 20</figureCitation>
|
||
) with median lobe of posterior margin as long as lateroposterior angles; end of median lobe slightly emarginate; four distinct, long, black macrosetae situated close to posterior sternite margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328B7E8FF99DF763956F8D0FF5CFD06" lastPageId="8" lastPageNumber="147" pageId="7" pageNumber="146" type="distribution">
|
||
<paragraph id="8B8DE463FF99DF793956F8D0FAFFF884" blockId="7.[824,1437,767,1976]" pageId="7" pageNumber="146">
|
||
<emphasis id="B9463871FF99DF793956F8D0FC7AF8A5" bold="true" box="[847,988,1852,1876]" pageId="7" pageNumber="146">Distribution</emphasis>
|
||
:
|
||
<taxonomicName id="4C329FE0FF99DF7939F4F8D1FB6BF8A4" authorityName="Frisch & Mainda" authorityYear="2022" box="[1005,1229,1853,1877]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="potamides">
|
||
<emphasis id="B9463871FF99DF7939F4F8D1FB6BF8A4" box="[1005,1229,1853,1877]" italics="true" pageId="7" pageNumber="146">Scopaeus potamides</emphasis>
|
||
</taxonomicName>
|
||
is hitherto known only from Kolombangara Island,
|
||
<collectingCountry id="F325A4F3FF99DF793E86F8B1FAF3F884" box="[1183,1365,1885,1909]" name="Solomon Islands" pageId="7" pageNumber="146">Solomon Islands</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B8DE463FF99DF763956F892FF5CFD06" blockId="7.[824,1437,767,1976]" lastBlockId="8.[151,762,241,1055]" lastPageId="8" lastPageNumber="147" pageId="7" pageNumber="146">
|
||
<emphasis id="B9463871FF99DF793956F892FB8CF867" bold="true" box="[847,1066,1918,1942]" pageId="7" pageNumber="146">Comparative notes</emphasis>
|
||
: Within the
|
||
<taxonomicName id="4C329FE0FF99DF793EACF893FA3AF867" authorityName="Cameron" authorityYear="1938" box="[1205,1436,1918,1942]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="kokodanus">
|
||
<emphasis id="B9463871FF99DF793EACF893FA3AF867" box="[1205,1436,1918,1942]" italics="true" pageId="7" pageNumber="146">Scopaeus kokodanus</emphasis>
|
||
</taxonomicName>
|
||
species group,
|
||
<taxonomicName id="4C329FE0FF99DF7939E5F84CFB3EF846" authorityName="Frisch & Mainda" authorityYear="2022" box="[1020,1176,1951,1975]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="146" phylum="Arthropoda" rank="species" species="potamides">
|
||
<emphasis id="B9463871FF99DF7939E5F84CFB3EF846" box="[1020,1176,1951,1975]" italics="true" pageId="7" pageNumber="146">S. potamides</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1309F8E6FF99DF793EAEF873FAAEF846" box="[1207,1288,1951,1975]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="7" pageNumber="146">Fig. 4</figureCitation>
|
||
) is readily distinguished by the lighter coloring with reddish brown head and pronotum and unicolorous yellowish brown appendages. The species differs from
|
||
<taxonomicName id="4C329FE0FF96DF7638FCFED8FEA8FE9D" authorityName="Cameron" authorityYear="1938" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="kokodanus">
|
||
<emphasis id="B9463871FF96DF7638FCFED8FEA8FE9D" italics="true" pageId="8" pageNumber="147">S. kokodanus</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1309F8E6FF96DF763B07FEB8FEFAFE9D" box="[286,348,340,364]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="8" pageNumber="147">Fig. 1</figureCitation>
|
||
) and
|
||
<taxonomicName id="4C329FE0FF96DF763B84FEB9FDF3FE9D" authorityName="Frisch & Mainda" authorityYear="2022" box="[413,597,340,364]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="arfakmontium">
|
||
<emphasis id="B9463871FF96DF763B84FEB9FDF3FE9D" box="[413,597,340,364]" italics="true" pageId="8" pageNumber="147">S. arfakmontium</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="1309F8E6FF96DF76387CFEB8FD00FE9D" box="[613,678,340,364]" captionStart="Figures 1–4" captionStartId="2.[151,224,1930,1952]" captionTargetBox="[275,1286,249,1823]" captionTargetId="figure-24@2.[151,1398,230,1903]" captionTargetPageId="2" captionText="Figures 1–4. Habitus of Scopaeus kokodanus, Papua New Guinea, Morobe, Waritaian (1), S. arfakmontium, paratype, Indonesia, West Papua,Arfak Mts (2), S. balkei, paratype, Indonesia, Papua, Cyclops Mts (3), S. potamides, paratype, Solomon Islands, Kolombangara (4)." figureDoi="http://doi.org/10.5281/zenodo.13193964" httpUri="https://zenodo.org/record/13193964/files/figure.png" pageId="8" pageNumber="147">Fig. 2</figureCitation>
|
||
) by the round, non-pyriform head. It moreover differs from
|
||
<taxonomicName id="4C329FE0FF96DF7638FCFE99FE95FE5F" authorityName="Frisch & Mainda" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="arfakmontium">
|
||
<emphasis id="B9463871FF96DF7638FCFE99FE95FE5F" italics="true" pageId="8" pageNumber="147">S. arfakmontium</emphasis>
|
||
</taxonomicName>
|
||
by the smaller aedeagus (
|
||
<figureCitation id="1309F8E6FF96DF763874FE7AFD15FE5F" box="[621,691,406,430]" captionStart="Figures 5–7" captionStartId="4.[151,224,1907,1929]" captionTargetBox="[227,1323,999,1870]" captionTargetId="figure-374@4.[151,1398,985,1887]" captionTargetPageId="4" captionText="Figures 5–7. Aedeagus (in lateral view) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (5), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (6), S. potamides, holotype, Solomon Islands, Kolombangara (7). Abbreviations: dl – distal lobes, pb – phallobase." figureDoi="http://doi.org/10.5281/zenodo.13193968" httpUri="https://zenodo.org/record/13193968/files/figure.png" pageId="8" pageNumber="147">Fig. 7</figureCitation>
|
||
) with shorter lateral lobes, tapered apical lobes in dorsal view with characteristic shape in lateral view (
|
||
<figureCitation id="1309F8E6FF96DF763876FE34FD4BFE01" box="[623,749,472,496]" captionStart="Figures 8–16" captionStartId="5.[189,262,1908,1930]" captionTargetBox="[236,1378,299,1853]" captionTargetId="figure-16@5.[189,1436,227,1874]" captionTargetPageId="5" captionText="Figures 8–16. Apical portion of aedeagus in lateral (8, 11, 14), ventral (9, 12, 15), dorsal view (10, 13, 16) of Scopaeus arfakmontium, holotype, Indonesia, West Papua, Arfak Mts (8–10), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (11–13), S. potamides, holotype, Solomon Islands, Kolombangara (14–16). Abbreviations: al – apical lobes, dl – dorsal lobe, f – flagellum, ll – lateral lobe." figureDoi="http://doi.org/10.5281/zenodo.13193970" httpUri="https://zenodo.org/record/13193970/files/figure.png" pageId="8" pageNumber="147">Figs 14–16</figureCitation>
|
||
), round apex of the dorsal lobe and shorter median lobe of the posterior margin of abdominal sternite VIII, which is not longer than the lateroposterior angles of the sternite (
|
||
<figureCitation id="1309F8E6FF96DF763A86FDB7FF4BFD82" box="[159,237,603,627]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="8" pageNumber="147">Fig. 22</figureCitation>
|
||
) and lacks a clearly emarginate tip. In addition,
|
||
<taxonomicName id="4C329FE0FF96DF763A8EFD90FE91FD65" authorityName="Frisch & Mainda" authorityYear="2022" box="[151,311,636,660]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="potamides">
|
||
<emphasis id="B9463871FF96DF763A8EFD90FE91FD65" box="[151,311,636,660]" italics="true" pageId="8" pageNumber="147">S. potamides</emphasis>
|
||
</taxonomicName>
|
||
can be separated from both
|
||
<taxonomicName id="4C329FE0FF96DF7638FCFD90FE95FD44" authorityName="Frisch & Mainda" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="arfakmontium">
|
||
<emphasis id="B9463871FF96DF7638FCFD90FE95FD44" italics="true" pageId="8" pageNumber="147">S. arfakmontium</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C329FE0FF96DF763B7EFD71FE63FD44" authorityName="Frisch & Mainda" authorityYear="2022" box="[359,453,669,693]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="balkei">
|
||
<emphasis id="B9463871FF96DF763B7EFD71FE63FD44" box="[359,453,669,693]" italics="true" pageId="8" pageNumber="147">S. balkei</emphasis>
|
||
</taxonomicName>
|
||
by the posteriorly narrowed asetose median depression of abdominal sternite VII (
|
||
<figureCitation id="1309F8E6FF96DF763A86FD33FF48FD06" box="[159,238,735,759]" captionStart="Figures 17–24" captionStartId="6.[151,224,1885,1907]" captionTargetBox="[222,1362,228,1856]" captionTargetId="figure-24@6.[151,1398,224,1865]" captionTargetPageId="6" captionText="Figures 17–24. Abdominal sternite VII (17, 19, 21) and abdominal sternite VIII (18, 20, 22) of Scopaeus arfakmontium, holotype, Indonesia, West Papua,Arfak Mts (17, 18), S. balkei, holotype, Indonesia, Papua, Cyclops Mts (19, 20), S. potamides, paratype, Solomon Islands, Kolombangara (21, 22), S. kokodanus (23). Figs 23, 24: sperm pumps of (23) S. kokodanus, Papua New Guinea, Morobe, (24) S. balkei, paratype, Indonesia, Papua, Cyclops Mts." figureDoi="http://doi.org/10.5281/zenodo.13193972" httpUri="https://zenodo.org/record/13193972/files/figure.png" pageId="8" pageNumber="147">Fig. 21</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C328B7E8FF96DF763AB7FCECFD62FBEE" pageId="8" pageNumber="147" type="etymology">
|
||
<paragraph id="8B8DE463FF96DF763AB7FCECFD62FBEE" blockId="8.[151,762,241,1055]" pageId="8" pageNumber="147">
|
||
<emphasis id="B9463871FF96DF763AB7FCECFE8DFCE9" bold="true" box="[174,299,768,792]" pageId="8" pageNumber="147">Etymology</emphasis>
|
||
: With the epithet
|
||
<taxonomicName id="4C329FE0FF96DF763BE9FCECFDC1FCE9" authorityName="Frisch & Mainda" authorityYear="2022" box="[496,615,768,792]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="potamides">
|
||
‘
|
||
<emphasis id="B9463871FF96DF763BE1FCECFDC1FCE9" box="[504,615,768,792]" italics="true" pageId="8" pageNumber="147">potamides</emphasis>
|
||
</taxonomicName>
|
||
’ (greek noun Ποταμίδες, genitiv) reference is made to the potamids, the seductive river nymphs of the Greek mythology, because representatives of the genus
|
||
<taxonomicName id="4C329FE0FF96DF76382AFC8FFD3CFC8B" authorityName="Erichson" authorityYear="1839" box="[563,666,867,890]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9463871FF96DF76382AFC8FFD3CFC8B" box="[563,666,867,890]" italics="true" pageId="8" pageNumber="147">Scopaeus</emphasis>
|
||
</taxonomicName>
|
||
are also inhabitants of streams. As every stream is said to have its own potamid according to the Greek mythology, we dedicate
|
||
<taxonomicName id="4C329FE0FF96DF763AE2FC29FE2FFC2C" authorityName="Frisch & Mainda" authorityYear="2022" box="[251,393,965,989]" class="Insecta" family="Staphylinidae" genus="Scopaeus" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="147" phylum="Arthropoda" rank="species" species="potamides">
|
||
<emphasis id="B9463871FF96DF763AE2FC29FE2FFC2C" box="[251,393,965,989]" italics="true" pageId="8" pageNumber="147">S. potamides</emphasis>
|
||
</taxonomicName>
|
||
to the nameless water nymph of the Kolombara River of Kolombaranga Island, the name of which moreover translates as ‘lord of the waters’.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |