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<document ID-DOI="10.1002/tax.604016" ID-GBIF-Dataset="02740591-e945-4cf1-88f5-00c115545489" ID-Zenodo-Dep="7590433" approvedBy="tatia" checkinTime="1675179877152" checkinUser="tatiana" docAuthor="Claßen-Bockhoff, Regine, Oliver, Edward G. H., Hall, Anthony V. &amp; Quint, Marcus" docDate="2019" docId="039B276CFFCBFFCDFCE3FBCDFF6FFEAA" docLanguage="en" docName="Taxon.60.4.1138-1155.pdf" docOrigin="TAXON 60 (4)" docSource="http://dx.doi.org/10.1002/tax.604016" docTitle="Thamnea Brongn." docType="treatment" docVersion="3" lastPageNumber="1145" masterDocId="FFA25F14FFCCFFC5FFBEFFFCFFD2FFF6" masterDocTitle="A new classification of the South African endemic family Bruniaceae based on molecular and morphological data" masterLastPageNumber="1155" masterPageNumber="1138" pageNumber="1145" updateTime="1675183703234" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>A new classification of the South African endemic family Bruniaceae based on molecular and morphological data</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Claßen-Bockhoff, Regine</mods:namePart>
<mods:affiliation>Institut für Spezielle Botanik, Johannes Gutenberg-Universität Mainz, Germany</mods:affiliation>
<mods:nameIdentifier type="email">classenb@uni-mainz.de</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Oliver, Edward G. H.</mods:namePart>
<mods:affiliation>Department of Botany &amp; Zoology, Stellenbosch University, South Africa</mods:affiliation>
<mods:nameIdentifier type="email">eoliver@sun.ac.za</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hall, Anthony V.</mods:namePart>
<mods:affiliation>Bolus Herbarium, University of Cape Town, South Africa</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Quint, Marcus</mods:namePart>
<mods:affiliation>Institut für Spezielle Botanik, Johannes Gutenberg-Universität Mainz, Germany</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>TAXON</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-01-02</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>60</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>4</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1138</mods:start>
<mods:end>1155</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.1002/tax.604016</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">10.1002/tax.604016</mods:identifier>
<mods:identifier type="GBIF-Dataset">02740591-e945-4cf1-88f5-00c115545489</mods:identifier>
<mods:identifier type="Zenodo-Dep">7590433</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.7590474" ID-Zenodo-Dep="7590474" LSID="urn:lsid:plazi:treatment:039B276CFFCBFFCDFCE3FBCDFF6FFEAA" httpUri="http://treatment.plazi.org/id/039B276CFFCBFFCDFCE3FBCDFF6FFEAA" lastPageId="8" lastPageNumber="1145" pageId="7" pageNumber="1145">
<subSubSection box="[861,1183,1072,1097]" pageId="7" pageNumber="1145" type="nomenclature">
<paragraph blockId="7.[860,1505,1072,1971]" box="[861,1183,1072,1097]" pageId="7" pageNumber="1145">
3.
<taxonomicName ID-CoL="7V86" ID-ENA="266476" authority="Sol. ex Brongn." authorityName="Brongn." box="[887,1183,1072,1097]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="genus">
<emphasis bold="true" box="[887,993,1072,1096]" italics="true" pageId="7" pageNumber="1145">Thamnea</emphasis>
Sol. ex Brongn.
</taxonomicName>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1145" type="reference_group">
<paragraph blockId="7.[860,1505,1072,1971]" pageId="7" pageNumber="1145">
in
<bibCitation journal="Ann. Sci. Nat. (Paris)" page="386" pageId="7" pageNumber="1145" year="1826">Ann. Sci. Nat. (Paris) 8: 386, t. 38. 1826</bibCitation>
, nom. cons. (McNeill &amp; al.,
<bibCitation page="405" pageId="7" pageNumber="1145" score="1.97023359760351E-6" volumeTitle="ICBN" year="2006">ICBN: 405. 2006</bibCitation>
) pro
<taxonomicName authority="P. Browne" authorityName="P. Browne" authorityYear="1756" box="[1021,1234,1135,1159]" class="Magnoliopsida" family="Salicaceae" genus="Thamnia" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="genus">
<emphasis box="[1021,1115,1135,1159]" italics="true" pageId="7" pageNumber="1145">Thamnia</emphasis>
P. Browne
</taxonomicName>
(
<taxonomicName box="[1250,1411,1135,1159]" family="Flacourtiaceae" kingdom="Plantae" order="Malpighiales" pageId="7" pageNumber="1145" rank="family">Flacourtiaceae</taxonomicName>
);
<bibCitation author="Pillans" journal="J. S. African Bot." page="143" pageId="7" pageNumber="1145" score="73.24581464374481" volume="13" year="1947">
Pillans in
<bibCitation box="[934,1263,1166,1191]" journal="J. S. African Bot." page="143" pageId="7" pageNumber="1145" score="73.24581464374481" volume="13" year="1947">J. S. African Bot. 13: 143. 1947</bibCitation>
</bibCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1145" type="type_taxon">
<paragraph blockId="7.[860,1505,1072,1971]" pageId="7" pageNumber="1145">
Type:
<taxonomicName class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="7" pageNumber="1145">Thamnea uniflora</emphasis>
(L.) Sol. ex Brongn
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection pageId="7" pageNumber="1145" type="reference_group">
<paragraph blockId="7.[860,1505,1072,1971]" pageId="7" pageNumber="1145">
=
<treatmentCitationGroup pageId="7" pageNumber="1145">
<treatmentCitation author="Engler &amp; Prantl" page="136" pageId="7" pageNumber="1145" year="1891">
<taxonomicName authority="Nied." authorityName="Nied." box="[885,1111,1229,1253]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="genus">
<emphasis box="[885,1047,1229,1253]" italics="true" pageId="7" pageNumber="1145">Pseudobaeckea</emphasis>
Nied.
</taxonomicName>
in Engler &amp; Prantl, Nat. Pflanzenfam. 2a: 136. 1891
</treatmentCitation>
p.p. quoad
<taxonomicName authority="Dummer." box="[1164,1350,1261,1284]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[1164,1243,1261,1284]" italics="true" pageId="7" pageNumber="1145">P. teres</emphasis>
Dümmer.
</taxonomicName>
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection lastPageId="8" lastPageNumber="1146" pageId="7" pageNumber="1145" type="description">
<paragraph blockId="7.[860,1505,1072,1971]" lastBlockId="8.[127,772,199,348]" lastPageId="8" lastPageNumber="1146" pageId="7" pageNumber="1145">
<emphasis box="[906,1055,1291,1315]" italics="true" pageId="7" pageNumber="1145">Growth form:</emphasis>
ericoid dwarf shrubs, prostrate, often in rock crevices; lignotubers present.
<emphasis box="[1236,1318,1323,1346]" italics="true" pageId="7" pageNumber="1145">Leaves:</emphasis>
± imbricate, sessile, slightly appressed; stipules minute (not seen in
<taxonomicName box="[1412,1488,1354,1377]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[1412,1488,1354,1377]" italics="true" pageId="7" pageNumber="1145">T. teres</emphasis>
</taxonomicName>
); hypostomatic (epistomatic in
<taxonomicName box="[1166,1240,1386,1409]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[1166,1240,1386,1409]" italics="true" pageId="7" pageNumber="1145">T. teres</emphasis>
</taxonomicName>
), stomata not surrounded by cuticular rim (except
<taxonomicName box="[1131,1211,1417,1440]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[1131,1211,1417,1440]" italics="true" pageId="7" pageNumber="1145">T. teres</emphasis>
</taxonomicName>
).
<emphasis box="[1235,1329,1416,1440]" italics="true" pageId="7" pageNumber="1145">Flowers:</emphasis>
solitary, sessile (except
<taxonomicName box="[942,1044,1447,1471]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="hirtella">
<emphasis box="[942,1044,1447,1471]" italics="true" pageId="7" pageNumber="1145">T. hirtella</emphasis>
</taxonomicName>
), on bracteate short shoots with 812 bracts (
<taxonomicName box="[869,948,1479,1502]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[869,948,1479,1502]" italics="true" pageId="7" pageNumber="1145">T. teres</emphasis>
</taxonomicName>
) or at top of leafy shoots subtended by uppermost involucre-like leaves (remaining species).
<emphasis box="[1293,1365,1510,1534]" italics="true" pageId="7" pageNumber="1145">Petals:</emphasis>
free, unguiculate, recurved, white, usuallywithpinklowerhalf (“claw”) or with red stains in lower half (
<taxonomicName box="[1173,1249,1573,1596]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[1173,1249,1573,1596]" italics="true" pageId="7" pageNumber="1145">T. teres</emphasis>
</taxonomicName>
), pale pink (
<taxonomicName authorityName="A. V. Hall" authorityYear="2019" baseAuthorityName="A. V. Hall" baseAuthorityYear="2019" box="[1384,1494,1572,1596]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="ustulata">
<emphasis box="[1384,1494,1572,1596]" italics="true" pageId="7" pageNumber="1145">T. ustulata</emphasis>
</taxonomicName>
) or completely pink (
<taxonomicName box="[1081,1236,1604,1627]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="massoniana">
<emphasis box="[1081,1236,1604,1627]" italics="true" pageId="7" pageNumber="1145">T. massoniana</emphasis>
</taxonomicName>
), with 2 separate, rather narrow, parallel ridges not fused at base.
<emphasis box="[1299,1365,1635,1658]" italics="true" pageId="7" pageNumber="1145">Scent:</emphasis>
occasionally present and sweet (
<taxonomicName box="[1068,1222,1666,1689]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="massoniana">
<emphasis box="[1068,1222,1666,1689]" italics="true" pageId="7" pageNumber="1145">T. massoniana</emphasis>
</taxonomicName>
,
<taxonomicName box="[1238,1384,1666,1690]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="diosmoides">
<emphasis box="[1238,1384,1666,1690]" italics="true" pageId="7" pageNumber="1145">T. diosmoides</emphasis>
</taxonomicName>
).
<emphasis box="[1407,1504,1666,1689]" italics="true" pageId="7" pageNumber="1145">Stamens:</emphasis>
included or shortly exserted; anthers linear or linear-oblong, thecae parallel, free only in basal part.
<emphasis box="[1260,1336,1728,1752]" italics="true" pageId="7" pageNumber="1145">Pollen:</emphasis>
3-colporate, except
<taxonomicName authorityName="A. V. Hall" authorityYear="2019" baseAuthorityName="A. V. Hall" baseAuthorityYear="2019" box="[911,1021,1759,1783]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="ustulata">
<emphasis box="[911,1021,1759,1783]" italics="true" pageId="7" pageNumber="1145">T. ustulata</emphasis>
</taxonomicName>
and
<taxonomicName box="[1070,1173,1759,1783]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="hirtella">
<emphasis box="[1070,1173,1759,1783]" italics="true" pageId="7" pageNumber="1145">T. hirtella</emphasis>
</taxonomicName>
with 4 or 5 pollen colpi, tectum granular.
<emphasis box="[962,1037,1791,1814]" italics="true" pageId="7" pageNumber="1145">Ovary:</emphasis>
inferior (up to half-inferior in
<taxonomicName box="[1358,1434,1791,1814]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[1358,1434,1791,1814]" italics="true" pageId="7" pageNumber="1145">T. teres</emphasis>
</taxonomicName>
), 1- or 2-locular with 28 ovules per locule, topped by a ring-shaped nectary (not in
<taxonomicName box="[1024,1157,1853,1877]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="thesioides">
<emphasis box="[1024,1157,1853,1877]" italics="true" pageId="7" pageNumber="1145">T. thesioides</emphasis>
</taxonomicName>
,
<taxonomicName box="[1170,1323,1854,1877]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="massoniana">
<emphasis box="[1170,1323,1854,1877]" italics="true" pageId="7" pageNumber="1145">T. massoniana</emphasis>
</taxonomicName>
, and
<taxonomicName box="[1382,1489,1853,1877]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis box="[1382,1489,1853,1877]" italics="true" pageId="7" pageNumber="1145">T. gracilis</emphasis>
</taxonomicName>
).
<emphasis box="[861,933,1884,1908]" italics="true" pageId="7" pageNumber="1145">Styles:</emphasis>
1 or 2, ± shortly exserted, united (free in
<taxonomicName box="[1408,1489,1885,1908]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="7" pageNumber="1145" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[1408,1489,1885,1908]" italics="true" pageId="7" pageNumber="1145">T. teres</emphasis>
</taxonomicName>
).
<emphasis box="[861,924,1916,1939]" italics="true" pageId="7" pageNumber="1145">Fruit:</emphasis>
1-seeded, dry, indehiscent.
<emphasis box="[1233,1323,1915,1939]" italics="true" pageId="7" pageNumber="1145">Habitat:</emphasis>
on well-drained soils at high altitudes (&gt;1000 m) with moisture-bearing clouds (
<taxonomicName box="[136,216,200,223]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="8" pageNumber="1146" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[136,216,200,223]" italics="true" pageId="8" pageNumber="1146">T. teres</emphasis>
</taxonomicName>
).
<emphasis box="[240,416,199,223]" italics="true" pageId="8" pageNumber="1146">Flowering time:</emphasis>
DecemberFebruary (
<taxonomicName class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="8" pageNumber="1146" phylum="Tracheophyta" rank="species" species="massoniana">
<emphasis italics="true" pageId="8" pageNumber="1146">T. massoniana</emphasis>
</taxonomicName>
), AprilJuly (
<taxonomicName box="[335,438,230,254]" class="Magnoliopsida" family="Bruniaceae" genus="Thamnea" kingdom="Plantae" order="Bruniales" pageId="8" pageNumber="1146" phylum="Tracheophyta" rank="species" species="hirtella">
<emphasis box="[335,438,230,254]" italics="true" pageId="8" pageNumber="1146">T. hirtella</emphasis>
</taxonomicName>
), OctoberNovember (
<taxonomicName box="[681,756,231,254]" class="Magnoliopsida" family="Bruniaceae" genus="Pseudobaeckea" kingdom="Plantae" order="Bruniales" pageId="8" pageNumber="1146" phylum="Tracheophyta" rank="species" species="teres">
<emphasis box="[681,756,231,254]" italics="true" pageId="8" pageNumber="1146">T. teres</emphasis>
</taxonomicName>
), or SeptemberNovember (remaining species).
<emphasis box="[611,698,262,285]" italics="true" pageId="8" pageNumber="1146">Species:</emphasis>
9 spp.; restricted tomountainslopesinsouth-westernpartof Western Cape.
</paragraph>
</subSubSection>
</treatment>
</document>