431 lines
84 KiB
XML
431 lines
84 KiB
XML
<document id="190691134161F9A0F1551CAFE80DF5D6" ID-CLB-Dataset="54924" ID-DOI="10.11646/zootaxa.5131.1.1" ID-GBIF-Dataset="1daf4875-bf0f-4fb9-b648-459a83357801" ID-ISSN="1175-5326" ID-Zenodo-Dep="6521113" ID-ZooBank="CF550031-D6A9-48A3-A953-A1BD40C72F5E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1651751834358" checkinUser="plazi" docAuthor="Grischenko, Andrei V., Gordon, Dennis P., Taylor, Paul D., Kuklinski, Piotr, Denisenko, Nina V., Spencer-Jones, Mary E. & Ostrovsky, Andrew N." docDate="2022" docId="038923740B013304FF73AB5F1B52FEF4" docLanguage="en" docName="zootaxa.5131.1.1.pdf" docOrigin="Zootaxa 5131 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5131.1.1" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Rhamphostomella obliqua Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky 2022, n. sp." docType="treatment" docVersion="9" lastPageNumber="58" masterDocId="FFB05B0C0B37333DFFE4AF611977FF90" masterDocTitle="Taxonomy, ecology and zoogeography of the Recent species of Rhamphostomella Lorenz, 1886 and Mixtoscutella n. gen. (Bryozoa, Cheilostomata)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="55" updateTime="1699329717204" updateUser="ExternalLinkService">
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<mods:title id="9ACAF2CD8DC9899B120E6B9CD4721241">Taxonomy, ecology and zoogeography of the Recent species of Rhamphostomella Lorenz, 1886 and Mixtoscutella n. gen. (Bryozoa, Cheilostomata)</mods:title>
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<mods:roleTerm id="3D56B98ED58F97BCEE7B0D2CAABD5525">Author</mods:roleTerm>
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<mods:namePart id="46BA51C7340EF7FCDBD85739AC767666">Grischenko, Andrei V.</mods:namePart>
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<mods:nameIdentifier id="E1C2FAD9A215A8EB1292E414CF632925" type="email">gat1971@mail.ru</mods:nameIdentifier>
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<mods:namePart id="121AA24A7A9873303AD91CFCA9C35CCA">Gordon, Dennis P.</mods:namePart>
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<mods:nameIdentifier id="F9BF5713F588E199C0AAB999E0163428" type="email">dennis.gordon@niwa.co.nz</mods:nameIdentifier>
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<mods:roleTerm id="D95BF0855A31B2F4B0D8F2C12CE0BCCC">Author</mods:roleTerm>
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<mods:namePart id="41EBF58933E4F9A5B25CF2CB190E3869">Taylor, Paul D.</mods:namePart>
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<mods:nameIdentifier id="226D6955C7B1BDB2A974639FD47B063E" type="email">p.taylor@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="C65764EDB1DD5AF4DEB0E78271B2B652">Kuklinski, Piotr</mods:namePart>
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<mods:nameIdentifier id="8F3A04D359FAC8CF31386C741E44EC4A" type="email">kuki@iopan.gda.pl</mods:nameIdentifier>
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<mods:namePart id="789257C63D91D98D269C8948704595E9">Denisenko, Nina V.</mods:namePart>
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<mods:nameIdentifier id="560ABF66C001992D99B12227282087A9" type="email">ndenisenko@zin.ru</mods:nameIdentifier>
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<mods:namePart id="257C469ADEC7B3D8E193EF6A8A339EF0">Spencer-Jones, Mary E.</mods:namePart>
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<mods:nameIdentifier id="5EE5312DD0A8D92D45CEDECBA6B63453" type="email">m.spencer-jones@nhm.ac.uk</mods:nameIdentifier>
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<mods:namePart id="5B079F0E8F61D013D1A1884050930412">Ostrovsky, Andrew N.</mods:namePart>
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<mods:nameIdentifier id="A7734580B766CD45A16D2603B14ED6C8" type="email">andrei.ostrovsky@univie.ac.at</mods:nameIdentifier>
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<mods:title id="C03DBC3256FEC00C1C9BC9FA5B3D571D">Zootaxa</mods:title>
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<mods:date id="8F26806A0B8319D70F1422D4648C8D5D">2022</mods:date>
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<mods:number id="15A3C37EA6A6B990FE29146D57530815">2022-05-02</mods:number>
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<mods:number id="013B23326A786F51553827B9CD88773B">5131</mods:number>
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<mods:url id="BF46C393252E0206E9FA6E1D4548556B">http://dx.doi.org/10.11646/zootaxa.5131.1.1</mods:url>
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<mods:classification id="3CDCD2BD7EED07D8284D73CA387E45EB">journal article</mods:classification>
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<mods:identifier id="572DFFA197A2CDF8F8BB6338EA9F85E8" type="DOI">10.11646/zootaxa.5131.1.1</mods:identifier>
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<mods:identifier id="A7B6B8D59B0B8C167FBB7B5E6B110FBC" type="Zenodo-Dep">6521113</mods:identifier>
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<mods:identifier id="E5107D40E546B6E294E7C21D2EEDD51F" type="ZooBank">CF550031-D6A9-48A3-A953-A1BD40C72F5E</mods:identifier>
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<treatment id="038923740B013304FF73AB5F1B52FEF4" ID-DOI="http://doi.org/10.5281/zenodo.6520697" ID-GBIF-Taxon="209285947" ID-Zenodo-Dep="6520697" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4" httpUri="http://treatment.plazi.org/id/038923740B013304FF73AB5F1B52FEF4" lastPageId="57" lastPageNumber="58" pageId="54" pageNumber="55">
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<subSubSection id="C33AC1E90B01330BFF73AB5F1B57FBC9" box="[151,544,1086,1113]" pageId="54" pageNumber="55" type="nomenclature">
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<paragraph id="8B9F92620B01330BFF73AB5F1B57FBC9" blockId="54.[151,544,1086,1150]" box="[151,544,1086,1113]" pageId="54" pageNumber="55">
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<heading id="D0D7250E0B01330BFF73AB5F1B57FBC9" bold="true" box="[151,544,1086,1113]" fontSize="11" level="2" pageId="54" pageNumber="55" reason="4">
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<emphasis id="B9544E700B01330BFF73AB5F1B57FBC9" bold="true" box="[151,544,1086,1113]" pageId="54" pageNumber="55">
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<taxonomicName id="4C20E9E10B01330BFF73AB5F18A1FBC9" authority="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky, 2022" authorityName="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky" authorityYear="2022" box="[151,470,1086,1113]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="54" pageNumber="55" phylum="Bryozoa" rank="species" species="obliqua" status="sp. nov.">
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<emphasis id="B9544E700B01330BFF73AB5F18A1FBC9" bold="true" box="[151,470,1086,1113]" italics="true" pageId="54" pageNumber="55">Rhamphostomella obliqua</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A267F30B0B01330BFE39AB5E1B57FBC9" box="[477,544,1087,1113]" pageId="54" pageNumber="55" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33AC1E90B01330BFF73AB02198DFBEE" box="[151,250,1123,1150]" pageId="54" pageNumber="55" type="description">
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<paragraph id="8B9F92620B01330BFF73AB02198DFBEE" blockId="54.[151,544,1086,1150]" box="[151,250,1123,1150]" pageId="54" pageNumber="55">
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(
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<figureCitation id="131B8EE70B01330BFF44AB021986FBEE" box="[160,241,1123,1150]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="54" pageNumber="55">Fig. 16</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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||
<subSubSection id="C33AC1E90B01330BFF73ABCB1D78F999" pageId="54" pageNumber="55" type="diagnosis">
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<paragraph id="8B9F92620B01330BFF73ABCB1D78F999" blockId="54.[151,1437,1194,1545]" pageId="54" pageNumber="55">
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<emphasis id="B9544E700B01330BFF73ABCB1865FB54" bold="true" box="[151,274,1194,1220]" pageId="54" pageNumber="55">Diagnosis.</emphasis>
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Colony encrusting, multiserial. Zooids very large, broadly hexagonal. Frontal shield thin-walled, moderately convex, finely granular. Interareolar ridges low, short, reaching sides of suboral avicularian cystid in distal half of zooid. Umbonuloid component large. Primary orifice bell-shaped, slightly longer than wide, with blunt, ill-defined, lateral condyles; proximal margin straight. Secondary orifice conforming to shape of primary orifice, cormidial, with low, thin-walled proximal peristome. No oral spines. Suboral avicularian cystid strongly elevated, with blunt apex, occupying one-quarter to one-third of frontal shield symmetrically, bulbous to conical, coarsely granular, strongly tilted distally, overhanging orifice, facing distolaterally to laterally. Rostrum elongate oval. Crossbar complete. No adventitious avicularia. Ovicells hyperstomial. Ectooecium smooth with sparse circular to slit-like pseudopores, no secondary calcification. Two mural pore chambers in distolateral wall and two multiporous septula in transverse walls. Basal surface of zooids fully calcified, flat, smooth.
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</paragraph>
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</subSubSection>
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<subSubSection id="C33AC1E90B01330BFF73A9571CEAF806" pageId="54" pageNumber="55" type="materials_examined">
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<paragraph id="8B9F92620B01330BFF73A9571B95F92D" blockId="54.[151,1437,1590,2013]" pageId="54" pageNumber="55">
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<emphasis id="B9544E700B01330BFF73A95718F7F9C0" bold="true" box="[151,384,1590,1616]" pageId="54" pageNumber="55">Material examined.</emphasis>
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<materialsCitation id="3B48983F0B01330BFE63A9561B95F92D" ID-GBIF-Occurrence="4027405462" collectingDate="2011-07-17" collectingMethod="Sigsbee trawl" collectionCode="ZIRAS, IMB" collectorName="A. P. Tsurpalo & A. V. Chernyshev" county="Pacific Ocean" elevation="0" latitude="47.056667" location="RV Akademik Oparin" longLatPrecision="112" longitude="152.245" municipality="Kuril Islands" pageId="54" pageNumber="55" specimenCode="1/50541" specimenCount="1" typeStatus="holotype">
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<emphasis id="B9544E700B01330BFE63A9561899F9C1" box="[391,494,1591,1617]" italics="true" pageId="54" pageNumber="55">
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<typeStatus id="549B2CC00B01330BFE63A9561899F9C1" box="[391,494,1591,1617]" pageId="54" pageNumber="55" type="holotype">Holotype</typeStatus>
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</emphasis>
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:
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<collectionCode id="ED310AA70B01330BFE18A9561B3AF9C1" box="[508,589,1591,1617]" collectionName="ZIRAS" pageId="54" pageNumber="55">ZIRAS</collectionCode>
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<specimenCode id="DB863A190B01330BFDB0A9571BC4F9C1" box="[596,691,1590,1617]" pageId="54" pageNumber="55">1/50541</specimenCode>
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, colony encrusting internal surface of broken shell of
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<taxonomicName id="4C20E9E10B01330BFAE9A9561CE0F9C1" box="[1293,1431,1591,1617]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinida" pageId="54" pageNumber="55" phylum="Mollusca" rank="species" species="undetermined">
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<emphasis id="B9544E700B01330BFAE9A9561C18F9C1" box="[1293,1391,1591,1617]" italics="true" pageId="54" pageNumber="55">Chlamys</emphasis>
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sp.
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||
</taxonomicName>
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||
,
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||
<collectionCode id="ED310AA70B01330BFF73A93A19BBF9E5" box="[151,204,1627,1653]" collectionName="IMB" pageId="54" pageNumber="55">IMB</collectionCode>
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Collection,
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<location id="8EFFC4B90B01330BFEBCA93A1B3BF9E5" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B01330BFEBCA93A1B3BF9E5" box="[344,588,1627,1653]" county="Pacific Ocean" latitude="47.056667" longLatPrecision="112" longitude="152.245" municipality="Kuril Islands" name="RV Akademik Oparin" pageId="54" pageNumber="55">
|
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RV
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<emphasis id="B9544E700B01330BFE60A93A1B3BF9E5" box="[388,588,1627,1653]" italics="true" pageId="54" pageNumber="55">Akademik Oparin</emphasis>
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</location>
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, 41st Expedition, Stn 31/26,
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||
<collectingDate id="EFDA4D4A0B01330BFC7CA93A1D5AF9E5" box="[920,1069,1627,1653]" pageId="54" pageNumber="55" value="2011-07-17">
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<date id="FF9EB4A20B01330BFC7CA93A1D5AF9E5" box="[920,1069,1627,1653]" pageId="54" pageNumber="55" value="1926-07-17">17 July 2011</date>
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</collectingDate>
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, eastward from
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<location id="8EFFC4B90B01330BFB00A93A1CEFF9E5" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B01330BFB00A93A1CEFF9E5" box="[1252,1432,1627,1653]" county="Pacific Ocean" latitude="47.056667" longLatPrecision="112" longitude="152.245" municipality="Kuril Islands" name="Simushir Island" pageId="54" pageNumber="55">Simushir Island</location>
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, middle
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<collectingMunicipality id="6BFB08180B01330BFF09A91E18F6F909" box="[237,385,1663,1689]" pageId="54" pageNumber="55">Kuril Islands</collectingMunicipality>
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,
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<collectingCounty id="62FEEAEE0B01330BFE69A91E1B5AF909" box="[397,557,1663,1689]" pageId="54" pageNumber="55">Pacific Ocean</collectingCounty>
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,
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,
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<geoCoordinate id="EE14F4A50B01330BFD26A9E11A31F909" box="[706,838,1663,1691]" degrees="152" direction="east" minutes="13.6" orientation="longitude" pageId="54" pageNumber="55" precision="92" value="152.22667">152°13.6ʹ E</geoCoordinate>
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||
– 47°03.4ʹ N, 152°14.7ʹ E, depth 82–
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<elevation id="000D75510B01330BFB15A91F1C4EF909" box="[1265,1337,1662,1689]" pageId="54" pageNumber="55">
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<quantity id="4CD83F870B01330BFB15A91F1C4EF909" box="[1265,1337,1662,1689]" metricMagnitude="1" metricUnit="m" metricValue="9.85" metricValueMax="11.5" metricValueMin="8.2" pageId="54" pageNumber="55" unit="m" value="98.5" valueMax="115.0" valueMin="82.0">115 m</quantity>
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</elevation>
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,
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||
<collectingMethod id="5261EA750B01330BFAA1A91E19A4F92D" pageId="54" pageNumber="55">Sigsbee trawl</collectingMethod>
|
||
, collectors
|
||
<collectorName id="26D5F7B40B01330BFEB5A9C2189FF92D" box="[337,488,1699,1725]" pageId="54" pageNumber="55">A.P. Tsurpalo</collectorName>
|
||
and
|
||
<collectorName id="26D5F7B40B01330BFDF9A9C21BA9F92D" box="[541,734,1699,1725]" pageId="54" pageNumber="55">A.V. Chernyshev</collectorName>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B01330BFF23A9A6185FF8E1" blockId="54.[151,1437,1590,2013]" pageId="54" pageNumber="55">
|
||
<materialsCitation id="3B48983F0B01330BFF23A9A6187AF8B9" ID-GBIF-Occurrence="4027405435" collectingDate="2011-07-17" collectionCode="NHMUK" collectorName="Pacific Ocean" country="American Samoa" county="Aleutian Islands" elevation="10" latitude="51.77" location="RV Norseman" longLatPrecision="108" longitude="-176.42667" municipality="Andreanof Islands" pageId="54" pageNumber="55" specimenCode="2013.10.21.2" specimenCount="1">
|
||
<collectionCode id="ED310AA70B01330BFF23A9A61846F971" box="[199,305,1735,1761]" collectionName="NHMUK" pageId="54" pageNumber="55">NHMUK</collectionCode>
|
||
<specimenCode id="DB863A190B01330BFEDFA9A618A6F971" box="[315,465,1735,1761]" pageId="54" pageNumber="55">2013.10.21.2</specimenCode>
|
||
, one colony,
|
||
<location id="8EFFC4B90B01330BFD88A9A61A7BF970" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B01330BFD88A9A61A7BF970" box="[620,780,1735,1761]" country="American Samoa" county="Aleutian Islands" latitude="51.77" longLatPrecision="108" longitude="-176.42667" municipality="Andreanof Islands" name="RV Norseman" pageId="54" pageNumber="55">
|
||
RV
|
||
<emphasis id="B9544E700B01330BFD7DA9A61A7BF970" box="[665,780,1735,1760]" italics="true" pageId="54" pageNumber="55">Norseman</emphasis>
|
||
</location>
|
||
, Stn AS–1,
|
||
<date id="FF9EB4A20B01330BFC7FA9A61D44F971" box="[923,1075,1735,1761]" pageId="54" pageNumber="55" value="2011-07-17">
|
||
<collectingDate id="EFDA4D4A0B01330BFC7FA9A61D44F971" box="[923,1075,1735,1761]" pageId="54" pageNumber="55" value="2011-07-17">17 July 2011</collectingDate>
|
||
</date>
|
||
, coastal waters of
|
||
<location id="8EFFC4B90B01330BFAEEA9A61CEFF971" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B01330BFAEEA9A61CEFF971" box="[1290,1432,1735,1761]" country="American Samoa" county="Aleutian Islands" latitude="51.77" longLatPrecision="108" longitude="-176.42667" municipality="Andreanof Islands" name="Adak Island" pageId="54" pageNumber="55">Adak Island</location>
|
||
,
|
||
<collectingMunicipality id="6BFB08180B01330BFF73A98A181CF895" box="[151,363,1771,1797]" pageId="54" pageNumber="55">Andreanof Islands</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62FEEAEE0B01330BFE93A98A1B44F895" box="[375,563,1771,1797]" pageId="54" pageNumber="55">Aleutian Islands</collectingCounty>
|
||
,
|
||
<collectorName id="26D5F7B40B01330BFDA5A98A1B95F895" box="[577,738,1771,1797]" pageId="54" pageNumber="55">Pacific Ocean</collectorName>
|
||
,
|
||
<geoCoordinate id="EE14F4A50B01330BFD14A98D1A18F895" box="[752,879,1771,1799]" degrees="51" direction="north" minutes="46.2" orientation="latitude" pageId="54" pageNumber="55" precision="92" value="51.77">51°46.2ʹ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE14F4A50B01330BFC99A98D1D79F895" box="[893,1038,1771,1799]" degrees="176" direction="west" minutes="25.6" orientation="longitude" pageId="54" pageNumber="55" precision="92" value="-176.42667">176°25.6ʹ W</geoCoordinate>
|
||
, depth
|
||
<quantity id="4CD83F870B01330BFB87A98A1DD6F895" box="[1123,1185,1771,1797]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="10.0">
|
||
<elevation id="000D75510B01330BFB87A98A1DD6F895" box="[1123,1185,1771,1797]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
,
|
||
<collectionCode id="ED310AA70B01330BFB4BA98A1C78F895" box="[1199,1295,1771,1797]" pageId="54" pageNumber="55">SCUBA</collectionCode>
|
||
, collector
|
||
<collectorName id="26D5F7B40B01330BFA6DA98A187AF8B9" pageId="54" pageNumber="55">P. Kuklinski.</collectorName>
|
||
</materialsCitation>
|
||
<materialsCitation id="3B48983F0B01330BFEF2A86E1DB2F8DD" ID-GBIF-Occurrence="4027405447" collectingDate="2011-07-17" collectionCode="NHMUK" collectorName="Pacific Ocean" country="American Samoa" county="Aleutian Islands" elevation="10" latitude="51.77" location="RV Norseman" longLatPrecision="108" longitude="-176.42667" municipality="Andreanof Islands" pageId="54" pageNumber="55" specimenCode="2013.10.21.2" specimenCount="1">
|
||
<collectionCode id="ED310AA70B01330BFEF2A86E18F7F8B9" box="[278,384,1807,1833]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" lsid="urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="54" pageNumber="55" type="Museum">NHMUK</collectionCode>
|
||
2013.10.21.8a, one colony,
|
||
<location id="8EFFC4B90B01330BFD21A86E1A12F8B8" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B01330BFD21A86E1A12F8B8" box="[709,869,1807,1833]" country="American Samoa" county="Aleutian Islands" latitude="51.77" longLatPrecision="108" longitude="-176.42667" municipality="Andreanof Islands" name="RV Norseman" pageId="54" pageNumber="55">
|
||
RV
|
||
<emphasis id="B9544E700B01330BFD16A86E1A12F8B8" box="[754,869,1807,1832]" italics="true" pageId="54" pageNumber="55">Norseman</emphasis>
|
||
</location>
|
||
, Stn AS–1,
|
||
<date id="FF9EB4A20B01330BFC17A86E1DFDF8B9" box="[1011,1162,1807,1833]" pageId="54" pageNumber="55" value="2011-07-17">
|
||
<collectingDate id="EFDA4D4A0B01330BFC17A86E1DFDF8B9" box="[1011,1162,1807,1833]" pageId="54" pageNumber="55" value="2011-07-17">17 July 2011</collectingDate>
|
||
</date>
|
||
, coastal waters of
|
||
<location id="8EFFC4B90B01330BFABBA86E19A9F8DD" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B01330BFABBA86E19A9F8DD" country="American Samoa" county="Aleutian Islands" latitude="51.77" longLatPrecision="108" longitude="-176.42667" municipality="Andreanof Islands" name="Adak Island" pageId="54" pageNumber="55">Adak Island</location>
|
||
,
|
||
<collectingMunicipality id="6BFB08180B01330BFF03A85218C0F8DD" box="[231,439,1843,1869]" pageId="54" pageNumber="55">Andreanof Islands</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62FEEAEE0B01330BFE24A8521B0FF8DD" box="[448,632,1843,1869]" pageId="54" pageNumber="55">Aleutian Islands</collectingCounty>
|
||
,
|
||
<collectorName id="26D5F7B40B01330BFD66A8521A57F8DD" box="[642,800,1843,1869]" pageId="54" pageNumber="55">Pacific Ocean</collectorName>
|
||
,
|
||
<geoCoordinate id="EE14F4A50B01330BFCCFA8551AD1F8DD" box="[811,934,1843,1871]" degrees="51" direction="north" minutes="46.2" orientation="latitude" pageId="54" pageNumber="55" precision="92" value="51.77">51°46.2ʹ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE14F4A50B01330BFC54A8551D4AF8DD" box="[944,1085,1843,1871]" degrees="176" direction="west" minutes="25.6" orientation="longitude" pageId="54" pageNumber="55" precision="92" value="-176.42667">176°25.6ʹ W</geoCoordinate>
|
||
, depth
|
||
<quantity id="4CD83F870B01330BFB6FA8521DB2F8DD" box="[1163,1221,1843,1869]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="10.0">
|
||
<elevation id="000D75510B01330BFB6FA8521DB2F8DD" box="[1163,1221,1843,1869]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="10.0">10 m</elevation>
|
||
</quantity>
|
||
</materialsCitation>
|
||
,
|
||
<collectionCode id="ED310AA70B01330BFB34A8521C47F8DD" box="[1232,1328,1843,1869]" pageId="54" pageNumber="55">SCUBA</collectionCode>
|
||
, collector
|
||
<collectorName id="26D5F7B40B01330BFF73A836185FF8E1" box="[151,296,1879,1905]" pageId="54" pageNumber="55">P. Kuklinski.</collectorName>
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B01330BFF23A81B1CEAF806" blockId="54.[151,1437,1590,2013]" box="[199,1437,1914,1942]" pageId="54" pageNumber="55">
|
||
<emphasis id="B9544E700B01330BFF23A81B18CDF804" bold="true" box="[199,442,1914,1940]" pageId="54" pageNumber="55">Additional material.</emphasis>
|
||
Three specimens. IMB Collection (2011) Stns 31/26, 64/54 (see Appendix 1 for details).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33AC1E90B01330BFF23A8FE18EBF84D" pageId="54" pageNumber="55" type="etymology">
|
||
<paragraph id="8B9F92620B01330BFF23A8FE18EBF84D" blockId="54.[151,1437,1590,2013]" pageId="54" pageNumber="55">
|
||
<emphasis id="B9544E700B01330BFF23A8FE1839F829" bold="true" box="[199,334,1951,1977]" pageId="54" pageNumber="55">Etymology.</emphasis>
|
||
The species name refers to the oblique position of the large subavicularian cystid, which is strongly angled over the orifice.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="DF5FC2EA0B00330AFF73A97D1B77F874" ID-DOI="http://doi.org/10.5281/zenodo.6521168" ID-Zenodo-Dep="6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="55" pageNumber="56" startId="55.[151,250,1564,1588]" targetBox="[246,1341,181,1540]" targetPageId="55">
|
||
<paragraph id="8B9F92620B00330AFF73A97D1B77F874" blockId="55.[151,1437,1564,2020]" pageId="55" pageNumber="56">
|
||
<emphasis id="B9544E700B00330AFF73A97D1854F9A4" bold="true" box="[151,291,1564,1588]" pageId="55" pageNumber="56">FIGURE 16.</emphasis>
|
||
<taxonomicName id="4C20E9E10B00330AFEC9A97D1B4CF9A4" authority="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky, 2022" authorityName="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky" authorityYear="2022" box="[301,571,1564,1588]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="55" pageNumber="56" phylum="Bryozoa" rank="species" species="obliqua" status="sp. nov.">
|
||
<emphasis id="B9544E700B00330AFEC9A97D1B4CF9A4" box="[301,571,1564,1588]" italics="true" pageId="55" pageNumber="56">Rhamphostomella obliqua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9544E700B00330AFDA1A97D1BF5F9A4" bold="true" box="[581,642,1564,1588]" pageId="55" pageNumber="56">
|
||
<taxonomicNameLabel id="A267F30B0B00330AFDA1A97D1BF5F9A4" box="[581,642,1564,1588]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
<emphasis id="B9544E700B00330AFD68A97D1B9EF9A4" box="[652,745,1564,1588]" italics="true" pageId="55" pageNumber="56">Holotype</emphasis>
|
||
, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E. Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C33AC1E90B0F3305FF23AFF61BF5FF45" pageId="56" pageNumber="57" type="materials_examined">
|
||
<paragraph id="8B9F92620B0F3305FF23AFF61BF5FF45" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
<materialsCitation id="3B48983F0B0F3305FF23AFF61BF5FF45" ID-GBIF-Occurrence="4027405495" county="Pacific Ocean" elevation="99" latitude="47.056667" location="Eastward" longLatPrecision="112" longitude="152.245" municipality="Kuril Islands" pageId="56" pageNumber="57" specimenCount="1" typeStatus="holotype">
|
||
<emphasis id="B9544E700B0F3305FF23AFF61810FF20" bold="true" box="[199,359,150,177]" pageId="56" pageNumber="57">
|
||
<typeStatus id="549B2CC00B0F3305FF23AFF61875FF21" box="[199,258,151,177]" pageId="56" pageNumber="57">Type</typeStatus>
|
||
locality.
|
||
</emphasis>
|
||
<location id="8EFFC4B90B0F3305FE94AFF618AFFF21" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B0F3305FE94AFF618AFFF21" box="[368,472,151,177]" county="Pacific Ocean" latitude="47.056667" longLatPrecision="112" longitude="152.245" municipality="Kuril Islands" name="Eastward" pageId="56" pageNumber="57">Eastward</location>
|
||
from
|
||
<location id="8EFFC4B90B0F3305FDC4AFF61BA2FF21" LSID="urn:lsid:plazi:treatment:038923740B013304FF73AB5F1B52FEF4:8EFFC4B90B0F3305FDC4AFF61BA2FF21" box="[544,725,151,177]" county="Pacific Ocean" latitude="47.056667" longLatPrecision="112" longitude="152.245" municipality="Kuril Islands" name="Simushir Island" pageId="56" pageNumber="57">Simushir Island</location>
|
||
, middle
|
||
<collectingMunicipality id="6BFB08180B0F3305FCDDAFF61AB8FF21" box="[825,975,151,177]" pageId="56" pageNumber="57">Kuril Islands</collectingMunicipality>
|
||
,
|
||
<collectingCounty id="62FEEAEE0B0F3305FC39AFF61D0AFF21" box="[989,1149,151,177]" pageId="56" pageNumber="57">Pacific Ocean</collectingCounty>
|
||
,
|
||
<geoCoordinate id="EE14F4A50B0F3305FB6FAFF91C7EFF21" box="[1163,1289,151,179]" degrees="47" direction="north" minutes="02.9" orientation="latitude" pageId="56" pageNumber="57" precision="92" value="47.048332">47°02.9ʹ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="EE14F4A50B0F3305FAF2AFF91CEBFF21" box="[1302,1436,151,179]" degrees="152" direction="east" minutes="13.6" orientation="longitude" pageId="56" pageNumber="57" precision="92" value="152.22667">152°13.6ʹ E</geoCoordinate>
|
||
– 47°03.4ʹ N, 152°14.7ʹ E, depth
|
||
<quantity id="4CD83F870B0F3305FDEFAFDB1B09FF45" box="[523,638,186,213]" metricMagnitude="1" metricUnit="m" metricValue="9.85" metricValueMax="11.5" metricValueMin="8.2" pageId="56" pageNumber="57" unit="m" value="98.5" valueMax="115.0" valueMin="82.0">
|
||
<elevation id="000D75510B0F3305FDEFAFDB1B09FF45" box="[523,638,186,213]" metricMagnitude="1" metricUnit="m" metricValue="9.85" metricValueMax="11.5" metricValueMin="8.2" pageId="56" pageNumber="57" unit="m" value="98.5" valueMax="115.0" valueMin="82.0">82–115 m</elevation>
|
||
</quantity>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33AC1E90B0F3305FF23AFBE1BCFF8FB" pageId="56" pageNumber="57" type="description">
|
||
<paragraph id="8B9F92620B0F3305FF23AFBE1A29FEF5" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
<emphasis id="B9544E700B0F3305FF23AFBE1809FF69" bold="true" box="[199,382,223,249]" pageId="56" pageNumber="57">Measurements.</emphasis>
|
||
ZIRAS 1/50541, Simushir Island, Kuril Islands, Pacific Ocean (
|
||
<figureCitation id="131B8EE70B0F3305FBB3AFBE1D9FFF6A" box="[1111,1256,223,250]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16A–M</figureCitation>
|
||
). ZL, 1.15–1.92 (1.50 ± 0.15). ZW, 0.65–0.95 (0.81 ± 0.08). ZD, 0.73–0.81 (
|
||
<emphasis id="B9544E700B0F3305FCDBAE621A3AFE8C" box="[831,845,259,284]" italics="true" pageId="56" pageNumber="57">n</emphasis>
|
||
= 2). OrL, 0.34–0.41 (0.38 ± 0.02). OrW, 0.33–0.39 (0.35 ± 0.02). OeL, 0.38–0.48 (0.43 ± 0.03) (
|
||
<emphasis id="B9544E700B0F3305FD7AAE461BDBFED0" box="[670,684,295,320]" italics="true" pageId="56" pageNumber="57">n</emphasis>
|
||
= 15). OeW, 0.45–0.55 (0.48 ± 0.03) (
|
||
<emphasis id="B9544E700B0F3305FB8FAE461D0EFED0" box="[1131,1145,295,320]" italics="true" pageId="56" pageNumber="57">n</emphasis>
|
||
= 15). Av(s)L, 0.21–0.43 (0.33 ± 0.05). P(m)N, 17–27 (22). P(oe)N, 13–21 (18) (
|
||
<emphasis id="B9544E700B0F3305FCE2AE2A1A63FEF4" box="[774,788,331,356]" italics="true" pageId="56" pageNumber="57">n</emphasis>
|
||
= 10).
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23AE0F1AB0FE64" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
<emphasis id="B9544E700B0F3305FF23AE0F182EFE18" bold="true" box="[199,345,366,392]" pageId="56" pageNumber="57">Description.</emphasis>
|
||
Colonies encrusting, multiserial, unilaminar (
|
||
<figureCitation id="131B8EE70B0F3305FC87AE0F1ABBFE19" box="[867,972,366,393]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16A</figureCitation>
|
||
), subcircular, deep brown to light brown when dry; maximal size observed 18 ×
|
||
<quantity id="4CD83F870B0F3305FD87AEF31BCFFE3D" box="[611,696,402,429]" metricMagnitude="-2" metricUnit="m" metricValue="2.0" pageId="56" pageNumber="57" unit="mm" value="20.0">20 mm</quantity>
|
||
. Zooids very large, broadly hexagonal, widest in midline, rarely elongate oval and tapering proximally, arranged in checkered pattern, demarcated by fine sutures between lateral and transverse zooidal walls; sutures less visible in older parts of colony.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23AE9F1A9FFC84" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
Frontal shield umbonuloid (
|
||
<figureCitation id="131B8EE70B0F3305FE1AAE9F1B0DFD89" box="[510,634,510,537]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16A, I</figureCitation>
|
||
), thin-walled, fragile, moderately convex, finely granulated, with single row of mostly elongate areolae along raised margins, separated by low, interareolar ridges; ridges normally less prominent in young zooids (
|
||
<figureCitation id="131B8EE70B0F3305FE32AD271B49FDF1" box="[470,574,582,609]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16A</figureCitation>
|
||
), though sometimes evident in them too (
|
||
<figureCitation id="131B8EE70B0F3305FBF6AD271D0EFDF1" box="[1042,1145,582,609]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16B</figureCitation>
|
||
). In older parts of colony, these ridges, when developed, often connecting to cystid of suboral avicularium and continuing to its apex (
|
||
<figureCitation id="131B8EE70B0F3305FA8BAD081987FD38" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16D, M</figureCitation>
|
||
), but in some instances not very prominent on avicularium (
|
||
<figureCitation id="131B8EE70B0F3305FC6CADEC1D78FD38" box="[904,1039,653,680]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16E–G</figureCitation>
|
||
). Interior of frontal shield (
|
||
<figureCitation id="131B8EE70B0F3305FADEADEC1CE3FD38" box="[1338,1428,653,680]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16I</figureCitation>
|
||
) with very fine ring scar (
|
||
<figureCitation id="131B8EE70B0F3305FE42ADD01B57FD5C" box="[422,544,689,716]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16I, L</figureCitation>
|
||
). Umbonuloid component occupying about 70% of length of frontal shield (68% in one measured zooid), with fine parallel lineation and accretionary banding. In cleaned specimens, semicircle of large pores (lower openings of areolar canals) evident proximal to ring scar.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23AC7C1B80FC33" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
Primary orifice (
|
||
<figureCitation id="131B8EE70B0F3305FE6CAC7C1B7CFCA7" box="[392,523,797,823]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16C, J</figureCitation>
|
||
) bell-shaped, sometimes semioval, slightly longer than wide; distal and lateral margins formed by upper terminal part of distal transverse wall bearing prominent shelf distally (
|
||
<figureCitation id="131B8EE70B0F3305FB09AC2019DFFCEF" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16A–C, E, F</figureCitation>
|
||
) and forming blunt, ill-defined condyles laterally (
|
||
<figureCitation id="131B8EE70B0F3305FD0FAC041A3BFC10" box="[747,844,869,896]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16J</figureCitation>
|
||
). Distal margin of orifice rounded, proximal margin straight, with broadly rounded proximolateral corners.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23ACCD1836FBC6" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
Secondary orifice (
|
||
<figureCitation id="131B8EE70B0F3305FE4CACCD1B45FC57" box="[424,562,940,967]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16A–E</figureCitation>
|
||
) conforming to shape of primary orifice, cormidial; distolateral curvature restricted by thickening of proximal and lateral vertical walls of daughter and neighbouring zooids; proximally bounded by low, thin-walled peristome formed by frontal shield and centrally incorporating cystid of suboral avicularium (
|
||
<figureCitation id="131B8EE70B0F3305FEC9AB7918A1FBA3" box="[301,470,1048,1075]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16D, F, H</figureCitation>
|
||
). In ovicellate zooids, peristome usually reaching proximolateral corners of ooecium. No oral spines.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23AB011ACDFA09" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
Сystid of suboral avicularium occupying distal one-quarter to one-third of zooidal frontal shield, situated mostly symmetrically relative to orifice, bulbous to conical, strongly elevated, coarsely and irregularly thickened on top; surface coarsely granulated, many granules look like sharp spinules, thus contrasting with rest of frontal shield, with 1–5 minute communication pores (
|
||
<figureCitation id="131B8EE70B0F3305FDF8ABAA1BBBFB76" box="[540,716,1227,1254]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16A–H, M</figureCitation>
|
||
). Avicularian cystid strongly angled distally, overhanging orifice in older parts of colony (
|
||
<figureCitation id="131B8EE70B0F3305FE4CAB8E1B78FA9A" box="[424,527,1263,1290]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16H</figureCitation>
|
||
), gradually tapering terminally, with raised vertical ridges uniting into conical tip. Avicularian frontal surface (rostral/postmandibular areas) situated on distolateral slope of cystid, usually crossing zooidal midline, sometimes to one side of it, facing distolaterally to laterally. Rostrum elongate oval, directed proximomedially to proximolaterally and frontally; palate lingulate, foramen elongate oval, bordered by narrow cryptocystal shelf; opesia more or less semicircular. Crossbar complete.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23AAC21881FA2C" blockId="56.[151,1437,150,2042]" box="[199,502,1442,1469]" pageId="56" pageNumber="57">No adventitious avicularia.</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23AAA71C3AF904" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
Ovicells hyperstomial in all parts of colony, ooecium never overgrown by secondary calcification. Ooecium formed by distal autozooid around crescentic slit with communication pore at bottom, situated in proximalmost part of frontal shield very close to distal margin of maternal primary orifice. Ooecial coelomic cavity connected to visceral coelom via communication canal opening on underside of proximal part of frontal shield of distal zooid as small, straight, slit-like communication pore situated halfway between transverse wall and ring scar (
|
||
<figureCitation id="131B8EE70B0F3305FAD5A9371CF8F9E1" box="[1329,1423,1622,1649]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16I</figureCitation>
|
||
). Ooecium with concave proximal margin. Ectooecium smooth, with sparse, circular to slit-like pseudopores.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23A9FC1D0EF96F" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
Zooids interconnected by two mural pore chambers (
|
||
<figureCitation id="131B8EE70B0F3305FCF1A9FC1A08F928" box="[789,895,1693,1720]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16M</figureCitation>
|
||
) in each distolateral wall. Communication pores spread across basal part of transverse walls either as horizontal “band” or forming two multiporous septula. In some zooids, transverse walls distally with two shallow recesses separated by median buttress.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23A8681895F8D7" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
Basal surface of zooids (
|
||
<figureCitation id="131B8EE70B0F3305FE38A8681B35F8B4" box="[476,578,1801,1828]" captionStart="FIGURE 16" captionStartId="55.[151,250,1564,1588]" captionTargetBox="[246,1341,181,1540]" captionTargetId="figure-17@55.[246,1341,181,1540]" captionTargetPageId="55" captionText="FIGURE 16. Rhamphostomella obliqua n. sp. Holotype, ZIRAS 1/50541 (Simushir Island, Kuril Islands, Pacific Ocean). A. Colony margin, showing young zooids with developing ooecia (arrows). B. Oblique view of young zooids, showing early stages of developing ooecia and avicularian cystids (interareolar ridges on the frontal shields are well evident). C. Zooids with fully developed suboral avicularia and early developmental stages of ooecia. D. Oblique view of zooids with fully-developed avicularia (some also with ooecia in early stage of development). E.Autozooids with developing ooecia in young part of colony (interareolar ridges on frontal shield have not yet developed). F. Ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). G. Group of ovicellate zooids in older part of colony (interareolar ridges on frontal shield weakly developed). H. Lateral view of distal half of ovicellate zooid. I. Interior of frontal shield, showing ring scar, areolae and ooecial communication slit (arrow) (openings surrounding ring scar are not pseudopores, but lower openings of areolae). J. Internal view of orifice, showing straight proximal margin and minute condyles. K. Basal surface of colony. L. Interior of frontal shield, showing ring scar and exterior wall microstructure of umbonuloid component. M. Lateral view of zooid, showing suboral avicularium and zooidal lateral wall with mural pore chambers. Scale bars: A, D, G, K, 500 μm; B, C, E, F, H, 250 μm; I, J, 100 μm; L, 50 μm; M, 200 μm." figureDoi="http://doi.org/10.5281/zenodo.6521168" httpUri="https://zenodo.org/record/6521168/files/figure.png" pageId="56" pageNumber="57">Fig. 16K</figureCitation>
|
||
) fully calcified, flat, smooth. Boundaries between some zooids recognizable by intermittent fine incisions.
|
||
</paragraph>
|
||
<paragraph id="8B9F92620B0F3305FF23A8301BCFF8FB" blockId="56.[151,1437,150,2042]" box="[199,696,1873,1899]" pageId="56" pageNumber="57">Ancestrula and early astogeny not observed.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33AC1E90B0F3305FF23A8141A45F86A" pageId="56" pageNumber="57" type="discussion">
|
||
<paragraph id="8B9F92620B0F3305FF23A8141A45F86A" blockId="56.[151,1437,150,2042]" pageId="56" pageNumber="57">
|
||
<emphasis id="B9544E700B0F3305FF23A814184CF81F" bold="true" box="[199,315,1909,1935]" pageId="56" pageNumber="57">Remarks.</emphasis>
|
||
<taxonomicName id="4C20E9E10B0F3305FEA6A81718C0F81F" authority="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky, 2022" authorityName="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky" authorityYear="2022" box="[322,439,1909,1935]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="56" pageNumber="57" phylum="Bryozoa" rank="species" species="obliqua" status="sp. nov.">
|
||
<emphasis id="B9544E700B0F3305FEA6A8171824F81F" box="[322,339,1910,1935]" italics="true" pageId="56" pageNumber="57">R</emphasis>
|
||
.
|
||
<emphasis id="B9544E700B0F3305FE85A81418C0F81F" box="[353,439,1909,1935]" italics="true" pageId="56" pageNumber="57">obliqua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9544E700B0F3305FE59A814188CF81F" bold="true" box="[445,507,1909,1935]" pageId="56" pageNumber="57">
|
||
<taxonomicNameLabel id="A267F30B0B0F3305FE59A814188CF81F" box="[445,507,1909,1935]" pageId="56" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is similar to
|
||
<taxonomicName id="4C20E9E10B0F3305FD69A8171B8FF81F" authority="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky, 2022" authorityName="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky" authorityYear="2022" box="[653,760,1909,1935]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="56" pageNumber="57" phylum="Bryozoa" rank="species" species="aspera" status="sp. nov.">
|
||
<emphasis id="B9544E700B0F3305FD69A8171BE9F81F" box="[653,670,1910,1935]" italics="true" pageId="56" pageNumber="57">R</emphasis>
|
||
.
|
||
<emphasis id="B9544E700B0F3305FD48A8171B8FF81F" box="[684,760,1910,1935]" italics="true" pageId="56" pageNumber="57">aspera</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9544E700B0F3305FD1BA8141A4AF81F" bold="true" box="[767,829,1909,1935]" pageId="56" pageNumber="57">
|
||
<taxonomicNameLabel id="A267F30B0B0F3305FD1BA8141A4AF81F" box="[767,829,1909,1935]" pageId="56" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, but differs from the latter in at least seven characters, described above (see Remarks for
|
||
<taxonomicName id="4C20E9E10B0F3305FDCFA8F81BEEF822" authority="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky, 2022" authorityName="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky" authorityYear="2022" box="[555,665,1945,1971]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="56" pageNumber="57" phylum="Bryozoa" rank="species" species="aspera" status="sp. nov.">
|
||
<emphasis id="B9544E700B0F3305FDCFA8F81B4BF822" box="[555,572,1945,1970]" italics="true" pageId="56" pageNumber="57">R</emphasis>
|
||
.
|
||
<emphasis id="B9544E700B0F3305FDA9A8F81BEEF822" box="[589,665,1945,1970]" italics="true" pageId="56" pageNumber="57">aspera</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9544E700B0F3305FD40A8F81B91F823" bold="true" box="[676,742,1945,1971]" pageId="56" pageNumber="57">
|
||
<taxonomicNameLabel id="A267F30B0B0F3305FD40A8F81B91F823" box="[676,742,1945,1971]" pageId="56" pageNumber="57" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
). This species also superficially resembles
|
||
<taxonomicName id="4C20E9E10B0F3305FB05A8F81C38F823" authorityName=": Androsova" authorityYear="1958" box="[1249,1359,1945,1971]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="56" pageNumber="57" phylum="Bryozoa" rank="species" species="scabra">
|
||
<emphasis id="B9544E700B0F3305FB05A8F81D85F822" box="[1249,1266,1945,1970]" italics="true" pageId="56" pageNumber="57">R</emphasis>
|
||
.
|
||
<emphasis id="B9544E700B0F3305FAE7A8F81C38F823" box="[1283,1359,1945,1971]" italics="true" pageId="56" pageNumber="57">scabra</emphasis>
|
||
</taxonomicName>
|
||
in the conical, prominent suboral avicularian mucro, laterally facing palatal foramen, and lingulate mandible, but differs from the latter in lacking additional adventitious avicularia.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33AC1E90B0E3304FF23AFF61B75FF45" pageId="57" pageNumber="58" type="biology_ecology">
|
||
<paragraph id="8B9F92620B0E3304FF23AFF61B75FF45" blockId="57.[151,1436,150,357]" pageId="57" pageNumber="58">
|
||
<emphasis id="B9544E700B0E3304FF23AFF6185BFF21" bold="true" box="[199,300,151,177]" pageId="57" pageNumber="58">Ecology.</emphasis>
|
||
<taxonomicName id="4C20E9E10B0E3304FED7AFF61B2DFF21" authority="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky, 2022" authorityName="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky" authorityYear="2022" box="[307,602,151,177]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="57" pageNumber="58" phylum="Bryozoa" rank="species" species="obliqua" status="sp. nov.">
|
||
<emphasis id="B9544E700B0E3304FED7AFF61B2DFF21" box="[307,602,151,177]" italics="true" pageId="57" pageNumber="58">Rhamphostomella obliqua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9544E700B0E3304FD85AFF61BD7FF21" bold="true" box="[609,672,151,177]" pageId="57" pageNumber="58">
|
||
<taxonomicNameLabel id="A267F30B0B0E3304FD85AFF61BD7FF21" box="[609,672,151,177]" pageId="57" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is known from depths of
|
||
<quantity id="4CD83F870B0E3304FC25AFF71D45FF21" box="[961,1074,150,177]" metricMagnitude="2" metricUnit="m" metricValue="2.225" metricValueMax="4.35" metricValueMin="0.1" pageId="57" pageNumber="58" unit="m" value="222.5" valueMax="435.0" valueMin="10.0">10–435 m</quantity>
|
||
on pebbles and broken shells of the bivalve mollusc
|
||
<taxonomicName id="4C20E9E10B0E3304FE9DAFDB1B75FF45" box="[377,514,186,213]" class="Bivalvia" family="Pectinidae" genus="Chlamys" kingdom="Animalia" order="Pectinida" pageId="57" pageNumber="58" phylum="Mollusca" rank="species" species="undetermined">
|
||
<emphasis id="B9544E700B0E3304FE9DAFDB18ACFF44" box="[377,475,186,212]" italics="true" pageId="57" pageNumber="58">Chlamys</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C33AC1E90B0E3304FF23AFBF1B52FEF4" pageId="57" pageNumber="58" type="distribution">
|
||
<paragraph id="8B9F92620B0E3304FF23AFBF1B52FEF4" blockId="57.[151,1436,150,357]" pageId="57" pageNumber="58">
|
||
<emphasis id="B9544E700B0E3304FF23AFBF1816FF68" bold="true" box="[199,353,222,248]" pageId="57" pageNumber="58">Distribution.</emphasis>
|
||
The known distribution is based on three records, including one from the Sea of Okhotsk side of the middle to southern Kuril Islands, another from their Pacific side, and the third from the Pacific side of Adak Island, Andreanof Islands, Aleutian Islands.
|
||
<taxonomicName id="4C20E9E10B0E3304FD3CAE471D76FED0" authority="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky, 2022" authorityName="Grischenko & Gordon & Taylor & Kuklinski & Denisenko & Spencer-Jones & Ostrovsky" authorityYear="2022" box="[728,1025,294,320]" class="Gymnolaemata" family="Umbonulidae" genus="Rhamphostomella" kingdom="Animalia" order="Cheilostomatida" pageId="57" pageNumber="58" phylum="Bryozoa" rank="species" species="obliqua" status="sp. nov.">
|
||
<emphasis id="B9544E700B0E3304FD3CAE471D76FED0" box="[728,1025,294,320]" italics="true" pageId="57" pageNumber="58">Rhamphostomella obliqua</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9544E700B0E3304FBEEAE471D3CFED0" bold="true" box="[1034,1099,294,320]" pageId="57" pageNumber="58">
|
||
<taxonomicNameLabel id="A267F30B0B0E3304FBEEAE471D3CFED0" box="[1034,1099,294,320]" pageId="57" pageNumber="58" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is thus a Pacific high-boreal, sublittoral to upper bathyal species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |