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<document id="E6B4F836FE734AF8C07362E392B211CF" ID-CLB-Dataset="279635" ID-DOI="10.11646/zootaxa.5371.1.1" ID-GBIF-Dataset="572741c5-3248-4ea3-9229-ee389520e8ee" ID-ISSN="1175-5334" ID-Zenodo-Dep="10144644" ID-ZooBank="D60B50D1-280B-4403-9E5B-25C0704A43A1" IM.bibliography_approvedBy="carolina" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="carolina" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="carolina" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="carolina" checkinTime="1700160671039" checkinUser="plazi" docAuthor="Jałoszyński, Paweł" docDate="2023" docId="3E380C57FFFA4A5127ACB630FCA2E393" docLanguage="en" docName="zootaxa.5371.1.1.pdf" docOrigin="Zootaxa 5371 (1)" docSource="https://mapress.com/zt/article/download/zootaxa.5371.1.1/52271" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Scydmaenus (Choleropsis) geniculatus" docType="treatment" docVersion="2" lastPageNumber="19" masterDocId="C201742FFFF54A43273BB26CFFDFE652" masterDocTitle="Scydmaenus Latreille of Australia: Revision of species in subgenera Choleropsis Franz, Kingius Franz, Mascarensia Franz, Parallomicrus Franz, Scottiscydmaenus Franz, and description of Ascydmaenus subgen. n. (Coleoptera, Staphylinidae, Scydmaeninae)" masterLastPageNumber="75" masterPageNumber="1" pageNumber="16" updateTime="1701187519014" updateUser="ExternalLinkService">
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<mods:title id="162AB18616FBCA50A042DF6F55258574">Scydmaenus Latreille of Australia: Revision of species in subgenera Choleropsis Franz, Kingius Franz, Mascarensia Franz, Parallomicrus Franz, Scottiscydmaenus Franz, and description of Ascydmaenus subgen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<treatment id="3E380C57FFFA4A5127ACB630FCA2E393" LSID="urn:lsid:plazi:treatment:3E380C57FFFA4A5127ACB630FCA2E393" httpUri="http://treatment.plazi.org/id/3E380C57FFFA4A5127ACB630FCA2E393" lastPageId="18" lastPageNumber="19" pageId="15" pageNumber="16">
<subSubSection id="FE8BEECAFFFA4A4C27ACB630FD1AE225" box="[151,709,1116,1143]" pageId="15" pageNumber="16" type="nomenclature">
<paragraph id="B62EBD41FFFA4A4C27ACB630FD1AE225" blockId="15.[151,709,1116,1143]" box="[151,709,1116,1143]" pageId="15" pageNumber="16">
<heading id="ED660A2DFFFA4A4C27ACB630FD1AE225" bold="true" box="[151,709,1116,1143]" fontSize="11" level="2" pageId="15" pageNumber="16" reason="4">
<taxonomicName id="7191C6C2FFFA4A4C27ACB630FD1AE225" ID-CoL="79ZTP" authority="(King)" baseAuthorityName="King" box="[151,709,1116,1143]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="geniculatus" subGenus="Choleropsis">
<emphasis id="84E56153FFFA4A4C27ACB630FD1AE225" bold="true" box="[151,709,1116,1143]" pageId="15" pageNumber="16">
<emphasis id="84E56153FFFA4A4C27ACB630FEF0E225" bold="true" box="[151,303,1116,1143]" italics="true" pageId="15" pageNumber="16">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFFA4A4C267BB630FE10E225" bold="true" box="[320,463,1116,1143]" italics="true" pageId="15" pageNumber="16">Choleropsis</emphasis>
)
<emphasis id="84E56153FFFA4A4C26DBB630FDB3E225" bold="true" box="[480,620,1116,1143]" italics="true" pageId="15" pageNumber="16">geniculatus</emphasis>
(King)
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
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<paragraph id="B62EBD41FFFA4A4C27ACB6CDFD91E2E8" blockId="15.[151,776,1185,1338]" box="[151,590,1185,1210]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="96819A6FFFFA4A4C27ACB6CDFD91E2E8" box="[151,590,1185,1210]" pageId="15" pageNumber="16">
<treatmentCitation id="37309B50FFFA4A4C27ACB6CDFD95E2E8" author="King, R. L." box="[151,586,1185,1210]" page="98" pageId="15" pageNumber="16" year="1864">
<taxonomicName id="7191C6C2FFFA4A4C27ACB6CDFD95E2E8" authority="King, 1864: 98" authorityName="King" authorityPageNumber="98" authorityYear="1864" box="[151,586,1185,1210]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFFA4A4C27ACB6CDFE77E2EB" box="[151,424,1185,1209]" italics="true" pageId="15" pageNumber="16">Heterognathus geniculatus</emphasis>
<bibRefCitation id="D200C0B0FFFA4A4C2695B6CDFD95E2E8" author="King, R. L." box="[430,586,1185,1210]" pageId="15" pageNumber="16" pagination="91 - 99" refId="ref51270" refString="King, R. L. (1864) On the Scydmaenides of New South Wales. Transactions of the Entomological Society of New South Wales, 1, 91 - 99, pls. 6 - 7." type="journal article" year="1864">King, 1864: 98</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B62EBD41FFFA4A4C27ACB6ACFD22E28A" blockId="15.[151,776,1185,1338]" box="[151,765,1216,1240]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="96819A6FFFFA4A4C27ACB6ACFD22E28A" box="[151,765,1216,1240]" pageId="15" pageNumber="16">
<taxonomicName id="7191C6C2FFFA4A4C27ACB6ACFD8FE28A" ID-CoL="79ZTP" authority="(King)" baseAuthorityName="King" box="[151,592,1216,1240]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="geniculatus" subGenus="Cholerus">
<emphasis id="84E56153FFFA4A4C27ACB6ACFECBE28A" box="[151,276,1216,1240]" italics="true" pageId="15" pageNumber="16">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFFA4A4C2618B6ACFE5FE28A" box="[291,384,1216,1240]" italics="true" pageId="15" pageNumber="16">Cholerus</emphasis>
)
<emphasis id="84E56153FFFA4A4C26B4B6ACFDDBE28A" box="[399,516,1216,1240]" italics="true" pageId="15" pageNumber="16">geniculatus</emphasis>
(King)
</taxonomicName>
;
<treatmentCitation id="37309B50FFFA4A4C2560B6ACFD26E28A" author="Csiki, E." box="[603,761,1216,1240]" page="72" pageId="15" pageNumber="16" year="1919">
<bibRefCitation id="D200C0B0FFFA4A4C2560B6ACFD26E28A" author="Csiki, E." box="[603,761,1216,1240]" pageId="15" pageNumber="16" pagination="70" refId="ref49615" refString="Csiki, E. (1919) Pars 70: Scydmaenidae. In: Schenkling, S. &amp; Junk, W. (Eds.), Coleopterorum Catalogus. Vol. 7. W. Junk, Ber- lin, pp. 1 - 106." type="book chapter" year="1919">Csiki, 1919: 72</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="B62EBD41FFFA4A4C27ACB6B3FCD7E2A5" blockId="15.[151,776,1185,1338]" box="[151,776,1247,1271]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="96819A6FFFFA4A4C27ACB6B3FCD7E2A5" box="[151,776,1247,1271]" pageId="15" pageNumber="16">
<treatmentCitation id="37309B50FFFA4A4C27ACB6B3FDBCE2A5" author="Franz, H." box="[151,611,1247,1271]" page="288" pageId="15" pageNumber="16" year="1975">
<taxonomicName id="7191C6C2FFFA4A4C27ACB6B3FDBCE2A5" ID-CoL="4W3QW" authority="sensu Franz, 1975: 288" authorityName="sensu Franz" authorityPageNumber="288" authorityYear="1975" box="[151,611,1247,1271]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFFA4A4C27ACB6B3FEADE2A5" box="[151,370,1247,1271]" italics="true" pageId="15" pageNumber="16">Scydmaenus princeps</emphasis>
sensu
<bibRefCitation id="D200C0B0FFFA4A4C268CB6B3FDBCE2A5" author="Franz, H." box="[439,611,1247,1271]" pageId="15" pageNumber="16" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975: 288</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(misidentified).
</treatmentCitationGroup>
</paragraph>
<paragraph id="B62EBD41FFFA4A4C27ACB691FD21E347" blockId="15.[151,776,1185,1338]" box="[151,766,1277,1302]" pageId="15" pageNumber="16">
<treatmentCitationGroup id="96819A6FFFFA4A4C27ACB691FD21E347" box="[151,766,1277,1302]" pageId="15" pageNumber="16">
<treatmentCitation id="37309B50FFFA4A4C27ACB691FD86E344" author="Franz, H." box="[151,601,1277,1302]" page="292" pageId="15" pageNumber="16" year="1975">
<taxonomicName id="7191C6C2FFFA4A4C27ACB691FD86E344" ID-CoL="4W3PP" authority="sensu Franz, 1975: 292" authorityName="sensu Franz" authorityPageNumber="292" authorityYear="1975" box="[151,601,1277,1302]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="84E56153FFFA4A4C27ACB691FEB7E347" box="[151,360,1277,1301]" italics="true" pageId="15" pageNumber="16">Scydmaenus gracilis</emphasis>
sensu
<bibRefCitation id="D200C0B0FFFA4A4C2696B692FD86E344" author="Franz, H." box="[429,601,1277,1302]" pageId="15" pageNumber="16" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975: 292</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(misidentified).
</treatmentCitationGroup>
</paragraph>
<paragraph id="B62EBD41FFFA4A4C27ACB74CFDD7E368" blockId="15.[151,776,1185,1338]" box="[151,520,1311,1338]" pageId="15" pageNumber="16">
(
<figureCitation id="2EAAA1C4FFFA4A4C27A4B74CFEFFE368" box="[159,288,1312,1338]" captionStart-0="FIGURES 20–27" captionStart-1="FIGURES 28–29" captionStartId-0="16.[151,264,1861,1885]" captionStartId-1="17.[151,264,922,946]" captionTargetBox-0="[157,1430,562,1831]" captionTargetBox-1="[151,1435,181,898]" captionTargetId-0="figure-194@16.[151,1436,560,1837]" captionTargetId-1="figure-15@17.[151,1435,181,898]" captionTargetPageId-0="16" captionTargetPageId-1="17" captionText-0="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." captionText-1="FIGURES 28–29. Scydmaenus (Choleropsis) geniculatus (King), doubtful syntype male (SDEI), misidentified as S. princeps and used by Franz (1975) to redescribe S. princeps. Dorsal habitus (28) and anterior view (29). © K. Weissing (SDEI)." figureDoi-0="http://doi.org/10.5281/zenodo.10144656" figureDoi-1="http://doi.org/10.5281/zenodo.10144658" httpUri-0="https://zenodo.org/record/10144656/files/figure.png" httpUri-1="https://zenodo.org/record/10144658/files/figure.png" pageId="15" pageNumber="16">Figs 20–29</figureCitation>
,
<figureCitation id="2EAAA1C4FFFA4A4C2617B773FE95E368" box="[300,330,1311,1338]" captionStart="FIGURES 51–57" captionStartId="26.[151,264,1248,1272]" captionTargetBox="[178,1413,192,1220]" captionTargetId="figure-21@26.[164,1423,181,1224]" captionTargetPageId="26" captionText="FIGURES 51–57. Antennae of Choleropsis in dorsal view, males (51, 53–57) and female (52). Scydmaenus beechmonti Franz (51–52); S. geniculatus (King) (53); S. kurandae Franz (54); S. princeps (King) (55); S. curviclavatus sp. n. (56); S. kroombitanus sp. n. (57)." figureDoi="http://doi.org/10.5281/zenodo.10144668" httpUri="https://zenodo.org/record/10144668/files/figure.png" pageId="15" pageNumber="16">53</figureCitation>
,
<figureCitation id="2EAAA1C4FFFA4A4C266DB74CFE41E368" box="[342,414,1312,1338]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="15" pageNumber="16">60–61</figureCitation>
,
<figureCitation id="2EAAA1C4FFFA4A4C2691B74CFE17E368" box="[426,456,1312,1338]" captionStart="FIGURES 70–75" captionStartId="28.[151,264,1880,1904]" captionTargetBox="[151,1435,885,1857]" captionTargetId="figure-313@28.[151,1435,885,1857]" captionTargetPageId="28" captionText="FIGURES 70–75. Pterothorax of Choleropsis in ventral views, males. Scydmaenus beechmonti Franz (70); S. geniculatus (King) (71); S. kurandae Franz (72); S. princeps (King) (73); S. curviclavatus sp. n. (74); S. kroombitanus sp. n. (75)." figureDoi="http://doi.org/10.5281/zenodo.10144672" httpUri="https://zenodo.org/record/10144672/files/figure.png" pageId="15" pageNumber="16">71</figureCitation>
,
<figureCitation id="2EAAA1C4FFFA4A4C26EFB74CFE20E368" box="[468,511,1312,1338]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="15" pageNumber="16">191</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFFA4A4C27ACB704FC9FE15C" pageId="15" pageNumber="16" type="materials_examined">
<paragraph id="B62EBD41FFFA4A4C27ACB704FAF0E008" blockId="15.[151,1436,1383,2022]" pageId="15" pageNumber="16">
<emphasis id="84E56153FFFA4A4C27ACB704FE79E3D3" bold="true" box="[151,422,1383,1410]" pageId="15" pageNumber="16">Type material studied.</emphasis>
<materialsCitation id="06F9B71CFFFA4A4C2695B704FE95E3BC" collectionCode="AMS" country="Australia" location="King" pageId="15" pageNumber="16" specimenCount="2" typeStatus="lectotype">
<emphasis id="84E56153FFFA4A4C2695B704FC1BE3D3" bold="true" box="[430,964,1383,1410]" pageId="15" pageNumber="16">
<typeStatus id="692A03E3FFFA4A4C2695B704FDFBE3D0" box="[430,548,1384,1410]" pageId="15" pageNumber="16" type="lectotype">Lectotype</typeStatus>
(here designated) (
<collectingCountry id="CE86FDD1FFFA4A4C2430B70BFC67E3D3" box="[779,952,1383,1409]" name="Australia" pageId="15" pageNumber="16">AUSTRALIA</collectingCountry>
):
</emphasis>
<specimenCount id="A09776C8FFFA4A4C24F7B705FC3EE3D6" box="[972,993,1385,1412]" pageId="15" pageNumber="16">♂</specimenCount>
mounted on card with “
<specimenCount id="A09776C8FFFA4A4C23C2B705FAD4E3D6" box="[1273,1291,1385,1412]" pageId="15" pageNumber="16">♂</specimenCount>
” and “LECTOTYPE” written in pencil (
<figureCitation id="2EAAA1C4FFFA4A4C26E2B7E0FDF4E3F4" box="[473,555,1420,1446]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="15" pageNumber="16">Fig. 20</figureCitation>
), with labels illustrated in
<figureCitation id="2EAAA1C4FFFA4A4C246BB7E0FC6EE3F4" box="[848,945,1420,1446]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="15" pageNumber="16">Fig. 191</figureCitation>
: “K23218” [brownish, handwritten], “
<typeStatus id="692A03E3FFFA4A4C27ACB7DCFF02E398" box="[151,221,1456,1482]" pageId="15" pageNumber="16">SYNTYPE</typeStatus>
/
<taxonomicName id="7191C6C2FFFA4A4C27CCB7DCFE45E398" authorityName="King" authorityYear="1864" box="[247,410,1456,1482]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Heterognathus</taxonomicName>
/ geniculatus /
<location id="B34EEB9AFFFA4A4C2576B7DCFD57E398" LSID="urn:lsid:plazi:treatment:3E380C57FFFA4A5127ACB630FCA2E393:B34EEB9AFFFA4A4C2576B7DCFD57E398" box="[589,648,1456,1482]" country="Australia" name="King" pageId="15" pageNumber="16">King</location>
, 1863 {sic!}” [yellow, printed and handwritten], “K 197785” [white, printed] (
<collectionCode id="D0802584FFFA4A4C263BB7B8FE9DE3BC" box="[256,322,1492,1518]" collectionName="Australia, New South Wales, Sydney, Australian Museum" pageId="15" pageNumber="16">AMS</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="06F9B71CFFFA4A4C266DB7B8FD44E040" collectionCode="AMS" country="Australia" pageId="15" pageNumber="16" specimenCount="2" specimenCount-female="2" typeStatus="paralectotype">
<emphasis id="84E56153FFFA4A4C266DB7B8FDB3E3BF" bold="true" box="[342,620,1491,1518]" pageId="15" pageNumber="16">
<typeStatus id="692A03E3FFFA4A4C266DB7B8FDD9E3BC" box="[342,518,1492,1518]" pageId="15" pageNumber="16" type="paralectotype">Paralectotypes</typeStatus>
(3 exx.).
</emphasis>
<specimenCount id="A09776C8FFFA4A4C254FB7B8FD56E3BC" box="[628,649,1492,1518]" count="1" pageId="15" pageNumber="16" type="female">♀</specimenCount>
, on separate mounting card pinned below lectotype, with same labels (
<emphasis id="84E56153FFFA4A4C279AB794FDEAE043" bold="true" box="[161,565,1527,1554]" pageId="15" pageNumber="16">
but identity of
<specimenCount id="A09776C8FFFA4A4C2665B79BFE72E043" box="[350,429,1527,1553]" pageId="15" pageNumber="16" type="female">female</specimenCount>
uncertain!
</emphasis>
) (
<collectionCode id="D0802584FFFA4A4C256AB794FD4CE040" box="[593,659,1528,1554]" collectionName="Australia, New South Wales, Sydney, Australian Museum" pageId="15" pageNumber="16">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06F9B71CFFFA4A4C2590B795FAABE064" collectionCode="AMS" country="Australia" isTailEnumeration="true" location="King" municipality="Heterognathus" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="A09776C8FFFA4A4C2590B795FD1FE046" box="[683,704,1529,1556]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
, “K23218” [brownish, handwritten], “
<typeStatus id="692A03E3FFFA4A4C2344B794FADFE040" box="[1151,1280,1528,1554]" pageId="15" pageNumber="16" type="syntype">SYNTYPE</typeStatus>
/
<taxonomicName id="7191C6C2FFFA4A4C2225B794FF19E064" authorityName="King" authorityYear="1864" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">
<collectingMunicipality id="564A273BFFFA4A4C2225B794FF19E064" pageId="15" pageNumber="16">Heterognathus</collectingMunicipality>
</taxonomicName>
/ geniculatus /
<location id="B34EEB9AFFFA4A4C264FB470FE70E064" LSID="urn:lsid:plazi:treatment:3E380C57FFFA4A5127ACB630FCA2E393:B34EEB9AFFFA4A4C264FB470FE70E064" box="[372,431,1564,1590]" country="Australia" municipality="Heterognathus" name="King" pageId="15" pageNumber="16">King</location>
, 1863 {sic!}” [yellow, printed and handwritten], “K 197784” [white, printed] (
<collectionCode id="D0802584FFFA4A4C2212B470FAB4E064" box="[1321,1387,1564,1590]" collectionName="Australia, New South Wales, Sydney, Australian Museum" pageId="15" pageNumber="16">AMS</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="06F9B71CFFFA4A4C22BBB471FAF4E008" collectionCode="AMS" country="Australia" pageId="15" pageNumber="16" specimenCount="1" specimenCount-male="1" typeStatus="syntype">
<specimenCount id="A09776C8FFFA4A4C22BBB471FA4AE06A" box="[1408,1429,1565,1592]" pageId="15" pageNumber="16" type="male">♂</specimenCount>
, “K29219” [brownish, handwritten], “K 197787” [white, printed], “
<typeStatus id="692A03E3FFFA4A4C24B8B42CFBD9E008" box="[899,1030,1600,1626]" pageId="15" pageNumber="16" type="syntype">SYNTYPE</typeStatus>
” [yellow, printed] (
<collectionCode id="D0802584FFFA4A4C23DAB42CFAFCE008" box="[1249,1315,1600,1626]" collectionName="Australia, New South Wales, Sydney, Australian Museum" pageId="15" pageNumber="16">AMS</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="B62EBD41FFFA4A4C27FCB40FFC9FE15C" blockId="15.[151,1436,1383,2022]" pageId="15" pageNumber="16">
<emphasis id="84E56153FFFA4A4C27FCB40FFC48E02F" bold="true" box="[199,919,1635,1662]" pageId="15" pageNumber="16">
Additional material studied (7 exx.).
<collectingRegion id="745573A3FFFA4A4C2541B408FC4EE02F" box="[634,913,1635,1662]" country="Australia" name="New South Wales" pageId="15" pageNumber="16">NEW SOUTH WALES</collectingRegion>
:
</emphasis>
2 ♂♂, Illawarra (correctly identified, presumably by Lea) (
<collectionCode id="D0802584FFFA4A4C260FB4E4FEA2E0F0" box="[308,381,1672,1698]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="16">ANIC</collectionCode>
, on permanent loan from Macleay Museum, University of Sydney); 2 ♂♂,
<collectingRegion id="745573A3FFFA4A4C23F6B4E4FA47E0F0" box="[1229,1432,1672,1698]" country="Australia" name="New South Wales" pageId="15" pageNumber="16">New South Wales</collectingRegion>
, Sydney (
<collectionCode id="D0802584FFFA4A4C27C0B4C0FE9BE094" box="[251,324,1708,1734]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="16">ANIC</collectionCode>
, on permanent loan from Macleay Museum, University of Sydney, cPJ); 2 ♂♂,
<collectingRegion id="745573A3FFFA4A4C23F7B4C0FA46E094" box="[1228,1433,1708,1734]" country="Australia" name="New South Wales" pageId="15" pageNumber="16">New South Wales</collectingRegion>
, Ropes Creek (
<collectionCode id="D0802584FFFA4A4C260EB4BCFEA1E0B8" box="[309,382,1744,1770]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="15" pageNumber="16">ANIC</collectionCode>
, on permanent loan from Macleay Museum, University of Sydney); 1 ♂, misidentified as
<taxonomicName id="7191C6C2FFFA4A4C2251B4BCFEA4E15F" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFFA4A4C2251B4BCFEA4E15F" italics="true" pageId="15" pageNumber="16">Heterognathus princeps</emphasis>
</taxonomicName>
by Lea, Glenn Innes, leg. Lea (
<collectionCode id="D0802584FFFA4A4C25E6B498FCEBE15C" box="[733,820,1780,1806]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34982" lsid="urn:lsid:biocol.org:col:34982" name="South Australia Museum" pageId="15" pageNumber="16" type="Museum">SAMA</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFFA4A5327FCB57BFDF1E6AA" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16" type="discussion">
<paragraph id="B62EBD41FFFA4A5327FCB57BFDF1E6AA" blockId="15.[151,1436,1383,2022]" lastBlockId="16.[151,1437,150,537]" lastPageId="16" lastPageNumber="17" pageId="15" pageNumber="16">
<emphasis id="84E56153FFFA4A4C27FCB57BFCCEE160" bold="true" box="[199,785,1815,1842]" pageId="15" pageNumber="16">Specimens studied as high-quality photographs.</emphasis>
Doubtful
<typeStatus id="692A03E3FFFA4A4C24B7B574FC39E160" box="[908,998,1816,1842]" pageId="15" pageNumber="16" type="syntype">syntype</typeStatus>
, ♂ (
<figureCitation id="2EAAA1C4FFFA4A4C2327B574FB7CE160" box="[1052,1187,1816,1842]" captionStart="FIGURES 28–29" captionStartId="17.[151,264,922,946]" captionTargetBox="[151,1435,181,898]" captionTargetId="figure-15@17.[151,1435,181,898]" captionTargetPageId="17" captionText="FIGURES 28–29. Scydmaenus (Choleropsis) geniculatus (King), doubtful syntype male (SDEI), misidentified as S. princeps and used by Franz (1975) to redescribe S. princeps. Dorsal habitus (28) and anterior view (29). © K. Weissing (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.10144658" httpUri="https://zenodo.org/record/10144658/files/figure.png" pageId="15" pageNumber="16">Figs 28–29</figureCitation>
), “Heterocerus {sic!} / princeps / Type RLK / Paramatta” [brownish, handwritten],
<taxonomicName id="7191C6C2FFFA4A4C2461B550FC19E104" box="[858,966,1852,1878]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="princeps">“princeps</taxonomicName>
/ King / N- {unclear}” [brownish, handwritten], “C. Schaufuss 1930” [white, printed], “
<taxonomicName id="7191C6C2FFFA4A4C25FBB50CFC8FE128" authorityName="Jałoszyński" authorityYear="2023" box="[704,848,1888,1914]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
/ (Cholerus) /
<taxonomicName id="7191C6C2FFFA4A4C24C0B50CFB44E128" authority="King" authorityName="King" authorityYear="1864" box="[1019,1179,1888,1914]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="princeps">princeps King</taxonomicName>
/ det.H.Franz” [white, handwritten and printed], “
<taxonomicName id="7191C6C2FFFA4A4C26FFB5E8FDB5E1CC" authorityName="King" authorityYear="1864" box="[452,618,1924,1950]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="genus">Heterognathus</taxonomicName>
/
<taxonomicName id="7191C6C2FFFA4A4C25BBB5E8FCFFE1CC" authority="King" authorityName="King" authorityYear="1864" box="[640,800,1924,1950]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="princeps">princeps King</taxonomicName>
” [white, handwritten], “DEI Müncheberg / Col - 03300” [greenish, printed], “SDEI
<taxonomicName id="7191C6C2FFFA4A4C26F5B5C4FD96E190" authorityName="Linnaeus" authorityYear="1758" box="[462,585,1960,1986]" class="Magnoliopsida" higherTaxonomySource="GBIF" kingdom="Plantae" order="Coleoptera" pageId="15" pageNumber="16" phylum="Tracheophyta" rank="order">Coleoptera</taxonomicName>
/ # 303734” [white, printed], “
<typeStatus id="692A03E3FFFA4A4C2488B5C4FBFFE190" box="[947,1056,1960,1986]" pageId="15" pageNumber="16" type="syntype">Syntypus</typeStatus>
” [red, printed]
<emphasis id="84E56153FFFA4A4C23EEB5CBFA43E1B7" bold="true" pageId="15" pageNumber="16">
(misidentified as
<taxonomicName id="7191C6C2FFFA4A4C27ACB5A0FEFFE1B4" authorityName="King" authorityYear="1864" box="[151,288,1996,2022]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFFA4A4C27ACB5A0FEFFE1B4" bold="true" box="[151,288,1996,2022]" italics="true" pageId="15" pageNumber="16">H. princeps</emphasis>
</taxonomicName>
and used by
<bibRefCitation id="D200C0B0FFFA4A4C26F3B5A0FDBCE1B7" author="Franz, H." box="[456,611,1995,2022]" pageId="15" pageNumber="16" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz (1975)</bibRefCitation>
to redescribe
<taxonomicName id="7191C6C2FFFA4A4C242FB5A7FBCFE1B4" authorityName="sensu Franz" authorityYear="1975" box="[788,1040,1995,2022]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="15" pageNumber="16" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFFA4A4C242FB5A7FBCFE1B4" bold="true" box="[788,1040,1995,2022]" italics="true" pageId="15" pageNumber="16">Scydmaenus princeps</emphasis>
</taxonomicName>
; if this specimen is a
<typeStatus id="692A03E3FFFA4A4C2224B5A0FAA4E1B4" box="[1311,1403,1996,2022]" pageId="15" pageNumber="16" type="syntype">syntype</typeStatus>
of
</emphasis>
<emphasis id="84E56153FFE54A5327ACB2FAFA43E6E2" bold="true" box="[151,1436,150,177]" pageId="16" pageNumber="17">
<taxonomicName id="7191C6C2FFE54A5327ACB2FAFE73E6E2" authorityName="King" authorityYear="1864" box="[151,428,150,177]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE54A5327ACB2FAFE73E6E2" bold="true" box="[151,428,150,177]" italics="true" pageId="16" pageNumber="17">Heterognathus princeps</emphasis>
</taxonomicName>
, then it must be removed from the
<typeStatus id="692A03E3FFE54A532476B2FAFC5FE6E2" box="[845,896,150,176]" pageId="16" pageNumber="17">type</typeStatus>
series because of misidentification; the
<emphasis id="84E56153FFE54A5327ACB2D6FE10E6AA" bold="true" pageId="16" pageNumber="17">
handwriting and style of historical labels suggest that none of them is an original King’s label, so very likely this specimen is not a
<typeStatus id="692A03E3FFE54A532652B2B2FE18E6AA" box="[361,455,222,248]" pageId="16" pageNumber="17" type="syntype">syntype</typeStatus>
)
</emphasis>
</emphasis>
(
<collectionCode id="D0802584FFE54A5326E6B2B2FDFEE6AA" box="[477,545,222,248]" pageId="16" pageNumber="17">SDEI</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE54A5327FCB36EFE52E44B" pageId="16" pageNumber="17" type="diagnosis">
<paragraph id="B62EBD41FFE54A5327FCB36EFE52E44B" blockId="16.[151,1437,150,537]" pageId="16" pageNumber="17">
<emphasis id="84E56153FFE54A5327FCB36EFDB5E74E" bold="true" box="[199,618,258,284]" pageId="16" pageNumber="17">Revised diagnosis (based on male).</emphasis>
Antennomeres 9 and 10 subequal in width, 9 much shorter than 10 and strongly transverse (
<figureCitation id="2EAAA1C4FFE54A5326BBB34AFDD9E712" box="[384,518,294,320]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="16" pageNumber="17">Figs 60–61</figureCitation>
); antennomere 10 strongly elongate and with broad and deep lateral impression (
<figureCitation id="2EAAA1C4FFE54A5327A4B326FF2BE736" box="[159,244,330,356]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="16" pageNumber="17">Fig. 60</figureCitation>
); aedeagus in dorsal view (
<figureCitation id="2EAAA1C4FFE54A532518B326FD7DE736" box="[547,674,330,356]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="16" pageNumber="17">Figs 24, 27</figureCitation>
) with apical margin straight and as wide as about half of total width of median lobe, lateral subapical lobes narrowly subtriangular, each distinctly longer than broad and projecting strongly laterally and weakly proximally, in lateral view (
<figureCitation id="2EAAA1C4FFE54A532419B3FEFC7BE7FE" box="[802,932,402,429]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="16" pageNumber="17">Figs 25, 27</figureCitation>
) distal region of median lobe weakly broadened, with dorsal margin rounded in proximal half and concave in distal half; metaventrite (
<figureCitation id="2EAAA1C4FFE54A5323FEB3DAFAC6E782" box="[1221,1305,438,464]" captionStart="FIGURES 70–75" captionStartId="28.[151,264,1880,1904]" captionTargetBox="[151,1435,885,1857]" captionTargetId="figure-313@28.[151,1435,885,1857]" captionTargetPageId="28" captionText="FIGURES 70–75. Pterothorax of Choleropsis in ventral views, males. Scydmaenus beechmonti Franz (70); S. geniculatus (King) (71); S. kurandae Franz (72); S. princeps (King) (73); S. curviclavatus sp. n. (74); S. kroombitanus sp. n. (75)." figureDoi="http://doi.org/10.5281/zenodo.10144672" httpUri="https://zenodo.org/record/10144672/files/figure.png" pageId="16" pageNumber="17">Fig. 71</figureCitation>
) with deep median impression about as broad as half width of metaventrite and filled with short setae, but impression flanked by much longer setae.
</paragraph>
</subSubSection>
<caption id="E2EEEDC9FFE54A5327ACB529FD8AE1F7" ID-DOI="http://doi.org/10.5281/zenodo.10144656" ID-Zenodo-Dep="10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="16" pageNumber="17" startId="16.[151,264,1861,1885]" targetBox="[157,1430,562,1831]" targetPageId="16" targetType="figure">
<paragraph id="B62EBD41FFE54A5327ACB529FD8AE1F7" blockId="16.[151,1436,1861,1957]" pageId="16" pageNumber="17">
<emphasis id="84E56153FFE54A5327ACB529FE8AE10F" bold="true" box="[151,341,1861,1885]" pageId="16" pageNumber="17">FIGURES 20–27.</emphasis>
<taxonomicName id="7191C6C2FFE54A532660B529FCEEE10F" authority="(King)" baseAuthorityName="King" box="[347,817,1861,1885]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="17" phylum="Arthropoda" rank="species" species="geniculatus" subGenus="Choleropsis">
<emphasis id="84E56153FFE54A532660B529FE07E10F" box="[347,472,1861,1885]" italics="true" pageId="16" pageNumber="17">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFE54A5326DCB529FDBEE10F" box="[487,609,1861,1885]" italics="true" pageId="16" pageNumber="17">Choleropsis</emphasis>
)
<emphasis id="84E56153FFE54A53254BB529FD3AE10F" box="[624,741,1861,1885]" italics="true" pageId="16" pageNumber="17">geniculatus</emphasis>
(King)
</taxonomicName>
, lectotype male (
<emphasis id="84E56153FFE54A5324DBB529FBC0E10F" bold="true" box="[992,1055,1861,1885]" pageId="16" pageNumber="17">20–22</emphasis>
) and non-type male (
<emphasis id="84E56153FFE54A5323C0B529FAE5E10F" bold="true" box="[1275,1338,1861,1885]" pageId="16" pageNumber="17">23–27</emphasis>
). Mounting card with lectotype (
<emphasis id="84E56153FFE54A5326ABB505FE76E1D3" bold="true" box="[400,425,1897,1921]" pageId="16" pageNumber="17">20</emphasis>
); habitus in lateral view (
<emphasis id="84E56153FFE54A53258AB505FD15E1D3" bold="true" box="[689,714,1897,1921]" pageId="16" pageNumber="17">21</emphasis>
); head and antennae in lateral view (
<emphasis id="84E56153FFE54A532378B505FB83E1D3" bold="true" box="[1091,1116,1897,1921]" pageId="16" pageNumber="17">22</emphasis>
); dorsal habitus (
<emphasis id="84E56153FFE54A53222BB505FAF6E1D3" bold="true" box="[1296,1321,1897,1921]" pageId="16" pageNumber="17">23</emphasis>
); aedeagus in dorsal (
<emphasis id="84E56153FFE54A5327C5B5E1FEE2E1F7" bold="true" box="[254,317,1933,1957]" pageId="16" pageNumber="17">24, 26</emphasis>
) and lateral (
<emphasis id="84E56153FFE54A5326FEB5E1FDDBE1F7" bold="true" box="[453,516,1933,1957]" pageId="16" pageNumber="17">25, 27</emphasis>
) views.
</paragraph>
</caption>
<caption id="E2EEEDC9FFE44A5227ACB1F6FA96E584" ID-DOI="http://doi.org/10.5281/zenodo.10144658" ID-Zenodo-Dep="10144658" httpUri="https://zenodo.org/record/10144658/files/figure.png" pageId="17" pageNumber="18" startId="17.[151,264,922,946]" targetBox="[151,1435,181,898]" targetPageId="17" targetType="figure">
<paragraph id="B62EBD41FFE44A5227ACB1F6FA96E584" blockId="17.[151,1436,922,985]" pageId="17" pageNumber="18">
<emphasis id="84E56153FFE44A5227ACB1F6FE8AE5E0" bold="true" box="[151,341,922,946]" pageId="17" pageNumber="18">FIGURES 28–29.</emphasis>
<taxonomicName id="7191C6C2FFE44A522667B1F6FCEBE5E0" authority="(King)" baseAuthorityName="King" box="[348,820,922,946]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="geniculatus" subGenus="Choleropsis">
<emphasis id="84E56153FFE44A522667B1F6FE06E5E0" box="[348,473,922,946]" italics="true" pageId="17" pageNumber="18">Scydmaenus</emphasis>
(
<emphasis id="84E56153FFE44A5226D2B1F6FDBCE5E0" box="[489,611,922,946]" italics="true" pageId="17" pageNumber="18">Choleropsis</emphasis>
)
<emphasis id="84E56153FFE44A522549B1F6FD38E5E0" box="[626,743,922,946]" italics="true" pageId="17" pageNumber="18">geniculatus</emphasis>
(King)
</taxonomicName>
, doubtful syntype male (SDEI), misidentified as
<taxonomicName id="7191C6C2FFE44A522210B1F7FA43E5E0" authorityName="King" authorityYear="1864" box="[1323,1436,923,946]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE44A522210B1F7FA43E5E0" box="[1323,1436,923,946]" italics="true" pageId="17" pageNumber="18">S. princeps</emphasis>
</taxonomicName>
and used by
<bibRefCitation id="D200C0B0FFE44A52262CB1D3FE45E584" author="Franz, H." box="[279,410,958,982]" pageId="17" pageNumber="18" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz (1975)</bibRefCitation>
to redescribe
<taxonomicName id="7191C6C2FFE44A522512B1D3FD46E584" authorityName="King" authorityYear="1864" box="[553,665,959,982]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="18" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE44A522512B1D3FD46E584" box="[553,665,959,982]" italics="true" pageId="17" pageNumber="18">S. princeps</emphasis>
</taxonomicName>
. Dorsal habitus (
<emphasis id="84E56153FFE44A522473B1D2FCBEE584" bold="true" box="[840,865,958,982]" pageId="17" pageNumber="18">28</emphasis>
) and anterior view (
<emphasis id="84E56153FFE44A52230BB1D2FB96E584" bold="true" box="[1072,1097,958,982]" pageId="17" pageNumber="18">29</emphasis>
). © K. Weissing (SDEI).
</paragraph>
</caption>
<subSubSection id="FE8BEECAFFE44A5127FCB668FE7DE49F" lastPageId="18" lastPageNumber="19" pageId="17" pageNumber="18" type="description">
<paragraph id="B62EBD41FFE44A5227FCB668FB99E235" blockId="17.[151,1437,1028,2027]" pageId="17" pageNumber="18">
<emphasis id="84E56153FFE44A5227FCB668FEAAE24C" bold="true" box="[199,373,1028,1054]" pageId="17" pageNumber="18">Redescription.</emphasis>
Body of male (
<figureCitation id="2EAAA1C4FFE44A522512B668FD73E24C" box="[553,684,1028,1054]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="17" pageNumber="18">Figs 21, 23</figureCitation>
) slightly flattened, elongate and slender, BL 11.71–
<quantity id="716910A4FFE44A5223C0B668FABDE24D" box="[1275,1378,1028,1055]" metricMagnitude="-3" metricUnit="m" metricValue="1.98" pageId="17" pageNumber="18" unit="mm" value="1.98">1.98 mm</quantity>
; pigmentation uniformly light to moderately dark brown, appendages indistinctly lighter (difference best visible in darkest specimens); cuticle moderately glossy, covered with vestiture of yellowish setae.
</paragraph>
<paragraph id="B62EBD41FFE44A5227FCB61CFBC3E068" blockId="17.[151,1437,1028,2027]" pageId="17" pageNumber="18">
Head (
<figureCitation id="2EAAA1C4FFE44A52262AB61CFE4EE2D8" box="[273,401,1136,1162]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="17" pageNumber="18">Figs 21–23</figureCitation>
,
<figureCitation id="2EAAA1C4FFE44A5226A7B61CFE3CE2D8" box="[412,483,1136,1162]" captionStart="FIGURES 28–29" captionStartId="17.[151,264,922,946]" captionTargetBox="[151,1435,181,898]" captionTargetId="figure-15@17.[151,1435,181,898]" captionTargetPageId="17" captionText="FIGURES 28–29. Scydmaenus (Choleropsis) geniculatus (King), doubtful syntype male (SDEI), misidentified as S. princeps and used by Franz (1975) to redescribe S. princeps. Dorsal habitus (28) and anterior view (29). © K. Weissing (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.10144658" httpUri="https://zenodo.org/record/10144658/files/figure.png" pageId="17" pageNumber="18">28–29</figureCitation>
) in dorsal view transversely subrectangular, broadest at eyes, HL
<quantity id="716910A4FFE44A522384B61CFABCE2D9" box="[1215,1379,1136,1163]" metricMagnitude="-4" metricUnit="m" metricValue="3.4499999999999997" metricValueMax="3.8" metricValueMin="3.1" pageId="17" pageNumber="18" unit="mm" value="0.345" valueMax="0.38" valueMin="0.31">0.31–0.38 mm</quantity>
, HW
<quantity id="716910A4FFE44A5227ACB6F8FEE2E2FD" box="[151,317,1172,1199]" metricMagnitude="-4" metricUnit="m" metricValue="3.65" metricValueMax="4.0" metricValueMin="3.3" pageId="17" pageNumber="18" unit="mm" value="0.365" valueMax="0.4" valueMin="0.33">0.33–0.40 mm</quantity>
; vertex and frons confluent and weakly convex, posterior margin of vertex distinctly concave, arcuate; tempora slightly more than twice as long as length of eye in dorsal view; supraantennal tubercles indistinct; frons over antennal fossae broadly subtrapezoidal and with straight anterior margin. Eyes small, nearly semicircular in shape, indistinctly emarginate posteriorly and oblique in relation to long axis of head. Punctures on frons and vertex fine, inconspicuous; setae short, sparse, suberect, those on tempora as sparse as those on head dorsum. Genae as sparsely setose as frons and vertex.Antennae (
<figureCitation id="2EAAA1C4FFE44A52258EB724FCD3E330" box="[693,780,1352,1378]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="17" pageNumber="18">Figs 22</figureCitation>
,
<figureCitation id="2EAAA1C4FFE44A52242EB724FCECE330" box="[789,819,1352,1378]" captionStart="FIGURES 28–29" captionStartId="17.[151,264,922,946]" captionTargetBox="[151,1435,181,898]" captionTargetId="figure-15@17.[151,1435,181,898]" captionTargetPageId="17" captionText="FIGURES 28–29. Scydmaenus (Choleropsis) geniculatus (King), doubtful syntype male (SDEI), misidentified as S. princeps and used by Franz (1975) to redescribe S. princeps. Dorsal habitus (28) and anterior view (29). © K. Weissing (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.10144658" httpUri="https://zenodo.org/record/10144658/files/figure.png" pageId="17" pageNumber="18">29</figureCitation>
,
<figureCitation id="2EAAA1C4FFE44A522405B724FC83E331" box="[830,860,1352,1379]" captionStart="FIGURES 51–57" captionStartId="26.[151,264,1248,1272]" captionTargetBox="[178,1413,192,1220]" captionTargetId="figure-21@26.[164,1423,181,1224]" captionTargetPageId="26" captionText="FIGURES 51–57. Antennae of Choleropsis in dorsal view, males (51, 53–57) and female (52). Scydmaenus beechmonti Franz (51–52); S. geniculatus (King) (53); S. kurandae Franz (54); S. princeps (King) (55); S. curviclavatus sp. n. (56); S. kroombitanus sp. n. (57)." figureDoi="http://doi.org/10.5281/zenodo.10144668" httpUri="https://zenodo.org/record/10144668/files/figure.png" pageId="17" pageNumber="18">53</figureCitation>
,
<figureCitation id="2EAAA1C4FFE44A52245CB724FC70E330" box="[871,943,1352,1378]" captionStart="FIGURES 58–69" captionStartId="27.[151,264,1444,1468]" captionTargetBox="[159,1420,197,1409]" captionTargetId="figure-15@27.[151,1436,181,1421]" captionTargetPageId="27" captionText="FIGURES 58–69. Antennal clubs of Choleropsis in dorsal (58, 60, 62, 64, 66, 68) and outer lateral (59, 61, 63, 65, 67, 69) views, males. Scydmaenus beechmonti Franz (58–59); S. geniculatus (King) (60–61); S. kurandae Franz (62–63); S. princeps (King) (64–65); S. curviclavatus sp. n. (66–67); S. kroombitanus sp. n. (68–69)." figureDoi="http://doi.org/10.5281/zenodo.10144670" httpUri="https://zenodo.org/record/10144670/files/figure.png" pageId="17" pageNumber="18">60–61</figureCitation>
) long and slender, AnL
<quantity id="716910A4FFE44A52238EB724FA85E331" box="[1205,1370,1352,1379]" metricMagnitude="-3" metricUnit="m" metricValue="1.065" metricValueMax="1.08" metricValueMin="1.05" pageId="17" pageNumber="18" unit="mm" value="1.065" valueMax="1.08" valueMin="1.05">1.05–1.08 mm</quantity>
; three terminal antennomeres forming sharply delimited club; scape about 3 times as long as broad, distinctly broadening distally; pedicel twice as long as broad; antennomeres 3–4 each slightly elongate, 5 twice as long as broad, 6 nearly symmetrical and about as long as broad, 7 and 8 each distinctly transverse and strongly asymmetrical, 9 strongly asymmetrical, strongly transverse, with oblique lamina on distomesal margin demarcated by deep notch (best visible in
<figureCitation id="2EAAA1C4FFE44A5227D9B790FEECE044" box="[226,307,1532,1558]" captionStart="FIGURES 28–29" captionStartId="17.[151,264,922,946]" captionTargetBox="[151,1435,181,898]" captionTargetId="figure-15@17.[151,1435,181,898]" captionTargetPageId="17" captionText="FIGURES 28–29. Scydmaenus (Choleropsis) geniculatus (King), doubtful syntype male (SDEI), misidentified as S. princeps and used by Franz (1975) to redescribe S. princeps. Dorsal habitus (28) and anterior view (29). © K. Weissing (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.10144658" httpUri="https://zenodo.org/record/10144658/files/figure.png" pageId="17" pageNumber="18">Fig. 29</figureCitation>
), 10 nearly twice as long as 9 and subequal in width, strongly asymmetrical, with broad lateral impression; 11 only as long as 10, nearly twice as long as broad, slightly asymmetrical.
</paragraph>
<paragraph id="B62EBD41FFE44A5227FCB429FF08E165" blockId="17.[151,1437,1028,2027]" pageId="17" pageNumber="18">
Pronotum in dorsal view (
<figureCitation id="2EAAA1C4FFE44A5226D3B428FDE7E00C" box="[488,568,1604,1630]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="17" pageNumber="18">Fig. 23</figureCitation>
) elongate, broadest near anterior third, PL
<quantity id="716910A4FFE44A52232FB428FB67E00D" box="[1044,1208,1604,1631]" metricMagnitude="-4" metricUnit="m" metricValue="5.4" metricValueMax="5.8" metricValueMin="5.0" pageId="17" pageNumber="18" unit="mm" value="0.54" valueMax="0.58" valueMin="0.5">0.50–0.58 mm</quantity>
, PW
<quantity id="716910A4FFE44A5223C8B428FA47E00D" box="[1267,1432,1604,1631]" metricMagnitude="-4" metricUnit="m" metricValue="4.55" metricValueMax="4.8" metricValueMin="4.3" pageId="17" pageNumber="18" unit="mm" value="0.45499999999999996" valueMax="0.48" valueMin="0.43">0.43–0.48 mm</quantity>
; anterior and lateral margins confluent and rounded; posterior corners obtuse-angled and blunt; posterior margin nearly straight; base with narrow but distinct posterior marginal carina and with two pairs of small, distinct, sharply marked and relatively deep antebasal pits, each slightly to distinctly transverse. Pronotal disc covered with fine and inconspicuous punctures; setae similar to those on frons and vertex, short, moderately dense, suberect. Ventrally prothorax with nearly asetose and impunctate hypomera; basisternal region sparsely covered with short recumbent setae.
</paragraph>
<paragraph id="B62EBD41FFE44A5227FCB52CFA42E1F0" blockId="17.[151,1437,1028,2027]" pageId="17" pageNumber="18">
Elytra (
<figureCitation id="2EAAA1C4FFE44A522620B52CFEB4E108" box="[283,363,1856,1882]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="17" pageNumber="18">Fig. 23</figureCitation>
) oval, broadest slightly in front of middle, EL
<quantity id="716910A4FFE44A522448B52CFBC8E109" box="[883,1047,1856,1883]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" metricValueMax="10.3" metricValueMin="9.0" pageId="17" pageNumber="18" unit="mm" value="0.965" valueMax="1.03" valueMin="0.9">0.90–1.03 mm</quantity>
, EW
<quantity id="716910A4FFE44A52236FB52CFB26E109" box="[1108,1273,1856,1883]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="7.0" metricValueMin="6.3" pageId="17" pageNumber="18" unit="mm" value="0.665" valueMax="0.7" valueMin="0.63">0.63–0.70 mm</quantity>
, EI 1.44–1.56. Humeral calli weakly elevated; basal impression on each elytron barely marked; basal elytral foveae lacking; apices separately rounded. Punctures fine and inconspicuous; setae similar to those on pronotum. Hind wings not studied.
</paragraph>
<paragraph id="B62EBD41FFE44A5227FCB5C0FB05E1B9" blockId="17.[151,1437,1028,2027]" pageId="17" pageNumber="18">
Metaventrite (
<figureCitation id="2EAAA1C4FFE44A52265EB5C0FE69E194" box="[357,438,1964,1990]" captionStart="FIGURES 70–75" captionStartId="28.[151,264,1880,1904]" captionTargetBox="[151,1435,885,1857]" captionTargetId="figure-313@28.[151,1435,885,1857]" captionTargetPageId="28" captionText="FIGURES 70–75. Pterothorax of Choleropsis in ventral views, males. Scydmaenus beechmonti Franz (70); S. geniculatus (King) (71); S. kurandae Franz (72); S. princeps (King) (73); S. curviclavatus sp. n. (74); S. kroombitanus sp. n. (75)." figureDoi="http://doi.org/10.5281/zenodo.10144672" httpUri="https://zenodo.org/record/10144672/files/figure.png" pageId="17" pageNumber="18">Fig. 71</figureCitation>
) with large and deep longitudinal median impression about as broad as half width of ventrite, with diffuse lateral margins, filled with short setae and flanked by conspicuously long setae.
</paragraph>
<paragraph id="B62EBD41FFE74A5127FCB2FBFED2E7FF" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="19">
Legs (
<figureCitation id="2EAAA1C4FFE74A512635B2FBFEBFE6E3" box="[270,352,151,177]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="18" pageNumber="19">Fig. 23</figureCitation>
) long and slender; unmodified, except for distal penicillus on mesotibiae and weakly broadened protarsomere 1 and ventral tenent setae distributed on protarsomeres 1–3; protarsi short, tarsomeres 1–4 each about 1.5 × as long as broad, tarsomere 5 3.5 × as long as broad and slightly shorter than tarsomeres 2–4 combined; mesotarsi long, tarsomere 1 nearly 5 × as long as broad and subequal in length to tarsomeres 2–4 combined, tarsomeres 2–4 each about 1.5 × as long as broad, tarsomere 5 nearly 3.5 × as long as broad, subequal in length to tarsomeres 3–4 combined; metatarsi as long as mesotarsi, tarsomere 1 about 3.5 × as long as broad, tarsomeres 2–4 each elongate but decreasing in length distally, tarsomere 5 3.5 × as long as broad, subequal in length to tarsomeres 3–4 combined.
</paragraph>
<paragraph id="B62EBD41FFE74A5127FCB3DBFB77E4FB" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="19">
Aedeagus (
<figureCitation id="2EAAA1C4FFE74A51267DB3DBFE17E783" box="[326,456,439,465]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="18" pageNumber="19">Figs 24–27</figureCitation>
) elongate but not very slender, AeL
<quantity id="716910A4FFE74A512457B3DAFC0BE783" box="[876,980,438,465]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" pageId="18" pageNumber="19" unit="mm" value="0.35">0.35 mm</quantity>
, in dorsal view median lobe broadest at middle and indistinctly constricted in sub-basal and subapical regions; apical region subtrapezoidal, strongly tapering and with straight apical margin as wide as about half of total width of median lobe; lateral subapical lobes prominent and sharply demarcated from median lobe, each subtriangular, distinctly longer than wide, strongly projecting laterally and only slightly proximally; ostium situated in subapical third of median lobe, flagellum with abruptly broadened and asymmetrical vesicular structure; median lobe in lateral view with distal portion delimited by deep submedian constriction, its dorsal surface convex in proximal half and concave in distal half.
</paragraph>
<paragraph id="B62EBD41FFE74A5127FCB0DFFE7DE49F" blockId="18.[151,1437,151,1473]" box="[199,418,691,717]" pageId="18" pageNumber="19">
<emphasis id="84E56153FFE74A5127FCB0DFFEFAE49F" bold="true" box="[199,293,691,717]" pageId="18" pageNumber="19">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE74A5127FCB0BAFB2BE4A3" box="[199,1268,726,753]" pageId="18" pageNumber="19" type="distribution">
<paragraph id="B62EBD41FFE74A5127FCB0BAFB2BE4A3" blockId="18.[151,1437,151,1473]" box="[199,1268,726,753]" pageId="18" pageNumber="19">
<emphasis id="84E56153FFE74A5127FCB0BAFEBEE4A2" bold="true" box="[199,353,726,752]" pageId="18" pageNumber="19">Distribution.</emphasis>
<collectingRegion id="745573A3FFE74A51265CB0BBFE26E4A3" box="[359,505,727,753]" country="Australia" name="Western Australia" pageId="18" pageNumber="19">SE Australia</collectingRegion>
: CE and NE
<collectingRegion id="745573A3FFE74A5125B1B0BBFCD1E4A3" box="[650,782,727,753]" country="Australia" name="Queensland" pageId="18" pageNumber="19">Queensland</collectingRegion>
(most specimens from vicinity of Sydney).
</paragraph>
</subSubSection>
<subSubSection id="FE8BEECAFFE74A5127FCB097FCA2E393" pageId="18" pageNumber="19" type="discussion">
<paragraph id="B62EBD41FFE74A5127FCB097FBBBE297" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="19">
<emphasis id="84E56153FFE74A5127FCB097FEE4E547" bold="true" box="[199,315,763,789]" pageId="18" pageNumber="19">Remarks.</emphasis>
The name
<taxonomicName id="7191C6C2FFE74A51268DB097FDE8E547" box="[438,567,763,789]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE74A51268DB097FDE8E547" box="[438,567,763,789]" italics="true" pageId="18" pageNumber="19">geniculatus</emphasis>
</taxonomicName>
appears in
<bibRefCitation id="D200C0B0FFE74A51258DB097FC98E547" author="Franz, H." box="[694,839,762,789]" pageId="18" pageNumber="19" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz (1975)</bibRefCitation>
on p. 274, in a list of King’s species names originally combined with
<taxonomicName id="7191C6C2FFE74A51267EB173FE35E56B" authorityName="King" authorityYear="1864" box="[325,490,799,825]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="84E56153FFE74A51267EB173FE35E56B" box="[325,490,799,825]" italics="true" pageId="18" pageNumber="19">Heterognathus</emphasis>
</taxonomicName>
. Interestingly,
<bibRefCitation id="D200C0B0FFE74A5125AAB173FCFDE56B" author="Franz, H." box="[657,802,798,825]" pageId="18" pageNumber="19" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz (1975)</bibRefCitation>
did not redescribe this species under its own name, even though he redescribed it twice: on p. 288 misidentified as
<taxonomicName id="7191C6C2FFE74A51240AB12FFC70E50E" authorityName="King" authorityYear="1864" box="[817,943,835,861]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE74A51240AB12FFC70E50E" box="[817,943,835,861]" italics="true" pageId="18" pageNumber="19">S. princeps</emphasis>
</taxonomicName>
, and on p. 292 misidentified as
<taxonomicName id="7191C6C2FFE74A512219B12FFA49E50F" authorityName="sensu Franz" authorityYear="1975" box="[1314,1430,835,861]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="gracilis">
<emphasis id="84E56153FFE74A512219B12FFA49E50F" box="[1314,1430,835,861]" italics="true" pageId="18" pageNumber="19">S. gracilis</emphasis>
</taxonomicName>
. The ‘redescription’ of
<taxonomicName id="7191C6C2FFE74A5126AAB10BFD47E5D3" authority="sensu Franz" authorityName="Franz" box="[401,664,871,897]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE74A5126AAB10BFDD3E5D2" box="[401,524,871,897]" italics="true" pageId="18" pageNumber="19">S. princeps</emphasis>
sensu Franz
</taxonomicName>
was based on a single ‘syntype’ preserved in coll. Schaufuss (SDEI), which is a male (
<figureCitation id="2EAAA1C4FFE74A512669B1E7FE0EE5F7" box="[338,465,907,933]" captionStart="FIGURES 28–29" captionStartId="17.[151,264,922,946]" captionTargetBox="[151,1435,181,898]" captionTargetId="figure-15@17.[151,1435,181,898]" captionTargetPageId="17" captionText="FIGURES 28–29. Scydmaenus (Choleropsis) geniculatus (King), doubtful syntype male (SDEI), misidentified as S. princeps and used by Franz (1975) to redescribe S. princeps. Dorsal habitus (28) and anterior view (29). © K. Weissing (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.10144658" httpUri="https://zenodo.org/record/10144658/files/figure.png" pageId="18" pageNumber="19">Figs 28–29</figureCitation>
) that almost certainly was not a part of the type series (judging from labels). The antennal modification of this male does not agree with the description and illustration given by
<bibRefCitation id="D200C0B0FFE74A5123BBB1C3FADBE59B" author="King, R. L." box="[1152,1284,943,969]" pageId="18" pageNumber="19" pagination="91 - 99" refId="ref51270" refString="King, R. L. (1864) On the Scydmaenides of New South Wales. Transactions of the Entomological Society of New South Wales, 1, 91 - 99, pls. 6 - 7." type="journal article" year="1864">King (1864</bibRefCitation>
: pl. VII 2) for
<taxonomicName id="7191C6C2FFE74A5127ACB1BFFE7BE5BE" box="[151,420,979,1005]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE74A5127ACB1BFFE7BE5BE" box="[151,420,979,1005]" italics="true" pageId="18" pageNumber="19">Heterognathus princeps</emphasis>
</taxonomicName>
, which shows a deep lateral cavity in the antennomere 9, and the antennomere 10 about as long as broad with only a shallow longitudinal impression. The specimen from SDEI does not have such a cavity in the antennomere 9, and its antennomere 10 is distinctly elongate and has a deep lateral impression (
<figureCitation id="2EAAA1C4FFE74A512231B677FA80E267" box="[1290,1375,1051,1077]" captionStart="FIGURES 28–29" captionStartId="17.[151,264,922,946]" captionTargetBox="[151,1435,181,898]" captionTargetId="figure-15@17.[151,1435,181,898]" captionTargetPageId="17" captionText="FIGURES 28–29. Scydmaenus (Choleropsis) geniculatus (King), doubtful syntype male (SDEI), misidentified as S. princeps and used by Franz (1975) to redescribe S. princeps. Dorsal habitus (28) and anterior view (29). © K. Weissing (SDEI)." figureDoi="http://doi.org/10.5281/zenodo.10144658" httpUri="https://zenodo.org/record/10144658/files/figure.png" pageId="18" pageNumber="19">Fig. 29</figureCitation>
). It is
<taxonomicName id="7191C6C2FFE74A5127ACB653FEEAE20B" box="[151,309,1087,1113]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE74A5127ACB653FEEAE20B" box="[151,309,1087,1113]" italics="true" pageId="18" pageNumber="19">S. geniculatus</emphasis>
</taxonomicName>
, with identical antennal modifications as those in
<typeStatus id="692A03E3FFE74A51244BB653FC0DE20B" box="[880,978,1087,1113]" pageId="18" pageNumber="19" type="syntype">syntypes</typeStatus>
of
<taxonomicName id="7191C6C2FFE74A5124C2B653FAF8E20B" authorityName="King" authorityYear="1864" box="[1017,1319,1087,1113]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE74A5124C2B653FAF8E20B" box="[1017,1319,1087,1113]" italics="true" pageId="18" pageNumber="19">Heterognathus geniculatus</emphasis>
</taxonomicName>
preserved at AMS (
<figureCitation id="2EAAA1C4FFE74A5127C6B60FFE8DE22F" box="[253,338,1123,1149]" captionStart="FIGURES 20–27" captionStartId="16.[151,264,1861,1885]" captionTargetBox="[157,1430,562,1831]" captionTargetId="figure-194@16.[151,1436,560,1837]" captionTargetPageId="16" captionText="FIGURES 20–27. Scydmaenus (Choleropsis) geniculatus (King), lectotype male (20–22) and non-type male (23–27). Mounting card with lectotype (20); habitus in lateral view (21); head and antennae in lateral view (22); dorsal habitus (23); aedeagus in dorsal (24, 26) and lateral (25, 27) views." figureDoi="http://doi.org/10.5281/zenodo.10144656" httpUri="https://zenodo.org/record/10144656/files/figure.png" pageId="18" pageNumber="19">Fig. 22</figureCitation>
).
<taxonomicName id="7191C6C2FFE74A51265EB60FFDA7E22F" box="[357,632,1123,1149]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE74A51265EB60FFDA7E22F" box="[357,632,1123,1149]" italics="true" pageId="18" pageNumber="19">Scydmaenus geniculatus</emphasis>
</taxonomicName>
was also misidentified as
<taxonomicName id="7191C6C2FFE74A51249BB60FFBC3E22E" box="[928,1052,1123,1149]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="princeps">
<emphasis id="84E56153FFE74A51249BB60FFBC3E22E" box="[928,1052,1123,1149]" italics="true" pageId="18" pageNumber="19">S. princeps</emphasis>
</taxonomicName>
by Lea (1 ♂ from SAMA). It was certainly possible to confuse species of
<taxonomicName id="7191C6C2FFE74A512576B6EBFD08E2F3" authorityName="Jałoszyński" authorityYear="2023" box="[589,727,1159,1185]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="genus">
<emphasis id="84E56153FFE74A512576B6EBFD08E2F3" box="[589,727,1159,1185]" italics="true" pageId="18" pageNumber="19">Scydmaenus</emphasis>
</taxonomicName>
with modified antennal clubs in times when a stereomicroscope was not invented yet (King), or was still a rare instrument in hands of zoologists (Lea).
</paragraph>
<paragraph id="B62EBD41FFE74A5127FCB6A3FCA2E393" blockId="18.[151,1437,151,1473]" pageId="18" pageNumber="19">
<taxonomicName id="7191C6C2FFE74A5127FCB6A3FE07E2BB" box="[199,472,1231,1257]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE74A5127FCB6A3FE07E2BB" box="[199,472,1231,1257]" italics="true" pageId="18" pageNumber="19">Scydmaenus geniculatus</emphasis>
</taxonomicName>
clearly differs from all remaining species of
<taxonomicName id="7191C6C2FFE74A5124FCB6A3FB92E2BB" box="[967,1101,1231,1257]" pageId="18" pageNumber="21" rank="subGenus" subGenus="Choleropsis">
<emphasis id="84E56153FFE74A5124FCB6A3FB92E2BB" box="[967,1101,1231,1257]" italics="true" pageId="18" pageNumber="19">Choleropsis</emphasis>
</taxonomicName>
in the antennomere 9 strongly transverse, only half as long as antennomere 10, the antennomere 10 massive, strongly elongate, and with a deep and broad lateral impression. In all remaining species the antennomere 9 is at most as broad as long, only slightly shorter than 10, and the antennomere 10 is only slightly elongate. Only in
<taxonomicName id="7191C6C2FFE74A5124D1B757FBAEE307" authorityName="Franz" authorityYear="1975" box="[1002,1137,1339,1365]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="kurandae">
<emphasis id="84E56153FFE74A5124D1B757FBAEE307" box="[1002,1137,1339,1365]" italics="true" pageId="18" pageNumber="19">S. kurandae</emphasis>
</taxonomicName>
the antennomere 10 has a lateral impression, but it is much narrower than that in
<taxonomicName id="7191C6C2FFE74A512429B733FC6EE32B" box="[786,945,1375,1401]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE74A512429B733FC6EE32B" box="[786,945,1375,1401]" italics="true" pageId="18" pageNumber="19">S. geniculatus</emphasis>
</taxonomicName>
. The aedeagus of
<taxonomicName id="7191C6C2FFE74A5123BEB733FAFBE32B" box="[1157,1316,1375,1401]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="19" phylum="Arthropoda" rank="species" species="geniculatus">
<emphasis id="84E56153FFE74A5123BEB733FAFBE32B" box="[1157,1316,1375,1401]" italics="true" pageId="18" pageNumber="19">S. geniculatus</emphasis>
</taxonomicName>
is unique: no other species in this subgenus has the median lobe in dorsal view broadened at middle, and the subapical lateral lobes elongate subtriangular and directed laterally and proximally.
</paragraph>
</subSubSection>
</treatment>
</document>
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