584 lines
86 KiB
XML
584 lines
86 KiB
XML
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<mods:title id="983D3CD16CE8E424D56B50207246CBFD">Scydmaenus Latreille of Australia: Revision of species in subgenera Choleropsis Franz, Kingius Franz, Mascarensia Franz, Parallomicrus Franz, Scottiscydmaenus Franz, and description of Ascydmaenus subgen. n. (Coleoptera, Staphylinidae, Scydmaeninae)</mods:title>
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<mods:namePart id="1B78CE08A7FD018544173A18CEA53CEC">Jałoszyński, Paweł</mods:namePart>
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<paragraph id="B62EBD41FFDB4A6D27ACB3DAFD10E783" blockId="46.[151,719,438,465]" box="[151,719,438,465]" pageId="46" pageNumber="47">
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<taxonomicName id="7191C6C2FFDB4A6D27ACB3DAFD10E783" ID-CoL="4W3X3" authority="Franz" authorityName="Franz" box="[151,719,438,465]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="inflatitibia" subGenus="Parallomicrus">
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<emphasis id="84E56153FFDB4A6D27ACB3DAFD10E783" bold="true" box="[151,719,438,465]" pageId="46" pageNumber="47">
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<emphasis id="84E56153FFDB4A6D27ACB3DAFEF0E783" bold="true" box="[151,303,438,465]" italics="true" pageId="46" pageNumber="47">Scydmaenus</emphasis>
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(
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<emphasis id="84E56153FFDB4A6D267BB3DAFE32E783" bold="true" box="[320,493,438,465]" italics="true" pageId="46" pageNumber="47">Parallomicrus</emphasis>
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)
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<emphasis id="84E56153FFDB4A6D26C5B3DAFDA2E783" bold="true" box="[510,637,438,465]" italics="true" pageId="46" pageNumber="47">inflatitibia</emphasis>
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Franz
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="FE8BEECAFFDB4A6D27ACB397FEA5E465" pageId="46" pageNumber="47" type="reference_group">
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<paragraph id="B62EBD41FFDB4A6D27ACB397FD19E441" blockId="46.[151,710,507,567]" box="[151,710,507,531]" pageId="46" pageNumber="47">
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<treatmentCitationGroup id="96819A6FFFDB4A6D27ACB397FD19E441" box="[151,710,507,531]" pageId="46" pageNumber="47">
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<treatmentCitation id="37309B50FFDB4A6D27ACB397FD1DE441" author="Franz, H." box="[151,706,507,531]" page="284" pageId="46" pageNumber="47" year="1975">
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<taxonomicName id="7191C6C2FFDB4A6D27ACB397FD1DE441" ID-CoL="4W3X3" authority="Franz, 1975: 284" authorityName="Franz" authorityPageNumber="284" authorityYear="1975" box="[151,706,507,531]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="inflatitibia" subGenus="Allomicrus">
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||
<emphasis id="84E56153FFDB4A6D27ACB397FECBE441" box="[151,276,507,531]" italics="true" pageId="46" pageNumber="47">Scydmaenus</emphasis>
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||
(
|
||
<emphasis id="84E56153FFDB4A6D2618B397FE4DE441" box="[291,402,507,531]" italics="true" pageId="46" pageNumber="47">Allomicrus</emphasis>
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||
)
|
||
<emphasis id="84E56153FFDB4A6D269AB397FDD3E441" box="[417,524,507,531]" italics="true" pageId="46" pageNumber="47">inflatitibia</emphasis>
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||
<bibRefCitation id="D200C0B0FFDB4A6D2529B390FD1DE441" author="Franz, H." box="[530,706,507,531]" pageId="46" pageNumber="47" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975: 284</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph id="B62EBD41FFDB4A6D27ACB071FEA5E465" blockId="46.[151,710,507,567]" box="[151,378,541,567]" pageId="46" pageNumber="47">
|
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(
|
||
<figureCitation id="2EAAA1C4FFDB4A6D27A4B071FEE4E465" box="[159,315,541,567]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="46" pageNumber="47">Figs 118–124</figureCitation>
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,
|
||
<figureCitation id="2EAAA1C4FFDB4A6D267CB071FEADE465" box="[327,370,541,567]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="46" pageNumber="47">199</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="FE8BEECAFFDB4A6C27ACB009FDA4E687" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
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<paragraph id="B62EBD41FFDB4A6D27ACB009FDA4E505" blockId="46.[151,1436,613,856]" pageId="46" pageNumber="47">
|
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<emphasis id="84E56153FFDB4A6D27ACB009FE7BE42D" bold="true" box="[151,420,613,639]" pageId="46" pageNumber="47">Type material studied.</emphasis>
|
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<materialsCitation id="06F9B71CFFDB4A6D2697B009FEBEE55D" collectionCode="BNHM" collectorName="N. S. Wales" country="Australia" determinerName="A. M. Lea & H. Franz" location="Botany Bay" pageId="46" pageNumber="47" specimenCount="2" stateProvince="New South Wales" typeStatus="holotype">
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<emphasis id="84E56153FFDB4A6D2697B009FBD8E42D" bold="true" box="[428,1031,613,639]" pageId="46" pageNumber="47">
|
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<typeStatus id="692A03E3FFDB4A6D2697B009FDC6E42D" box="[428,537,613,639]" pageId="46" pageNumber="47" type="holotype">Holotype</typeStatus>
|
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(
|
||
<collectingCountry id="CE86FDD1FFDB4A6D2513B009FD0CE42D" box="[552,723,613,639]" name="Australia" pageId="46" pageNumber="47">AUSTRALIA</collectingCountry>
|
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:
|
||
<collectingRegion id="745573A3FFDB4A6D25E4B009FC25E42D" box="[735,1018,613,639]" country="Australia" name="New South Wales" pageId="46" pageNumber="47">NEW SOUTH WALES</collectingRegion>
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):
|
||
</emphasis>
|
||
♂ (
|
||
<figureCitation id="2EAAA1C4FFDB4A6D2308B009FB4AE42D" box="[1075,1173,613,639]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="46" pageNumber="47">Fig. 120</figureCitation>
|
||
), with labels illustrated in
|
||
<figureCitation id="2EAAA1C4FFDB4A6D278FB0E5FEC9E4F1" box="[180,278,649,675]" captionStart="FIGURES 189–205" captionStartId="71.[151,264,1644,1668]" captionTargetBox="[157,1429,187,1614]" captionTargetId="figure-15@71.[151,1436,181,1620]" captionTargetPageId="71" captionText="FIGURES 189–205. Historical labels of primary type specimens. Lectotype of Heterognathus gracilis King (189); holotype of Scydmaenus beechmonti Franz (190); lectotype of Heterognathus geniculatus King (191); holotype of Scydmaenus kurandae Franz (192); lectotype of Heterognathus princeps King (193); holotype of Scydmaenus marwillumbali Franz [© Lisha Jasper, (BM)] (194); lectotype of Heterognathus carinatus King (195); holotype of Scydmaenus formicarum Franz (196); holotype of Scydmaenus australiensis Franz (197); lectotype of Heterognathus myrmecophilus Lea (198); holotype of Scydmaenus inflatitibia Franz (199); holotype of Scydmaenus clarkianus Franz (200); lectotype of Scydmaenus optatus Sharp (201); holotype of Scydmaenus scotti Franz (202); lectotype of Scydmaenus swanensis Franz (203); holotype of Scydmaenus brachyponerophilus sp. n. (204); syntype of Heterognathus armitagei King [© Natalie Tees (AMS)] (205)." figureDoi="http://doi.org/10.5281/zenodo.10144723" httpUri="https://zenodo.org/record/10144723/files/figure.png" pageId="46" pageNumber="47">Fig. 199</figureCitation>
|
||
: “
|
||
<collectorName id="1B64D897FFDB4A6D2617B0E5FE79E4F1" box="[300,422,649,675]" pageId="46" pageNumber="47">N.S.Wales</collectorName>
|
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, ‘
|
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<location id="B34EEB9AFFDB4A6D26F9B0E6FD98E4F6" LSID="urn:lsid:plazi:treatment:3E380C57FFDB4A7227ACB3DAFA43E7ED:B34EEB9AFFDB4A6D26F9B0E6FD98E4F6" box="[450,583,650,676]" country="Australia" name="Botany Bay" pageId="46" pageNumber="47" stateProvince="New South Wales">Botany Bay</location>
|
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/ 1901. 42” [white with pink line below middle, handwritten], “
|
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<taxonomicName id="7191C6C2FFDB4A6D2227B0E5FF19E495" authorityName="King" authorityYear="1864" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Heterognathus</taxonomicName>
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/
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<determinerName id="43F25185FFDB4A6D27E6B0C2FE9DE49A" box="[221,322,686,712]" pageId="46" pageNumber="47">A.M.Lea</determinerName>
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det.” [white, handwritten and printed], “
|
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<taxonomicName id="7191C6C2FFDB4A6D2429B0C1FC7DE495" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[786,930,685,711]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Scydmaenus</taxonomicName>
|
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/ (
|
||
<taxonomicName id="7191C6C2FFDB4A6D24F8B0C1FBB5E495" authorityName="King" authorityYear="1864" box="[963,1130,685,711]" class="Insecta" family="Staphylinidae" genus="Heterognathus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="genus">Heterognathus</taxonomicName>
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) / inflatitibia / det.
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<determinerName id="43F25185FFDB4A6D227BB0C1FA43E495" box="[1344,1436,685,711]" pageId="46" pageNumber="47">H.Franz</determinerName>
|
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m.” [white, handwritten and printed], “Typus” [red, handwritten], “Holo- / type” [white circle with red margin, printed] (
|
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<collectionCode id="D0802584FFDB4A6D27C4B099FE86E55D" box="[255,345,757,783]" country="China" httpUri="http://grbio.org/cool/p1gh-na36" name="Beijing Natural History Museum" pageId="46" pageNumber="47">BNHM</collectionCode>
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)
|
||
</materialsCitation>
|
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.
|
||
<materialsCitation id="06F9B71CFFDB4A6D2657B099FDA8E505" collectionCode="BNHM" collectorName="N. S. Wales" country="Australia" location="Botany Bay" pageId="46" pageNumber="47" specimenCount="1" specimenCount-female="1" stateProvince="New South Wales" typeStatus="paratype">
|
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<emphasis id="84E56153FFDB4A6D2657B099FE3DE55D" bold="true" box="[364,482,757,783]" pageId="46" pageNumber="47">
|
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<typeStatus id="692A03E3FFDB4A6D2657B099FE02E55D" box="[364,477,757,783]" pageId="46" pageNumber="47" type="paratype">Paratype</typeStatus>
|
||
:
|
||
</emphasis>
|
||
<specimenCount id="A09776C8FFDB4A6D26D2B09AFE21E542" box="[489,510,758,784]" count="1" pageId="46" pageNumber="47" type="female">♀</specimenCount>
|
||
, “
|
||
<collectorName id="1B64D897FFDB4A6D252CB099FD4EE55D" box="[535,657,757,783]" pageId="46" pageNumber="47">N.S.Wales</collectorName>
|
||
, ‘
|
||
<location id="B34EEB9AFFDB4A6D2597B09AFCEEE542" LSID="urn:lsid:plazi:treatment:3E380C57FFDB4A7227ACB3DAFA43E7ED:B34EEB9AFFDB4A6D2597B09AFCEEE542" box="[684,817,758,784]" country="Australia" name="Botany Bay" pageId="46" pageNumber="47" stateProvince="New South Wales">Botany Bay</location>
|
||
/ 1901. 42” [white with pink line below middle, handwritten], “Para- / type” [white circle with yellow margin] {unlabelled by Franz but listed in original description; identity uncertain, see Remarks} (
|
||
<collectionCode id="D0802584FFDB4A6D252EB151FDB0E505" box="[533,623,829,855]" country="China" httpUri="http://grbio.org/cool/p1gh-na36" name="Beijing Natural History Museum" pageId="46" pageNumber="47">BNHM</collectionCode>
|
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)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<caption id="E2EEEDC9FFDB4A6D27ACB413FB18E0E9" ID-DOI="http://doi.org/10.5281/zenodo.10144697" ID-Zenodo-Dep="10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="46" pageNumber="47" startId="46.[151,264,1663,1687]" targetBox="[157,1435,885,1633]" targetPageId="46" targetType="figure">
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<paragraph id="B62EBD41FFDB4A6D27ACB413FB18E0E9" blockId="46.[151,1436,1663,1723]" pageId="46" pageNumber="47">
|
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<emphasis id="84E56153FFDB4A6D27ACB413FEB4E0C5" bold="true" box="[151,363,1663,1687]" pageId="46" pageNumber="47">FIGURES 118–124.</emphasis>
|
||
<taxonomicName id="7191C6C2FFDB4A6D264BB413FC97E0C5" authority="Franz" authorityName="Franz" box="[368,840,1663,1687]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="inflatitibia" subGenus="Parallomicrus">
|
||
<emphasis id="84E56153FFDB4A6D264BB413FE32E0C5" box="[368,493,1663,1687]" italics="true" pageId="46" pageNumber="47">Scydmaenus</emphasis>
|
||
(
|
||
<emphasis id="84E56153FFDB4A6D26C0B413FD52E0C5" box="[507,653,1663,1687]" italics="true" pageId="46" pageNumber="47">Parallomicrus</emphasis>
|
||
)
|
||
<emphasis id="84E56153FFDB4A6D25A0B413FCD9E0C5" box="[667,774,1663,1687]" italics="true" pageId="46" pageNumber="47">inflatitibia</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
, male holotype. Right hind tibia in dorsal (
|
||
<emphasis id="84E56153FFDB4A6D23CFB413FAC6E0C5" bold="true" box="[1268,1305,1663,1687]" pageId="46" pageNumber="47">118</emphasis>
|
||
) and slightly outer lateral (
|
||
<emphasis id="84E56153FFDB4A6D261BB4CFFE9AE0E9" bold="true" box="[288,325,1699,1723]" pageId="46" pageNumber="47">119</emphasis>
|
||
) views; dorsal habitus (
|
||
<emphasis id="84E56153FFDB4A6D2503B4CFFD81E0E9" bold="true" box="[568,606,1699,1723]" pageId="46" pageNumber="47">120</emphasis>
|
||
); aedeagus in dorsal (
|
||
<emphasis id="84E56153FFDB4A6D2406B4CFFC49E0E9" bold="true" box="[829,918,1699,1723]" pageId="46" pageNumber="47">121, 123</emphasis>
|
||
) and lateral (
|
||
<emphasis id="84E56153FFDB4A6D2325B4CFFBA9E0E9" bold="true" box="[1054,1142,1699,1723]" pageId="46" pageNumber="47">122, 124</emphasis>
|
||
) views.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="B62EBD41FFDB4A6D27FCB485FACFE156" blockId="46.[151,1436,1769,2012]" box="[199,1296,1769,1798]" pageId="46" pageNumber="47">
|
||
<emphasis id="84E56153FFDB4A6D27FCB485FCE3E151" bold="true" box="[199,828,1769,1796]" pageId="46" pageNumber="47">
|
||
Additional material studied.
|
||
<collectingRegion id="745573A3FFDB4A6D2524B486FCE8E151" box="[543,823,1769,1796]" country="Australia" name="New South Wales" pageId="46" pageNumber="47">NEW SOUTH WALES</collectingRegion>
|
||
:
|
||
</emphasis>
|
||
♂, Acacia Plateau, J. Armstrong (
|
||
<collectionCode id="D0802584FFDB4A6D2382B486FADBE156" box="[1209,1284,1770,1796]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:32981" lsid="urn:lsid:biocol.org:col:32981" name="Australian National Insect Collection" pageId="46" pageNumber="47">ANIC</collectionCode>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFDB4A6C27FCB561FDA4E687" blockId="46.[151,1436,1769,2012]" lastBlockId="47.[151,1437,151,1941]" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
|
||
<emphasis id="84E56153FFDB4A6D27FCB561FE44E175" bold="true" box="[199,411,1805,1831]" pageId="46" pageNumber="47">Revised diagnosis</emphasis>
|
||
. In male antennomeres 3–4 each about 1.5 × as long as broad, 5 twice as long as broad, 6 1.5 × as long as broad, antennomeres 9 and 10 each 1.4 × as long as broad, and antennomere 11 less than twice as long as broad, with triangularly expanded angulate outer lateral margin (
|
||
<figureCitation id="2EAAA1C4FFDB4A6D248FB539FBCCE122" box="[948,1043,1877,1904]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="46" pageNumber="47">Fig. 120</figureCitation>
|
||
, right antenna); metatibiae strongly modified (
|
||
<figureCitation id="2EAAA1C4FFDB4A6D2635B515FE71E1C6" box="[270,430,1913,1940]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="46" pageNumber="47">Figs 118–120</figureCitation>
|
||
), each medially broadened and with lateral (outer) elongate deep concavity surrounded by sharp margins and filled with setae, apical region constricted, narrowed and curved; pronotum with two pairs of antebasal pits; aedeagus in dorsal view (
|
||
<figureCitation id="2EAAA1C4FFDB4A6D2545B5ADFCC3E18E" box="[638,796,1985,2012]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="46" pageNumber="47">Figs 121, 123</figureCitation>
|
||
) with broad apex of median lobe, slightly narrower than half of widest (subapical) site of median lobe, flanked by weakly elongate, subtriangular weakly sclerotized lateral subapical lobes; median lobe lacking setae.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE8BEECAFFDA4A6C27FCB2B2FD95E17B" pageId="47" pageNumber="48" type="description">
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB2B2FE29E713" blockId="47.[151,1437,151,1941]" pageId="47" pageNumber="48">
|
||
<emphasis id="84E56153FFDA4A6C27FCB2B2FEAAE6AA" bold="true" box="[199,373,222,248]" pageId="47" pageNumber="48">Redescription.</emphasis>
|
||
Body in male (
|
||
<figureCitation id="2EAAA1C4FFDA4A6C251FB2B3FD5BE6AB" box="[548,644,223,249]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="47" pageNumber="48">Fig. 120</figureCitation>
|
||
) slightly flattened, elongate and relatively slender, BL
|
||
<quantity id="716910A4FFDA4A6C23CAB2B2FA47E6AB" box="[1265,1432,222,249]" metricMagnitude="-3" metricUnit="m" metricValue="1.78" metricValueMax="1.81" metricValueMin="1.75" pageId="47" pageNumber="48" unit="mm" value="1.78" valueMax="1.81" valueMin="1.75">1.75–1.81 mm</quantity>
|
||
; pigmentation uniformly light brown (including appendages); cuticle moderately glossy, covered with vestiture of setae slightly lighter than body.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB327FDEBE50F" blockId="47.[151,1437,151,1941]" pageId="47" pageNumber="48">
|
||
Head (Fi. 120) in dorsal view indistinctly transverse and somewhat subhexagonal, broadest at eyes, HL
|
||
<quantity id="716910A4FFDA4A6C2266B327FF20E7DB" metricMagnitude="-4" metricUnit="m" metricValue="3.2" metricValueMax="3.4" metricValueMin="3.0" pageId="47" pageNumber="48" unit="mm" value="0.32" valueMax="0.34" valueMin="0.3">0.30– 0.34 mm</quantity>
|
||
, HW
|
||
<quantity id="716910A4FFDA4A6C267FB302FE32E7DB" box="[324,493,366,393]" metricMagnitude="-4" metricUnit="m" metricValue="3.3999999999999995" metricValueMax="3.5" metricValueMin="3.3" pageId="47" pageNumber="48" unit="mm" value="0.33999999999999997" valueMax="0.35" valueMin="0.33">0.33–0.35 mm</quantity>
|
||
; vertex and frons confluent and weakly convex, posterior margin of vertex evenly arcuate and anteriorly convex; tempora about 2.5 × as long as length of eye in dorsal view; supraantennal tubercles indistinct; frons over antennal fossae broadly subtriangular and with rounded anterior margin. Eyes small, weakly oval, not emarginate posteriorly and oblique in relation to long axis of head. Punctures on frons and vertex fine, inconspicuous; setae (including those on tempora) short, sparse, nearly recumbent. Genae as sparsely setose as frons and vertex. Anterior (exposed) region of head capsule demarcated from neck region by short abrupt impression around occipital constriction, anterior margin of gular plate on neck region with indistinct, narrow anteriorly-directed projection. Antennae (
|
||
<figureCitation id="2EAAA1C4FFDA4A6C26E7B007FDE4E4D7" box="[476,571,619,645]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="47" pageNumber="48">Fig. 120</figureCitation>
|
||
) long and slender, AnL 1.00–
|
||
<quantity id="716910A4FFDA4A6C24B1B007FC2FE4D7" box="[906,1008,619,645]" metricMagnitude="-3" metricUnit="m" metricValue="1.03" pageId="47" pageNumber="48" unit="mm" value="1.03">1.03 mm</quantity>
|
||
; three terminal antennomeres forming indistinctly delimited club; scape 2.5 × as long as broad, distinctly broadening distally; pedicel 1.8 × as long as broad; antennomeres 3–4 each about 1.5 × as long as broad, 5 twice as long as broad, 6 1.5 × as long as broad, 7 and 8 each distinctly asymmetrical and each about as long as broad, 9 1.4 × as long as broad and barrel-shaped, 10 1.4 × as long as broad and distinctly broadening distally, 11 much shorter than 9 and 10 combined, about 1.6 × as long as broad, distinctly asymmetrical, with broadly subtriangular, angulate expansion on proximal region of outer margin (visible in right antenna in
|
||
<figureCitation id="2EAAA1C4FFDA4A6C26FDB12FFDF8E50F" box="[454,551,835,861]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="47" pageNumber="48">Fig. 120</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB10BFE8BE22F" blockId="47.[151,1437,151,1941]" pageId="47" pageNumber="48">
|
||
Pronotum in dorsal view (
|
||
<figureCitation id="2EAAA1C4FFDA4A6C26C8B10BFD8AE5D3" box="[499,597,871,897]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="47" pageNumber="48">Fig. 120</figureCitation>
|
||
) distinctly elongate, broadest between middle and anterior third, PL
|
||
<quantity id="716910A4FFDA4A6C2250B10AFF1AE5F7" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="47" pageNumber="48" unit="mm" value="0.5">0.50 mm</quantity>
|
||
, PW
|
||
<quantity id="716910A4FFDA4A6C27C4B1E7FE7BE5F7" box="[255,420,907,933]" metricMagnitude="-4" metricUnit="m" metricValue="4.05" metricValueMax="4.1" metricValueMin="4.0" pageId="47" pageNumber="48" unit="mm" value="0.405" valueMax="0.41" valueMin="0.4">0.40–0.41 mm</quantity>
|
||
; anterior margin arcuate and laterally confluent with rounded lateral margins, so that anterior corners are not marked; posterior corners obtuse-angled and blunt; posterior margin nearly straight at middle and bent anteriorly at sides; base with very narrow and indistinct posterior marginal carina, with two pairs of distinct, small and slightly transverse pits. Pronotal disc covered with fine, shallow and unremarkable sparse punctures; setae similar to those on head, moderately dense and long, suberect. Ventrally prothorax with nearly asetose and impunctate hypomera and basisternal region only slightly longer than procoxal rests, sparsely covered with moderately long recumbent setae.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB6EBFBAFE35F" blockId="47.[151,1437,151,1941]" pageId="47" pageNumber="48">
|
||
Elytra (
|
||
<figureCitation id="2EAAA1C4FFDA4A6C2626B6EBFE5AE2F3" box="[285,389,1159,1185]" captionStart="FIGURES 109–110" captionStartId="42.[151,264,1481,1505]" captionTargetBox="[177,1411,381,1457]" captionTargetId="figure-94@42.[177,1411,381,1457]" captionTargetPageId="42" captionText="FIGURES 109–110. Scydmaenus (Parallomicrus) myrmecobius Csiki, lectotype male. Head, pronotum and elytral base in dorsal view (109); head and prothorax in ventral view (110). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mst, mediostipes; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144691" httpUri="https://zenodo.org/record/10144691/files/figure.png" pageId="47" pageNumber="48">Figs 109</figureCitation>
|
||
,
|
||
<figureCitation id="2EAAA1C4FFDA4A6C26A8B6EBFE62E2F3" box="[403,445,1159,1185]" captionStart="FIGURES 113–117" captionStartId="44.[151,264,1175,1199]" captionTargetBox="[157,1430,459,1146]" captionTargetId="figure-144@44.[151,1436,453,1152]" captionTargetPageId="44" captionText="FIGURES 113–117. Scydmaenus (Parallomicrus) myrmecobius Csiki, lectotype male. Dorsal habitus (113); aedeagus in dorsal (114, 116) and lateral (115, 117) views." figureDoi="http://doi.org/10.5281/zenodo.10144695" httpUri="https://zenodo.org/record/10144695/files/figure.png" pageId="47" pageNumber="48">113</figureCitation>
|
||
) oval, broadest slightly in front of middle, EL
|
||
<quantity id="716910A4FFDA4A6C24DAB6EAFB56E2F3" box="[993,1161,1158,1185]" metricMagnitude="-4" metricUnit="m" metricValue="9.65" metricValueMax="9.8" metricValueMin="9.5" pageId="47" pageNumber="48" unit="mm" value="0.965" valueMax="0.98" valueMin="0.95">0.95–0.98 mm</quantity>
|
||
, EW
|
||
<quantity id="716910A4FFDA4A6C23F7B6EAFAACE2F3" box="[1228,1395,1158,1185]" metricMagnitude="-4" metricUnit="m" metricValue="6.65" metricValueMax="6.8" metricValueMin="6.5" pageId="47" pageNumber="48" unit="mm" value="0.665" valueMax="0.68" valueMin="0.65">0.65–0.68 mm</quantity>
|
||
, EI 1.41–1.50. Humeral calli small but distinctly elevated and each mesally demarcated by shallow and transverse basal elytral impression; basal elytral foveae lacking; apices separately rounded. Elytral punctures fine and inconspicuous; setae similar to those on pronotum, moderately dense. Hind wings fully developed.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB77BFF00E07F" blockId="47.[151,1437,151,1941]" pageId="47" pageNumber="48">
|
||
Legs (
|
||
<figureCitation id="2EAAA1C4FFDA4A6C2637B77BFEB5E363" box="[268,362,1303,1329]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="47" pageNumber="48">Fig. 120</figureCitation>
|
||
) long and slender. Metatibia (
|
||
<figureCitation id="2EAAA1C4FFDA4A6C2589B77BFC93E363" box="[690,844,1303,1329]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="47" pageNumber="48">Figs 118–120</figureCitation>
|
||
) strongly modified, medially broadened, with strongly elongate oval cavity on lateral (outer) surface demarcated by sharp edges and filled with setae, distal 1/4 rapidly narrowed, almost constricted and distinctly curved. Protarsi with tenent setae on tarsomeres 1–3, protarsomere 1 moderately strongly broadened, weakly elongate, 2–4 each almost as long as broad, 5 about 2.5 × as long as broad; mesotarsi longer than protarsi, mesotarsomere 1 about 3 × as long as broad, tarsomeres 2–4 each weakly elongate, tarsomere 5 about 2.5 × as long as broad; metatarsi slightly longer than mesotarsi, metatarsomere 1 about 3 × as long as broad, tarsomeres 2–4 each distinctly elongate, but decreasing in length distally, tarsomere 5 about 3 × as long as broad.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB45BFE48E157" blockId="47.[151,1437,151,1941]" pageId="47" pageNumber="48">
|
||
Aedeagus (
|
||
<figureCitation id="2EAAA1C4FFDA4A6C267EB45BFE3EE003" box="[325,481,1591,1617]" captionStart="FIGURES 118–124" captionStartId="46.[151,264,1663,1687]" captionTargetBox="[157,1435,885,1633]" captionTargetId="figure-244@46.[151,1436,879,1639]" captionTargetPageId="46" captionText="FIGURES 118–124. Scydmaenus (Parallomicrus) inflatitibia Franz, male holotype. Right hind tibia in dorsal (118) and slightly outer lateral (119) views; dorsal habitus (120); aedeagus in dorsal (121, 123) and lateral (122, 124) views." figureDoi="http://doi.org/10.5281/zenodo.10144697" httpUri="https://zenodo.org/record/10144697/files/figure.png" pageId="47" pageNumber="48">Figs 121–122</figureCitation>
|
||
) elongate but not very slender, AeL
|
||
<quantity id="716910A4FFDA4A6C2441B45BFC3EE003" box="[890,993,1591,1617]" metricMagnitude="-4" metricUnit="m" metricValue="3.8" pageId="47" pageNumber="48" unit="mm" value="0.38">0.38 mm</quantity>
|
||
, in dorsal view median lobe (excluding lateral subapical lobes) broadest in distal 1/3, narrowing both distally and proximally, but basal region again broadened, apex slightly less than half as wide as total width of median lobe, apical margin indistinctly concave; lateral subapical lobes large, each subtriangular and slightly longer than wide, projecting laterally; flagellum broadened in proximal region to form asymmetrical coils; median lobe lacking setae; ostium situated in distal third of median lobe, far from its apex.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB563FD95E17B" blockId="47.[151,1437,151,1941]" box="[199,586,1807,1833]" pageId="47" pageNumber="48">
|
||
<emphasis id="84E56153FFDA4A6C27FCB563FEFAE17B" bold="true" box="[199,293,1807,1833]" pageId="47" pageNumber="48">Female.</emphasis>
|
||
Unknown (see Remarks).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE8BEECAFFDA4A6C27FCB55EFC86E11F" box="[199,857,1842,1869]" pageId="47" pageNumber="48" type="distribution">
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB55EFC86E11F" blockId="47.[151,1437,151,1941]" box="[199,857,1842,1869]" pageId="47" pageNumber="48">
|
||
<emphasis id="84E56153FFDA4A6C27FCB55EFEBEE11E" bold="true" box="[199,353,1842,1868]" pageId="47" pageNumber="48">Distribution.</emphasis>
|
||
<collectingRegion id="745573A3FFDA4A6C265CB55FFE26E11F" box="[359,505,1843,1869]" country="Australia" name="Western Australia" pageId="47" pageNumber="48">SE Australia</collectingRegion>
|
||
: CE and NE
|
||
<collectingRegion id="745573A3FFDA4A6C25B1B55FFC8BE11F" box="[650,852,1843,1869]" country="Australia" name="New South Wales" pageId="47" pageNumber="48">New South Wales</collectingRegion>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE8BEECAFFDA4A7327FCB53BFD9EE5D3" lastPageId="48" lastPageNumber="49" pageId="47" pageNumber="48" type="discussion">
|
||
<paragraph id="B62EBD41FFDA4A6C27FCB53BFBB4E1C7" blockId="47.[151,1437,151,1941]" pageId="47" pageNumber="48">
|
||
<emphasis id="84E56153FFDA4A6C27FCB53BFEE4E123" bold="true" box="[199,315,1879,1905]" pageId="47" pageNumber="48">Remarks.</emphasis>
|
||
The identity of the only known
|
||
<typeStatus id="692A03E3FFDA4A6C2596B53BFCD2E123" box="[685,781,1879,1905]" pageId="47" pageNumber="48" type="paratype">paratype</typeStatus>
|
||
female is uncertain. This specimen is remarkably smaller than the
|
||
<typeStatus id="692A03E3FFDA4A6C27C3B517FE85E1C7" box="[248,346,1915,1941]" pageId="47" pageNumber="48" type="holotype">holotype</typeStatus>
|
||
male (BL
|
||
<quantity id="716910A4FFDA4A6C26EAB517FDEAE1C7" box="[465,565,1915,1941]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="47" pageNumber="48" unit="mm" value="1.6">1.60 mm</quantity>
|
||
vs. 1.75), and it may belong to a different species.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFC54A7327FCB2FBFBF4E547" blockId="48.[151,1437,151,789]" pageId="48" pageNumber="49">
|
||
<taxonomicName id="7191C6C2FFC54A7327FCB2FBFE10E6E3" box="[199,463,151,177]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="inflatitibia">
|
||
<emphasis id="84E56153FFC54A7327FCB2FBFE10E6E3" box="[199,463,151,177]" italics="true" pageId="48" pageNumber="49">Scydmaenus inflatitibia</emphasis>
|
||
</taxonomicName>
|
||
combines some characters typical of
|
||
<taxonomicName id="7191C6C2FFC54A73244FB2FBFBC9E6E3" box="[884,1046,151,177]" pageId="48" pageNumber="42" rank="subGenus" subGenus="Parallomicrus">
|
||
<emphasis id="84E56153FFC54A73244FB2FBFBC9E6E3" box="[884,1046,151,177]" italics="true" pageId="48" pageNumber="49">Parallomicrus</emphasis>
|
||
</taxonomicName>
|
||
(fused metanepisterna),
|
||
<taxonomicName id="7191C6C2FFC54A732211B2FBFF6AE686" pageId="48" pageNumber="21" rank="subGenus" subGenus="Choleropsis">
|
||
<emphasis id="84E56153FFC54A732211B2FBFF6AE686" italics="true" pageId="48" pageNumber="49">Choleropsis</emphasis>
|
||
</taxonomicName>
|
||
(two pairs of antebasal pronotal pits), and
|
||
<taxonomicName id="7191C6C2FFC54A7325B6B2D7FCDCE687" authorityName="Franz" authorityYear="1975" box="[653,771,187,213]" class="Insecta" family="Staphylinidae" genus="Corbulifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="84E56153FFC54A7325B6B2D7FCDCE687" box="[653,771,187,213]" italics="true" pageId="48" pageNumber="49">Corbulifer</emphasis>
|
||
</taxonomicName>
|
||
(modified metatibia in males). It remarkably differs from
|
||
<taxonomicName id="7191C6C2FFC54A7322BCB2D7FEF3E6AB" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="myrmecobius">
|
||
<emphasis id="84E56153FFC54A7322BCB2D7FEF3E6AB" italics="true" pageId="48" pageNumber="49">S. myrmecobius</emphasis>
|
||
</taxonomicName>
|
||
, especially in aedeagal features. Although the aedeagus within the large genus
|
||
<taxonomicName id="7191C6C2FFC54A73239CB2B3FAEEE6AB" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1191,1329,223,249]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="84E56153FFC54A73239CB2B3FAEEE6AB" box="[1191,1329,223,249]" italics="true" pageId="48" pageNumber="49">Scydmaenus</emphasis>
|
||
</taxonomicName>
|
||
is poorly studied, its general structure is usually stable within subgenera. For instance, it was demonstrated that despite a wide spectrum of metatibial modifications in
|
||
<taxonomicName id="7191C6C2FFC54A732560B34BFD0EE713" authorityName="Franz" authorityYear="1975" box="[603,721,295,321]" class="Insecta" family="Staphylinidae" genus="Corbulifer" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="84E56153FFC54A732560B34BFD0EE713" box="[603,721,295,321]" italics="true" pageId="48" pageNumber="49">Corbulifer</emphasis>
|
||
</taxonomicName>
|
||
, the structure of the aedeagus is very similar in all species, i.e., all of them, independently of the shape of the median lobe, share long setae and asymmetrical proximal ‘chambers’ of the flagellum (
|
||
<bibRefCitation id="D200C0B0FFC54A732662B303FDFFE7DB" author="Jaloszynski, P." box="[345,544,367,393]" pageId="48" pageNumber="49" pagination="505 - 531" refId="ref50945" refString="Jaloszynski, P. (2018) " Monster Scydmaenus " of Australia: revision of subgenus Corbulifer Franz (Coleoptera: Staphylinidae: Scydmaeninae). Zootaxa, 4472 (3), 505 - 531. https: // doi. org / 10.11646 / zootaxa. 4472.3.5" type="journal article" year="2018">Jałoszyński 2018</bibRefCitation>
|
||
). All species of
|
||
<taxonomicName id="7191C6C2FFC54A7325E9B303FC87E7DB" box="[722,856,367,393]" pageId="48" pageNumber="21" rank="subGenus" subGenus="Choleropsis">
|
||
<emphasis id="84E56153FFC54A7325E9B303FC87E7DB" box="[722,856,367,393]" italics="true" pageId="48" pageNumber="49">Choleropsis</emphasis>
|
||
</taxonomicName>
|
||
treated in the present study also have asymmetrical proximal flagellar coils, whereas two known species of
|
||
<taxonomicName id="7191C6C2FFC54A732435B3FFFCBAE7FE" box="[782,869,403,428]" pageId="48" pageNumber="30" rank="subGenus" subGenus="Kingius">
|
||
<emphasis id="84E56153FFC54A732435B3FFFCBAE7FE" box="[782,869,403,428]" italics="true" pageId="48" pageNumber="49">Kingius</emphasis>
|
||
</taxonomicName>
|
||
, four Australian species of
|
||
<taxonomicName id="7191C6C2FFC54A7323A5B3FFFABAE7FF" box="[1182,1381,403,429]" pageId="48" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC54A7323A5B3FFFABAE7FF" box="[1182,1381,403,429]" italics="true" pageId="48" pageNumber="49">Scottiscydmaenus</emphasis>
|
||
</taxonomicName>
|
||
, and two species of the newly described
|
||
<taxonomicName id="7191C6C2FFC54A73250CB3DBFD10E783" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[567,719,439,465]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="subGenus" status="subgen. nov." subGenus="Ascydmaenus">
|
||
<emphasis id="84E56153FFC54A73250CB3DBFD10E783" box="[567,719,439,465]" italics="true" pageId="48" pageNumber="49">Ascydmaenus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9FD6DC28FFC54A7325E2B3DBFC91E783" box="[729,846,439,465]" pageId="48" pageNumber="49" rank="subGenus">subgen. n.</taxonomicNameLabel>
|
||
share not coils, but symmetrical expansions of the proximal flagellar region, which appear as three consecutive ‘chambers’ demarcated by constrictions.
|
||
<taxonomicName id="7191C6C2FFC54A732229B3B7FE0FE44B" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="australiensis" subGenus="Mascarensia">
|
||
<emphasis id="84E56153FFC54A732229B3B7FA43E7A7" box="[1298,1436,475,501]" italics="true" pageId="48" pageNumber="49">Scydmaenus</emphasis>
|
||
(
|
||
<emphasis id="84E56153FFC54A73279AB393FEEEE44A" box="[161,305,511,536]" italics="true" pageId="48" pageNumber="49">Mascarensia</emphasis>
|
||
)
|
||
<emphasis id="84E56153FFC54A73267AB393FE0FE44B" box="[321,464,511,537]" italics="true" pageId="48" pageNumber="49">australiensis</emphasis>
|
||
</taxonomicName>
|
||
has strongly asymmetrical, broad coils, and even though the previously studied
|
||
<taxonomicName id="7191C6C2FFC54A732265B393FD44E46F" authority="Franz, 1976" authorityName="Franz" authorityYear="1976" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="kasuganus" subGenus="Mascarensia">
|
||
<emphasis id="84E56153FFC54A732265B393FF32E46E" italics="true" pageId="48" pageNumber="49">Scydmaenus</emphasis>
|
||
(
|
||
<emphasis id="84E56153FFC54A7327C7B04FFE53E46E" box="[252,396,547,572]" italics="true" pageId="48" pageNumber="49">Mascarensia</emphasis>
|
||
)
|
||
<emphasis id="84E56153FFC54A7326A1B04FFDCFE46F" box="[410,528,547,573]" italics="true" pageId="48" pageNumber="49">kasuganus</emphasis>
|
||
<bibRefCitation id="D200C0B0FFC54A73252DB04FFD44E46F" author="Franz, H." box="[534,667,547,573]" pageId="48" pageNumber="49" pagination="51 - 60" refId="ref50099" refString="Franz, H. (1976) Neue Scydmaeniden aus Japan, sowie Bemerkungen zu bekannten Arten. Entomologische Ergebnisse einer auf Einladung der Internationalen Vereinigung fur Vegetationskunde nach Japan durchgefuhrten Forschungsreise. Entomologische Blatter fur Biologie und Systematik der Kafer, 72, 51 - 60." type="journal article" year="1976">Franz, 1976</bibRefCitation>
|
||
</taxonomicName>
|
||
has a clearly different shape of the median lobe, its proximal flagellar region is also strongly asymmetrical in the dorsal view (
|
||
<bibRefCitation id="D200C0B0FFC54A7325C4B02BFC1AE433" author="Jaloszynski, P." box="[767,965,583,609]" pageId="48" pageNumber="49" pagination="38 - 48" refId="ref51044" refString="Jaloszynski, P. (2022) Rediscovery and re (re) description of Scydmaenus kasuganus Franz, the only Japanese species of the subgenus Mascarensia Franz (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 5093 (1), 38 - 48. https: // doi. org / 10.11646 / zootaxa. 5093.1.2" type="journal article" year="2022">Jałoszyński 2022</bibRefCitation>
|
||
). Two species of
|
||
<taxonomicName id="7191C6C2FFC54A7323BAB02BFAD4E433" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[1153,1291,583,609]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="84E56153FFC54A7323BAB02BFAD4E433" box="[1153,1291,583,609]" italics="true" pageId="48" pageNumber="49">Scydmaenus</emphasis>
|
||
</taxonomicName>
|
||
belonging in the subgenus
|
||
<taxonomicName id="7191C6C2FFC54A732615B007FE3BE4D7" box="[302,484,619,645]" pageId="48" pageNumber="49" rank="subGenus" subGenus="Geoscydmaenus">
|
||
<emphasis id="84E56153FFC54A732615B007FE3BE4D7" box="[302,484,619,645]" italics="true" pageId="48" pageNumber="49">Geoscydmaenus</emphasis>
|
||
</taxonomicName>
|
||
and inhabiting
|
||
<collectingCountry id="CE86FDD1FFC54A7325AAB007FCC7E4D7" box="[657,792,619,645]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
|
||
have symmetrical proximal flagellar regions (
|
||
<bibRefCitation id="D200C0B0FFC54A732222B007FF3DE4FB" author="Jaloszynski, P." pageId="48" pageNumber="49" pagination="265 - 274" refId="ref50898" refString="Jaloszynski, P. (2016 b) Review of Scydmaenus (Geoscydmaenus) of Madagascar (Coleoptera, Staphylinidae, Scydmaeninae). Zootaxa, 4205 (3), 265 - 274. https: // doi. org / 10.11646 / zootaxa. 4205.3.6" type="journal article" year="2016">Jałoszyński 2016b</bibRefCitation>
|
||
). These observations suggest that groups of closely related species have similar structures of the flagellum. However, in the Australian
|
||
<taxonomicName id="7191C6C2FFC54A7326F4B0DFFDAEE49F" box="[463,625,691,717]" pageId="48" pageNumber="42" rank="subGenus" subGenus="Parallomicrus">
|
||
<emphasis id="84E56153FFC54A7326F4B0DFFDAEE49F" box="[463,625,691,717]" italics="true" pageId="48" pageNumber="49">Parallomicrus</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7191C6C2FFC54A7325BBB0DFFCEDE49F" box="[640,818,691,717]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="myrmecobius">
|
||
<emphasis id="84E56153FFC54A7325BBB0DFFCEDE49F" box="[640,818,691,717]" italics="true" pageId="48" pageNumber="49">S. myrmecobius</emphasis>
|
||
</taxonomicName>
|
||
has symmetrical ‘chambers’, whereas in
|
||
<taxonomicName id="7191C6C2FFC54A732232B0DFFA43E49F" box="[1289,1436,691,717]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="inflatitibia">
|
||
<emphasis id="84E56153FFC54A732232B0DFFA43E49F" box="[1289,1436,691,717]" italics="true" pageId="48" pageNumber="49">S. inflatitibia</emphasis>
|
||
</taxonomicName>
|
||
they form a loose spiral. Because external features (pronotal pits, male secondary sexual modifications) are also different in these two species, in future they may be placed in separate subgenera.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFC54A7327ACB10AFD9EE5D3" blockId="48.[151,577,870,897]" box="[151,577,870,897]" pageId="48" pageNumber="49">
|
||
<emphasis id="84E56153FFC54A7327ACB10AFD9EE5D3" bold="true" box="[151,577,870,897]" pageId="48" pageNumber="49">
|
||
Subgenus
|
||
<taxonomicName id="7191C6C2FFC54A73262CB10AFD9EE5D3" authority="Franz" authorityName="Franz" box="[279,577,870,897]" pageId="48" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC54A73262CB10AFE31E5D3" bold="true" box="[279,494,870,897]" italics="true" pageId="48" pageNumber="49">Scottiscydmaenus</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE8BEECAFFC54A7327ACB1C7FED9E5B0" pageId="48" pageNumber="49" type="reference_group">
|
||
<paragraph id="B62EBD41FFC54A7327ACB1C7FED9E5B0" blockId="48.[151,1437,939,994]" pageId="48" pageNumber="49">
|
||
<treatmentCitationGroup id="96819A6FFFC54A7327ACB1C7FED9E5B0" pageId="48" pageNumber="49">
|
||
<treatmentCitation id="37309B50FFC54A7327ACB1C7FDD4E591" author="Franz, H." box="[151,523,939,963]" page="277" pageId="48" pageNumber="49" year="1975">
|
||
<taxonomicName id="7191C6C2FFC54A7327ACB1C7FDD4E591" ID-CoL="8XSD2" authority="Franz, 1975: 277" authorityName="Franz" authorityYear="1975" box="[151,523,939,963]" kingdom="Animalia" pageId="48" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC54A7327ACB1C7FE94E591" box="[151,331,939,963]" italics="true" pageId="48" pageNumber="49">Scottiscydmaenus</emphasis>
|
||
<bibRefCitation id="D200C0B0FFC54A73266DB1C0FDD4E591" author="Franz, H." box="[342,523,939,963]" pageId="48" pageNumber="49" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz, 1975: 277</bibRefCitation>
|
||
</taxonomicName>
|
||
</treatmentCitation>
|
||
(as subgenus of
|
||
<taxonomicName id="7191C6C2FFC54A7325FDB1C7FC9CE591" ID-CoL="8VS3H" baseAuthorityName="Franz" baseAuthorityYear="1975" box="[710,835,939,963]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="84E56153FFC54A7325FDB1C7FC9CE591" box="[710,835,939,963]" italics="true" pageId="48" pageNumber="49">Scydmaenus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
<typeStatus id="692A03E3FFC54A732466B1C0FC50E596" box="[861,911,940,964]" pageId="48" pageNumber="49">Type</typeStatus>
|
||
species:
|
||
<taxonomicName id="7191C6C2FFC54A7324CFB1C7FABEE591" ID-CoL="4W3XF" authority="Scott, 1922" authorityName="Scott" authorityYear="1922" box="[1012,1377,939,963]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="lodoiceae">
|
||
<emphasis id="84E56153FFC54A7324CFB1C7FB01E591" box="[1012,1246,939,963]" italics="true" pageId="48" pageNumber="49">Scydmaenus lodoiceae</emphasis>
|
||
<bibRefCitation id="D200C0B0FFC54A7323D2B1C7FABEE591" author="Scott, H." box="[1257,1377,939,963]" pageId="48" pageNumber="49" pagination="195 - 260" refId="ref51964" refString="Scott, H. (1922) No. IV. - Coleoptera: Scydmaenidae, Scaphidiidae, Phalacridae, Cucujidae (Supplement), Lathridiidae, Mycetophagidae (including Propalticus), Bostrychidae, Lyctidae. Transactions of the Linnean Society of London, Series 2: Zoology, 18, 195 - 260. https: // doi. org / 10.1111 / j. 1096 - 3642.1922. tb 00550. x" type="journal article" year="1922">Scott, 1922</bibRefCitation>
|
||
</taxonomicName>
|
||
(des. orig.).
|
||
</treatmentCitationGroup>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE8BEECAFFC54A7327ACB67CFDB6E350" pageId="48" pageNumber="49" type="diagnosis">
|
||
<paragraph id="B62EBD41FFC54A7327ACB67CFDB6E350" blockId="48.[151,1437,1040,2038]" pageId="48" pageNumber="49">
|
||
<emphasis id="84E56153FFC54A7327ACB67CFECDE278" bold="true" box="[151,274,1040,1066]" pageId="48" pageNumber="49">Diagnosis.</emphasis>
|
||
<bibRefCitation id="D200C0B0FFC54A732621B67DFE73E279" author="Franz, H." box="[282,428,1040,1067]" pageId="48" pageNumber="49" pagination="1 - 312" refId="ref50046" refString="Franz, H. (1975) Revision der Scydmaeniden von Australien, Neuseeland und den benachbarten Inseln. Denkschriften der Osterreichische Akademie der Wissenschaften, Mathematisch-Naturwissenschaftliche Klasse, 118, 1 - 312. https: // doi. org / 10.1007 / 978 - 3 - 7091 - 5642 - 1" type="journal article" year="1975">Franz (1975)</bibRefCitation>
|
||
defined
|
||
<taxonomicName id="7191C6C2FFC54A73252AB67CFD07E278" box="[529,728,1040,1066]" pageId="48" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC54A73252AB67CFD07E278" box="[529,728,1040,1066]" italics="true" pageId="48" pageNumber="49">Scottiscydmaenus</emphasis>
|
||
</taxonomicName>
|
||
by the following characters: (1) antennae with slender trimerous clubs with elongate antennomeres 9 (
|
||
<figureCitation id="2EAAA1C4FFC54A732554B658FD0CE21C" box="[623,723,1076,1102]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 127</figureCitation>
|
||
); (2) the pronotum strongly convex, lacking antebasal pits and a longitudinal carina (
|
||
<figureCitation id="2EAAA1C4FFC54A7326AEB634FE25E221" box="[405,506,1112,1139]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="48" pageNumber="49">Fig. 125</figureCitation>
|
||
); (3) elytra strongly convex, lacking humeral calli and humeral folds (
|
||
<figureCitation id="2EAAA1C4FFC54A732211B634FA50E220" box="[1322,1423,1112,1138]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="48" pageNumber="49">Fig. 130</figureCitation>
|
||
); (4) metanepisterna not demarcated from the metaventrite (
|
||
<figureCitation id="2EAAA1C4FFC54A73240FB610FC46E2C4" box="[820,921,1148,1174]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 128</figureCitation>
|
||
); (5) the profemur in males with two ventral carinae, anterior one distally with a broad and blunt projection (
|
||
<figureCitation id="2EAAA1C4FFC54A73245AB6CCFC1CE2E8" box="[865,963,1184,1210]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 129</figureCitation>
|
||
); and (6) protibia in males distally weakly broadened, laterally flattened. The latter character was here found rather problematic, as the protibiae in all studied species are almost unmodified (
|
||
<figureCitation id="2EAAA1C4FFC54A7326C1B684FD82E350" box="[506,605,1256,1282]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 129</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE8BEECAFFC54A7327FCB760FDB4E1A4" pageId="48" pageNumber="49" type="discussion">
|
||
<paragraph id="B62EBD41FFC54A7327FCB760FD36E0E0" blockId="48.[151,1437,1040,2038]" pageId="48" pageNumber="49">
|
||
<emphasis id="84E56153FFC54A7327FCB760FEE4E374" bold="true" box="[199,315,1292,1318]" pageId="48" pageNumber="49">Remarks.</emphasis>
|
||
The status of
|
||
<taxonomicName id="7191C6C2FFC54A7326E2B760FD7FE374" box="[473,672,1292,1318]" pageId="48" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC54A7326E2B760FD7FE374" box="[473,672,1292,1318]" italics="true" pageId="48" pageNumber="49">Scottiscydmaenus</emphasis>
|
||
</taxonomicName>
|
||
as a separate subgenus will require verification. The Australian species share similarly modified profemora in males (with a distal anterior/dorsal tooth (
|
||
<figureCitation id="2EAAA1C4FFC54A73235BB75CFB62E318" box="[1120,1213,1328,1354]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 129</figureCitation>
|
||
)—the anterior/dorsal terminology refers to the femur observed in the position as in
|
||
<figureCitation id="2EAAA1C4FFC54A7324BFB738FC38E33C" box="[900,999,1364,1390]" captionStart="FIGURES 130–134" captionStartId="52.[151,264,1084,1108]" captionTargetBox="[157,1430,187,1054]" captionTargetId="figure-21@52.[151,1436,181,1060]" captionTargetPageId="52" captionText="FIGURES 130–134. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Dorsal habitus (130); aedeagus in dorsal (131, 133) and lateral (132, 134) views." figureDoi="http://doi.org/10.5281/zenodo.10144703" httpUri="https://zenodo.org/record/10144703/files/figure.png" pageId="48" pageNumber="49">Fig. 130</figureCitation>
|
||
); antennae with conspicuously slender proximal flagellomeres and also a slender, elongate antennomere 9, so that the club appears as dimerous (
|
||
<figureCitation id="2EAAA1C4FFC54A732216B714FA50E3C0" box="[1325,1423,1400,1426]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 127</figureCitation>
|
||
); barely broadened protarsi in males with only sparse, poorly visible tenent setae (
|
||
<figureCitation id="2EAAA1C4FFC54A73232BB7F0FBAEE3E4" box="[1040,1137,1436,1462]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 129</figureCitation>
|
||
); pronota lacking pits (
|
||
<figureCitation id="2EAAA1C4FFC54A732254B7F0FF1AE389" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="48" pageNumber="49">Fig. 125</figureCitation>
|
||
); elytra with reduced humeral structures (humeral calli, humeral folds, and basal impressions) due to lack of wings (
|
||
<figureCitation id="2EAAA1C4FFC54A7327D1B788FE93E3AD" box="[234,332,1508,1535]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="48" pageNumber="49">Fig. 125</figureCitation>
|
||
); lack of lateral submental lobes (
|
||
<figureCitation id="2EAAA1C4FFC54A7325FCB788FCF5E3AC" box="[711,810,1508,1534]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="48" pageNumber="49">Fig. 126</figureCitation>
|
||
); the basisternal region of the prosternum distinctly, but not much longer than procoxal rests (
|
||
<figureCitation id="2EAAA1C4FFC54A73250DB464FD47E070" box="[566,664,1544,1570]" captionStart="FIGURES 125–126" captionStartId="50.[151,264,1744,1768]" captionTargetBox="[151,1436,181,1721]" captionTargetId="figure-21@50.[151,1436,181,1721]" captionTargetPageId="50" captionText="FIGURES 125–126. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Head, pronotum and elytral base in dorsal view (125); head and prothorax in ventral view (126). Abbreviations: bst, basistipes; bstr, basisternal region of prosternum; gen, gena; gp, gular plate; hr, hypomeral ridge; hyr, hypostomal ridge; lp, labial palp; md, mandible; mn, mentum; mxp, maxillary palp; occ, occipital constriction; plb, prelabium; ppf, palpifer; ptp, posterior tentorial pit; smn, submentum." figureDoi="http://doi.org/10.5281/zenodo.10144699" httpUri="https://zenodo.org/record/10144699/files/figure.png" pageId="48" pageNumber="49">Fig. 126</figureCitation>
|
||
); a narrow, almost carinate mesoventral intermesocoxal process (
|
||
<figureCitation id="2EAAA1C4FFC54A732254B464FF1AE014" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 128</figureCitation>
|
||
); metaventral intermetacoxal process about as wide as one metacoxa (
|
||
<figureCitation id="2EAAA1C4FFC54A7324F1B440FBF4E014" box="[970,1067,1580,1606]" captionStart="FIGURES 127–129" captionStartId="51.[151,264,1361,1385]" captionTargetBox="[151,1436,181,1337]" captionTargetId="figure-15@51.[151,1436,181,1337]" captionTargetPageId="51" captionText="FIGURES 127–129. Scydmaenus (Scottiscydmaenus) clarkianus Franz, holotype male. Distal region of right antenna in dorsal view (127); pterothorax and base of abdomen in ventral view (128); distal region of right fore leg in dorsal view (129). Abbreviations: aest3, metanepisternum; amvp, anterior metaventral process; a7‒11, antennomere 7‒11; msvp, mesoventral process; mtvp, metaventral process; pcr, procoxal rest; prp, prepectus." figureDoi="http://doi.org/10.5281/zenodo.10144701" httpUri="https://zenodo.org/record/10144701/files/figure.png" pageId="48" pageNumber="49">Fig. 128</figureCitation>
|
||
); and the aedeagus in dorsal view narrowing distally, in lateral view weakly curved, lacking lateral subapical lobes, with dorsodistal minute setae and symmetrical ‘chambers’ of the proximal flagellar region (e.g.,
|
||
<figureCitation id="2EAAA1C4FFC54A732466B418FC23E0DC" box="[861,1020,1652,1678]" captionStart="FIGURES 135–144" captionStartId="54.[151,264,1919,1943]" captionTargetBox="[157,1430,187,1889]" captionTargetId="figure-21@54.[151,1436,181,1895]" captionTargetPageId="54" captionText="FIGURES 135–144. Scydmaenus (Scottiscydmaenus) optatus Sharp, holotype male (135–139), and holotype male of Scydmaenus scotti Franz (140–144). Dorsal habitus (135, 140); aedeagus in dorsal (136, 138, 141, 143) and lateral (137, 139, 142, 144) views." figureDoi="http://doi.org/10.5281/zenodo.10144705" httpUri="https://zenodo.org/record/10144705/files/figure.png" pageId="48" pageNumber="49">Figs 136–139</figureCitation>
|
||
). However, it is unclear if this set of characters is sufficient to define a separate subgenus.
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFC54A7327FCB4D0FDB4E1A4" blockId="48.[151,1437,1040,2038]" pageId="48" pageNumber="49">
|
||
<emphasis id="84E56153FFC54A7327FCB4D0FD4AE084" bold="true" box="[199,661,1724,1750]" pageId="48" pageNumber="49">Composition, distribution and biology.</emphasis>
|
||
<taxonomicName id="7191C6C2FFC54A7325A6B4D0FCBBE084" box="[669,868,1724,1750]" pageId="48" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC54A7325A6B4D0FCBBE084" box="[669,868,1724,1750]" italics="true" pageId="48" pageNumber="49">Scottiscydmaenus</emphasis>
|
||
</taxonomicName>
|
||
includes six valid species names, but two of them were found to be synonymous during the present study, and one more new species is described. This subgenus is distributed in
|
||
<collectingCountry id="CE86FDD1FFC54A732601B568FE1CE14C" box="[314,451,1796,1822]" name="Madagascar" pageId="48" pageNumber="49">Madagascar</collectingCountry>
|
||
,
|
||
<collectingCountry id="CE86FDD1FFC54A7326E9B568FD95E14C" box="[466,586,1796,1822]" name="Seychelles" pageId="48" pageNumber="49">Seychelles</collectingCountry>
|
||
(Praslin), and continental
|
||
<collectingCountry id="CE86FDD1FFC54A732444B568FC36E14C" box="[895,1001,1796,1822]" name="Australia" pageId="48" pageNumber="49">Australia</collectingCountry>
|
||
. The Australian species are relatively variable in the body length and proportions of body parts; the most stable features are the shape of the head and the profemoral tooth in males.
|
||
<taxonomicName id="7191C6C2FFC54A7326CEB520FC97E135" authority="Franz" authorityName="Franz" box="[501,840,1868,1895]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="clarkianus">
|
||
<emphasis id="84E56153FFC54A7326CEB520FD22E134" box="[501,765,1868,1894]" italics="true" pageId="48" pageNumber="49">Scydmaenus clarkianus</emphasis>
|
||
Franz
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="7191C6C2FFC54A73246FB520FB8DE134" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[852,1106,1868,1894]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="brachyponerophilus" status="sp. n.">
|
||
<emphasis id="84E56153FFC54A73246FB520FB8DE134" box="[852,1106,1868,1894]" italics="true" pageId="48" pageNumber="49">S. brachyponerophilus</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="84E56153FFC54A732362B520FB46E134" bold="true" box="[1113,1177,1868,1894]" pageId="48" pageNumber="49">
|
||
<taxonomicNameLabel id="9FD6DC28FFC54A732362B520FB46E134" box="[1113,1177,1868,1894]" pageId="48" pageNumber="49" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="7191C6C2FFC54A7323EAB520FABFE134" box="[1233,1376,1868,1894]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="swanensis">
|
||
<emphasis id="84E56153FFC54A7323EAB520FABFE134" box="[1233,1376,1868,1894]" italics="true" pageId="48" pageNumber="49">S. swanensis</emphasis>
|
||
</taxonomicName>
|
||
have been collected from colonies of the ant
|
||
<taxonomicName id="7191C6C2FFC54A732569B51CFC54E1D8" authority="(Mayr)" baseAuthorityName="Mayr" box="[594,907,1904,1930]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="lutea">
|
||
<emphasis id="84E56153FFC54A732569B51CFCEDE1D8" box="[594,818,1904,1930]" italics="true" pageId="48" pageNumber="49">Brachyponera lutea</emphasis>
|
||
(Mayr)
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="7191C6C2FFC54A7324ACB51CFB91E1D8" authority="Sharp" authorityName="Sharp" box="[919,1102,1904,1930]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="optatus">
|
||
<emphasis id="84E56153FFC54A7324ACB51CFBDAE1D8" box="[919,1029,1904,1930]" italics="true" pageId="48" pageNumber="49">S. optatus</emphasis>
|
||
Sharp
|
||
</taxonomicName>
|
||
from colonies of
|
||
<taxonomicName id="7191C6C2FFC54A73222EB51DFAB2E1D8" box="[1301,1389,1904,1930]" class="Insecta" family="Formicidae" genus="Brachyponera" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hymenoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="lutea">
|
||
<emphasis id="84E56153FFC54A73222EB51DFAB2E1D8" box="[1301,1389,1904,1930]" italics="true" pageId="48" pageNumber="49">B. lutea</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="7191C6C2FFC54A7327ACB5F8FDD0E1FC" authority="(Andre)" baseAuthorityName="Andre" baseAuthorityYear="1896" box="[151,527,1940,1966]" class="Insecta" family="Cerambycidae" genus="Rythidoponera" kingdom="Animalia" order="Coleoptera" pageId="48" pageNumber="49" phylum="Arthropoda" rank="species" species="victoriae">
|
||
<emphasis id="84E56153FFC54A7327ACB5F8FE75E1FC" box="[151,426,1940,1966]" italics="true" pageId="48" pageNumber="49">Rythidoponera victoriae</emphasis>
|
||
(André)
|
||
</taxonomicName>
|
||
. It is possible that the relatively large variation in body size and proportions of body parts found among the Australian
|
||
<taxonomicName id="7191C6C2FFC54A732562B5D4FCFFE180" box="[601,800,1976,2002]" pageId="48" pageNumber="49" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC54A732562B5D4FCFFE180" box="[601,800,1976,2002]" italics="true" pageId="48" pageNumber="49">Scottiscydmaenus</emphasis>
|
||
</taxonomicName>
|
||
is due to a partial or complete genetic isolation of local populations living in fragmented habitats.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="FE8BEECAFFC44A7227ACB2FBFA43E7ED" pageId="49" pageNumber="50" type="key">
|
||
<paragraph id="B62EBD41FFC44A7227ACB2FBFB43E6E3" blockId="49.[151,1180,151,178]" box="[151,1180,151,178]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7227ACB2FBFB43E6E3" bold="true" box="[151,1180,151,178]" pageId="49" pageNumber="50">
|
||
Identification key to males of Australian species of
|
||
<taxonomicName id="7191C6C2FFC44A722428B2FBFB43E6E3" ID-CoL="8XSDD" authorityName="Franz" authorityYear="1975" box="[787,1180,151,178]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="subGenus" subGenus="Scottiscydmaenus">
|
||
<emphasis id="84E56153FFC44A722428B2FBFC74E6E0" bold="true" box="[787,939,151,178]" italics="true" pageId="49" pageNumber="50">Scydmaenus</emphasis>
|
||
(
|
||
<emphasis id="84E56153FFC44A722480B2FBFB4DE6E0" bold="true" box="[955,1170,151,178]" italics="true" pageId="49" pageNumber="50">Scottiscydmaenus</emphasis>
|
||
)
|
||
</taxonomicName>
|
||
</emphasis>
|
||
</paragraph>
|
||
<key id="C200EA70FFC44A7227ACB2B0FA43E7ED" pageId="49" pageNumber="50">
|
||
<keyStep id="0D65A5E4FFC44A7227ACB2B0FA43E75C" pageId="49" pageNumber="50">
|
||
<paragraph id="B62EBD41FFC44A7227ACB2B0FA43E6A3" blockId="49.[151,1437,220,447]" box="[151,1436,220,242]" pageId="49" pageNumber="50">
|
||
<keyLead id="0D601E74FFC44A7227ACB2B0FA43E6A3" box="[151,1436,220,242]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7227ACB2B0FF7CE6A3" bold="true" box="[151,163,220,241]" pageId="49" pageNumber="50">1</emphasis>
|
||
Head (together with mouthparts) clearly elongate...........................................................
|
||
<emphasis id="84E56153FFC44A7222ABB2B0FA43E6A3" bold="true" box="[1424,1436,220,241]" pageId="49" pageNumber="50">2</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFC44A7227ACB296FA43E75C" blockId="49.[151,1437,220,447]" box="[151,1436,249,271]" pageId="49" pageNumber="50">
|
||
<keyLead id="0D601E74FFC44A7227ACB296FA43E75C" box="[151,1436,249,271]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7227ACB296FF40E75D" bold="true" box="[151,159,250,271]" pageId="49" pageNumber="50">-</emphasis>
|
||
Head about as long as broad............................................................................
|
||
<emphasis id="84E56153FFC44A7222ABB295FA43E75C" bold="true" box="[1424,1436,249,270]" pageId="49" pageNumber="50">3</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="0D65A5E4FFC44A7227ACB37BFA43E71B" pageId="49" pageNumber="50">
|
||
<paragraph id="B62EBD41FFC44A7227ACB37BFA43E77E" blockId="49.[151,1437,220,447]" box="[151,1436,278,301]" pageId="49" pageNumber="50">
|
||
<keyLead id="0D601E74FFC44A7227ACB37BFA43E77E" box="[151,1436,278,301]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7227ACB37BFF7CE77E" bold="true" box="[151,163,279,300]" pageId="49" pageNumber="50">2</emphasis>
|
||
Profemoral tooth sharp-angled and elongate; BL>
|
||
<quantity id="716910A4FFC44A7225F0B37BFCDDE77E" box="[715,770,279,300]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="49" pageNumber="50" unit="mm" value="2.0">2 mm</quantity>
|
||
..................... ..
|
||
<emphasis id="84E56153FFC44A72232CB37AFA43E77E" bold="true" box="[1047,1436,278,300]" pageId="49" pageNumber="50">
|
||
<taxonomicName id="7191C6C2FFC44A72232CB37AFABFE77E" authority="Jałoszyński, 2023" authorityName="Jałoszyński" authorityYear="2023" box="[1047,1376,278,300]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="brachyponerophilus" status="sp. nov.">
|
||
<emphasis id="84E56153FFC44A72232CB37AFABFE77E" bold="true" box="[1047,1376,278,300]" italics="true" pageId="49" pageNumber="50">Scydmaenus brachyponerophilus</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="9FD6DC28FFC44A72225DB37BFA43E77E" box="[1382,1436,279,300]" pageId="49" pageNumber="50" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFC44A7227ACB358FA43E71B" blockId="49.[151,1437,220,447]" box="[151,1436,308,330]" pageId="49" pageNumber="50">
|
||
<keyLead id="0D601E74FFC44A7227ACB358FA43E71B" box="[151,1436,308,330]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7227ACB358FF40E71B" bold="true" box="[151,159,308,329]" pageId="49" pageNumber="50">-</emphasis>
|
||
Profemoral tooth at most right-angled, broader than long or as long as broad; BL <
|
||
<quantity id="716910A4FFC44A72233AB358FBE7E718" box="[1025,1080,308,330]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" pageId="49" pageNumber="50" unit="mm" value="2.0">2 mm</quantity>
|
||
...
|
||
<taxonomicName id="7191C6C2FFC44A722355B358FA43E71B" ID-CoL="4W3XD" authority="Franz" authorityName="Franz" box="[1134,1436,308,330]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="clarkianus">
|
||
<emphasis id="84E56153FFC44A722355B358FA43E71B" bold="true" box="[1134,1436,308,330]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A722355B358FA86E718" bold="true" box="[1134,1369,308,330]" italics="true" pageId="49" pageNumber="50">Scydmaenus clarkianus</emphasis>
|
||
Franz
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
<keyStep id="0D65A5E4FFC44A7227ACB33DFA43E7ED" pageId="49" pageNumber="50">
|
||
<paragraph id="B62EBD41FFC44A7227ACB33DFA42E7D6" blockId="49.[151,1437,220,447]" pageId="49" pageNumber="50">
|
||
<keyLead id="0D601E74FFC44A7227ACB33DFA42E7D6" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7227ACB33DFF7CE734" bold="true" box="[151,163,337,358]" pageId="49" pageNumber="50">3</emphasis>
|
||
Length of temple in dorsal view equal to about 3 × length of eye; body slender; aedeagus in dorsal view with subapical broadening as wide as sub-basal broadening.................................................
|
||
<taxonomicName id="7191C6C2FFC44A7223B6B302FA42E7D6" ID-CoL="4W3XG" authority="Sharp" authorityName="Sharp" box="[1165,1437,366,388]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="optatus">
|
||
<emphasis id="84E56153FFC44A7223B6B302FA42E7D6" bold="true" box="[1165,1437,366,388]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7223B6B302FA87E7D6" bold="true" box="[1165,1368,366,388]" italics="true" pageId="49" pageNumber="50">Scydmaenus optatus</emphasis>
|
||
Sharp
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
<paragraph id="B62EBD41FFC44A7227ACB3E0FA43E7ED" blockId="49.[151,1437,220,447]" pageId="49" pageNumber="50">
|
||
<keyLead id="0D601E74FFC44A7227ACB3E0FA43E7ED" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A7227ACB3E0FF40E7F3" bold="true" box="[151,159,396,417]" pageId="49" pageNumber="50">-</emphasis>
|
||
Length of temple in dorsal view equal to about 2 × length of eye; body robust; aedeagus in dorsal view with subapical broadening narrower than sub-basal narrowing............................................
|
||
<taxonomicName id="7191C6C2FFC44A722348B3C5FA43E7ED" ID-CoL="4W3XK" authority="Franz" authorityName="Franz" box="[1139,1436,425,447]" class="Insecta" family="Staphylinidae" genus="Scydmaenus" kingdom="Animalia" order="Coleoptera" pageId="49" pageNumber="50" phylum="Arthropoda" rank="species" species="swanensis">
|
||
<emphasis id="84E56153FFC44A722348B3C5FA43E7ED" bold="true" box="[1139,1436,425,447]" pageId="49" pageNumber="50">
|
||
<emphasis id="84E56153FFC44A722348B3C5FA86E7EC" bold="true" box="[1139,1369,425,447]" italics="true" pageId="49" pageNumber="50">Scydmaenus swanensis</emphasis>
|
||
Franz
|
||
</emphasis>
|
||
</taxonomicName>
|
||
</keyLead>
|
||
</paragraph>
|
||
</keyStep>
|
||
</key>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |