410 lines
82 KiB
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410 lines
82 KiB
XML
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<mods:title id="23F8227880BE13C73C21BCC7EC31D5E6">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:affiliation id="FD450513A2FAEC7666C0E0FBEBB6BE06">Faculty of Biology, VNU University of Science, Vietnam National University, Hanoi, Nguyen Trai, Thanh Xuan, Hanoi, Vietnam.</mods:affiliation>
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<taxonomicName id="4BF74D18FFFD291A3A05FCCC74490C83" ID-CoL="4SNHH" authority="Tran & Yang, 2006" authorityName="Tran & Yang" authorityYear="2006" box="[538,1050,894,922]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="anderseni">
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<emphasis id="BE83EA89FFFD291A3A05FCCC73610C83" bold="true" box="[538,818,894,921]" italics="true" pageId="33" pageNumber="34">Rhyacobates anderseni</emphasis>
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<bibRefCitation id="E8664B6AFFFD291A3B26FC3274490C83" author="Tran & Yang" box="[825,1050,895,922]" firstAuthor="Tran" pageId="33" pageNumber="34" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang, 2006</bibRefCitation>
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<paragraph id="8C48369BFFFD291A3A65FC1073EA0CA6" blockId="33.[538,1050,894,956]" box="[634,953,929,956]" pageId="33" pageNumber="34">
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<figureCitation id="14CC2A1EFFFD291A3A65FC1072800CA6" box="[634,723,930,956]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="33" pageNumber="34">Figs 3F</figureCitation>
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,
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<figureCitation id="14CC2A1EFFFD291A3AC0FC1073500CA6" box="[735,771,930,956]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="33" pageNumber="34">4E</figureCitation>
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,
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<figureCitation id="14CC2A1EFFFD291A3B11FC1373610CA6" box="[782,818,929,956]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="33" pageNumber="34">5E</figureCitation>
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,
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<figureCitation id="14CC2A1EFFFD291A3B21FC1073320CA6" box="[830,865,930,956]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="33" pageNumber="34">6E</figureCitation>
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,
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<figureCitation id="14CC2A1EFFFD291A3B72FC1073C30CA6" box="[877,912,930,956]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="33" pageNumber="34">8E</figureCitation>
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,
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<figureCitation id="14CC2A1EFFFD291A3B83FC1073EA0CA6" box="[924,953,930,956]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">23</figureCitation>
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<taxonomicName id="4BF74D18FFFD291A38A2FC5073430CE6" ID-CoL="4SNHH" authority="Tran & Yang, 2006: 14 - 16" authorityName="Tran & Yang" authorityPageNumber="14 - 16" authorityYear="2006" box="[189,784,994,1021]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="anderseni">
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<emphasis id="BE83EA89FFFD291A38A2FC50719D0CE6" box="[189,462,994,1020]" italics="true" pageId="33" pageNumber="34">Rhyacobates anderseni</emphasis>
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<bibRefCitation id="E8664B6AFFFD291A39CAFC5173430CE6" author="Tran & Yang" box="[469,784,994,1021]" firstAuthor="Tran" page="14 - 16" pageId="33" pageNumber="34" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang, 2006: 14–16</bibRefCitation>
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, figs 7–16, 27 (original description).
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<taxonomicName id="4BF74D18FFFD291A38A2FB91719D0B27" ID-CoL="4SNHH" authorityName="Tran & Yang" authorityYear="2006" box="[189,462,1059,1085]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="33" pageNumber="34" phylum="Arthropoda" rank="species" species="anderseni">
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<emphasis id="BE83EA89FFFD291A38A2FB91719D0B27" box="[189,462,1059,1085]" italics="true" pageId="33" pageNumber="34">Rhyacobates anderseni</emphasis>
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–
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<treatmentCitation id="0D56108AFFFD291A39F4FB9673720B24" ID-DOI="http://doi.org/10.5281/zenodo.4509714" ID-Zenodo-Dep="4509714" author="Tran A. D. & Nguyen X. Q." box="[491,801,1059,1086]" httpUri="http://treatment.plazi.org/id/4F67DC7C68699F6DFF00F8A318F6FD2F" page="513" pageId="33" pageNumber="34" year="2016">
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<bibRefCitation id="E8664B6AFFFD291A39F4FB9673720B24" author="Tran & Nguyen" box="[491,801,1059,1086]" firstAuthor="Tran" page="513" pageId="33" pageNumber="34" pagination="501 - 516" refId="ref33477" refString="Tran A. D. & Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran & Nguyen 2016: 513</bibRefCitation>
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(with remarks).
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<heading id="D70081F7FFFD291A38A2FBD671650B64" bold="true" box="[189,310,1124,1150]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
|
||
<emphasis id="BE83EA89FFFD291A38A2FBD671650B64" bold="true" box="[189,310,1124,1150]" pageId="33" pageNumber="34">Diagnosis</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFD291A38A2FB207273091D" blockId="33.[188,1401,1170,1544]" pageId="33" pageNumber="34">
|
||
Body length of females 6.79–7.30, of males 6.00–6.20. Both sexes: mesonotum mainly black with a median brownish-yellow stripe (broader in female, narrower in male); metanotum chiefly blackish, without yellow markings (
|
||
<figureCitation id="14CC2A1EFFFD291A39E5FB6A720B0BE8" box="[506,600,1240,1266]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="33" pageNumber="34">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFD291A3A78FB6A72CD0BE8" box="[615,670,1240,1266]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">23A</figureCitation>
|
||
). Female: hind margin of metanotum with a pointed median process extending over abdominal tergum I (
|
||
<figureCitation id="14CC2A1EFFFD291A3AF4FB4873D20A0E" box="[747,897,1274,1300]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">Fig. 23C–D</figureCitation>
|
||
); pronotum mainly black with a median brownish-yellow spot; posterior margin of abdominal segment VII with four processes, dorsally with a long process terminating each connexivum (
|
||
<figureCitation id="14CC2A1EFFFD291A3AF6FAF2732B0A40" box="[745,888,1344,1370]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">Fig. 23F–G</figureCitation>
|
||
), laterally with a pair of pointed processes (
|
||
<figureCitation id="14CC2A1EFFFD291A38D7FAD071060A66" box="[200,341,1378,1404]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">Fig. 23F, H</figureCitation>
|
||
), ventrally almost truncate, without median process (
|
||
<figureCitation id="14CC2A1EFFFD291A3BCCFAD074100A66" box="[979,1091,1378,1404]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">Fig. 23H</figureCitation>
|
||
). Male: middle trochanter without spines; middle femur with scattered small spines, not arranged in distinct row (
|
||
<figureCitation id="14CC2A1EFFFD291A3CB0FA37754F0A85" box="[1199,1308,1413,1439]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">Fig. 23K</figureCitation>
|
||
); length of middle tibia ca 1.8–1.9 times length of hind tibia; proctiger with rounded lobes laterally (
|
||
<figureCitation id="14CC2A1EFFFD291A3D0FFA1A75230AD8" box="[1296,1392,1448,1474]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="33" pageNumber="34">Figs 6E</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFD291A38A0FA7870A10AFE" box="[191,242,1482,1508]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">23L</figureCitation>
|
||
); paramere relatively stout and evenly curved, middle part thickened, distal part tapering towards rounded apex (
|
||
<figureCitation id="14CC2A1EFFFD291A3972FA5F7198091D" box="[365,459,1517,1543]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="33" pageNumber="34">Figs 8E</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFD291A39C8FA5F7240091D" box="[471,531,1517,1543]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="33" pageNumber="34">23M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFFD291A38A2F99C731608FC" pageId="33" pageNumber="34" type="materials_examined">
|
||
<paragraph id="8C48369BFFFD291A38A2F99C71F90952" blockId="33.[189,426,1582,1608]" box="[189,426,1582,1608]" pageId="33" pageNumber="34">
|
||
<heading id="D70081F7FFFD291A38A2F99C71F90952" bold="true" box="[189,426,1582,1608]" fontSize="11" level="3" pageId="33" pageNumber="34" reason="3">
|
||
<emphasis id="BE83EA89FFFD291A38A2F99C71F90952" bold="true" box="[189,426,1582,1608]" pageId="33" pageNumber="34">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFD291A38F5F9EE71FD096C" blockId="33.[189,483,1628,1689]" box="[234,430,1628,1654]" pageId="33" pageNumber="34">
|
||
<emphasis id="BE83EA89FFFD291A38F5F9EE71FD096C" bold="true" box="[234,430,1628,1654]" pageId="33" pageNumber="34">
|
||
<typeStatus id="534C8839FFFD291A38F5F9EE7174096C" box="[234,295,1628,1654]" pageId="33" pageNumber="34">Type</typeStatus>
|
||
specimens
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFD291A38A2F9CD71B00982" blockId="33.[189,483,1628,1689]" box="[189,483,1662,1689]" pageId="33" pageNumber="34">
|
||
See
|
||
<bibRefCitation id="E8664B6AFFFD291A38F1F9CD718C0982" author="Tran & Yang" box="[238,479,1662,1689]" firstAuthor="Tran" pageId="33" pageNumber="34" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang (2006)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFD291A38F5F97271B709C3" blockId="33.[189,1399,1727,1823]" box="[234,484,1727,1754]" pageId="33" pageNumber="34">
|
||
<emphasis id="BE83EA89FFFD291A38F5F97271B709C3" bold="true" box="[234,484,1727,1754]" pageId="33" pageNumber="34">Non-type specimens</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFD291A38A2F95073FC0805" blockId="33.[189,1399,1727,1823]" pageId="33" pageNumber="34">
|
||
<materialsCitation id="3C9F3CC6FFFD291A38A2F95073F80805" ID-GBIF-Occurrence="4429869318" collectingDate="2016-07-30" collectionCode="NKUM" collectorName="Zhen Ye" country="China" elevation="879" latitude="22.551138" location="Cai-yang River" longLatPrecision="1" longitude="101.087395" municipality="Jing-hong City" pageId="33" pageNumber="34" specimenCount="3" specimenCount-female="3" stateProvince="Yunnan">
|
||
<collectingCountry id="F4E0760BFFFD291A38A2F950714B09E6" box="[189,280,1762,1788]" name="China" pageId="33" pageNumber="34">CHINA</collectingCountry>
|
||
• 5 ♁♁,
|
||
<specimenCount id="9AF1FD12FFFD291A3990F950718709E7" box="[399,468,1762,1789]" pageId="33" pageNumber="34" type="female">3 ♀♀</specimenCount>
|
||
(apterous);
|
||
<collectingRegion id="4E33F879FFFD291A3A78F951736509E6" box="[615,822,1762,1789]" country="China" name="Yunnan" pageId="33" pageNumber="34">Yunnan Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6C2CACE1FFFD291A3B5AF95073AA09E6" box="[837,1017,1762,1788]" pageId="33" pageNumber="34">Jing-hong City</collectingMunicipality>
|
||
,
|
||
<location id="89286040FFFD291A3C17F950749009E6" LSID="urn:lsid:plazi:treatment:045E878DFFFD291F3A05FCCC714B0E96:89286040FFFD291A3C17F950749009E6" box="[1032,1219,1762,1788]" country="China" latitude="22.551138" longLatPrecision="1" longitude="101.087395" municipality="Jing-hong City" name="Cai-yang River" pageId="33" pageNumber="34" stateProvince="Yunnan">Cai-yang River</location>
|
||
;
|
||
<geoCoordinate id="E9C3505CFFFD291A3CCDF953752109E7" box="[1234,1394,1761,1789]" degrees="22" direction="north" minutes="33" orientation="latitude" pageId="33" pageNumber="34" precision="1" seconds="4.1" value="22.551138">22°33′4.1″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFFD291A38A2F8B171350805" box="[189,358,1795,1823]" degrees="101" direction="east" minutes="5" orientation="longitude" pageId="33" pageNumber="34" precision="1" seconds="14.6" value="101.087395">101°5′14.6″ E</geoCoordinate>
|
||
;
|
||
<elevation id="07DAD1A8FFFD291A396DF8B771AF0805" box="[370,508,1797,1823]" metricMagnitude="2" metricUnit="m" metricValue="8.79" pageId="33" pageNumber="34" unit="m" value="879.0">
|
||
<quantity id="4B0F9B7EFFFD291A396DF8B771EF0805" box="[370,444,1797,1823]" metricMagnitude="2" metricUnit="m" metricValue="8.79" pageId="33" pageNumber="34" unit="m" value="879.0">879 m</quantity>
|
||
a.s.l.
|
||
</elevation>
|
||
;
|
||
<date id="F849105BFFFD291A3A15F8B772CE0805" box="[522,669,1797,1823]" pageId="33" pageNumber="34" value="2016-07-30">
|
||
<collectingDate id="E80DE9B3FFFD291A3A15F8B772CE0805" box="[522,669,1797,1823]" pageId="33" pageNumber="34" value="2016-07-30">30 Jul. 2016</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFFD291A3AB6F8B773580805" box="[681,779,1797,1823]" pageId="33" pageNumber="34">Zhen Ye</collectorName>
|
||
leg.;
|
||
<collectionCode id="EAE6AE5EFFFD291A3B52F8B773F80805" box="[845,939,1797,1823]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="33" pageNumber="34">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFD291A38A2F8F4749B08BC" blockId="33.[189,1398,1860,1958]" pageId="33" pageNumber="34">
|
||
<materialsCitation id="3C9F3CC6FFFD291A38A2F8F471C708BC" ID-GBIF-Occurrence="4429869322" collectingDate="2022-04-21" collectionCode="ZVNU" collectorName="A. D. Tran" country="Vietnam" latitude="18.292084" location="Khe Nam Cham" longLatPrecision="1" longitude="105.355194" municipality="Vu Quang National Park" pageId="33" pageNumber="34" specimenCount="1" stateProvince="Ha Tinh">
|
||
<collectingCountry id="F4E0760BFFFD291A38A2F8F47117087A" box="[189,324,1862,1888]" name="Vietnam" pageId="33" pageNumber="34">VIETNAM</collectingCountry>
|
||
• 1 ♁ (apterous), 1 ♁ (macropterous);
|
||
<collectingRegion id="4E33F879FFFD291A3B1EF8F47382087A" box="[769,977,1862,1888]" country="Vietnam" name="Ha Tinh" pageId="33" pageNumber="34">Hà Tĩnh Province</collectingRegion>
|
||
,
|
||
<collectingMunicipality id="6C2CACE1FFFD291A3BC4F8F474AE087A" box="[987,1277,1862,1888]" pageId="33" pageNumber="34">Vũ Quang National Park</collectingMunicipality>
|
||
,
|
||
<location id="89286040FFFD291A3D18F8F471500898" LSID="urn:lsid:plazi:treatment:045E878DFFFD291F3A05FCCC714B0E96:89286040FFFD291A3D18F8F471500898" country="Vietnam" latitude="18.292084" longLatPrecision="1" longitude="105.355194" municipality="Vu Quang National Park" name="Khe Nam Cham" pageId="33" pageNumber="34" stateProvince="Ha Tinh">Khe Nam Châm</location>
|
||
stream, site #1, upstream;
|
||
<geoCoordinate id="E9C3505CFFFD291A3A56F8D572A40899" box="[585,759,1895,1923]" degrees="18" direction="north" minutes="17" orientation="latitude" pageId="33" pageNumber="34" precision="1" seconds="31.5" value="18.292084">18°17′31.5″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFFD291A3B1AF8D573EC0899" box="[773,959,1895,1923]" degrees="105" direction="east" minutes="21" orientation="longitude" pageId="33" pageNumber="34" precision="1" seconds="18.7" value="105.355194">105°21′18.7″ E</geoCoordinate>
|
||
;
|
||
<date id="F849105BFFFD291A3BD1F8DB743D0899" box="[974,1134,1897,1923]" pageId="33" pageNumber="34" value="2022-04-21">
|
||
<collectingDate id="E80DE9B3FFFD291A3BD1F8DB743D0899" box="[974,1134,1897,1923]" pageId="33" pageNumber="34" value="2022-04-21">21 Apr. 2022</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFFD291A3C64F8DB74A00899" box="[1147,1267,1897,1923]" pageId="33" pageNumber="34">A.D. Tran</collectorName>
|
||
<emphasis id="BE83EA89FFFD291A3CE2F8DB75690898" box="[1277,1338,1896,1922]" italics="true" pageId="33" pageNumber="34">et al.</emphasis>
|
||
leg.; TAD2208;
|
||
<collectionCode id="EAE6AE5EFFFD291A395CF83E71C708BC" box="[323,404,1932,1958]" pageId="33" pageNumber="34">ZVNU</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3C9F3CC6FFFD291A39B1F839749608BC" ID-GBIF-Occurrence="4429869306" box="[430,1221,1931,1958]" collectingDate="2022-04-21" collectionCode="NKUM" collectorName="A. D. Tran" country="Vietnam" latitude="18.292084" location="Khe Nam Cham" longLatPrecision="1" longitude="105.355194" municipality="Vu Quang National Park" pageId="33" pageNumber="34" specimenCount="1" specimenCount-female="1" stateProvince="Ha Tinh">
|
||
<specimenCount id="9AF1FD12FFFD291A39B1F839718908BC" box="[430,474,1931,1958]" pageId="33" pageNumber="34" type="female">1 ♀</specimenCount>
|
||
(macropterous); same collection data as for preceding;
|
||
<collectionCode id="EAE6AE5EFFFD291A3C78F83E749608BC" box="[1127,1221,1932,1958]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="33" pageNumber="34">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFD291A38A2F87E731608FC" blockId="33.[189,837,1996,2023]" box="[189,837,1996,2023]" pageId="33" pageNumber="34">
|
||
GPS data of previous records: see
|
||
<bibRefCitation id="E8664B6AFFFD291A3A4FF87F731208FC" author="Tran & Yang" box="[592,833,1996,2023]" firstAuthor="Tran" pageId="33" pageNumber="34" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang (2006)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="D8886613FFFE291938A2F949749308F3" ID-DOI="http://doi.org/10.5281/zenodo.8385686" ID-Zenodo-Dep="8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="34" pageNumber="35" startId="34.[189,232,1787,1813]" targetBox="[282,1322,280,1711]" targetPageId="34">
|
||
<paragraph id="8C48369BFFFE291938A2F949749308F3" blockId="34.[189,1399,1787,2025]" pageId="34" pageNumber="35">
|
||
<emphasis id="BE83EA89FFFE291938A2F949714D080C" bold="true" box="[189,286,1787,1814]" pageId="34" pageNumber="35">Fig. 23.</emphasis>
|
||
Morphological features of
|
||
<taxonomicName id="4BF74D18FFFE29193A72F94E7428080C" authority="Tran & Yang, 2006" authorityName="Tran & Yang" authorityYear="2006" box="[621,1147,1788,1814]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="34" pageNumber="35" phylum="Arthropoda" rank="species" species="anderseni">
|
||
<emphasis id="BE83EA89FFFE29193A72F94E73D1080C" box="[621,898,1788,1814]" italics="true" pageId="34" pageNumber="35">Rhyacobates anderseni</emphasis>
|
||
<bibRefCitation id="E8664B6AFFFE29193B93F94E7428080C" author="Tran & Yang" box="[908,1147,1788,1814]" firstAuthor="Tran" pageId="34" pageNumber="35" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
.
|
||
<emphasis id="BE83EA89FFFE29193C95F94974CC080F" bold="true" box="[1162,1183,1787,1813]" pageId="34" pageNumber="35">A</emphasis>
|
||
. Body of female, dorsal view.
|
||
<emphasis id="BE83EA89FFFE2919394BF8AD71340823" bold="true" box="[340,359,1823,1849]" pageId="34" pageNumber="35">B</emphasis>
|
||
. Body of female, excluding head and prothorax, ventral view.
|
||
<emphasis id="BE83EA89FFFE29193C53F8AD74320823" bold="true" box="[1100,1121,1823,1849]" pageId="34" pageNumber="35">C</emphasis>
|
||
. Metathorax of female, dorsal view.
|
||
<emphasis id="BE83EA89FFFE29193945F8F0713C0846" bold="true" box="[346,367,1858,1884]" pageId="34" pageNumber="35">D</emphasis>
|
||
. Metathorax of female, lateral view.
|
||
<emphasis id="BE83EA89FFFE29193B26F8F0731E0846" bold="true" box="[825,845,1858,1884]" pageId="34" pageNumber="35">E</emphasis>
|
||
. Body of female, lateral view.
|
||
<emphasis id="BE83EA89FFFE29193CD0F8F074B20846" bold="true" box="[1231,1249,1858,1884]" pageId="34" pageNumber="35">F</emphasis>
|
||
. Abdominal end of female, lateral view.
|
||
<emphasis id="BE83EA89FFFE29193A0FF8D772740865" bold="true" box="[528,551,1893,1919]" pageId="34" pageNumber="35">G</emphasis>
|
||
. Abdominal end of female, dorsal view.
|
||
<emphasis id="BE83EA89FFFE29193C0AF8D7747F0865" bold="true" box="[1045,1068,1893,1919]" pageId="34" pageNumber="35">H</emphasis>
|
||
. Abdominal end of female, ventral view.
|
||
<emphasis id="BE83EA89FFFE2919397DF83B713E08B9" bold="true" box="[354,365,1929,1955]" pageId="34" pageNumber="35">I</emphasis>
|
||
. Body of male, dorsal view.
|
||
<emphasis id="BE83EA89FFFE29193ACCF83B72B108B9" bold="true" box="[723,738,1929,1955]" pageId="34" pageNumber="35">J</emphasis>
|
||
. Body of male, excluding head and prothorax, ventral view.
|
||
<emphasis id="BE83EA89FFFE2919391CF81E714908DC" bold="true" box="[259,282,1964,1990]" pageId="34" pageNumber="35">K</emphasis>
|
||
. Basal part of right middle leg of male, ventral view.
|
||
<emphasis id="BE83EA89FFFE29193B94F81E73CC08DC" bold="true" box="[907,927,1964,1990]" pageId="34" pageNumber="35">L</emphasis>
|
||
. Proctiger of male, dorsal view.
|
||
<emphasis id="BE83EA89FFFE29193D04F81E756408DC" bold="true" box="[1307,1335,1964,1990]" pageId="34" pageNumber="35">M</emphasis>
|
||
. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="C4ED6510FFFF291838A2FEBA73BA09CD" pageId="35" pageNumber="36" type="description">
|
||
<paragraph id="8C48369BFFFF291838A2FEBA71AE0E38" blockId="35.[189,509,264,290]" box="[189,509,264,290]" pageId="35" pageNumber="36">
|
||
<heading id="D70081F7FFFF291838A2FEBA71AE0E38" bold="true" box="[189,509,264,290]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
|
||
<emphasis id="BE83EA89FFFF291838A2FEBA71AE0E38" bold="true" box="[189,509,264,290]" pageId="35" pageNumber="36">Supplemental description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838F5FE8571EB0E4B" blockId="35.[186,1399,311,549]" box="[234,440,311,337]" pageId="35" pageNumber="36">
|
||
<emphasis id="BE83EA89FFFF291838F5FE8571EB0E4B" bold="true" box="[234,440,311,337]" pageId="35" pageNumber="36">Apterous female</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A2FEE9757B0D3E" blockId="35.[186,1399,311,549]" pageId="35" pageNumber="36">
|
||
<smallCapsWord id="8AAEA047FFFF291838A2FEE971380E69" baselines="366,366" box="[189,363,347,373]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="35" pageNumber="36">MEASUREMENTS</smallCapsWord>
|
||
. Body length 6.79–7.30, width 2.00–2.60, head width 1.32, interocular width 0.58, eye length (dorsal view) 0.59; relative lengths of antennal segments I–IV: 3:0.86: 1.21: 0.74; pronotum: length 0.73, width 1.47; mesonotum: length 2.36, width 2.54; metanotum: length 0.88, width 1.98; abdomen length (ventral view) 2.88; abdominal sternum VII: length 1.23, width 1.18; abdominal mediotergite I: length 0.15, width 0.96; relative lengths of leg segments (femur:tibia: tarsal segment I:tarsal segment II): fore leg: 3.03: 2.35:1.17:0.83, middle leg: 8.75: 5.35:2.63:0.45, hind leg: 8.85:3.08:0.15: 0.21.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A2FDFE75790DD5" blockId="35.[189,1398,588,720]" pageId="35" pageNumber="36">
|
||
<smallCapsWord id="8AAEA047FFFF291838A2FDFE711B0D7E" baselines="607,607" box="[189,328,588,614]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Coloration" pageId="35" pageNumber="36">COLORATION</smallCapsWord>
|
||
. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum mainly black with a median brownish-yellow stripe. Metanotum chiefly blackish, without yellow markings (
|
||
<figureCitation id="14CC2A1EFFFF29183AD4FD20737A0DB6" box="[715,809,658,684]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="35" pageNumber="36">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFF29183B26FD2073230DB6" box="[825,880,658,684]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">23A</figureCitation>
|
||
). Connexivum dorsally blackish. Venter of female chiefly blackish with a median yellowish spot (
|
||
<figureCitation id="14CC2A1EFFFF29183B23FD0773FB0DD5" box="[828,936,693,719]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23B</figureCitation>
|
||
). Abdominal venter light yellow.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A2FD4572220B1D" blockId="35.[187,1400,759,1032]" pageId="35" pageNumber="36">
|
||
<smallCapsWord id="8AAEA047FFFF291838A2FD45717F0C15" baselines="778,778" box="[189,300,759,785]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Abdomen" pageId="35" pageNumber="36">ABDOMEN</smallCapsWord>
|
||
. Hind margin of metanotum produced as a median protuberance over abdominal tergum I (
|
||
<figureCitation id="14CC2A1EFFFF291838D9FCA871060C2E" box="[198,341,794,820]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23C–D</figureCitation>
|
||
). Abdomen relatively short, nearly straight, moderately curved dorsad posteriorly (
|
||
<figureCitation id="14CC2A1EFFFF29183D0BFCA875210C2F" box="[1300,1394,794,821]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="35" pageNumber="36">Figs 5E</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFF291838A3FC8F70BC0C4D" box="[188,239,829,855]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">23E</figureCitation>
|
||
). Connexivum on abdominal segments I–VI dorsally blackish, forming a straight line in dorsal view (
|
||
<figureCitation id="14CC2A1EFFFF29183917FCD271350C60" box="[264,358,864,890]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="35" pageNumber="36">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFF2918396AFCD271FF0C60" box="[373,428,864,890]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">23A</figureCitation>
|
||
). Abdominal segment VII elongate, nearly as long as three preceding abdominal segments together (
|
||
<figureCitation id="14CC2A1EFFFF291839BBFC31725D0C84" box="[420,526,899,926]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23E</figureCitation>
|
||
), completely enclosing genital segments (
|
||
<figureCitation id="14CC2A1EFFFF29183BE2FC31740A0C84" box="[1021,1113,899,926]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="35" pageNumber="36">Figs 3F</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFF29183C79FC3674CE0C84" box="[1126,1181,900,926]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">23G</figureCitation>
|
||
). Posterior margin of abdominal segment VII with four processes, dorsally with a long, slender process terminating each connexivum (
|
||
<figureCitation id="14CC2A1EFFFF29183942FC7871940CFE" box="[349,455,970,996]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23G</figureCitation>
|
||
), laterally with a pair of pointed processes (
|
||
<figureCitation id="14CC2A1EFFFF29183BA8FC78746F0CFE" box="[951,1084,970,996]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23F, H</figureCitation>
|
||
), ventrally almost truncate, without a median process (
|
||
<figureCitation id="14CC2A1EFFFF291839E9FC5F72370B1D" box="[502,612,1005,1031]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23H</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838F5FB9C71F20B53" blockId="35.[186,1400,1070,1309]" box="[234,417,1070,1097]" pageId="35" pageNumber="36">
|
||
<emphasis id="BE83EA89FFFF291838F5FB9C71F20B53" bold="true" box="[234,417,1070,1097]" pageId="35" pageNumber="36">Apterous male</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A2FBE0757B0A07" blockId="35.[186,1400,1070,1309]" pageId="35" pageNumber="36">
|
||
<smallCapsWord id="8AAEA047FFFF291838A2FBE071380B70" baselines="1125,1125" box="[189,363,1106,1132]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Measurements" pageId="35" pageNumber="36">MEASUREMENTS</smallCapsWord>
|
||
. Body length 6.00–6.20, width 1.70–1.90, head width 1.21, interocular width 0.55, eye length (dorsal view) 0.53; relative lengths of antennal segments I–IV: 2.84: 0.89:1.14:0.78; pronotum: length 0.68, width 1.31; mesonotum: length2.13,width 1.85; metanotum:length 0.64, width 1.65;abdomen length (ventral view) 1.94; abdominal sternum VII: length 0.38, width 0.59; abdominal mediotergite I: length 0.19, width 0.68; relative lengths of leg segments (femur:tibia: tarsal segment I:tarsal segment II): fore leg: 3.04: 2.44:0.89:0.54, middle leg: 8.80: 4.86:2.38:0.36, hind leg: 8.75:2.62:0.11: 0.15.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A6FAF173470AF0" blockId="35.[185,1399,1347,1515]" pageId="35" pageNumber="36">
|
||
C OLORATION. Median black spot of head posteriorly bifurcate. Pronotum mainly black with a median brownish-yellow spot. Mesonotum mainly black with a median brownish-yellow stripe. Metanotum chiefly blackish, without yellow markings (
|
||
<figureCitation id="14CC2A1EFFFF29183AA4FA3B73490ABE" box="[699,794,1417,1444]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="35" pageNumber="36">Figs 4E</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFF29183B39FA3873020ABE" box="[806,849,1418,1444]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">23I</figureCitation>
|
||
). Connexivum dorsally blackish. Metasternum anteriorly blackish and posteriorly light yellow (
|
||
<figureCitation id="14CC2A1EFFFF29183B19FA1F733E0ADD" box="[774,877,1453,1479]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23J</figureCitation>
|
||
). Abdominal venter anteriorly blackish and posteriorly light yellow or completely light yellow.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A2F9A071270955" blockId="35.[187,1398,1553,1615]" pageId="35" pageNumber="36">
|
||
<smallCapsWord id="8AAEA047FFFF291838A2F9A070B80930" baselines="1573,1573" box="[189,235,1554,1580]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Leg" pageId="35" pageNumber="36">LEG</smallCapsWord>
|
||
. Middle trochanter without spines; middle femur with scattered small spines, not arranged in distinct row (
|
||
<figureCitation id="14CC2A1EFFFF291838E6F98771340955" box="[249,359,1589,1615]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A2F9C473BA09CD" blockId="35.[189,1399,1654,1751]" pageId="35" pageNumber="36">
|
||
<smallCapsWord id="8AAEA047FFFF291838A2F9C471620995" baselines="1673,1674" box="[189,305,1654,1680]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Genitalia" pageId="35" pageNumber="36">GENITALIA</smallCapsWord>
|
||
. Abdominal segment VIII ventro-laterally impressed. Pygophore large, ovate. Proctiger with rounded lobes laterally (
|
||
<figureCitation id="14CC2A1EFFFF29183A04F92B722809A9" box="[539,635,1689,1715]" captionStart="Fig" captionStartId="8.[189,232,1892,1918]" captionTargetBox="[229,1341,570,1825]" captionTargetId="figure-92@8.[189,1398,537,1851]" captionTargetPageId="8" captionText="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016. G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="35" pageNumber="36">Figs 6E</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFF29183A96F92B72EF09A9" box="[649,700,1689,1715]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">23L</figureCitation>
|
||
). Paramere relatively stout and evenly curved, middle part thickened, distal part tapering towards rounded apex (
|
||
<figureCitation id="14CC2A1EFFFF29183B2AF90E73C709CD" box="[821,916,1724,1751]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen & Chen, 1995. J. R. recurvus Andersen & Chen, 1995. K. R. scorpio Andersen & Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="35" pageNumber="36">Figs 8E</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFFF29183BBFF90F738F09CD" box="[928,988,1725,1751]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">23M</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFFF291838A2F94C731A085C" box="[189,841,1790,1863]" pageId="35" pageNumber="36" type="distribution">
|
||
<paragraph id="8C48369BFFFF291838A2F94C71050802" blockId="35.[189,342,1790,1816]" box="[189,342,1790,1816]" pageId="35" pageNumber="36">
|
||
<heading id="D70081F7FFFF291838A2F94C71050802" bold="true" box="[189,342,1790,1816]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
|
||
<emphasis id="BE83EA89FFFF291838A2F94C71050802" bold="true" box="[189,342,1790,1816]" pageId="35" pageNumber="36">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291838A2F89E731A085C" blockId="35.[189,841,1836,1863]" box="[189,841,1836,1863]" pageId="35" pageNumber="36">
|
||
<collectingCountry id="F4E0760BFFFF291838A2F89E715B085C" box="[189,264,1836,1862]" name="China" pageId="35" pageNumber="36">China</collectingCountry>
|
||
:
|
||
<collectingRegion id="4E33F879FFFF2918390EF89F713C085D" box="[273,367,1837,1863]" country="China" name="Yunnan" pageId="35" pageNumber="36">Yunnan</collectingRegion>
|
||
;
|
||
<collectingCountry id="F4E0760BFFFF29183965F89E71B1085C" box="[378,482,1836,1862]" name="Vietnam" pageId="35" pageNumber="36">Vietnam</collectingCountry>
|
||
:
|
||
<collectingRegion id="4E33F879FFFF291839F2F89F721E085C" box="[493,589,1836,1863]" country="Vietnam" name="Ha Tinh" pageId="35" pageNumber="36">Hà Tĩnh</collectingRegion>
|
||
(
|
||
<bibRefCitation id="E8664B6AFFFF29183A43F89F736E085C" author="Tran & Yang" box="[604,829,1836,1863]" firstAuthor="Tran" pageId="35" pageNumber="36" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang 2006</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFFF291F38A2F8DC714B0E96" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36" type="discussion">
|
||
<paragraph id="8C48369BFFFF291838A2F8DC71F90892" blockId="35.[189,426,1902,1928]" box="[189,426,1902,1928]" pageId="35" pageNumber="36">
|
||
<heading id="D70081F7FFFF291838A2F8DC71F90892" bold="true" box="[189,426,1902,1928]" fontSize="11" level="3" pageId="35" pageNumber="36" reason="3">
|
||
<emphasis id="BE83EA89FFFF291838A2F8DC71F90892" bold="true" box="[189,426,1902,1928]" pageId="35" pageNumber="36">Comparative notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFFF291F38A2F82E714B0E96" blockId="35.[189,1398,1948,2010]" lastBlockId="36.[188,1399,265,396]" lastPageId="36" lastPageNumber="37" pageId="35" pageNumber="36">
|
||
<taxonomicName id="4BF74D18FFFF291838A2F82E718008AC" authorityName="Tran & Yang" authorityYear="2006" box="[189,467,1948,1974]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="35" pageNumber="36" phylum="Arthropoda" rank="species" species="anderseni">
|
||
<emphasis id="BE83EA89FFFF291838A2F82E718008AC" box="[189,467,1948,1974]" italics="true" pageId="35" pageNumber="36">Rhyacobates anderseni</emphasis>
|
||
</taxonomicName>
|
||
is distinct from all congeners in having a median process on the posterior margin of the metanotum in the female (
|
||
<figureCitation id="14CC2A1EFFFF29183A81F80D735808C0" box="[670,779,1983,2010]" captionStart="Fig" captionStartId="34.[189,232,1787,1813]" captionTargetBox="[282,1322,280,1711]" captionTargetId="figure-15@34.[221,1367,265,1739]" captionTargetPageId="34" captionText="Fig. 23. Morphological features of Rhyacobates anderseni Tran & Yang, 2006. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Metathorax of female, dorsal view. D. Metathorax of female, lateral view. E. Body of female, lateral view. F. Abdominal end of female, lateral view. G. Abdominal end of female, dorsal view. H. Abdominal end of female, ventral view. I. Body of male, dorsal view. J. Body of male, excluding head and prothorax, ventral view. K. Basal part of right middle leg of male, ventral view. L. Proctiger of male, dorsal view. M. Left paramere, lateral aspect, two different views. Scale bars: A–K = 1 mm; L–M = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385686" httpUri="https://zenodo.org/record/8385686/files/figure.png" pageId="35" pageNumber="36">Fig. 23D</figureCitation>
|
||
), which is present in three other ptilomerine genera, i.e.,
|
||
<taxonomicName id="4BF74D18FFF8291F38EBFEBB71D60E39" authorityName="Zettel & Chen" authorityYear="1996" box="[244,389,265,291]" class="Insecta" family="Gerridae" genus="Andersenius" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BE83EA89FFF8291F38EBFEBB71D60E39" box="[244,389,265,291]" italics="true" pageId="36" pageNumber="37">Andersenius</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4BF74D18FFF8291F3987FEBB72720E39" authorityName="Esaki" authorityYear="1930" box="[408,545,265,291]" class="Insecta" family="Gerridae" genus="Pleciobates" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BE83EA89FFF8291F3987FEBB72720E39" box="[408,545,265,291]" italics="true" pageId="36" pageNumber="37">Pleciobates</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFF8291F3A7DFEBB73D10E39" authority="Distant, 1910" authorityName="Distant" authorityYear="1910" box="[610,898,265,291]" class="Insecta" family="Gerridae" genus="Jucundus" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BE83EA89FFF8291F3A7DFEBB72820E39" box="[610,721,265,291]" italics="true" pageId="36" pageNumber="37">Jucundus</emphasis>
|
||
Distant, 1910
|
||
</taxonomicName>
|
||
. However, this species matches all other characteristics of
|
||
<taxonomicName id="4BF74D18FFF8291F3997FE9E724D0E5C" authorityName="Esaki" authorityYear="1923" box="[392,542,300,326]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="36" pageNumber="37" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="BE83EA89FFF8291F3997FE9E724D0E5C" box="[392,542,300,326]" italics="true" pageId="36" pageNumber="37">Rhyacobates</emphasis>
|
||
</taxonomicName>
|
||
defined by
|
||
<bibRefCitation id="E8664B6AFFF8291F3ABBFE9E739F0E5D" author="Andersen & Chen" box="[676,972,300,327]" firstAuthor="Andersen" pageId="36" pageNumber="37" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. & Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen & Chen (1995)</bibRefCitation>
|
||
, as discussed by
|
||
<bibRefCitation id="E8664B6AFFF8291F3C90FE9F70B40E73" author="Tran & Yang" firstAuthor="Tran" page="16" pageId="36" pageNumber="37" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang (2006: 16)</bibRefCitation>
|
||
. Future phylogenetic studies using molecular data may help to resolve the taxonomic position of this species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |