275 lines
51 KiB
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275 lines
51 KiB
XML
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<mods:title id="C83FF6E25108E1AA4957FBBA4FD2986F">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:namePart id="6E4BC386F40F670434A40D0B1095C48A">Leng, Zhaoqi</mods:namePart>
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<mods:affiliation id="B8D7A174E11ED2F50C4672293CB4E388">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:nameIdentifier id="34DDD6A5FDC2520A16BF6FE609F3A8F2" type="email">zhaoqileng0712@gmail.com</mods:nameIdentifier>
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<mods:namePart id="40BD42E8D4BF445C37C96B8C7239CF99">Tran, Anh Duc</mods:namePart>
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<mods:affiliation id="45A9DE060A8E905476061C581299E0D8">Faculty of Biology, VNU University of Science, Vietnam National University, Hanoi, Nguyen Trai, Thanh Xuan, Hanoi, Vietnam.</mods:affiliation>
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<mods:affiliation id="982CB2B4CDD75B9E451EF44999C01D19">Institute of Entomology, College of Life Sciences, Nankai University, Tianjin, 300071, China.</mods:affiliation>
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<mods:title id="ED2683DD4811814C595229114884A1A0">European Journal of Taxonomy</mods:title>
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<mods:date id="BD7B43BD01A1B098B870BB93EC23B7A6">2023</mods:date>
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<treatment id="045E878DFFF229143A35FB0A72A90B4E" ID-DOI="http://doi.org/10.5281/zenodo.10063667" ID-GBIF-Taxon="214246585" ID-Zenodo-Dep="10063667" LSID="urn:lsid:plazi:treatment:045E878DFFF229143A35FB0A72A90B4E" httpUri="http://treatment.plazi.org/id/045E878DFFF229143A35FB0A72A90B4E" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47">
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<heading id="D70081F7FFF229153A35FB0A74590BC8" box="[554,1034,1208,1235]" centered="true" fontSize="11" level="2" pageId="46" pageNumber="47" reason="2">
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<taxonomicName id="4BF74D18FFF229153A35FB0A74590BC8" ID-CoL="4SNHP" authority="Tran & Yang, 2006" authorityName="Tran & Yang" authorityYear="2006" box="[554,1034,1208,1235]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gongvo">
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<emphasis id="BE83EA89FFF229153A35FB0A73710BC9" bold="true" box="[554,802,1208,1235]" italics="true" pageId="46" pageNumber="47">Rhyacobates gongvo</emphasis>
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<bibRefCitation id="E8664B6AFFF229153B36FB0B74590BC8" author="Tran & Yang" box="[809,1034,1208,1235]" firstAuthor="Tran" pageId="46" pageNumber="47" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang, 2006</bibRefCitation>
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</taxonomicName>
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</heading>
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</paragraph>
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<paragraph id="8C48369BFFF229153ABAFB6973C30BEC" blockId="46.[554,1034,1208,1270]" box="[677,912,1243,1270]" pageId="46" pageNumber="47">
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<figureCitation id="14CC2A1EFFF229153ABAFB6973570BEC" box="[677,772,1243,1270]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="46" pageNumber="47">Figs 3K</figureCitation>
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,
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<figureCitation id="14CC2A1EFFF229153B0EFB6E736B0BEC" box="[785,824,1244,1270]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="46" pageNumber="47">4H</figureCitation>
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,
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<figureCitation id="14CC2A1EFFF229153B5BFB6973330BEC" box="[836,864,1243,1270]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="46" pageNumber="47">5J</figureCitation>
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,
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<figureCitation id="14CC2A1EFFF229153B73FB6E73C30BEC" box="[876,912,1244,1270]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran & Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen & Chen, 1995. D. R. recurvus Andersen & Chen, 1995. E. R. scorpio Andersen & Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran & Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="46" pageNumber="47">7A</figureCitation>
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</paragraph>
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<subSubSection id="C4ED6510FFF2291538A2FAAF74190A62" pageId="46" pageNumber="47" type="reference_group">
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<paragraph id="8C48369BFFF2291538A2FAAF74FD0A2D" blockId="46.[189,1198,1309,1336]" box="[189,1198,1309,1336]" pageId="46" pageNumber="47">
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<taxonomicName id="4BF74D18FFF2291538A2FAAF72A00A2D" ID-CoL="4SNHP" authority="Tran & Yang, 2006: 16 - 19" authorityName="Tran & Yang" authorityPageNumber="16 - 19" authorityYear="2006" box="[189,755,1309,1335]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gongvo">
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<emphasis id="BE83EA89FFF2291538A2FAAF71E30A2D" box="[189,432,1309,1335]" italics="true" pageId="46" pageNumber="47">Rhyacobates gongvo</emphasis>
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<bibRefCitation id="E8664B6AFFF2291539A8FAAF72A00A2D" author="Tran & Yang" box="[439,755,1309,1335]" firstAuthor="Tran" page="16 - 19" pageId="46" pageNumber="47" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang, 2006: 16–19</bibRefCitation>
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</taxonomicName>
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, figs 17–25, 28 (original description).
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</paragraph>
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<paragraph id="8C48369BFFF2291538A2FAEC74190A62" blockId="46.[189,1098,1374,1401]" box="[189,1098,1374,1401]" pageId="46" pageNumber="47">
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<treatmentCitationGroup id="ACE711B5FFF2291538A2FAEC74190A62" box="[189,1098,1374,1401]" pageId="46" pageNumber="47">
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<taxonomicName id="4BF74D18FFF2291538A2FAEC71E30A62" ID-CoL="4SNHP" authorityName="Tran & Yang" authorityYear="2006" box="[189,432,1374,1400]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="46" pageNumber="47" phylum="Arthropoda" rank="species" species="gongvo">
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<emphasis id="BE83EA89FFF2291538A2FAEC71E30A62" box="[189,432,1374,1400]" italics="true" pageId="46" pageNumber="47">Rhyacobates gongvo</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="0D56108AFFF2291539D2FAED73540A63" ID-DOI="http://doi.org/10.5281/zenodo.4509716" ID-Zenodo-Dep="4509716" author="Tran A. D. & Nguyen X. Q." box="[461,775,1374,1401]" httpUri="http://treatment.plazi.org/id/4F67DC7C686E9F6DFF00FD70182DF8ED" page="513" pageId="46" pageNumber="47" year="2016">
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<bibRefCitation id="E8664B6AFFF2291539D2FAED73540A63" author="Tran & Nguyen" box="[461,775,1374,1401]" firstAuthor="Tran" page="513" pageId="46" pageNumber="47" pagination="501 - 516" refId="ref33477" refString="Tran A. D. & Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran & Nguyen 2016: 513</bibRefCitation>
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, figs 41–42 (with remarks).
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</paragraph>
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<subSubSection id="C4ED6510FFF2291538A2FA2D74C4083C" pageId="46" pageNumber="47" type="diagnosis">
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<paragraph id="8C48369BFFF2291538A2FA2D71650AA3" blockId="46.[189,310,1439,1465]" box="[189,310,1439,1465]" pageId="46" pageNumber="47">
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<heading id="D70081F7FFF2291538A2FA2D71650AA3" bold="true" box="[189,310,1439,1465]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
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<emphasis id="BE83EA89FFF2291538A2FA2D71650AA3" bold="true" box="[189,310,1439,1465]" pageId="46" pageNumber="47">Diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8C48369BFFF2291538A2FA7C74C4083C" blockId="46.[188,1401,1486,1830]" pageId="46" pageNumber="47">
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Body length of apterous females 7.80–8.30, of apterous males 6.20–6.50. Color (
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<figureCitation id="14CC2A1EFFF229153C92FA7C74A30AF2" box="[1165,1264,1486,1512]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="46" pageNumber="47">Figs 3K</figureCitation>
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,
|
||
<figureCitation id="14CC2A1EFFF229153D1FFA7C757B0AF2" box="[1280,1320,1486,1512]" captionStart="Fig" captionStartId="6.[189,232,1787,1813]" captionTargetBox="[257,1326,279,1736]" captionTargetId="figure-14@6.[226,1362,264,1746]" captionTargetPageId="6" captionText="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995. E. R. anderseni Tran & Yang, 2006. F. R. chinensis Hungerford & Matsuda, 1959. G. R. edentatus Andersen & Chen, 1995. H. R. gongvo Tran & Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen & Chen, 1995. K. R. recurvus Andersen & Chen, 1995. L. R. scorpio Andersen & Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="46" pageNumber="47">4H</figureCitation>
|
||
): head yellow with black markings; pronotum: of female with a large subtriangular yellow marking, of male with subovate yellow spot; in apterous form, mesonotum with yellow median stripe; metanotum: of female completely black, without yellow markings, of male chiefly black, usually with very thin yellow median stripe. Female: abdomen elongate and straight (
|
||
<figureCitation id="14CC2A1EFFF229153B4DF9E973FB096C" box="[850,936,1627,1654]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="46" pageNumber="47">Fig. 5J</figureCitation>
|
||
); ventral length of pregenital abdomen about 0.4 times body length; posterior part of abdominal segment VII slightly depressed dorso-ventrally; connexival processes of segment VII long, straight, flat (
|
||
<figureCitation id="14CC2A1EFFF229153B75F913739E09A6" box="[874,973,1697,1724]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="46" pageNumber="47">Fig. 3K</figureCitation>
|
||
); sternum VII not totally enclosing genital segments, posterior margin straight, without process (
|
||
<figureCitation id="14CC2A1EFFF229153BBBF977745609C5" box="[932,1029,1733,1759]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="46" pageNumber="47">Fig. 3K</figureCitation>
|
||
). Male: length of middle tibia ca 2.1 times length of hind tibia; pygophore simple; proctiger with angular lobes laterally (
|
||
<figureCitation id="14CC2A1EFFF229153D18F95A753A0818" box="[1287,1385,1768,1794]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran & Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen & Chen, 1995. D. R. recurvus Andersen & Chen, 1995. E. R. scorpio Andersen & Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran & Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="46" pageNumber="47">Fig. 7A</figureCitation>
|
||
); paramere slender, strongly curved at basal third, then tapering towards narrow apex.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFF2291438A2F8FE74320ED7" lastPageId="47" lastPageNumber="48" pageId="46" pageNumber="47" type="materials_examined">
|
||
<paragraph id="8C48369BFFF2291538A2F8FE71F9087C" blockId="46.[189,426,1868,1894]" box="[189,426,1868,1894]" pageId="46" pageNumber="47">
|
||
<heading id="D70081F7FFF2291538A2F8FE71F9087C" bold="true" box="[189,426,1868,1894]" fontSize="11" level="3" pageId="46" pageNumber="47" reason="3">
|
||
<emphasis id="BE83EA89FFF2291538A2F8FE71F9087C" bold="true" box="[189,426,1868,1894]" pageId="46" pageNumber="47">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFF2291538F5F8C971FD088F" blockId="46.[189,483,1915,1977]" box="[234,430,1915,1941]" pageId="46" pageNumber="47">
|
||
<emphasis id="BE83EA89FFF2291538F5F8C971FD088F" bold="true" box="[234,430,1915,1941]" pageId="46" pageNumber="47">
|
||
<typeStatus id="534C8839FFF2291538F5F8C97174088F" box="[234,295,1915,1941]" pageId="46" pageNumber="47">Type</typeStatus>
|
||
specimens
|
||
</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFF2291538A2F82C71B008A2" blockId="46.[189,483,1915,1977]" box="[189,483,1950,1977]" pageId="46" pageNumber="47">
|
||
See
|
||
<bibRefCitation id="E8664B6AFFF2291538F1F82D718C08A2" author="Tran & Yang" box="[238,479,1950,1977]" firstAuthor="Tran" pageId="46" pageNumber="47" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang (2006)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFF3291438F5FEBB71B70E38" blockId="47.[189,1398,264,397]" box="[234,484,264,291]" pageId="47" pageNumber="48">
|
||
<emphasis id="BE83EA89FFF3291438F5FEBB71B70E38" bold="true" box="[234,484,264,291]" pageId="47" pageNumber="48">Non-type specimens</emphasis>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFF3291438A2FE9E733B0E97" blockId="47.[189,1398,264,397]" pageId="47" pageNumber="48">
|
||
<materialsCitation id="3C9F3CC6FFF3291438A2FE9E757F0E73" ID-GBIF-Occurrence="4429869310" collectingDate="2020-10-27" collectionCode="ZVNU" collectorName="A. D. Tran" country="Vietnam" county="Sa Pa" latitude="22.325806" location="Ho" longLatPrecision="1" longitude="103.83136" municipality="Cat Cat" pageId="47" pageNumber="48" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai">
|
||
<collectingCountry id="F4E0760BFFF3291438A2FE9E71170E5C" box="[189,324,300,326]" name="Vietnam" pageId="47" pageNumber="48">VIETNAM</collectingCountry>
|
||
• 1 ♁,
|
||
<specimenCount id="9AF1FD12FFF32914398CFE9E71ED0E5C" box="[403,446,300,326]" pageId="47" pageNumber="48" type="female">1 ♀</specimenCount>
|
||
(apterous);
|
||
<collectingRegion id="4E33F879FFF329143A56FE9E73440E5C" box="[585,791,300,326]" country="Vietnam" name="Lao Cai" pageId="47" pageNumber="48">Lào Cai Province</collectingRegion>
|
||
,
|
||
<collectingCounty id="65294E17FFF329143B3DFE9E73360E5C" box="[802,869,300,326]" pageId="47" pageNumber="48">Sa Pa</collectingCounty>
|
||
,
|
||
<collectingMunicipality id="6C2CACE1FFF329143B6FFE9E73980E5C" box="[880,971,300,326]" pageId="47" pageNumber="48">Cát Cát</collectingMunicipality>
|
||
,
|
||
<location id="89286040FFF329143BCAFE9E73AA0E5C" LSID="urn:lsid:plazi:treatment:045E878DFFF229143A35FB0A72A90B4E:89286040FFF329143BCAFE9E73AA0E5C" box="[981,1017,300,326]" country="Vietnam" county="Sa Pa" latitude="22.325806" longLatPrecision="1" longitude="103.83136" municipality="Cat Cat" name="Ho" pageId="47" pageNumber="48" stateProvince="Lao Cai">Ho</location>
|
||
stream (feeder stream of Mường Hoa stream);
|
||
<geoCoordinate id="E9C3505CFFF32914397FFEFF725D0E73" box="[352,526,333,361]" degrees="22" direction="north" minutes="19" orientation="latitude" pageId="47" pageNumber="48" precision="1" seconds="32.9" value="22.325806">22°19′32.9″ N</geoCoordinate>
|
||
,
|
||
<geoCoordinate id="E9C3505CFFF329143A04FEFF72870E73" box="[539,724,333,361]" degrees="103" direction="east" minutes="49" orientation="longitude" pageId="47" pageNumber="48" precision="1" seconds="52.9" value="103.83136">103°49′52.9″ E</geoCoordinate>
|
||
;
|
||
<date id="F849105BFFF329143AFCFEFD73D10E73" box="[739,898,335,361]" pageId="47" pageNumber="48" value="2020-10-27">
|
||
<collectingDate id="E80DE9B3FFF329143AFCFEFD73D10E73" box="[739,898,335,361]" pageId="47" pageNumber="48" value="2020-10-27">27 Oct. 2020</collectingDate>
|
||
</date>
|
||
;
|
||
<collectorName id="2102534DFFF329143B91FEFD74550E73" box="[910,1030,335,361]" pageId="47" pageNumber="48">A.D. Tran</collectorName>
|
||
leg.; TAD20-23;
|
||
<collectionCode id="EAE6AE5EFFF329143CC4FEFD757F0E73" box="[1243,1324,335,361]" pageId="47" pageNumber="48">ZVNU</collectionCode>
|
||
</materialsCitation>
|
||
•
|
||
<materialsCitation id="3C9F3CC6FFF329143D56FEFD73360E97" ID-GBIF-Occurrence="4429869315" collectingDate="2020-10-27" collectionCode="NKUM" collectorName="A. D. Tran" country="Vietnam" county="Sa Pa" latitude="22.325806" location="Ho" longLatPrecision="1" longitude="103.83136" municipality="Cat Cat" pageId="47" pageNumber="48" specimenCount="1" specimenCount-female="1" stateProvince="Lao Cai">
|
||
<specimenCount id="9AF1FD12FFF329143D56FEFD75250E73" box="[1353,1398,335,361]" pageId="47" pageNumber="48" type="female">1 ♀</specimenCount>
|
||
(apterous); same collection data as for preceding;
|
||
<collectionCode id="EAE6AE5EFFF329143B18FEC173360E97" box="[775,869,371,397]" country="China" httpUri="http://grbio.org/cool/hp11-qr98" name="Nankai University" pageId="47" pageNumber="48">NKUM</collectionCode>
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFF3291438A2FE0674320ED7" blockId="47.[189,1121,435,462]" box="[189,1121,435,462]" pageId="47" pageNumber="48">
|
||
GPS data of previous records: see
|
||
<bibRefCitation id="E8664B6AFFF329143A4FFE0673130ED7" author="Tran & Yang" box="[592,832,435,462]" firstAuthor="Tran" pageId="47" pageNumber="48" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang (2006)</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="E8664B6AFFF329143B53FE06740D0ED7" author="Tran & Nguyen" box="[844,1118,435,462]" firstAuthor="Tran" pageId="47" pageNumber="48" pagination="501 - 516" refId="ref33477" refString="Tran A. D. & Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran & Nguyen (2016)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFF3291438A2FE4773C00D27" box="[189,915,501,574]" pageId="47" pageNumber="48" type="distribution">
|
||
<paragraph id="8C48369BFFF3291438A2FE4771050D15" blockId="47.[189,342,501,527]" box="[189,342,501,527]" pageId="47" pageNumber="48">
|
||
<heading id="D70081F7FFF3291438A2FE4771050D15" bold="true" box="[189,342,501,527]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
|
||
<emphasis id="BE83EA89FFF3291438A2FE4771050D15" bold="true" box="[189,342,501,527]" pageId="47" pageNumber="48">Distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFF3291438A2FD9173C00D27" blockId="47.[189,915,547,574]" box="[189,915,547,574]" pageId="47" pageNumber="48">
|
||
<collectingCountry id="F4E0760BFFF3291438A2FD9171760D27" box="[189,293,547,573]" name="Vietnam" pageId="47" pageNumber="48">Vietnam</collectingCountry>
|
||
:
|
||
<collectingRegion id="4E33F879FFF32914392FFD9671DD0D27" box="[304,398,547,574]" country="Vietnam" name="Lao Cai" pageId="47" pageNumber="48">Lào Cai</collectingRegion>
|
||
(
|
||
<bibRefCitation id="E8664B6AFFF329143982FD9672280D27" author="Tran & Yang" box="[413,635,547,574]" firstAuthor="Tran" pageId="47" pageNumber="48" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang 2006</bibRefCitation>
|
||
;
|
||
<bibRefCitation id="E8664B6AFFF329143A98FD9673D40D27" author="Tran & Nguyen" box="[647,903,547,574]" firstAuthor="Tran" pageId="47" pageNumber="48" pagination="501 - 516" refId="ref33477" refString="Tran A. D. & Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran & Nguyen 2016</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C4ED6510FFF3291438A2FDD772A90B4E" pageId="47" pageNumber="48" type="discussion">
|
||
<paragraph id="8C48369BFFF3291438A2FDD771F90D65" blockId="47.[189,426,613,639]" box="[189,426,613,639]" pageId="47" pageNumber="48">
|
||
<heading id="D70081F7FFF3291438A2FDD771F90D65" bold="true" box="[189,426,613,639]" fontSize="11" level="3" pageId="47" pageNumber="48" reason="3">
|
||
<emphasis id="BE83EA89FFF3291438A2FDD771F90D65" bold="true" box="[189,426,613,639]" pageId="47" pageNumber="48">Comparative notes</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8C48369BFFF3291438A2FD2172A90B4E" blockId="47.[188,1401,659,1108]" pageId="47" pageNumber="48">
|
||
This species is most similar to
|
||
<taxonomicName id="4BF74D18FFF329143A35FD2672FD0DB7" authorityName="Andersen & Chen" authorityYear="1995" box="[554,686,659,685]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="malaisei">
|
||
<emphasis id="BE83EA89FFF329143A35FD2672FD0DB7" box="[554,686,659,685]" italics="true" pageId="47" pageNumber="48">R. malaisei</emphasis>
|
||
</taxonomicName>
|
||
, which was discussed by
|
||
<bibRefCitation id="E8664B6AFFF329143BC0FD2675790DB4" author="Tran & Yang" box="[991,1322,659,686]" firstAuthor="Tran" page="18 - 19" pageId="47" pageNumber="48" pagination="11 - 20" refId="ref33532" refString="Tran A. D. & Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran & Yang (2006: 18–19)</bibRefCitation>
|
||
. Here, we provide further comparative notes between the two species.
|
||
<taxonomicName id="4BF74D18FFF329143BA0FD0574E70DCA" authorityName="Tran & Yang" authorityYear="2006" box="[959,1204,695,721]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gongvo">
|
||
<emphasis id="BE83EA89FFF329143BA0FD0574E70DCA" box="[959,1204,695,721]" italics="true" pageId="47" pageNumber="48">Rhyacobates gongvo</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4BF74D18FFF329143CEDFD05752B0DCB" authorityName="Andersen & Chen" authorityYear="1995" box="[1266,1400,695,721]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="malaisei">
|
||
<emphasis id="BE83EA89FFF329143CEDFD05752B0DCB" box="[1266,1400,695,721]" italics="true" pageId="47" pageNumber="48">R. malaisei</emphasis>
|
||
</taxonomicName>
|
||
share the following characteristics of the female: (1) the metanotum is completely black, without yellow markings (
|
||
<figureCitation id="14CC2A1EFFF32914398EFD4F724B0C0D" box="[401,536,765,791]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">Fig. 3K, M</figureCitation>
|
||
); (2) the abdomen is relatively straight and elongate; (3) the connexiva of abdominal segments I–VI are blackish, forming straight lines in dorsal view (
|
||
<figureCitation id="14CC2A1EFFF329143C25FC9274ED0C20" box="[1082,1214,800,826]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">Fig. 3K, M</figureCitation>
|
||
); (4) abdominal segment VII has a pair of pointed or angular lateral processes (
|
||
<figureCitation id="14CC2A1EFFF329143B92FCF173BE0C44" box="[909,1005,835,862]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">Figs 3K</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFF329143BE6FCF174670C44" box="[1017,1076,835,862]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen & Chen, 1995.E. R. anderseni Tran & Yang, 2006. F. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford & Matsuda, 1959. H. R. constrictus Tran & Nguyen, 2016. I. R. edentatus Andersen & Chen, 1995. J. R. gongvo Tran & Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen & Chen, 1995. M. R. recurvus Andersen & Chen, 1995. N. R. scorpio Andersen & Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="47" pageNumber="48">5J, L</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFF329143C20FCF674250C44" box="[1087,1142,836,862]" captionStart="Fig" captionStartId="52.[189,232,1788,1814]" captionTargetBox="[295,1291,280,1715]" captionTargetId="figure-15@52.[261,1327,265,1739]" captionTargetPageId="52" captionText="Fig. 30. Morphological features of Rhyacobates malaisei Andersen & Chen, 1995. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D–E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385700" httpUri="https://zenodo.org/record/8385700/files/figure.png" pageId="47" pageNumber="48">30G</figureCitation>
|
||
); and (5) the posterior margin of sternum VII is truncate or slightly sunken, without a median process (
|
||
<figureCitation id="14CC2A1EFFF329143C8FFCD574AF0C9B" box="[1168,1276,871,897]" captionStart="Fig" captionStartId="52.[189,232,1788,1814]" captionTargetBox="[295,1291,280,1715]" captionTargetId="figure-15@52.[261,1327,265,1739]" captionTargetPageId="52" captionText="Fig. 30. Morphological features of Rhyacobates malaisei Andersen & Chen, 1995. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D–E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385700" httpUri="https://zenodo.org/record/8385700/files/figure.png" pageId="47" pageNumber="48">Fig. 30F</figureCitation>
|
||
). The key differences between these two species are as follows: in the female of
|
||
<taxonomicName id="4BF74D18FFF329143C00FC3974CA0CBE" authorityName="Tran & Yang" authorityYear="2006" box="[1055,1177,907,932]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gongvo">
|
||
<emphasis id="BE83EA89FFF329143C00FC3974CA0CBE" box="[1055,1177,907,932]" italics="true" pageId="47" pageNumber="48">R. gongvo</emphasis>
|
||
</taxonomicName>
|
||
, the connexiva of abdominal segment VII do not meet each other over the dorsum (
|
||
<figureCitation id="14CC2A1EFFF329143BCBFC1F74660CDD" box="[980,1077,941,967]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">Fig. 3K</figureCitation>
|
||
), whereas in the female of
|
||
<taxonomicName id="4BF74D18FFF3291438A0FC6371110CF0" authorityName="Andersen & Chen" authorityYear="1995" box="[191,322,976,1002]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="malaisei">
|
||
<emphasis id="BE83EA89FFF3291438A0FC6371110CF0" box="[191,322,976,1002]" italics="true" pageId="47" pageNumber="48">R. malaisei</emphasis>
|
||
</taxonomicName>
|
||
, the connexiva of abdominal segment VII are more developed, curved mesad and meeting over the middle of the abdominal dorsum (
|
||
<figureCitation id="14CC2A1EFFF329143AADFC41734B0B14" box="[690,792,1011,1038]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen & Chen, 1995. F. R. anderseni Tran & Yang, 2006. G. R. angustus Tran & Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford & Matsuda, 1959. I. R. constrictus Tran & Nguyen, 2016. J. R. edentatus Andersen & Chen, 1995. K. R. gongvo Tran & Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen & Chen, 1995. N. R. recurvus Andersen & Chen, 1995. O. R. scorpio Andersen & Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran & Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="47" pageNumber="48">Figs 3M</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFF329143B3BFC46732F0B14" box="[804,892,1012,1038]" captionStart="Fig" captionStartId="52.[189,232,1788,1814]" captionTargetBox="[295,1291,280,1715]" captionTargetId="figure-15@52.[261,1327,265,1739]" captionTargetPageId="52" captionText="Fig. 30. Morphological features of Rhyacobates malaisei Andersen & Chen, 1995. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D–E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385700" httpUri="https://zenodo.org/record/8385700/files/figure.png" pageId="47" pageNumber="48">30D–E</figureCitation>
|
||
). In the male of
|
||
<taxonomicName id="4BF74D18FFF329143C27FC4674FD0B17" authorityName="Tran & Yang" authorityYear="2006" box="[1080,1198,1012,1037]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="gongvo">
|
||
<emphasis id="BE83EA89FFF329143C27FC4674FD0B17" box="[1080,1198,1012,1037]" italics="true" pageId="47" pageNumber="48">R. gongvo</emphasis>
|
||
</taxonomicName>
|
||
, the lateral lobes of the proctiger are angular (
|
||
<figureCitation id="14CC2A1EFFF329143A0EFBA572230B2B" box="[529,624,1047,1073]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran & Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen & Chen, 1995. D. R. recurvus Andersen & Chen, 1995. E. R. scorpio Andersen & Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran & Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="47" pageNumber="48">Fig. 7A</figureCitation>
|
||
), whereas in
|
||
<taxonomicName id="4BF74D18FFF329143B11FBA573C10B2B" authorityName="Andersen & Chen" authorityYear="1995" box="[782,914,1047,1073]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="47" pageNumber="48" phylum="Arthropoda" rank="species" species="malaisei">
|
||
<emphasis id="BE83EA89FFF329143B11FBA573C10B2B" box="[782,914,1047,1073]" italics="true" pageId="47" pageNumber="48">R. malaisei</emphasis>
|
||
</taxonomicName>
|
||
, each lateral lobe of the proctiger has an angular process directed ventrad (
|
||
<figureCitation id="14CC2A1EFFF329143A55FB8872FA0B4E" box="[586,681,1082,1108]" captionStart="Fig" captionStartId="9.[190,233,1892,1918]" captionTargetBox="[196,1386,1025,1826]" captionTargetId="figure-221@9.[189,1398,1007,1850]" captionTargetPageId="9" captionText="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran & Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen & Chen, 1995. D. R. recurvus Andersen & Chen, 1995. E. R. scorpio Andersen & Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran & Nguyen, 2016. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385648" httpUri="https://zenodo.org/record/8385648/files/figure.png" pageId="47" pageNumber="48">Figs 7C</figureCitation>
|
||
,
|
||
<figureCitation id="14CC2A1EFFF329143AA9FB8872BE0B4E" box="[694,749,1082,1108]" captionStart="Fig" captionStartId="52.[189,232,1788,1814]" captionTargetBox="[295,1291,280,1715]" captionTargetId="figure-15@52.[261,1327,265,1739]" captionTargetPageId="52" captionText="Fig. 30. Morphological features of Rhyacobates malaisei Andersen & Chen, 1995. A. Body of female, dorsal view. B. Body of female, excluding head and prothorax, ventral view. C. Body of female, lateral view. D–E. Abdominal end of female, dorsal view. F. Abdominal end of female, ventral view. G. Abdominal end of female, lateral view. H. Body of male, dorsal view. I. Body of male, excluding head and prothorax, ventral view. J. Basal part of right middle leg of male, ventral view. K. Proctiger of male, dorsal view. L. Left paramere, lateral aspect, two different views. Scale bars: A–J = 1 mm; K–L = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385700" httpUri="https://zenodo.org/record/8385700/files/figure.png" pageId="47" pageNumber="48">30K</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |