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<mods:title id="A50A9313F57F5B52F518529ABB664F03">Taxonomic review of Rhyacobates Esaki, 1923, with descriptions of three new species (Hemiptera: Heteroptera: Gerridae)</mods:title>
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<mods:namePart id="F9079265D8D6EAB003AFD87A5F749C93">Leng, Zhaoqi</mods:namePart>
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Genus
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<bibRefCitation id="E8664B6AFFDE29393B51FBF9738A0B7C" author="Esaki" box="[846,985,1099,1126]" firstAuthor="Esaki" pageId="2" pageNumber="3" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki, 1923</bibRefCitation>
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<taxonomicName id="4BF74D18FFDE293938A2FB3F72770BBD" ID-CoL="79GN" authority="Esaki, 1923: 387" authorityName="Esaki" authorityPageNumber="387" authorityYear="1923" box="[189,548,1165,1191]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDE293938A2FB3F71000BBD" box="[189,339,1165,1191]" italics="true" pageId="2" pageNumber="3">Rhyacobates</emphasis>
<bibRefCitation id="E8664B6AFFDE29393945FB3F72770BBD" author="Esaki" box="[346,548,1165,1191]" firstAuthor="Esaki" page="387" pageId="2" pageNumber="3" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki, 1923: 387</bibRefCitation>
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.
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<treatmentCitation id="0D56108AFFDE293938A2FB0272630BD0" author="Lundblad O. M." box="[189,560,1200,1226]" page="22" pageId="2" pageNumber="3" year="1934">
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<emphasis id="BE83EA89FFDE293938A2FB0271100BD0" box="[189,323,1200,1226]" italics="true" pageId="2" pageNumber="3">Esakobates</emphasis>
<bibRefCitation id="E8664B6AFFDE29393955FB0272630BD0" author="Lundblad" box="[330,560,1200,1226]" firstAuthor="Lundblad" page="22" pageId="2" pageNumber="3" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934: 22</bibRefCitation>
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(syn.
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<bibRefCitation id="E8664B6AFFDE29393A66FB0273AC0BD0" author="Hungerford &amp; Matsuda" box="[633,1023,1200,1227]" firstAuthor="Hungerford" page="69" pageId="2" pageNumber="3" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda 1959: 69</bibRefCitation>
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).
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species
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<taxonomicName id="4BF74D18FFDE293938A2FA9272390A20" ID-CoL="4SNHW" authority="Esaki, 1923" authorityName="Esaki" authorityYear="1923" box="[189,618,1312,1338]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="BE83EA89FFDE293938A2FA9271870A20" box="[189,468,1312,1338]" italics="true" pageId="2" pageNumber="3">Rhyacobates takahashii</emphasis>
<bibRefCitation id="E8664B6AFFDE293939C3FA9272390A20" author="Esaki" box="[476,618,1312,1338]" firstAuthor="Esaki" pageId="2" pageNumber="3" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki, 1923</bibRefCitation>
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.
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<emphasis id="BE83EA89FFDE293938A2FAD371650A61" bold="true" box="[189,310,1377,1403]" pageId="2" pageNumber="3">Diagnosis</emphasis>
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Medium-sized (
<figureCitation id="14CC2A1EFFDE29393969FA22718C0AB0" box="[374,479,1424,1450]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[189,232,1963,1989]" captionStartId-1="4.[189,232,1892,1918]" captionTargetBox-0="[236,1357,570,1892]" captionTargetBox-1="[336,1266,297,1835]" captionTargetId-0="figure-106@3.[189,1398,528,1922]" captionTargetId-1="figure-14@4.[255,1333,265,1851]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Rhyacobates scorpio Andersen &amp; Chen, 1995. Apterous female, dorsal habitus. Scale bar = 3 mm." captionText-1="Fig. 2. Habitus of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise. A. R. abdominalisAndersen &amp; Chen, 1995.B. R. chinensis Hungerford &amp; Matsuda, 1959.C. R. lundbladi (Hungerford, 1957). D. R. scorpio Andersen &amp; Chen, 1995. E. R. svenhedini (Lundblad, 1934). F. R. takahashii Esaki, 1923, dealated macropterous female. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385632" figureDoi-1="http://doi.org/10.5281/zenodo.8385634" httpUri-0="https://zenodo.org/record/8385632/files/figure.png" httpUri-1="https://zenodo.org/record/8385634/files/figure.png" pageId="2" pageNumber="3">Figs 12</figureCitation>
), females 6.7912.21, males 5.428.41, females distinctly larger than males in the same species (
<figureCitation id="14CC2A1EFFDE29393991FA0171AA0AD4" box="[398,505,1459,1486]" captionStart-0="Fig" captionStart-1="Fig" captionStart-2="Fig" captionStartId-0="5.[189,232,1716,1742]" captionStartId-1="6.[189,232,1787,1813]" captionStartId-2="7.[189,232,1743,1769]" captionTargetBox-0="[256,1333,273,1657]" captionTargetBox-1="[257,1326,279,1736]" captionTargetBox-2="[200,1390,762,1681]" captionTargetId-0="figure-12@5.[228,1359,265,1677]" captionTargetId-1="figure-14@6.[226,1362,264,1746]" captionTargetId-2="figure-118@7.[189,1398,726,1709]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionTargetPageId-2="7" captionText-0="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." captionText-1="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. gongvo Tran &amp; Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen &amp; Chen, 1995. K. R. recurvus Andersen &amp; Chen, 1995. L. R. scorpio Andersen &amp; Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." captionText-2="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385638" figureDoi-1="http://doi.org/10.5281/zenodo.8385642" figureDoi-2="http://doi.org/10.5281/zenodo.8385644" httpUri-0="https://zenodo.org/record/8385638/files/figure.png" httpUri-1="https://zenodo.org/record/8385642/files/figure.png" httpUri-2="https://zenodo.org/record/8385644/files/figure.png" pageId="2" pageNumber="3">Figs 35</figureCitation>
). Dorsum mainly blackish with silvery pubescence, pronotum with a median yellowish marking (
<figureCitation id="14CC2A1EFFDE293939ADFA6472710AEB" box="[434,546,1494,1521]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="5.[189,232,1716,1742]" captionStartId-1="6.[189,232,1787,1813]" captionTargetBox-0="[256,1333,273,1657]" captionTargetBox-1="[257,1326,279,1736]" captionTargetId-0="figure-12@5.[228,1359,265,1677]" captionTargetId-1="figure-14@6.[226,1362,264,1746]" captionTargetPageId-0="5" captionTargetPageId-1="6" captionText-0="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." captionText-1="Fig. 4. Photographs of bodies of Rhyacobates spp. in dorsal view (males), apterous form if not stated otherwise.A. R. bui sp. nov.B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. gongvo Tran &amp; Yang, 2006. I. R. lundbladi (Hungerford, 1957). J. R. malaisei Andersen &amp; Chen, 1995. K. R. recurvus Andersen &amp; Chen, 1995. L. R. scorpio Andersen &amp; Chen, 1995. M. R. svenhedini (Lundblad, 1934), apterous form. N. R. svenhedini, macropterous form. O. R. takahashii Esaki, 1923, dealated macropterous form. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385638" figureDoi-1="http://doi.org/10.5281/zenodo.8385642" httpUri-0="https://zenodo.org/record/8385638/files/figure.png" httpUri-1="https://zenodo.org/record/8385642/files/figure.png" pageId="2" pageNumber="3">Figs 34</figureCitation>
). Antennal tubercles pronounced, angularly produced in dorsal view; antennal segment I longer than other three segments combined, segment II shorter than segment III, segment IV curved, with whitish groove at distal two-fifths (
<figureCitation id="14CC2A1EFFDE29393B9CF9AF73BD092D" box="[899,1006,1565,1591]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[189,232,1963,1989]" captionStartId-1="4.[189,232,1892,1918]" captionTargetBox-0="[236,1357,570,1892]" captionTargetBox-1="[336,1266,297,1835]" captionTargetId-0="figure-106@3.[189,1398,528,1922]" captionTargetId-1="figure-14@4.[255,1333,265,1851]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Rhyacobates scorpio Andersen &amp; Chen, 1995. Apterous female, dorsal habitus. Scale bar = 3 mm." captionText-1="Fig. 2. Habitus of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise. A. R. abdominalisAndersen &amp; Chen, 1995.B. R. chinensis Hungerford &amp; Matsuda, 1959.C. R. lundbladi (Hungerford, 1957). D. R. scorpio Andersen &amp; Chen, 1995. E. R. svenhedini (Lundblad, 1934). F. R. takahashii Esaki, 1923, dealated macropterous female. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385632" figureDoi-1="http://doi.org/10.5281/zenodo.8385634" httpUri-0="https://zenodo.org/record/8385632/files/figure.png" httpUri-1="https://zenodo.org/record/8385634/files/figure.png" pageId="2" pageNumber="3">Figs 12</figureCitation>
). Fore femur slender, subapically without tooth on ventral surface. Middle femur with black spines along proximal three quarters of ventral margin, but usually not in distinct row. Middle coxa without apical spine, not elongate. Middle and hind tarsi without claws (
<figureCitation id="14CC2A1EFFDE293939ABF93471AF09BB" box="[436,508,1670,1697]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[236,1357,570,1892]" captionTargetId="figure-106@3.[189,1398,528,1922]" captionTargetPageId="3" captionText="Fig. 1. Rhyacobates scorpio Andersen &amp; Chen, 1995. Apterous female, dorsal habitus. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385632" httpUri="https://zenodo.org/record/8385632/files/figure.png" pageId="2" pageNumber="3">Fig. 1</figureCitation>
). Female: posterior abdominal segments curved dorsad or nearly straight (
<figureCitation id="14CC2A1EFFDE293938D9F918715C09DE" box="[198,271,1705,1732]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="2" pageNumber="3">Fig. 5</figureCitation>
); segment VII usually modified, length of sternum VII about twice the length of sternum VI. Female gonocoxa directed caudad, usually completely withdrawn into sternum VII. Male genitalia: pygophore simple, without lateral process; proctiger laterally produced into rounded or angular lobes (
<figureCitation id="14CC2A1EFFDE293938D8F8A171600837" box="[199,307,1811,1837]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="8.[189,232,1892,1918]" captionStartId-1="9.[190,233,1892,1918]" captionTargetBox-0="[229,1341,570,1825]" captionTargetBox-1="[196,1386,1025,1826]" captionTargetId-0="figure-92@8.[189,1398,537,1851]" captionTargetId-1="figure-221@9.[189,1398,1007,1850]" captionTargetPageId-0="8" captionTargetPageId-1="9" captionText-0="Fig. 6. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016. G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. Scale bar = 0.2 mm." captionText-1="Fig. 7. Proctiger of Rhyacobates spp. in dorsal view (males). A. R. gongvo Tran &amp; Yang, 2006. B. R. undbladi (Hungerford, 1957). C. R. malaisei Andersen &amp; Chen, 1995. D. R. recurvus Andersen &amp; Chen, 1995. E. R. scorpio Andersen &amp; Chen, 1995. F. R. svenhedini (Lundblad, 1934). G. R. takahashii Esaki, 1923. H. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385646" figureDoi-1="http://doi.org/10.5281/zenodo.8385648" httpUri-0="https://zenodo.org/record/8385646/files/figure.png" httpUri-1="https://zenodo.org/record/8385648/files/figure.png" pageId="2" pageNumber="3">Figs 67</figureCitation>
); paramere long and curved dorsad, without long setae (
<figureCitation id="14CC2A1EFFDE29393BDAF8A1745F0837" box="[965,1036,1811,1837]" captionStart="Fig" captionStartId="10.[189,232,1822,1848]" captionTargetBox="[241,1331,291,1744]" captionTargetId="figure-14@10.[218,1369,265,1777]" captionTargetPageId="10" captionText="Fig. 8. Paramere of Rhyacobates spp. in lateral view. A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov. D. R. abdominalis Andersen &amp; Chen, 1995. E. R. anderseni Tran &amp; Yang, 2006. F. R. chinensis Hungerford &amp; Matsuda, 1959. G. R. edentatus Andersen &amp; Chen, 1995. H. R. lundbladi (Hungerford, 1957). I. R. malaisei Andersen &amp; Chen, 1995. J. R. recurvus Andersen &amp; Chen, 1995. K. R. scorpio Andersen &amp; Chen, 1995. L. R. svenhedini (Lundblad, 1934). M. R. takahashii Esaki, 1923. Scale bar = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.8385652" httpUri="https://zenodo.org/record/8385652/files/figure.png" pageId="2" pageNumber="3">Fig. 8</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFDE293938A2F8E671F90875" blockId="2.[189,426,1876,1903]" box="[189,426,1876,1903]" pageId="2" pageNumber="3">
<heading id="D70081F7FFDE293938A2F8E671F90875" bold="true" box="[189,426,1876,1903]" fontSize="11" level="3" pageId="2" pageNumber="3" reason="3">
<emphasis id="BE83EA89FFDE293938A2F8E671F90875" bold="true" box="[189,426,1876,1903]" pageId="2" pageNumber="3">Comparative notes</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFDE293838A2F831735D0EC9" blockId="2.[189,1399,1923,2020]" lastBlockId="3.[189,1399,265,467]" lastPageId="3" lastPageNumber="4" pageId="2" pageNumber="3">
The differences between
<taxonomicName id="4BF74D18FFDE293939FAF83172280887" authorityName="Esaki" authorityYear="1923" box="[485,635,1923,1949]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDE293939FAF83172280887" box="[485,635,1923,1949]" italics="true" pageId="2" pageNumber="3">Rhyacobates</emphasis>
</taxonomicName>
and its closely related genera (i.e.,
<taxonomicName id="4BF74D18FFDE29393C03F83174F90887" authorityName="Bianchi" authorityYear="1896" box="[1052,1194,1923,1949]" class="Insecta" family="Gerridae" genus="Heterobates" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDE29393C03F83174F90887" box="[1052,1194,1923,1949]" italics="true" pageId="2" pageNumber="3">Heterobates</emphasis>
</taxonomicName>
and
<taxonomicName id="4BF74D18FFDE29393CFBF831753E0887" authorityName="Esaki" authorityYear="1930" box="[1252,1389,1923,1949]" class="Insecta" family="Gerridae" genus="Pleciobates" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDE29393CFBF831753E0887" box="[1252,1389,1923,1949]" italics="true" pageId="2" pageNumber="3">Pleciobates</emphasis>
</taxonomicName>
) were summarized by
<bibRefCitation id="E8664B6AFFDE293939A4F81472BF08DB" author="Andersen &amp; Chen" box="[443,748,1958,1985]" firstAuthor="Andersen" pageId="2" pageNumber="3" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen (1995)</bibRefCitation>
. Three genera described after the study of
<bibRefCitation id="E8664B6AFFDE29393CF8F814710808FE" author="Andersen &amp; Chen" firstAuthor="Andersen" pageId="2" pageNumber="3" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen (1995)</bibRefCitation>
,
<taxonomicName id="4BF74D18FFDE29393972F87871AD08FE" authorityName="Zettel &amp; Chen" authorityYear="1996" box="[365,510,1994,2020]" class="Insecta" family="Gerridae" genus="Andersenius" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDE29393972F87871AD08FE" box="[365,510,1994,2020]" italics="true" pageId="2" pageNumber="3">Andersenius</emphasis>
</taxonomicName>
,
<taxonomicName id="4BF74D18FFDE29393A0CF87872F708FE" authorityName="Chen, Nieser &amp; Wattanachaiyingcharoen" authorityYear="2002" box="[531,676,1994,2020]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDE29393A0CF87872F708FE" box="[531,676,1994,2020]" italics="true" pageId="2" pageNumber="3">Pleciogonus</emphasis>
</taxonomicName>
and
<taxonomicName id="4BF74D18FFDE29393AF4F878732B08FE" authorityName="Zettel" authorityYear="2009" box="[747,888,1994,2020]" class="Insecta" family="Gerridae" genus="Celerobates" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDE29393AF4F878732B08FE" box="[747,888,1994,2020]" italics="true" pageId="2" pageNumber="3">Celerobates</emphasis>
</taxonomicName>
are also closely related to
<emphasis id="BE83EA89FFDE29393CC6F878752408FE" box="[1241,1399,1994,2020]" italics="true" pageId="2" pageNumber="3">
<taxonomicName id="4BF74D18FFDE29393CC6F878752008FE" authorityName="Esaki" authorityYear="1923" box="[1241,1395,1994,2020]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="2" pageNumber="3" phylum="Arthropoda" rank="genus">Rhyacobates</taxonomicName>
.
</emphasis>
However,
<taxonomicName id="4BF74D18FFDF29383928FEBB719E0E39" authorityName="Esaki" authorityYear="1923" box="[311,461,265,291]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDF29383928FEBB719E0E39" box="[311,461,265,291]" italics="true" pageId="3" pageNumber="4">Rhyacobates</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4BF74D18FFDF29383B0EFEBB73F10E39" authorityName="Zettel &amp; Chen" authorityYear="1996" box="[785,930,265,291]" class="Insecta" family="Gerridae" genus="Andersenius" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDF29383B0EFEBB73F10E39" box="[785,930,265,291]" italics="true" pageId="3" pageNumber="4">Andersenius</emphasis>
</taxonomicName>
by the hind coxa, which is shorter than wide in the former. In
<taxonomicName id="4BF74D18FFDF293839D5FE9E72080E5C" authorityName="Zettel &amp; Chen" authorityYear="1996" box="[458,603,300,326]" class="Insecta" family="Gerridae" genus="Andersenius" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDF293839D5FE9E72080E5C" box="[458,603,300,326]" italics="true" pageId="3" pageNumber="4">Andersenius</emphasis>
</taxonomicName>
, the hind coxa is distinctly longer than wide, i.e., 3.54.0 times as long as wide in the female and 1.22.0 times as long as wide in the male.
<taxonomicName id="4BF74D18FFDF29383C51FEFD74B70E73" authorityName="Esaki" authorityYear="1923" box="[1102,1252,335,361]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDF29383C51FEFD74B70E73" box="[1102,1252,335,361]" italics="true" pageId="3" pageNumber="4">Rhyacobates</emphasis>
</taxonomicName>
can also be distinguished from
<taxonomicName id="4BF74D18FFDF293839B9FEC072640E96" authorityName="Chen, Nieser &amp; Wattanachaiyingcharoen" authorityYear="2002" box="[422,567,370,396]" class="Insecta" family="Gerridae" genus="Pleciogonus" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDF293839B9FEC072640E96" box="[422,567,370,396]" italics="true" pageId="3" pageNumber="4">Pleciogonus</emphasis>
</taxonomicName>
by connexival segment VI of the female, which is simple, without a long caudal process (
<figureCitation id="14CC2A1EFFDF293839A8FE2472710EAA" box="[439,546,406,432]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="3.[189,232,1963,1989]" captionStartId-1="4.[189,232,1892,1918]" captionTargetBox-0="[236,1357,570,1892]" captionTargetBox-1="[336,1266,297,1835]" captionTargetId-0="figure-106@3.[189,1398,528,1922]" captionTargetId-1="figure-14@4.[255,1333,265,1851]" captionTargetPageId-0="3" captionTargetPageId-1="4" captionText-0="Fig. 1. Rhyacobates scorpio Andersen &amp; Chen, 1995. Apterous female, dorsal habitus. Scale bar = 3 mm." captionText-1="Fig. 2. Habitus of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise. A. R. abdominalisAndersen &amp; Chen, 1995.B. R. chinensis Hungerford &amp; Matsuda, 1959.C. R. lundbladi (Hungerford, 1957). D. R. scorpio Andersen &amp; Chen, 1995. E. R. svenhedini (Lundblad, 1934). F. R. takahashii Esaki, 1923, dealated macropterous female. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.8385632" figureDoi-1="http://doi.org/10.5281/zenodo.8385634" httpUri-0="https://zenodo.org/record/8385632/files/figure.png" httpUri-1="https://zenodo.org/record/8385634/files/figure.png" pageId="3" pageNumber="4">Figs 12</figureCitation>
).
<taxonomicName id="4BF74D18FFDF29383A25FE2472830EAA" authorityName="Esaki" authorityYear="1923" box="[570,720,406,432]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDF29383A25FE2472830EAA" box="[570,720,406,432]" italics="true" pageId="3" pageNumber="4">Rhyacobates</emphasis>
</taxonomicName>
can be distinguished from
<taxonomicName id="4BF74D18FFDF29383C0CFE2474F30EAA" authorityName="Zettel" authorityYear="2009" box="[1043,1184,406,432]" class="Insecta" family="Gerridae" genus="Celerobates" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDF29383C0CFE2474F30EAA" box="[1043,1184,406,432]" italics="true" pageId="3" pageNumber="4">Celerobates</emphasis>
</taxonomicName>
by the absence of distinct claws in the middle and hind tarsi (
<figureCitation id="14CC2A1EFFDF29383AA7FE0B72AE0EC9" box="[696,765,441,467]" captionStart="Fig" captionStartId="3.[189,232,1963,1989]" captionTargetBox="[236,1357,570,1892]" captionTargetId="figure-106@3.[189,1398,528,1922]" captionTargetPageId="3" captionText="Fig. 1. Rhyacobates scorpio Andersen &amp; Chen, 1995. Apterous female, dorsal habitus. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385632" httpUri="https://zenodo.org/record/8385632/files/figure.png" pageId="3" pageNumber="4">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="D8886613FFDF293838A2F819715B08F3" ID-DOI="http://doi.org/10.5281/zenodo.8385632" ID-Zenodo-Dep="8385632" httpUri="https://zenodo.org/record/8385632/files/figure.png" pageId="3" pageNumber="4" startId="3.[189,232,1963,1989]" targetBox="[236,1357,570,1892]" targetPageId="3">
<paragraph id="8C48369BFFDF293838A2F819715B08F3" blockId="3.[189,1399,1963,2025]" pageId="3" pageNumber="4">
<emphasis id="BE83EA89FFDF293838A2F819715C08DF" bold="true" box="[189,271,1963,1989]" pageId="3" pageNumber="4">Fig. 1.</emphasis>
<taxonomicName id="4BF74D18FFDF29383904F819731A08DC" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[283,841,1963,1990]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="3" pageNumber="4" phylum="Arthropoda" rank="species" species="scorpio">
<emphasis id="BE83EA89FFDF29383904F819724708DF" box="[283,532,1963,1989]" italics="true" pageId="3" pageNumber="4">Rhyacobates scorpio</emphasis>
<bibRefCitation id="E8664B6AFFDF29383A02F819731A08DC" author="Andersen &amp; Chen" box="[541,841,1963,1990]" firstAuthor="Andersen" pageId="3" pageNumber="4" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
. Apterous female, dorsal habitus. Scale bar = 3 mm.
</paragraph>
</caption>
<caption id="D8886613FFD8293F38A2F8D6740D08F3" ID-DOI="http://doi.org/10.5281/zenodo.8385634" ID-Zenodo-Dep="8385634" httpUri="https://zenodo.org/record/8385634/files/figure.png" pageId="4" pageNumber="5" startId="4.[189,232,1892,1918]" targetBox="[336,1266,297,1835]" targetPageId="4">
<paragraph id="8C48369BFFD8293F38A2F8D6740D08F3" blockId="4.[189,1399,1892,2025]" pageId="4" pageNumber="5">
<emphasis id="BE83EA89FFD8293F38A2F8D6715D0865" bold="true" box="[189,270,1892,1919]" pageId="4" pageNumber="5">Fig. 2.</emphasis>
Habitus of
<taxonomicName id="4BF74D18FFD8293F39BEF8D772220865" box="[417,625,1893,1919]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BE83EA89FFD8293F39BEF8D772640865" box="[417,567,1893,1919]" italics="true" pageId="4" pageNumber="5">Rhyacobates</emphasis>
spp.
</taxonomicName>
in dorsal view (females), apterous form if not stated otherwise.
<emphasis id="BE83EA89FFD8293F38A2F83A708108B8" bold="true" box="[189,210,1928,1954]" pageId="4" pageNumber="5">A</emphasis>
.
<taxonomicName id="4BF74D18FFD8293F38C0F83B72F808B9" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[223,683,1928,1955]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="BE83EA89FFD8293F38C0F83B71DD08B8" box="[223,398,1928,1954]" italics="true" pageId="4" pageNumber="5">R. abdominalis</emphasis>
<bibRefCitation id="E8664B6AFFD8293F398EF83A72F808B9" author="Andersen &amp; Chen" box="[401,683,1928,1955]" firstAuthor="Andersen" pageId="4" pageNumber="5" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD8293F3AAEF83A729608B8" bold="true" box="[689,709,1928,1954]" pageId="4" pageNumber="5">B</emphasis>
.
<taxonomicName id="4BF74D18FFD8293F3ACBF83B74EE08B9" authority="Hungerford &amp; Matsuda, 1959" authorityName="Hungerford &amp; Matsuda" authorityYear="1959" box="[724,1213,1928,1955]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="BE83EA89FFD8293F3ACBF83B730C08B8" box="[724,863,1928,1954]" italics="true" pageId="4" pageNumber="5">R. chinensis</emphasis>
<bibRefCitation id="E8664B6AFFD8293F3B7BF83A74EE08B9" author="Hungerford &amp; Matsuda" box="[868,1213,1928,1955]" firstAuthor="Hungerford" pageId="4" pageNumber="5" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda, 1959</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD8293F3CDBF83A748A08B8" bold="true" box="[1220,1241,1928,1954]" pageId="4" pageNumber="5">C</emphasis>
.
<taxonomicName id="4BF74D18FFD8293F3CF7F83B71FE08DC" authority="(Hungerford, 1957)" baseAuthorityName="Hungerford" baseAuthorityYear="1957" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFD8293F3CF7F83B752508B8" box="[1256,1398,1928,1954]" italics="true" pageId="4" pageNumber="5">R. lundbladi</emphasis>
(
<bibRefCitation id="E8664B6AFFD8293F38DAF81971F708DC" author="Hungerford" box="[197,420,1963,1990]" firstAuthor="Hungerford" pageId="4" pageNumber="5" pagination="33 - 36" refId="ref33146" refString="Hungerford H. B. 1957. A new gerrid from China (Gerridae, Hemiptera). Journal of the Kansas Entomological Society 30 (2): 33 - 36." type="journal article" year="1957">Hungerford, 1957</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFD8293F39A0F819718708DF" bold="true" box="[447,468,1963,1989]" pageId="4" pageNumber="5">D</emphasis>
.
<taxonomicName id="4BF74D18FFD8293F39F5F81E73FF08DC" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[490,940,1963,1990]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="scorpio">
<emphasis id="BE83EA89FFD8293F39F5F81E723908DF" box="[490,618,1964,1989]" italics="true" pageId="4" pageNumber="5">R. scorpio</emphasis>
<bibRefCitation id="E8664B6AFFD8293F3A69F81973FF08DC" author="Andersen &amp; Chen" box="[630,940,1963,1990]" firstAuthor="Andersen" pageId="4" pageNumber="5" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD8293F3BA1F819738108DF" bold="true" box="[958,978,1963,1989]" pageId="4" pageNumber="5">E</emphasis>
.
<taxonomicName id="4BF74D18FFD8293F3BF8F81E752008DF" authority="(Lundblad, 1934)" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[999,1395,1963,1989]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFD8293F3BF8F81E74DF08DF" box="[999,1164,1963,1989]" italics="true" pageId="4" pageNumber="5">R. svenhedini</emphasis>
(
<bibRefCitation id="E8664B6AFFD8293F3CBCF819753908DF" author="Lundblad" box="[1187,1386,1963,1989]" firstAuthor="Lundblad" pageId="4" pageNumber="5" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFD8293F38A2F87C709C08F2" bold="true" box="[189,207,1998,2024]" pageId="4" pageNumber="5">F</emphasis>
.
<taxonomicName id="4BF74D18FFD8293F38C1F87D725E08F2" authority="Esaki, 1923" authorityName="Esaki" authorityYear="1923" box="[222,525,1998,2024]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="BE83EA89FFD8293F38C1F87D712B08F2" box="[222,376,1998,2024]" italics="true" pageId="4" pageNumber="5">R. takahashii</emphasis>
<bibRefCitation id="E8664B6AFFD8293F399FF87C725E08F2" author="Esaki" box="[384,525,1998,2024]" firstAuthor="Esaki" pageId="4" pageNumber="5" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki, 1923</bibRefCitation>
</taxonomicName>
, dealated macropterous female. Scale bar = 3 mm.
</paragraph>
</caption>
<caption id="D8886613FFD9293E38A2F906721A08F3" ID-DOI="http://doi.org/10.5281/zenodo.8385638" ID-Zenodo-Dep="8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="5" pageNumber="6" startId="5.[189,232,1716,1742]" targetBox="[256,1333,273,1657]" targetPageId="5">
<paragraph id="8C48369BFFD9293E38A2F906721A08F3" blockId="5.[187,1399,1716,2025]" pageId="5" pageNumber="6">
<emphasis id="BE83EA89FFD9293E38A2F906715809D5" bold="true" box="[189,267,1716,1743]" pageId="5" pageNumber="6">Fig. 3.</emphasis>
Photographs of bodies of
<taxonomicName id="4BF74D18FFD9293E3A5EF907735D09D5" box="[577,782,1717,1743]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BE83EA89FFD9293E3A5EF907728409D5" box="[577,727,1717,1743]" italics="true" pageId="5" pageNumber="6">Rhyacobates</emphasis>
spp.
</taxonomicName>
in dorsal view (females), apterous form if not stated otherwise.
<emphasis id="BE83EA89FFD9293E3925F96A711C09E8" bold="true" box="[314,335,1752,1778]" pageId="5" pageNumber="6">A</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3943F96B71F309E8" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[348,416,1752,1778]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="bui" status="sp. nov.">
<emphasis id="BE83EA89FFD9293E3943F96B71F309E8" box="[348,416,1752,1778]" italics="true" pageId="5" pageNumber="6">R. bui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFD9293E39B9F96A71AC09E8" box="[422,511,1752,1778]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFD9293E3A1CF96A724409E8" bold="true" box="[515,535,1752,1778]" pageId="5" pageNumber="6">B</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3A3AF96B72E509E8" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[549,694,1752,1778]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="BE83EA89FFD9293E3A3AF96B72E509E8" box="[549,694,1752,1778]" italics="true" pageId="5" pageNumber="6">R. elongatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFD9293E3AA4F96A734709E8" box="[699,788,1752,1778]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFD9293E3B07F96A737E09E8" bold="true" box="[792,813,1752,1778]" pageId="5" pageNumber="6">C</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3B24F96B73E809E8" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[827,955,1752,1778]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
<emphasis id="BE83EA89FFD9293E3B24F96B73E809E8" box="[827,955,1752,1778]" italics="true" pageId="5" pageNumber="6">R. turgidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFD9293E3BDEF96A744F09E8" box="[961,1052,1752,1778]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, apterous form.
<emphasis id="BE83EA89FFD9293E3CCCF96A74BB09E8" bold="true" box="[1235,1256,1752,1778]" pageId="5" pageNumber="6">D</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3CE8F96B752409E8" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[1271,1399,1752,1778]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
<emphasis id="BE83EA89FFD9293E3CE8F96B752409E8" box="[1271,1399,1752,1778]" italics="true" pageId="5" pageNumber="6">R. turgidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFD9293E38A2F94E714B080C" box="[189,280,1788,1814]" pageId="5" pageNumber="6" rank="species">sp. nov.</taxonomicNameLabel>
, dealated macropterous form.
<emphasis id="BE83EA89FFD9293E3A6CF94972D4080F" bold="true" box="[627,647,1787,1813]" pageId="5" pageNumber="6">E</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3A8AF94E7437080C" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[661,1124,1787,1814]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="BE83EA89FFD9293E3A8AF94E7316080F" box="[661,837,1787,1813]" italics="true" pageId="5" pageNumber="6">R. abdominalis</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3B56F9497437080C" author="Andersen &amp; Chen" box="[841,1124,1787,1814]" firstAuthor="Andersen" pageId="5" pageNumber="6" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3C73F949742D080F" bold="true" box="[1132,1150,1787,1813]" pageId="5" pageNumber="6">F</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3C93F94E71150822" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="BE83EA89FFD9293E3C93F94E754C080F" box="[1164,1311,1787,1813]" italics="true" pageId="5" pageNumber="6">R. anderseni</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3D3BF94E71150822" author="Tran &amp; Yang" firstAuthor="Tran" pageId="5" pageNumber="6" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3948F8AC713F0822" bold="true" box="[343,364,1822,1848]" pageId="5" pageNumber="6">G</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3962F8AD73480822" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[381,795,1822,1849]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="BE83EA89FFD9293E3962F8AD72540822" box="[381,519,1823,1848]" italics="true" pageId="5" pageNumber="6">R. angustus</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3A0FF8AD73480822" author="Tran &amp; Nguyen" box="[528,795,1822,1849]" firstAuthor="Tran" pageId="5" pageNumber="6" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
, holotype (ZVNU).
<emphasis id="BE83EA89FFD9293E3C12F8AC74770822" bold="true" box="[1037,1060,1822,1848]" pageId="5" pageNumber="6">H</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3C2AF8AD71250846" authority="Hungerford &amp; Matsuda, 1959" authorityName="Hungerford &amp; Matsuda" authorityYear="1959" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="BE83EA89FFD9293E3C2AF8AD74970822" box="[1077,1220,1822,1848]" italics="true" pageId="5" pageNumber="6">R. chinensis</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3CD1F8AC71250846" author="Hungerford &amp; Matsuda" firstAuthor="Hungerford" pageId="5" pageNumber="6" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda, 1959</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E399AF8F071C30846" bold="true" box="[389,400,1858,1884]" pageId="5" pageNumber="6">I</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E39BBF8F0733E0841" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[420,877,1857,1884]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="BE83EA89FFD9293E39BBF8F072180841" box="[420,587,1858,1883]" italics="true" pageId="5" pageNumber="6">R. constrictus</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3A48F8F0733E0841" author="Tran &amp; Nguyen" box="[599,877,1857,1884]" firstAuthor="Tran" pageId="5" pageNumber="6" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3B62F8F073DF0846" bold="true" box="[893,908,1858,1884]" pageId="5" pageNumber="6">J</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3BBFF8F075210846" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[928,1394,1857,1884]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="edentatus">
<emphasis id="BE83EA89FFD9293E3BBFF8F074650841" box="[928,1078,1857,1883]" italics="true" pageId="5" pageNumber="6">R. edentatus</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3C5DF8F375210846" author="Andersen &amp; Chen" box="[1090,1394,1857,1884]" firstAuthor="Andersen" pageId="5" pageNumber="6" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E38A2F8D770870865" bold="true" box="[189,212,1893,1919]" pageId="5" pageNumber="6">K</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E38FEF8D4726F0865" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" box="[225,572,1893,1919]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="gongvo">
<emphasis id="BE83EA89FFD9293E38FEF8D471050865" box="[225,342,1894,1919]" italics="true" pageId="5" pageNumber="6">R. gongvo</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3942F8D7726F0865" author="Tran &amp; Yang" box="[349,572,1893,1919]" firstAuthor="Tran" pageId="5" pageNumber="6" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3A5AF8D7720A0865" bold="true" box="[581,601,1893,1919]" pageId="5" pageNumber="6">L</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3A78F8D473B6089A" authority="(Hungerford, 1957)" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[615,997,1893,1920]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFD9293E3A78F8D472A50865" box="[615,758,1893,1919]" italics="true" pageId="5" pageNumber="6">R. lundbladi</emphasis>
(
<bibRefCitation id="E8664B6AFFD9293E3B1BF8D7738F089A" author="Hungerford" box="[772,988,1893,1920]" firstAuthor="Hungerford" pageId="5" pageNumber="6" pagination="33 - 36" refId="ref33146" refString="Hungerford H. B. 1957. A new gerrid from China (Gerridae, Hemiptera). Journal of the Kansas Entomological Society 30 (2): 33 - 36." type="journal article" year="1957">Hungerford, 1957</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3BF2F8D7745A0865" bold="true" box="[1005,1033,1893,1919]" pageId="5" pageNumber="6">M</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3C08F8D470A908B9" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="BE83EA89FFD9293E3C08F8D474CA0865" box="[1047,1177,1893,1919]" italics="true" pageId="5" pageNumber="6">R. malaisei</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3C81F8D770A908B9" author="Andersen &amp; Chen" firstAuthor="Andersen" pageId="5" pageNumber="6" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3918F83A714F08B8" bold="true" box="[263,284,1928,1954]" pageId="5" pageNumber="6">N</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3934F83B728A08B9" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[299,729,1928,1955]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="recurvus">
<emphasis id="BE83EA89FFD9293E3934F83B71E108B8" box="[299,434,1929,1954]" italics="true" pageId="5" pageNumber="6">R. recurvus</emphasis>
<bibRefCitation id="E8664B6AFFD9293E39A7F83A728A08B9" author="Andersen &amp; Chen" box="[440,729,1928,1955]" firstAuthor="Andersen" pageId="5" pageNumber="6" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3AFAF83A72A808B8" bold="true" box="[741,763,1928,1954]" pageId="5" pageNumber="6">O</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3B14F83B74F808B9" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[779,1195,1928,1955]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="scorpio">
<emphasis id="BE83EA89FFD9293E3B14F83B73D708B8" box="[779,900,1929,1954]" italics="true" pageId="5" pageNumber="6">R. scorpio</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3B95F83A74F808B9" author="Andersen &amp; Chen" box="[906,1195,1928,1955]" firstAuthor="Andersen" pageId="5" pageNumber="6" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E3CA8F83A749A08B8" bold="true" box="[1207,1225,1928,1954]" pageId="5" pageNumber="6">P</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3CC7F83B71C008DF" authority="(Lundblad, 1934)" baseAuthorityName="Lundblad" baseAuthorityYear="1934" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFD9293E3CC7F83B752508B8" box="[1240,1398,1928,1954]" italics="true" pageId="5" pageNumber="6">R. svenhedini</emphasis>
(
<bibRefCitation id="E8664B6AFFD9293E38DAF81971DA08DF" author="Lundblad" box="[197,393,1963,1989]" firstAuthor="Lundblad" pageId="5" pageNumber="6" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFD9293E39BDF81971EA08DF" bold="true" box="[418,441,1963,1989]" pageId="5" pageNumber="6">Q</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E39D3F81E735B08DF" authority="Esaki, 1923" authorityName="Esaki" authorityYear="1923" box="[460,776,1963,1989]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="BE83EA89FFD9293E39D3F81E723808DF" box="[460,619,1963,1989]" italics="true" pageId="5" pageNumber="6">R. takahashii</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3A68F819735B08DF" author="Esaki" box="[631,776,1963,1989]" firstAuthor="Esaki" pageId="5" pageNumber="6" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki, 1923</bibRefCitation>
</taxonomicName>
, dealated macropterous form.
<emphasis id="BE83EA89FFD9293E3C60F81974C708DF" bold="true" box="[1151,1172,1963,1989]" pageId="5" pageNumber="6">R</emphasis>
.
<taxonomicName id="4BF74D18FFD9293E3CB8F81E713408F2" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="zetteli">
<emphasis id="BE83EA89FFD9293E3CB8F81E754108DF" box="[1191,1298,1963,1989]" italics="true" pageId="5" pageNumber="6">R. zetteli</emphasis>
<bibRefCitation id="E8664B6AFFD9293E3D02F81E713408F2" author="Tran &amp; Nguyen" firstAuthor="Tran" pageId="5" pageNumber="6" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
. Scale bar = 3 mm.
</paragraph>
</caption>
<caption id="D8886613FFDA293D38A2F949756808F3" ID-DOI="http://doi.org/10.5281/zenodo.8385642" ID-Zenodo-Dep="8385642" httpUri="https://zenodo.org/record/8385642/files/figure.png" pageId="6" pageNumber="7" startId="6.[189,232,1787,1813]" targetBox="[257,1326,279,1736]" targetPageId="6">
<paragraph id="8C48369BFFDA293D38A2F949756808F3" blockId="6.[185,1399,1787,2025]" pageId="6" pageNumber="7">
<emphasis id="BE83EA89FFDA293D38A2F949715E080F" bold="true" box="[189,269,1787,1813]" pageId="6" pageNumber="7">Fig. 4.</emphasis>
Photographs of bodies of
<taxonomicName id="4BF74D18FFDA293D3A54F949734B080C" box="[587,792,1787,1814]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BE83EA89FFDA293D3A54F94972B2080F" box="[587,737,1787,1813]" italics="true" pageId="6" pageNumber="7">Rhyacobates</emphasis>
spp.
</taxonomicName>
in dorsal view (males), apterous form if not stated otherwise.
<emphasis id="BE83EA89FFDA293D3925F8AC711C0822" bold="true" box="[314,335,1822,1848]" pageId="6" pageNumber="7">A</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3944F8AD71CC0822" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[347,415,1822,1848]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="bui" status="sp. nov.">
<emphasis id="BE83EA89FFDA293D3944F8AD71CC0822" box="[347,415,1822,1848]" italics="true" pageId="6" pageNumber="7">R. bui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFDA293D39BBF8AD71AF0823" box="[420,508,1823,1849]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFDA293D39E0F8AC72400822" bold="true" box="[511,531,1822,1848]" pageId="6" pageNumber="7">B</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3A3EF8AD72E20822" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[545,689,1822,1848]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="BE83EA89FFDA293D3A3EF8AD72E20822" box="[545,689,1822,1848]" italics="true" pageId="6" pageNumber="7">R. elongatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFDA293D3AA9F8AD735C0823" box="[694,783,1823,1849]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFDA293D3B0EF8AC73750822" bold="true" box="[785,806,1822,1848]" pageId="6" pageNumber="7">C</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3B2AF8AD73E70822" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[821,948,1822,1848]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
<emphasis id="BE83EA89FFDA293D3B2AF8AD73E70822" box="[821,948,1822,1848]" italics="true" pageId="6" pageNumber="7">R. turgidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFDA293D3BA6F8AD74420823" box="[953,1041,1823,1849]" pageId="6" pageNumber="7" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFDA293D3C0BF8AC747A0822" bold="true" box="[1044,1065,1822,1848]" pageId="6" pageNumber="7">D</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3C28F8AD71190846" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="BE83EA89FFDA293D3C28F8AD74B40822" box="[1079,1255,1822,1848]" italics="true" pageId="6" pageNumber="7">R. abdominalis</emphasis>
<bibRefCitation id="E8664B6AFFDA293D3CF5F8AC71190846" author="Andersen &amp; Chen" firstAuthor="Andersen" pageId="6" pageNumber="7" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D3940F8F071200846" bold="true" box="[351,371,1858,1884]" pageId="6" pageNumber="7">E</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3997F8F073750841" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" box="[392,806,1857,1884]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="BE83EA89FFDA293D3997F8F072700841" box="[392,547,1857,1883]" italics="true" pageId="6" pageNumber="7">R. anderseni</emphasis>
<bibRefCitation id="E8664B6AFFDA293D3A30F8F073750841" author="Tran &amp; Yang" box="[559,806,1857,1884]" firstAuthor="Tran" pageId="6" pageNumber="7" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D3B28F8F0731A0846" bold="true" box="[823,841,1858,1884]" pageId="6" pageNumber="7">F</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3B41F8F075210846" authority="Hungerford &amp; Matsuda, 1959" authorityName="Hungerford &amp; Matsuda" authorityYear="1959" box="[862,1394,1857,1884]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="BE83EA89FFDA293D3B41F8F073A10841" box="[862,1010,1857,1883]" italics="true" pageId="6" pageNumber="7">R. chinensis</emphasis>
<bibRefCitation id="E8664B6AFFDA293D3C1FF8F375210846" author="Hungerford &amp; Matsuda" box="[1024,1394,1857,1884]" firstAuthor="Hungerford" pageId="6" pageNumber="7" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda, 1959</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D38A2F8D670870864" bold="true" box="[189,212,1892,1918]" pageId="6" pageNumber="7">G</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D38FDF8D472C6089A" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[226,661,1893,1920]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="edentatus">
<emphasis id="BE83EA89FFDA293D38FDF8D471220865" box="[226,369,1893,1919]" italics="true" pageId="6" pageNumber="7">R. edentatus</emphasis>
<bibRefCitation id="E8664B6AFFDA293D3968F8D772C6089A" author="Andersen &amp; Chen" box="[375,661,1893,1920]" firstAuthor="Andersen" pageId="6" pageNumber="7" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D3A81F8D772E60865" bold="true" box="[670,693,1893,1919]" pageId="6" pageNumber="7">H</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3ADBF8D474720865" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" box="[708,1057,1893,1919]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="gongvo">
<emphasis id="BE83EA89FFDA293D3ADBF8D473690865" box="[708,826,1894,1919]" italics="true" pageId="6" pageNumber="7">R. gongvo</emphasis>
<bibRefCitation id="E8664B6AFFDA293D3B5FF8D774720865" author="Tran &amp; Yang" box="[832,1057,1893,1919]" firstAuthor="Tran" pageId="6" pageNumber="7" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D3C32F8D7746B0865" bold="true" box="[1069,1080,1893,1919]" pageId="6" pageNumber="7">I</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3C59F8D4715608B9" authority="(Hungerford, 1957)" baseAuthorityName="Hungerford" baseAuthorityYear="1957" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFDA293D3C59F8D474850865" box="[1094,1238,1893,1919]" italics="true" pageId="6" pageNumber="7">R. lundbladi</emphasis>
(
<bibRefCitation id="E8664B6AFFDA293D3CFCF8D770AF08B9" author="Hungerford" firstAuthor="Hungerford" pageId="6" pageNumber="7" pagination="33 - 36" refId="ref33146" refString="Hungerford H. B. 1957. A new gerrid from China (Gerridae, Hemiptera). Journal of the Kansas Entomological Society 30 (2): 33 - 36." type="journal article" year="1957">Hungerford, 1957</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D390DF83A717208B8" bold="true" box="[274,289,1928,1954]" pageId="6" pageNumber="7">J</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D392EF83B72B008B9" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[305,739,1928,1955]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="BE83EA89FFDA293D392EF83B71E508B8" box="[305,438,1928,1954]" italics="true" pageId="6" pageNumber="7">R. malaisei</emphasis>
<bibRefCitation id="E8664B6AFFDA293D39A1F83A72B008B9" author="Andersen &amp; Chen" box="[446,739,1928,1955]" firstAuthor="Andersen" pageId="6" pageNumber="7" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D3AEEF83A735508B8" bold="true" box="[753,774,1928,1954]" pageId="6" pageNumber="7">K</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3B08F83B749808B9" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[791,1227,1928,1955]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="recurvus">
<emphasis id="BE83EA89FFDA293D3B08F83B73CC08B8" box="[791,927,1929,1954]" italics="true" pageId="6" pageNumber="7">R. recurvus</emphasis>
<bibRefCitation id="E8664B6AFFDA293D3BB9F83A749808B9" author="Andersen &amp; Chen" box="[934,1227,1928,1955]" firstAuthor="Andersen" pageId="6" pageNumber="7" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D3CC5F83A74BF08B8" bold="true" box="[1242,1260,1928,1954]" pageId="6" pageNumber="7">L</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3CE3F83B71A608DC" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="scorpio">
<emphasis id="BE83EA89FFDA293D3CE3F83B752508B8" box="[1276,1398,1929,1954]" italics="true" pageId="6" pageNumber="7">R. scorpio</emphasis>
<bibRefCitation id="E8664B6AFFDA293D38A2F81971A608DC" author="Andersen &amp; Chen" box="[189,501,1963,1990]" firstAuthor="Andersen" pageId="6" pageNumber="7" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDA293D3A17F819727808DF" bold="true" box="[520,555,1963,1989]" pageId="6" pageNumber="7">M.</emphasis>
<taxonomicName id="4BF74D18FFDA293D3A24F81E739B08DF" authority="(Lundblad, 1934)" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[571,968,1963,1989]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFDA293D3A24F81E72B308DF" box="[571,736,1963,1989]" italics="true" pageId="6" pageNumber="7">R. svenhedini</emphasis>
(
<bibRefCitation id="E8664B6AFFDA293D3AE7F81973EC08DF" author="Lundblad" box="[760,959,1963,1989]" firstAuthor="Lundblad" pageId="6" pageNumber="7" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934</bibRefCitation>
)
</taxonomicName>
, apterous form.
<emphasis id="BE83EA89FFDA293D3C82F81974E108DF" bold="true" box="[1181,1202,1963,1989]" pageId="6" pageNumber="7">N</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D3CD6F81E753C08DF" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[1225,1391,1963,1989]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFDA293D3CD6F81E753C08DF" box="[1225,1391,1963,1989]" italics="true" pageId="6" pageNumber="7">R. svenhedini</emphasis>
</taxonomicName>
, macropterous form.
<emphasis id="BE83EA89FFDA293D39B3F87C719008F2" bold="true" box="[428,451,1998,2024]" pageId="6" pageNumber="7">O</emphasis>
.
<taxonomicName id="4BF74D18FFDA293D39CEF87D735308F2" authority="Esaki, 1923" authorityName="Esaki" authorityYear="1923" box="[465,768,1998,2024]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="BE83EA89FFDA293D39CEF87D723F08F2" box="[465,620,1998,2024]" italics="true" pageId="6" pageNumber="7">R. takahashii</emphasis>
<bibRefCitation id="E8664B6AFFDA293D3A6CF87C735308F2" author="Esaki" box="[627,768,1998,2024]" firstAuthor="Esaki" pageId="6" pageNumber="7" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki, 1923</bibRefCitation>
</taxonomicName>
, dealated macropterous form. Scale bar = 3 mm.
</paragraph>
</caption>
<subSubSection id="C4ED6510FFDB293C38A2FEBA74470E6E" pageId="7" pageNumber="8" type="distribution">
<paragraph id="8C48369BFFDB293C38A2FEBA71050E38" blockId="7.[189,342,264,290]" box="[189,342,264,290]" pageId="7" pageNumber="8">
<heading id="D70081F7FFDB293C38A2FEBA71050E38" bold="true" box="[189,342,264,290]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="BE83EA89FFDB293C38A2FEBA71050E38" bold="true" box="[189,342,264,290]" pageId="7" pageNumber="8">Distribution</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFDB293C38A2FE8574470E6E" blockId="7.[189,1398,311,373]" pageId="7" pageNumber="8">
The genus
<taxonomicName id="4BF74D18FFDB293C395DFE85718B0E4B" authorityName="Esaki" authorityYear="1923" box="[322,472,311,337]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDB293C395DFE85718B0E4B" box="[322,472,311,337]" italics="true" pageId="7" pageNumber="8">Rhyacobates</emphasis>
</taxonomicName>
is distributed from eastern Asia (the Korean Peninsula, mainland
<collectingCountry id="F4E0760BFFDB293C3CEAFE85756D0E4B" box="[1269,1342,311,337]" name="China" pageId="7" pageNumber="8">China</collectingCountry>
, and
<collectingCountry id="F4E0760BFFDB293C38A0FEE871350E6E" box="[191,358,346,372]" name="Taiwan Island" pageId="7" pageNumber="8">Taiwan Island</collectingCountry>
) to Indochina (
<collectingCountry id="F4E0760BFFDB293C3A07FEE872DD0E6E" box="[536,654,346,372]" name="Myanmar" pageId="7" pageNumber="8">Myanmar</collectingCountry>
,
<collectingCountry id="F4E0760BFFDB293C3A86FEE873570E6E" box="[665,772,346,372]" name="Thailand" pageId="7" pageNumber="8">Thailand</collectingCountry>
, and
<collectingCountry id="F4E0760BFFDB293C3B5EFEE873F50E6E" box="[833,934,346,372]" name="Vietnam" pageId="7" pageNumber="8">Vietnam</collectingCountry>
) (
<figureCitation id="14CC2A1EFFDB293C3BA1FEE874550E6E" box="[958,1030,346,372]" captionStart="Fig" captionStartId="11.[189,232,1998,2024]" captionTargetBox="[192,1391,472,1939]" captionTargetId="figure-51@11.[189,1398,456,1947]" captionTargetPageId="11" captionText="Fig. 9. Distribution map of Rhyacobates spp." figureDoi="http://doi.org/10.5281/zenodo.8385654" httpUri="https://zenodo.org/record/8385654/files/figure.png" pageId="7" pageNumber="8">Fig. 9</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C4ED6510FFDB293038A2FE29720F0E73" lastPageId="11" lastPageNumber="12" pageId="7" pageNumber="8" type="biology_ecology">
<paragraph id="8C48369BFFDB293C38A2FE2971E50EAC" blockId="7.[189,438,411,438]" box="[189,438,411,438]" pageId="7" pageNumber="8">
<heading id="D70081F7FFDB293C38A2FE2971E50EAC" bold="true" box="[189,438,411,438]" fontSize="11" level="3" pageId="7" pageNumber="8" reason="3">
<emphasis id="BE83EA89FFDB293C38A2FE2971E50EAC" bold="true" box="[189,438,411,438]" pageId="7" pageNumber="8">Biology and ecology</emphasis>
</heading>
</paragraph>
<paragraph id="8C48369BFFDB293C38F5FE7871050EFE" blockId="7.[189,1400,458,661]" box="[234,342,458,484]" pageId="7" pageNumber="8">
<emphasis id="BE83EA89FFDB293C38F5FE7871050EFE" bold="true" box="[234,342,458,484]" pageId="7" pageNumber="8">Habitats</emphasis>
</paragraph>
<paragraph id="8C48369BFFDB293C38A2FE5F740B0D8E" blockId="7.[189,1400,458,661]" pageId="7" pageNumber="8">
Species of
<taxonomicName id="4BF74D18FFDB293C395CFE5F718A0D1D" authorityName="Esaki" authorityYear="1923" box="[323,473,493,519]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDB293C395CFE5F718A0D1D" box="[323,473,493,519]" italics="true" pageId="7" pageNumber="8">Rhyacobates</emphasis>
</taxonomicName>
inhabit foot-hill and mountainous streams, rivers and sometimes pools. Most species are found only in running water with relatively cool temperatures (
<figureCitation id="14CC2A1EFFDB293C3C2AFDA374940D31" box="[1077,1223,529,555]" captionStart="Fig" captionStartId="12.[189,232,1928,1954]" captionTargetBox="[243,1344,265,1883]" captionTargetId="figure-14@12.[243,1344,265,1883]" captionTargetPageId="12" captionText="Fig. 10. Habitat of Rhyacobates spp. A. Habitat of R. chinensis Hungerford &amp; Matsuda, 1959, a muddy mountain stream. B. Habitat of R. chinensis Hungerford &amp; Matsuda, 1959, a muddy river near the mountain. C. Habitat of R. recurvus Andersen &amp; Chen, 1995, a mountain stream with emerged rocks." figureDoi="http://doi.org/10.5281/zenodo.8385658" httpUri="https://zenodo.org/record/8385658/files/figure.png" pageId="7" pageNumber="8">Fig. 10AC</figureCitation>
). Elevations of habitats have been recorded from
<quantity id="4B0F9B7EFFDB293C3A45FD8673530D54" box="[602,768,564,590]" metricMagnitude="3" metricUnit="m" metricValue="1.0315" metricValueMax="2.041" metricValueMin="0.022" pageId="7" pageNumber="8" unit="m" value="1031.5" valueMax="2041.0" valueMin="22.0">22 to 2041 m</quantity>
, but mostly between 500 and
<quantity id="4B0F9B7EFFDB293C3C71FD86749E0D54" box="[1134,1229,564,590]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="7" pageNumber="8" unit="m" value="1000.0">1000 m</quantity>
. Streams with emergent rocks in the current are typical habitats of
<taxonomicName id="4BF74D18FFDB293C3B30FDE573900D6B" authorityName="Esaki" authorityYear="1923" box="[815,963,599,625]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFDB293C3B30FDE573900D6B" box="[815,963,599,625]" italics="true" pageId="7" pageNumber="8">Rhyacobates</emphasis>
</taxonomicName>
, as the rocks offer necessary resting areas for these skaters (
<bibRefCitation id="E8664B6AFFDB293C39CFFDC8720B0D8E" author="Esaki" box="[464,600,634,660]" firstAuthor="Esaki" pageId="7" pageNumber="8" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki 1923</bibRefCitation>
;
<bibRefCitation id="E8664B6AFFDB293C3A7BFDC973120D8E" author="Tran &amp; Yang" box="[612,833,634,661]" firstAuthor="Tran" pageId="7" pageNumber="8" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang 2006</bibRefCitation>
;
<bibRefCitation id="E8664B6AFFDB293C3B53FDC9741F0D8E" author="Tran &amp; Nguyen" box="[844,1100,634,661]" firstAuthor="Tran" pageId="7" pageNumber="8" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen 2016</bibRefCitation>
).
</paragraph>
<caption id="D8886613FFDB293C38A2F97D718C08FB" ID-DOI="http://doi.org/10.5281/zenodo.8385644" ID-Zenodo-Dep="8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="7" pageNumber="8" startId="7.[189,232,1743,1769]" targetBox="[200,1390,762,1681]" targetPageId="7">
<paragraph id="8C48369BFFDB293C38A2F97D718C08FB" blockId="7.[189,1399,1743,2017]" pageId="7" pageNumber="8">
<emphasis id="BE83EA89FFDB293C38A2F97D715809F0" bold="true" box="[189,267,1743,1770]" pageId="7" pageNumber="8">Fig. 5.</emphasis>
Photographs of bodies of
<taxonomicName id="4BF74D18FFDB293C3A5FF962735D09F0" box="[576,782,1744,1770]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BE83EA89FFDB293C3A5FF962728509F0" box="[576,726,1744,1770]" italics="true" pageId="7" pageNumber="8">Rhyacobates</emphasis>
spp.
</taxonomicName>
in lateral view (apterous females).
<emphasis id="BE83EA89FFDB293C3CB4F97D749309F3" bold="true" box="[1195,1216,1743,1769]" pageId="7" pageNumber="8">A</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3CD0F962754709F0" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[1231,1300,1744,1770]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="bui" status="sp. nov.">
<emphasis id="BE83EA89FFDB293C3CD0F962754709F0" box="[1231,1300,1744,1770]" italics="true" pageId="7" pageNumber="8">R. bui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFDB293C3D03F962752409F0" box="[1308,1399,1744,1770]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFDB293C38A2F94170820817" bold="true" box="[189,209,1779,1805]" pageId="7" pageNumber="8">B</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C38C2F946713E0817" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[221,365,1779,1805]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="BE83EA89FFDB293C38C2F946713E0817" box="[221,365,1779,1805]" italics="true" pageId="7" pageNumber="8">R. elongatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFDB293C396DF94171990817" box="[370,458,1779,1805]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFDB293C39D3F94171B20817" bold="true" box="[460,481,1779,1805]" pageId="7" pageNumber="8">C</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C39F0F946723C0817" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[495,623,1779,1805]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
<emphasis id="BE83EA89FFDB293C39F0F946723C0817" box="[495,623,1779,1805]" italics="true" pageId="7" pageNumber="8">R. turgidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFDB293C3A6CF94172980817" box="[627,715,1779,1805]" pageId="7" pageNumber="8" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFDB293C3AD2F94172B10817" bold="true" box="[717,738,1779,1805]" pageId="7" pageNumber="8">D</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3AEEF94674EF0814" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[753,1212,1779,1806]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="BE83EA89FFDB293C3AEEF94673F30817" box="[753,928,1779,1805]" italics="true" pageId="7" pageNumber="8">R. abdominalis</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3BBCF94174EF0814" author="Andersen &amp; Chen" box="[931,1212,1779,1806]" firstAuthor="Andersen" pageId="7" pageNumber="8" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3CDDF94174850817" bold="true" box="[1218,1238,1779,1805]" pageId="7" pageNumber="8">E</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3CFAF94671EB082A" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="BE83EA89FFDB293C3CFAF94675240817" box="[1253,1399,1779,1805]" italics="true" pageId="7" pageNumber="8">R. anderseni</emphasis>
<bibRefCitation id="E8664B6AFFDB293C38A2F8A571EB082A" author="Tran &amp; Yang" box="[189,440,1814,1841]" firstAuthor="Tran" pageId="7" pageNumber="8" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C39D4F8A4718E082A" bold="true" box="[459,477,1814,1840]" pageId="7" pageNumber="8">F</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C39EBF8A573FC082A" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[500,943,1814,1841]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="BE83EA89FFDB293C39EBF8A572D7082A" box="[500,644,1815,1840]" italics="true" pageId="7" pageNumber="8">R. angustus</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3A8CF8A573FC082A" author="Tran &amp; Nguyen" box="[659,943,1814,1841]" firstAuthor="Tran" pageId="7" pageNumber="8" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
, holotype (ZVNU).
<emphasis id="BE83EA89FFDB293C3CABF8A47498082A" bold="true" box="[1204,1227,1814,1840]" pageId="7" pageNumber="8">G</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3CFEF8A57274084E" authority="Hungerford &amp; Matsuda, 1959" authorityName="Hungerford &amp; Matsuda" authorityYear="1959" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="BE83EA89FFDB293C3CFEF8A57525082A" box="[1249,1398,1814,1840]" italics="true" pageId="7" pageNumber="8">R. chinensis</emphasis>
<bibRefCitation id="E8664B6AFFDB293C38A2F88B7274084E" author="Hungerford &amp; Matsuda" box="[189,551,1849,1876]" firstAuthor="Hungerford" pageId="7" pageNumber="8" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda, 1959</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3A2AF88B721F0849" bold="true" box="[565,588,1849,1875]" pageId="7" pageNumber="8">H</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3A41F888744F0849" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[606,1052,1849,1876]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="BE83EA89FFDB293C3A41F88873500849" box="[606,771,1850,1875]" italics="true" pageId="7" pageNumber="8">R. constrictus</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3B12F888744F0849" author="Tran &amp; Nguyen" box="[781,1052,1849,1876]" firstAuthor="Tran" pageId="7" pageNumber="8" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3C35F88B74660849" bold="true" box="[1066,1077,1849,1875]" pageId="7" pageNumber="8">I</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3C58F888711E086D" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="edentatus">
<emphasis id="BE83EA89FFDB293C3C58F88874880849" box="[1095,1243,1849,1875]" italics="true" pageId="7" pageNumber="8">R. edentatus</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3CFAF88B711E086D" author="Andersen &amp; Chen" firstAuthor="Andersen" pageId="7" pageNumber="8" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3944F8EF7139086D" bold="true" box="[347,362,1885,1911]" pageId="7" pageNumber="8">J</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3963F8EF72A0086C" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" box="[380,755,1884,1911]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="gongvo">
<emphasis id="BE83EA89FFDB293C3963F8EF71A4086C" box="[380,503,1885,1910]" italics="true" pageId="7" pageNumber="8">R. gongvo</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3A1CF8EF72A0086C" author="Tran &amp; Yang" box="[515,755,1884,1911]" firstAuthor="Tran" pageId="7" pageNumber="8" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3B1DF8EF734A086D" bold="true" box="[770,793,1885,1911]" pageId="7" pageNumber="8">K</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3B34F8EF74EA086D" authority="(Hungerford, 1957)" baseAuthorityName="Hungerford" baseAuthorityYear="1957" box="[811,1209,1884,1911]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="lundbladi">
<emphasis id="BE83EA89FFDB293C3B34F8EF7393086C" box="[811,960,1884,1910]" italics="true" pageId="7" pageNumber="8">R. lundbladi</emphasis>
(
<bibRefCitation id="E8664B6AFFDB293C3BCBF8EE74E3086D" author="Hungerford" box="[980,1200,1884,1911]" firstAuthor="Hungerford" pageId="7" pageNumber="8" pagination="33 - 36" refId="ref33146" refString="Hungerford H. B. 1957. A new gerrid from China (Gerridae, Hemiptera). Journal of the Kansas Entomological Society 30 (2): 33 - 36." type="journal article" year="1957">Hungerford, 1957</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3CD7F8EF748F086D" bold="true" box="[1224,1244,1885,1911]" pageId="7" pageNumber="8">L</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3CF1F8EF71B50880" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="malaisei">
<emphasis id="BE83EA89FFDB293C3CF1F8EF7525086C" box="[1262,1398,1884,1910]" italics="true" pageId="7" pageNumber="8">R. malaisei</emphasis>
<bibRefCitation id="E8664B6AFFDB293C38A2F83271B50880" author="Andersen &amp; Chen" box="[189,486,1919,1946]" firstAuthor="Andersen" pageId="7" pageNumber="8" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C39EBF83272430880" bold="true" box="[500,528,1920,1946]" pageId="7" pageNumber="8">M</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3A3DF832738F0880" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[546,988,1919,1946]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="recurvus">
<emphasis id="BE83EA89FFDB293C3A3DF83272F80883" box="[546,683,1920,1945]" italics="true" pageId="7" pageNumber="8">R. recurvus</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3AACF832738F0880" author="Andersen &amp; Chen" box="[691,988,1919,1946]" firstAuthor="Andersen" pageId="7" pageNumber="8" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3BF4F83274530880" bold="true" box="[1003,1024,1920,1946]" pageId="7" pageNumber="8">N</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3C0EF83270A808A4" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="scorpio">
<emphasis id="BE83EA89FFDB293C3C0EF83274DE0883" box="[1041,1165,1920,1945]" italics="true" pageId="7" pageNumber="8">R. scorpio</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3C8AF83270A808A4" author="Andersen &amp; Chen" firstAuthor="Andersen" pageId="7" pageNumber="8" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3917F811714C08A7" bold="true" box="[264,287,1955,1981]" pageId="7" pageNumber="8">O</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3930F81672F808A7" authority="(Lundblad, 1934)" baseAuthorityName="Lundblad" baseAuthorityYear="1934" box="[303,683,1955,1981]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="svenhedini">
<emphasis id="BE83EA89FFDB293C3930F816719D08A7" box="[303,462,1955,1981]" italics="true" pageId="7" pageNumber="8">R. svenhedini</emphasis>
(
<bibRefCitation id="E8664B6AFFDB293C39FFF81172F208A7" author="Lundblad" box="[480,673,1955,1981]" firstAuthor="Lundblad" pageId="7" pageNumber="8" pagination="1 - 31" refId="ref33236" refString="Lundblad O. M. 1934. Schwedisch-Chinesische wissenschafliche Expedition nach den nordwestlichen Provinzen Chinas. Arkiv f ˆ r Zoologi 27 A (14): 1 - 31." type="journal article" year="1934">Lundblad, 1934</bibRefCitation>
)
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3AA6F811729A08A7" bold="true" box="[697,713,1955,1981]" pageId="7" pageNumber="8">P</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3AC6F816745D08A7" authority="Esaki, 1923" authorityName="Esaki" authorityYear="1923" box="[729,1038,1955,1981]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="takahashii">
<emphasis id="BE83EA89FFDB293C3AC6F816732508A7" box="[729,886,1955,1981]" italics="true" pageId="7" pageNumber="8">R. takahashii</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3B60F811745D08A7" author="Esaki" box="[895,1038,1955,1981]" firstAuthor="Esaki" pageId="7" pageNumber="8" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki, 1923</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFDB293C3C02F811746108A7" bold="true" box="[1053,1074,1955,1981]" pageId="7" pageNumber="8">Q</emphasis>
.
<taxonomicName id="4BF74D18FFDB293C3C5DF81670AE08FA" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="8" phylum="Arthropoda" rank="species" species="zetteli">
<emphasis id="BE83EA89FFDB293C3C5DF81674F808A7" box="[1090,1195,1955,1981]" italics="true" pageId="7" pageNumber="8">R. zetteli</emphasis>
<bibRefCitation id="E8664B6AFFDB293C3CACF81170AE08FA" author="Tran &amp; Nguyen" firstAuthor="Tran" pageId="7" pageNumber="8" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
. Scale bar = 3 mm.
</paragraph>
</caption>
<paragraph id="8C48369BFFD4293338F5FEBA71DF0E38" blockId="8.[189,1400,264,467]" box="[234,396,264,290]" pageId="8" pageNumber="9">
<emphasis id="BE83EA89FFD4293338F5FEBA71DF0E38" bold="true" box="[234,396,264,290]" pageId="8" pageNumber="9">Living forms</emphasis>
</paragraph>
<paragraph id="8C48369BFFD4293338A2FE9E74D40EC9" blockId="8.[189,1400,264,467]" pageId="8" pageNumber="9">
Usually, most adults found in the populations are the apterous form (
<figureCitation id="14CC2A1EFFD429333BE9FE9E74DB0E5C" box="[1014,1160,300,326]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[221,1366,265,1852]" captionTargetId="figure-11@13.[221,1366,265,1852]" captionTargetPageId="13" captionText="Fig. 11. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, live habitus in situ. A. A group of nymphs and adults resting on a riverside rock. B. Individual of apterous female resting on rock. C. Individual of apterous male striding on the water surface. D. A group of nymphs and adults shifting on the water surface when disturbed. E. Nymphs resting on rock. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385660" httpUri="https://zenodo.org/record/8385660/files/figure.png" pageId="8" pageNumber="9">Fig. 11AD</figureCitation>
). However, in some rare situations, a population may completely consist of macropterous and dealated form, e.g., of
<taxonomicName id="4BF74D18FFD4293338A2FEC1711D0E96" authorityName="Hungerford &amp; Matsuda" authorityYear="1959" box="[189,334,370,396]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="BE83EA89FFD4293338A2FEC1711D0E96" box="[189,334,370,396]" italics="true" pageId="8" pageNumber="9">R. chinensis</emphasis>
</taxonomicName>
(
<figureCitation id="14CC2A1EFFD42933397EFEC0719C0E96" box="[353,463,370,396]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[247,1341,265,1811]" captionTargetId="figure-15@14.[247,1341,265,1811]" captionTargetPageId="14" captionText="Fig. 12. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. A. Dealated macropterous male mating with apterous female, lateral view. B. Dealated macropterous male mating with dealated macropterous female, dorsal view. C. Dealated macropterous male mating with apterous female, dorsal view. D. Apterous male mating with apterous female on the water surface, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385662" httpUri="https://zenodo.org/record/8385662/files/figure.png" pageId="8" pageNumber="9">Fig. 12B</figureCitation>
).
<bibRefCitation id="E8664B6AFFD4293339F8FEC172AD0E96" author="Tran &amp; Nguyen" box="[487,766,370,397]" firstAuthor="Tran" pageId="8" pageNumber="9" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen (2016)</bibRefCitation>
reported one macropterous specimen (with dealated wings) of
<taxonomicName id="4BF74D18FFD429333928FE2471CC0EB5" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[311,415,405,431]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="zetteli">
<emphasis id="BE83EA89FFD429333928FE2471CC0EB5" box="[311,415,405,431]" italics="true" pageId="8" pageNumber="9">R. zetteli</emphasis>
</taxonomicName>
inhabiting an unshaded, tiny water flow ca
<quantity id="4B0F9B7EFFD429333BB9FE2773A10EAA" box="[934,1010,405,432]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="8" pageNumber="9" unit="m" value="500.0">500 m</quantity>
away from the population in the main stream, indicating that the macropterous form might have good flying ability.
</paragraph>
<caption id="D8886613FFD4293338A2F8D674C408F3" ID-DOI="http://doi.org/10.5281/zenodo.8385646" ID-Zenodo-Dep="8385646" httpUri="https://zenodo.org/record/8385646/files/figure.png" pageId="8" pageNumber="9" startId="8.[189,232,1892,1918]" targetBox="[229,1341,570,1825]" targetPageId="8">
<paragraph id="8C48369BFFD4293338A2F8D674C408F3" blockId="8.[189,1399,1892,2025]" pageId="8" pageNumber="9">
<emphasis id="BE83EA89FFD4293338A2F8D671580865" bold="true" box="[189,267,1892,1919]" pageId="8" pageNumber="9">Fig. 6.</emphasis>
Proctiger of
<taxonomicName id="4BF74D18FFD4293339BAF8D772200865" box="[421,627,1893,1919]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="BE83EA89FFD4293339BAF8D772680865" box="[421,571,1893,1919]" italics="true" pageId="8" pageNumber="9">Rhyacobates</emphasis>
spp.
</taxonomicName>
in dorsal view (males).
<emphasis id="BE83EA89FFD429333B91F8D673F00864" bold="true" box="[910,931,1892,1918]" pageId="8" pageNumber="9">A</emphasis>
.
<taxonomicName id="4BF74D18FFD429333BAEF8D473A40865" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[945,1015,1893,1919]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="bui" status="sp. nov.">
<emphasis id="BE83EA89FFD429333BAEF8D473A40865" box="[945,1015,1893,1919]" italics="true" pageId="8" pageNumber="9">R. bui</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFD429333BE0F8D7740A0865" box="[1023,1113,1893,1919]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFD429333C7FF8D774270865" bold="true" box="[1120,1140,1893,1919]" pageId="8" pageNumber="9">B</emphasis>
.
<taxonomicName id="4BF74D18FFD429333C9DF8D475460865" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[1154,1301,1893,1919]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="elongatus" status="sp. nov.">
<emphasis id="BE83EA89FFD429333C9DF8D475460865" box="[1154,1301,1893,1919]" italics="true" pageId="8" pageNumber="9">R. elongatus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFD429333D03F8D775240865" box="[1308,1399,1893,1919]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFD4293338A2F83A708108B8" bold="true" box="[189,210,1928,1954]" pageId="8" pageNumber="9">C</emphasis>
.
<taxonomicName id="4BF74D18FFD4293338FEF83B713108B8" authority="Leng &amp; Tran, 2023" authorityName="Leng &amp; Tran" authorityYear="2023" box="[225,354,1928,1954]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="turgidus" status="sp. nov.">
<emphasis id="BE83EA89FFD4293338FEF83B713108B8" box="[225,354,1928,1954]" italics="true" pageId="8" pageNumber="9">R. turgidus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A5B057F2FFD429333976F83A719708B8" box="[361,452,1928,1954]" pageId="8" pageNumber="9" rank="species">sp. nov.</taxonomicNameLabel>
<emphasis id="BE83EA89FFD4293339D4F83A71B308B8" bold="true" box="[459,480,1928,1954]" pageId="8" pageNumber="9">D</emphasis>
.
<taxonomicName id="4BF74D18FFD4293339F0F83B739508B9" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[495,966,1928,1955]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="abdominalis">
<emphasis id="BE83EA89FFD4293339F0F83B72F308B8" box="[495,672,1928,1954]" italics="true" pageId="8" pageNumber="9">R. abdominalis</emphasis>
<bibRefCitation id="E8664B6AFFD429333AB9F83A739508B9" author="Andersen &amp; Chen" box="[678,966,1928,1955]" firstAuthor="Andersen" pageId="8" pageNumber="9" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD429333BCEF83A73B608B8" bold="true" box="[977,997,1928,1954]" pageId="8" pageNumber="9">E</emphasis>
.
<taxonomicName id="4BF74D18FFD429333BEBF83B752108B8" authority="Tran &amp; Yang, 2006" authorityName="Tran &amp; Yang" authorityYear="2006" box="[1012,1394,1928,1954]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="anderseni">
<emphasis id="BE83EA89FFD429333BEBF83B74DB08B8" box="[1012,1160,1928,1954]" italics="true" pageId="8" pageNumber="9">R. anderseni</emphasis>
<bibRefCitation id="E8664B6AFFD429333C90F83A752108B8" author="Tran &amp; Yang" box="[1167,1394,1928,1954]" firstAuthor="Tran" pageId="8" pageNumber="9" pagination="11 - 20" refId="ref33532" refString="Tran A. D. &amp; Yang C. M. 2006. New species of the water strider genera Eotrechus Kirkaldy and Rhyacobates Esaki (Heteroptera: Gerridae) from Vietnam. Raffles Bulletin of Zoology 54 (1): 11 - 20." type="journal article" year="2006">Tran &amp; Yang, 2006</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD4293338A2F819709C08DF" bold="true" box="[189,207,1963,1989]" pageId="8" pageNumber="9">F</emphasis>
.
<taxonomicName id="4BF74D18FFD4293338C1F81E722408DF" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" box="[222,631,1963,1990]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="angustus">
<emphasis id="BE83EA89FFD4293338C1F81E713B08DF" box="[222,360,1964,1989]" italics="true" pageId="8" pageNumber="9">R. angustus</emphasis>
<bibRefCitation id="E8664B6AFFD429333970F81E722408DF" author="Tran &amp; Nguyen" box="[367,631,1963,1990]" firstAuthor="Tran" pageId="8" pageNumber="9" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD429333A9BF81972C908DF" bold="true" box="[644,666,1963,1989]" pageId="8" pageNumber="9">G</emphasis>
.
<taxonomicName id="4BF74D18FFD429333AB6F81E74F108DC" authority="Hungerford &amp; Matsuda, 1959" authorityName="Hungerford &amp; Matsuda" authorityYear="1959" box="[681,1186,1963,1990]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="chinensis">
<emphasis id="BE83EA89FFD429333AB6F81E736B08DF" box="[681,824,1963,1989]" italics="true" pageId="8" pageNumber="9">R. chinensis</emphasis>
<bibRefCitation id="E8664B6AFFD429333B5FF81974F108DC" author="Hungerford &amp; Matsuda" box="[832,1186,1963,1990]" firstAuthor="Hungerford" pageId="8" pageNumber="9" pagination="67 - 92" refId="ref33179" refString="Hungerford H. B. &amp; Matsuda R. 1959. Synonymy of the genera Rhyacobates Esaki, 1923 and Esakobates Lundblad, 1934, and a description of a new species of Rhyacobates from China (Hemiptera: Gerridae). Journal of the Kansas Entomological Society 32 (1): 67 - 92." type="journal article" year="1959">Hungerford &amp; Matsuda, 1959</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD429333CB0F819749608DF" bold="true" box="[1199,1221,1963,1989]" pageId="8" pageNumber="9">H</emphasis>
.
<taxonomicName id="4BF74D18FFD429333CCBF81E719108F2" authority="Tran &amp; Nguyen, 2016" authorityName="Tran &amp; Nguyen" authorityYear="2016" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="constrictus">
<emphasis id="BE83EA89FFD429333CCBF81E752408DF" box="[1236,1399,1964,1989]" italics="true" pageId="8" pageNumber="9">R. constrictus</emphasis>
<bibRefCitation id="E8664B6AFFD4293338A2F87D719108F2" author="Tran &amp; Nguyen" box="[189,450,1998,2025]" firstAuthor="Tran" pageId="8" pageNumber="9" pagination="501 - 516" refId="ref33477" refString="Tran A. D. &amp; Nguyen X. Q. 2016. Three new species of the water strider genus Rhyacobates Esaki, 1923 (Hemiptera: Gerridae) from Vietnam. Zootaxa 4121 (5): 501 - 516. https: // doi. org / 10.11646 / zootaxa. 4121.5.1" type="journal article" year="2016">Tran &amp; Nguyen, 2016</bibRefCitation>
</taxonomicName>
.
<emphasis id="BE83EA89FFD4293339D1F87C718A08F2" bold="true" box="[462,473,1998,2024]" pageId="8" pageNumber="9">I</emphasis>
.
<taxonomicName id="4BF74D18FFD4293339F7F87D73CC08F3" authority="Andersen &amp; Chen, 1995" authorityName="Andersen &amp; Chen" authorityYear="1995" box="[488,927,1998,2025]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="8" pageNumber="9" phylum="Arthropoda" rank="species" species="edentatus">
<emphasis id="BE83EA89FFD4293339F7F87D722B08F2" box="[488,632,1998,2024]" italics="true" pageId="8" pageNumber="9">R. edentatus</emphasis>
<bibRefCitation id="E8664B6AFFD429333A61F87C73CC08F3" author="Andersen &amp; Chen" box="[638,927,1998,2025]" firstAuthor="Andersen" pageId="8" pageNumber="9" pagination="51 - 67" refId="ref32941" refString="Andersen N. M. &amp; Chen P. P. 1995. A taxonomic revision of the Ptilomerine genus Rhyacobates Esaki (Hemiptera: Gerridae), with five new species from China and adjacent countries. Tijdschrift voor Entomologie 138: 51 - 67." type="journal article" year="1995">Andersen &amp; Chen, 1995</bibRefCitation>
</taxonomicName>
. Scale bar = 0.2 mm.
</paragraph>
</caption>
<paragraph id="8C48369BFFD5293238F5FEBA71820E38" blockId="9.[189,1399,264,502]" box="[234,465,264,290]" pageId="9" pageNumber="10">
<emphasis id="BE83EA89FFD5293238F5FEBA71820E38" bold="true" box="[234,465,264,290]" pageId="9" pageNumber="10">Perching behavior</emphasis>
</paragraph>
<paragraph id="8C48369BFFD5293238A2FE9E72650EEC" blockId="9.[189,1399,264,502]" pageId="9" pageNumber="10">
Although most species of
<taxonomicName id="4BF74D18FFD5293239E6FE9E72DC0E5C" authorityName="Esaki" authorityYear="1923" box="[505,655,300,326]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFD5293239E6FE9E72DC0E5C" box="[505,655,300,326]" italics="true" pageId="9" pageNumber="10">Rhyacobates</emphasis>
</taxonomicName>
can stride on torrent or fast-running water, they spend plenty of time resting on waterside rocks (
<figureCitation id="14CC2A1EFFD529323A7BFEFD734B0E73" box="[612,792,335,361]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[221,1366,265,1852]" captionTargetId="figure-11@13.[221,1366,265,1852]" captionTargetPageId="13" captionText="Fig. 11. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, live habitus in situ. A. A group of nymphs and adults resting on a riverside rock. B. Individual of apterous female resting on rock. C. Individual of apterous male striding on the water surface. D. A group of nymphs and adults shifting on the water surface when disturbed. E. Nymphs resting on rock. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385660" httpUri="https://zenodo.org/record/8385660/files/figure.png" pageId="9" pageNumber="10">Figs 11AB, E</figureCitation>
,
<figureCitation id="14CC2A1EFFD529323B39FEFD732D0E73" box="[806,894,335,361]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[247,1341,265,1811]" captionTargetId="figure-15@14.[247,1341,265,1811]" captionTargetPageId="14" captionText="Fig. 12. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. A. Dealated macropterous male mating with apterous female, lateral view. B. Dealated macropterous male mating with dealated macropterous female, dorsal view. C. Dealated macropterous male mating with apterous female, dorsal view. D. Apterous male mating with apterous female on the water surface, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385662" httpUri="https://zenodo.org/record/8385662/files/figure.png" pageId="9" pageNumber="10">12AC</figureCitation>
,
<figureCitation id="14CC2A1EFFD529323B94FEFD73F90E73" box="[907,938,335,361]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[255,1332,265,1881]" captionTargetId="figure-11@15.[255,1332,265,1881]" captionTargetPageId="15" captionText="Fig. 13. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. Apterous male mating and seizing the dealated macropterous female using its fore leg on a riverside rock, while the female is feeding on its prey." figureDoi="http://doi.org/10.5281/zenodo.8385664" httpUri="https://zenodo.org/record/8385664/files/figure.png" pageId="9" pageNumber="10">13</figureCitation>
,
<figureCitation id="14CC2A1EFFD529323BA6FEFD74400E73" box="[953,1043,335,361]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[233,1355,265,1881]" captionTargetId="figure-15@16.[233,1355,265,1881]" captionTargetPageId="16" captionText="Fig. 14. Rhyacobates recurvus Andersen &amp; Chen, 1995, live habitus in situ. A. Nnymphs and adult resting on a rock over a vortex, dorsal view. BC. Apterous male mating with apterous female on a riverside rock, dorsal view. D. A nymph resting on a riverside rock, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385666" httpUri="https://zenodo.org/record/8385666/files/figure.png" pageId="9" pageNumber="10">14AD</figureCitation>
), which was first observed by
<bibRefCitation id="E8664B6AFFD5293238A2FEC071040E96" author="Esaki" box="[189,343,370,396]" firstAuthor="Esaki" pageId="9" pageNumber="10" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki (1923)</bibRefCitation>
. They are very alert and when potential enemies (predators or larger animals) are detected nearby, they will jump into the water and stride irregularly at an extremely fast speed (
<figureCitation id="14CC2A1EFFD529323CABFE2775730EB5" box="[1204,1312,405,431]" captionStart="Fig" captionStartId="13.[189,232,1892,1918]" captionTargetBox="[221,1366,265,1852]" captionTargetId="figure-11@13.[221,1366,265,1852]" captionTargetPageId="13" captionText="Fig. 11. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, live habitus in situ. A. A group of nymphs and adults resting on a riverside rock. B. Individual of apterous female resting on rock. C. Individual of apterous male striding on the water surface. D. A group of nymphs and adults shifting on the water surface when disturbed. E. Nymphs resting on rock. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385660" httpUri="https://zenodo.org/record/8385660/files/figure.png" pageId="9" pageNumber="10">Fig. 11D</figureCitation>
).
<bibRefCitation id="E8664B6AFFD529323D2AFE27715F0EC9" author="Esaki" firstAuthor="Esaki" pageId="9" pageNumber="10" pagination="387 - 391" refId="ref33092" refString="Esaki T. 1923. An interesting new water strider from Formosa. Philippine Journal of Science 22 (4): 387 - 391." type="journal article" year="1923">Esaki (1923)</bibRefCitation>
hypothesized that this swift gliding may cause disturbance of the water and make the insects unrecognizable to the predators.
</paragraph>
<paragraph id="8C48369BFFD5293238F5FDAF718C0D2D" blockId="9.[189,1399,541,708]" box="[234,479,541,567]" pageId="9" pageNumber="10">
<emphasis id="BE83EA89FFD5293238F5FDAF718C0D2D" bold="true" box="[234,479,541,567]" pageId="9" pageNumber="10">Predatory behavior</emphasis>
</paragraph>
<paragraph id="8C48369BFFD5293238A2FDF271ED0DDE" blockId="9.[189,1399,541,708]" pageId="9" pageNumber="10">
Apparently, individuals of
<taxonomicName id="4BF74D18FFD529323A1EFDF272C40D40" authorityName="Esaki" authorityYear="1923" box="[513,663,576,602]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFD529323A1EFDF272C40D40" box="[513,663,576,602]" italics="true" pageId="9" pageNumber="10">Rhyacobates</emphasis>
</taxonomicName>
access the water surface and stride against the torrent when they are searching for food. We have observed that they can rapidly locate living insects floating nearby. They sometimes jump onto waterside rocks soon after they catch their prey, where they can feed on it undisturbed (
<figureCitation id="14CC2A1EFFD529323946FD1871E30DDE" box="[345,432,682,708]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[255,1332,265,1881]" captionTargetId="figure-11@15.[255,1332,265,1881]" captionTargetPageId="15" captionText="Fig. 13. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. Apterous male mating and seizing the dealated macropterous female using its fore leg on a riverside rock, while the female is feeding on its prey." figureDoi="http://doi.org/10.5281/zenodo.8385664" httpUri="https://zenodo.org/record/8385664/files/figure.png" pageId="9" pageNumber="10">Fig. 13</figureCitation>
).
</paragraph>
<paragraph id="8C48369BFFD5293038F5FD59720F0E73" blockId="9.[189,1399,747,915]" lastBlockId="11.[188,1399,265,362]" lastPageId="11" lastPageNumber="12" pageId="9" pageNumber="10">
<emphasis id="BE83EA89FFD5293238F5FD5971EF0C1F" bold="true" box="[234,444,747,773]" pageId="9" pageNumber="10">Mating behavior</emphasis>
During mating, the male jumps onto the dorsum of a female, hugging the female with its fore legs (
<figureCitation id="14CC2A1EFFD5293238DAFC80710F0C56" box="[197,348,818,844]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[247,1341,265,1811]" captionTargetId="figure-15@14.[247,1341,265,1811]" captionTargetPageId="14" captionText="Fig. 12. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. A. Dealated macropterous male mating with apterous female, lateral view. B. Dealated macropterous male mating with dealated macropterous female, dorsal view. C. Dealated macropterous male mating with apterous female, dorsal view. D. Apterous male mating with apterous female on the water surface, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385662" httpUri="https://zenodo.org/record/8385662/files/figure.png" pageId="9" pageNumber="10">Figs 12AD</figureCitation>
,
<figureCitation id="14CC2A1EFFD529323976FC8071DB0C56" box="[361,392,818,844]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[255,1332,265,1881]" captionTargetId="figure-11@15.[255,1332,265,1881]" captionTargetPageId="15" captionText="Fig. 13. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. Apterous male mating and seizing the dealated macropterous female using its fore leg on a riverside rock, while the female is feeding on its prey." figureDoi="http://doi.org/10.5281/zenodo.8385664" httpUri="https://zenodo.org/record/8385664/files/figure.png" pageId="9" pageNumber="10">13</figureCitation>
,
<figureCitation id="14CC2A1EFFD529323988FC8071BC0C56" box="[407,495,818,844]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[233,1355,265,1881]" captionTargetId="figure-15@16.[233,1355,265,1881]" captionTargetPageId="16" captionText="Fig. 14. Rhyacobates recurvus Andersen &amp; Chen, 1995, live habitus in situ. A. Nnymphs and adult resting on a rock over a vortex, dorsal view. BC. Apterous male mating with apterous female on a riverside rock, dorsal view. D. A nymph resting on a riverside rock, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385666" httpUri="https://zenodo.org/record/8385666/files/figure.png" pageId="9" pageNumber="10">14BC</figureCitation>
). The entire mating process may happen while skating on the water surface (
<figureCitation id="14CC2A1EFFD5293238D7FCE771600C75" box="[200,307,853,879]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[247,1341,265,1811]" captionTargetId="figure-15@14.[247,1341,265,1811]" captionTargetPageId="14" captionText="Fig. 12. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. A. Dealated macropterous male mating with apterous female, lateral view. B. Dealated macropterous male mating with dealated macropterous female, dorsal view. C. Dealated macropterous male mating with apterous female, dorsal view. D. Apterous male mating with apterous female on the water surface, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385662" httpUri="https://zenodo.org/record/8385662/files/figure.png" pageId="9" pageNumber="10">Fig. 12D</figureCitation>
) or on waterside rocks (
<figureCitation id="14CC2A1EFFD529323A51FCE772B20C75" box="[590,737,853,879]" captionStart="Fig" captionStartId="14.[189,232,1857,1883]" captionTargetBox="[247,1341,265,1811]" captionTargetId="figure-15@14.[247,1341,265,1811]" captionTargetPageId="14" captionText="Fig. 12. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. A. Dealated macropterous male mating with apterous female, lateral view. B. Dealated macropterous male mating with dealated macropterous female, dorsal view. C. Dealated macropterous male mating with apterous female, dorsal view. D. Apterous male mating with apterous female on the water surface, dorsal view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385662" httpUri="https://zenodo.org/record/8385662/files/figure.png" pageId="9" pageNumber="10">Figs 12AC</figureCitation>
,
<figureCitation id="14CC2A1EFFD529323AF2FCE7735F0C75" box="[749,780,853,879]" captionStart="Fig" captionStartId="15.[189,232,1928,1954]" captionTargetBox="[255,1332,265,1881]" captionTargetId="figure-11@15.[255,1332,265,1881]" captionTargetPageId="15" captionText="Fig. 13. Rhyacobates chinensis Hungerford &amp; Matsuda, 1959, in copulation. Apterous male mating and seizing the dealated macropterous female using its fore leg on a riverside rock, while the female is feeding on its prey." figureDoi="http://doi.org/10.5281/zenodo.8385664" httpUri="https://zenodo.org/record/8385664/files/figure.png" pageId="9" pageNumber="10">13</figureCitation>
,
<figureCitation id="14CC2A1EFFD529323B05FCE773220C75" box="[794,881,853,879]" captionStart="Fig" captionStartId="16.[189,232,1928,1954]" captionTargetBox="[233,1355,265,1881]" captionTargetId="figure-15@16.[233,1355,265,1881]" captionTargetPageId="16" captionText="Fig. 14. Rhyacobates recurvus Andersen &amp; Chen, 1995, live habitus in situ. A. Nnymphs and adult resting on a rock over a vortex, dorsal view. BC. Apterous male mating with apterous female on a riverside rock, dorsal view. D. A nymph resting on a riverside rock, lateral view. Images not to scale." figureDoi="http://doi.org/10.5281/zenodo.8385666" httpUri="https://zenodo.org/record/8385666/files/figure.png" pageId="9" pageNumber="10">14BC</figureCitation>
). The adults of
<taxonomicName id="4BF74D18FFD529323C39FCE774EF0C75" authorityName="Esaki" authorityYear="1923" box="[1062,1212,853,879]" class="Insecta" family="Gerridae" genus="Rhyacobates" kingdom="Animalia" order="Hemiptera" pageId="9" pageNumber="10" phylum="Arthropoda" rank="genus">
<emphasis id="BE83EA89FFD529323C39FCE774EF0C75" box="[1062,1212,853,879]" italics="true" pageId="9" pageNumber="10">Rhyacobates</emphasis>
</taxonomicName>
are often found as copulating pairs; they tend to remain connected even after being captured, seemingly too unwilling to separate from their mates. Abdominal segment VII of the female is elongate and highly modified, and the genital segments are often withdrawn into it (
<figureCitation id="14CC2A1EFFD729303AEEFE9E73680E5C" box="[753,827,300,326]" captionStart="Fig" captionStartId="5.[189,232,1716,1742]" captionTargetBox="[256,1333,273,1657]" captionTargetId="figure-12@5.[228,1359,265,1677]" captionTargetPageId="5" captionText="Fig. 3. Photographs of bodies of Rhyacobates spp. in dorsal view (females), apterous form if not stated otherwise.A. R. bui sp. nov. B. R. elongatus sp. nov. C. R. turgidus sp. nov., apterous form.D. R. turgidus sp. nov., dealated macropterous form.E. R. abdominalis Andersen &amp; Chen, 1995. F. R. anderseni Tran &amp; Yang, 2006. G. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). H. R. chinensis Hungerford &amp; Matsuda, 1959. I. R. constrictus Tran &amp; Nguyen, 2016. J. R. edentatus Andersen &amp; Chen, 1995. K. R. gongvo Tran &amp; Yang, 2006. L. R. lundbladi (Hungerford, 1957). M. R. malaisei Andersen &amp; Chen, 1995. N. R. recurvus Andersen &amp; Chen, 1995. O. R. scorpio Andersen &amp; Chen, 1995. P. R. svenhedini (Lundblad, 1934). Q. R. takahashii Esaki, 1923, dealated macropterous form. R. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385638" httpUri="https://zenodo.org/record/8385638/files/figure.png" pageId="11" pageNumber="12">Figs 3</figureCitation>
,
<figureCitation id="14CC2A1EFFD729303B58FE9E730B0E5D" box="[839,856,300,327]" captionStart="Fig" captionStartId="7.[189,232,1743,1769]" captionTargetBox="[200,1390,762,1681]" captionTargetId="figure-118@7.[189,1398,726,1709]" captionTargetPageId="7" captionText="Fig. 5. Photographs of bodies of Rhyacobates spp. in lateral view (apterous females). A. R. bui sp. nov. B. R. elongatus sp. nov.C. R. turgidus sp. nov.D. R. abdominalisAndersen &amp; Chen, 1995.E. R. anderseni Tran &amp; Yang, 2006. F. R. angustus Tran &amp; Nguyen, 2016, holotype (ZVNU). G. R. chinensis Hungerford &amp; Matsuda, 1959. H. R. constrictus Tran &amp; Nguyen, 2016. I. R. edentatus Andersen &amp; Chen, 1995. J. R. gongvo Tran &amp; Yang, 2006. K. R. lundbladi (Hungerford, 1957). L. R. malaisei Andersen &amp; Chen, 1995. M. R. recurvus Andersen &amp; Chen, 1995. N. R. scorpio Andersen &amp; Chen, 1995. O. R. svenhedini (Lundblad, 1934). P. R. takahashii Esaki, 1923. Q. R. zetteli Tran &amp; Nguyen, 2016. Scale bar = 3 mm." figureDoi="http://doi.org/10.5281/zenodo.8385644" httpUri="https://zenodo.org/record/8385644/files/figure.png" pageId="11" pageNumber="12">5</figureCitation>
). We hypothesize that this structure might help females reject an unwanted mating.
</paragraph>
</subSubSection>
</treatment>
</document>