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<document id="47D7E665AE19A087BBE54762FF7095B1" ID-CLB-Dataset="4890" ID-DOI="10.11646/zootaxa.3884.2.2" ID-GBIF-Dataset="15d9cc85-4968-4b16-9752-55c63984dc4d" ID-ISSN="1175-5326" ID-Zenodo-Dep="4951550" ID-ZooBank="62CBBF94-20A9-4813-9B0F-C3507E143F77" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1623727098464" checkinUser="felipe" docAuthor="Shavrin, Alexey V." docDate="2014" docId="03BB87CA411EE207FF1C7968F0F0FEDF" docLanguage="en" docName="zt03884p140.pdf" docOrigin="Zootaxa 3884 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Lesteva Shavrin 2014" docType="key" docVersion="7" lastPageNumber="139" masterDocId="FF82FFB2410EE216FF8B7D28F56CFFA5" masterDocTitle="Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) of Iran and adjacent territories" masterLastPageNumber="140" masterPageNumber="122" pageNumber="138" updateTime="1699016201036" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="A53E05C2075DEB1D06D266BA0CBDA4C0">Review of the genus Lesteva Latreille, 1797 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) of Iran and adjacent territories</mods:title>
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<mods:namePart id="4D8CEC81FE28358277ECB3BBD02C638A">Shavrin, Alexey V.</mods:namePart>
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<mods:date id="A589F548E4E9B023F99219EB49A2D18B">2014</mods:date>
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<treatment id="03BB87CA411EE207FF1C7968F0F0FEDF" ID-DOI="http://doi.org/10.5281/zenodo.4952518" ID-GBIF-Taxon="184849561" ID-Zenodo-Dep="4952518" LSID="urn:lsid:plazi:treatment:03BB87CA411EE207FF1C7968F0F0FEDF" httpUri="http://treatment.plazi.org/id/03BB87CA411EE207FF1C7968F0F0FEDF" lastPageId="17" lastPageNumber="139" pageId="16" pageNumber="138">
<subSubSection id="C3086557411EE206FF1C7968F74BFBFF" box="[151,551,1087,1114]" pageId="16" pageNumber="138" type="nomenclature">
<paragraph id="8BAD36DC411EE206FF1C7968F74BFBFF" blockId="16.[151,551,1087,1114]" box="[151,551,1087,1114]" pageId="16" pageNumber="138">
<heading id="D0E581B0411EE206FF1C7968F74BFBFF" bold="true" box="[151,551,1087,1114]" fontSize="11" level="1" pageId="16" pageNumber="138" reason="1">
<emphasis id="B966EACE411EE206FF1C7968F74BFBFF" bold="true" box="[151,551,1087,1114]" pageId="16" pageNumber="138">
Key to species of
<taxonomicName id="4C124D5F411EE206FEE67917F4A4FBFC" ID-CoL="8VRSJ" authorityName="Shavrin" authorityYear="2014" box="[365,456,1087,1113]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="genus">
<emphasis id="B966EACE411EE206FEE67917F4A4FBFC" bold="true" box="[365,456,1087,1113]" italics="true" pageId="16" pageNumber="138">Lesteva</emphasis>
</taxonomicName>
of
<collectingCountry id="F305764C411EE206FE7B7968F74BFBFF" box="[496,551,1088,1114]" name="Iran" pageId="16" pageNumber="138">Iran</collectingCountry>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C3086557411EE207FF1C79AEF0F0FEDF" lastPageId="17" lastPageNumber="139" pageId="16" pageNumber="138" type="key">
<keyStep id="30E62E79411EE206FF1C79AEF0F0FA8B" pageId="16" pageNumber="138">
<paragraph id="8BAD36DC411EE206FF1C79AEF0F7FB56" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C79AEF0F7FB56" pageId="16" pageNumber="138">
1 Pronotum bordered along antero-lateral sides. Antennomeres wide, antennae moderately short (
<figureCitation id="13292A59411EE206FB0C79AEF1A3FB3E" box="[1159,1231,1158,1179]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 11</figureCitation>
). Punctation of forebody very coarse. Abdomen glossy. Posterior tarsi distinctly less than half of tibiae. Body length: 3.2−4.0 mm. Aedeagus short, with median lobe strongly converging toward apex, with wide parameres significantly longer than apex of aedeagus (
<bibRefCitation id="EF834B2D411EE206FADB79E9F42FFB56" author="Zanetti, A." pageId="16" pageNumber="138" pagination="1 - 472" refId="ref15163" refString="Zanetti, A. (1987) Coleoptera. Staphylinidae. Omaliinae. Fauna d'Italia, 25, i - xii + 1 - 472. [Calderini, Bologna]" type="journal article" year="1987">Zanetti, 1987: 341</bibRefCitation>
, Fig. e, f).........................................................
<taxonomicName id="4C124D5F411EE206FBD179F6F0F7FB56" ID-CoL="722SZ" authority="Erichson" authorityName="Erichson. Scale" authorityYear="1839" box="[1114,1435,1245,1267]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="species" species="punctata" subGenus="Lestevidia">
<emphasis id="B966EACE411EE206FBD179F6F10BFB56" box="[1114,1127,1246,1267]" italics="true" pageId="16" pageNumber="138">L</emphasis>
. (
<emphasis id="B966EACE411EE206FBF779F5F1B2FB56" box="[1148,1246,1245,1267]" italics="true" pageId="16" pageNumber="138">Lestevidia</emphasis>
)
<emphasis id="B966EACE411EE206FB6779F6F02CFB56" box="[1260,1344,1246,1267]" italics="true" pageId="16" pageNumber="138">punctata</emphasis>
Erichson
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAD36DC411EE206FF1C79D4F0F0FA8B" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C79D4F0F0FA8B" pageId="16" pageNumber="138">- Pronotum not bordered at sides. Antennomeres longer. Punctation of forebody finer. Abdomen dull. Posterior tarsi half long as tibiae. Aedeagus longer and narrower, with wider median lobe and narrower parameres............................ 2</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E62E79411EE206FF1C781EF0F7FA64" pageId="16" pageNumber="138">
<paragraph id="8BAD36DC411EE206FF1C781EF0F0FA23" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C781EF0F0FA23" pageId="16" pageNumber="138">
2 Punctation of forebody coarser. Antenna as in
<figureCitation id="13292A59411EE206FD23781EF78FFAEE" box="[680,739,1334,1355]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 7</figureCitation>
. Tarsomere V of metatarsi longer than tarsomere I. Elytra with straight hind margins. Head and pronotum dark brown to black; elytra dark brown to black, frequently with red brown lateral and hind margins. Body length:
<quantity id="4CEA9B39411EE206FE377859F75FFA23" box="[444,563,1393,1414]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="4.7" metricValueMin="3.3" pageId="16" pageNumber="138" unit="mm" value="4.0" valueMax="4.7" valueMin="3.3">3.3−4.7 mm</quantity>
. Aedeagus with median keel (
<figureCitation id="13292A59411EE206FCD87859F6D2FA23" box="[851,958,1393,1414]" captionStart="FIGURES 33–41" captionStartId="10.[151,264,1937,1959]" captionTargetBox="[238,1334,189,1900]" captionTargetId="figure-64@10.[216,1367,180,1919]" captionTargetPageId="10" captionText="FIGURES 33–41. Aedeagus of Lesteva longoelytrata longoelytrata (Goeze) (33–35: Kandavan (Iran), 36–38: Azish-Tau (Russia), 39–41: Yeniköy, paratype of L. soror Smetana (Turkey)): 33, 36, 39—parameral view, 34, 37, 40—lateral view, 35, 38, 41—parameral surface of median lobe of aedeagus. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4951582" httpUri="https://zenodo.org/record/4951582/files/figure.png" pageId="16" pageNumber="138">Fig. 33−41</figureCitation>
)..
<taxonomicName id="4C124D5F411EE206FC657859F1AEFA23" box="[1006,1218,1392,1414]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="species" species="longoelytata">
<emphasis id="B966EACE411EE206FC657859F697FA23" box="[1006,1019,1393,1414]" italics="true" pageId="16" pageNumber="138">L</emphasis>
. (s.str.)
<emphasis id="B966EACE411EE206FBC17858F1AEFA23" box="[1098,1218,1392,1414]" italics="true" pageId="16" pageNumber="138">longoelytata</emphasis>
</taxonomicName>
<taxonomicName id="4C124D5F411EE206FB4C7858F0F0FA23" ID-CoL="6PR3D" authority="(Goeze)" baseAuthorityName="Goeze" baseAuthorityYear="1777" box="[1223,1436,1392,1414]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="species" species="longoelytrata">
<emphasis id="B966EACE411EE206FB4C7858F024FA23" box="[1223,1352,1392,1414]" italics="true" pageId="16" pageNumber="138">longoelytrata</emphasis>
(Goeze)
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAD36DC411EE206FF1C78A6F0F7FA64" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C78A6F0F7FA64" pageId="16" pageNumber="138">- Punctation of forebody significantly finer. Tarsomere V of metatarsi shorter or as long as tarsomere I. Elytra with widely rounded hind margins. Aedeagus without median keel........................................................ 3</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E62E79411EE206FF1C78E1F0F7F9F6" pageId="16" pageNumber="138">
<paragraph id="8BAD36DC411EE206FF1C78E1F0FAF9BC" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C78E1F0FAF9BC" pageId="16" pageNumber="138">
3 Antennomeres significantly longer (
<figureCitation id="13292A59411EE206FDB778E1F71BFA7B" box="[572,631,1481,1502]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 4</figureCitation>
). Forebody and abdomen brown; antennae, mouthparts, ocelli, legs and apical margins of abdominal tergites VII−VIII yellow brown; tarsi and antennomere XI yellow. Body larger:
<quantity id="4CEA9B39411EE206FB2F78CDF192FA5E" box="[1188,1278,1509,1531]" metricMagnitude="-3" metricUnit="m" metricValue="5.75" pageId="16" pageNumber="138" unit="mm" value="5.75">5.75 mm</quantity>
. Aedeagus as in
<figureCitation id="13292A59411EE206FF6A7B2CF1DDF9BC" box="[225,1201,1540,1561]" captionStart="FIGURES 1–3" captionStartId="2.[151,264,1273,1295]" captionTargetBox="[217,1372,185,1232]" captionTargetId="figure-324@2.[196,1396,174,1257]" captionTargetPageId="2" captionText="FIGURES 1–3. Lesteva antennata sp.n.: 1—aedeagus, parameral view, 2—aedeagus, lateral view, 3—aedeagus, parameral surface of median lobe. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4951554" httpUri="https://zenodo.org/record/4951554/files/figure.png" pageId="16" pageNumber="138">
Figs. 1–3........................................................................
<taxonomicName id="4C124D5F411EE206FB2F7B2CF1DDF9BC" ID-CoL="6Q33J" authority="Shavrin, 2014" authorityName="Shavrin" authorityYear="2014" box="[1188,1201,1540,1561]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="species" species="antennata" status="sp. nov.">
<emphasis id="B966EACE411EE206FB2F7B2CF1DDF9BC" box="[1188,1201,1540,1561]" italics="true" pageId="16" pageNumber="138">L</emphasis>
</taxonomicName>
</figureCitation>
. (s.str.)
<emphasis id="B966EACE411EE206FA8B7B2CF00CF9BC" box="[1280,1376,1540,1561]" italics="true" pageId="16" pageNumber="138">antennata</emphasis>
<emphasis id="B966EACE411EE206FAED7B2BF0FAF9BC" bold="true" box="[1382,1430,1539,1561]" pageId="16" pageNumber="138">
<taxonomicNameLabel id="A25557B5411EE206FAED7B2BF0FAF9BC" box="[1382,1430,1539,1561]" pageId="16" pageNumber="138" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</keyLead>
</paragraph>
<paragraph id="8BAD36DC411EE206FF1C7B09F0F7F9F6" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C7B09F0F7F9F6" pageId="16" pageNumber="138">- Antennomeres shorter. Forebody darker (brown to black), sometimes with yellow to red brown maculae on each elytron. Body smaller............................................................................................. 4</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E62E79411EE206FF1C7B74F0F7F96C" pageId="16" pageNumber="138">
<paragraph id="8BAD36DC411EE206FF1C7B74F0F0F92B" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C7B74F0F0F92B" pageId="16" pageNumber="138">
4 Punctation of head and abdomen denser and smaller. Dark brown, legs red brown. Antenna as in
<figureCitation id="13292A59411EE206FB1D7B74F1BDF9D4" box="[1174,1233,1628,1649]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 9</figureCitation>
. Aedeagus short and wide (
<bibRefCitation id="EF834B2D411EE206FE977B51F4A1F92B" author="Jarrige, J." box="[284,461,1657,1678]" pageId="16" pageNumber="138" pagination="483 - 502" refId="ref14240" refString="Jarrige, J. (1971) Contribution a la faune de L'Iran. 21. Coleopteres Brachelytra. Annales de la Societe Entomologique de France, 7 (2), 483 - 502. http: // dx. doi. org / 10.5962 / bhl. title. 8757" type="journal article" year="1971">Jarrige, 1971: 487</bibRefCitation>
,
<figureCitation id="13292A59411EE206FE5C7B51F77CF92B" box="[471,528,1657,1678]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 6</figureCitation>
)......................................................
<taxonomicName id="4C124D5F411EE206FB2F7B51F0F0F92B" ID-CoL="7233D" authority="Jarrige" authorityName="Jarrige" authorityYear="1971" box="[1188,1436,1657,1678]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="species" species="morvani">
<emphasis id="B966EACE411EE206FB2F7B51F1D9F92B" box="[1188,1205,1657,1678]" italics="true" pageId="16" pageNumber="138">L</emphasis>
. (s.str.)
<emphasis id="B966EACE411EE206FA8F7B51F038F92B" box="[1284,1364,1657,1678]" italics="true" pageId="16" pageNumber="138">morvani</emphasis>
Jarrige
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BAD36DC411EE206FF1C7BBEF0F7F96C" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C7BBEF0F7F96C" pageId="16" pageNumber="138">- Punctation of head and abdomen sparser, punctures moderately larger. Brown to black, with or without yellow-brown maculae on each elytron. Aedeagus longer and narrower............................................................. 5</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E62E79411EE206FF1C7BF9F0F7F8A6" pageId="16" pageNumber="138">
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<keyLead id="30E395E9411EE206FF1C7BF9F0F7F943" box="[151,1435,1745,1766]" pageId="16" pageNumber="138">5 Tarsomere I of metatarsus as long as the combined length of tarsomeres II−III..................................... 6</keyLead>
</paragraph>
<paragraph id="8BAD36DC411EE206FF1C7BC6F0F7F8A6" blockId="16.[151,1437,1158,2030]" box="[151,1435,1774,1795]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C7BC6F0F7F8A6" box="[151,1435,1774,1795]" pageId="16" pageNumber="138">- Tarsomere I of metatarsus shorter than the combined length of tarsomeres II−III................................... 7</keyLead>
</paragraph>
</keyStep>
<keyStep id="30E62E79411EE206FF1C7A24F0F0F874" pageId="16" pageNumber="138">
<paragraph id="8BAD36DC411EE206FF1C7A24F0F0F8FE" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C7A24F0F0F8FE" pageId="16" pageNumber="138">
6 Apex of aedeagus evenly narrowed in apical third; parameres significantly longer than apex of aedeagus (
<figureCitation id="13292A59411EE206FB6B7A24F03AF884" box="[1248,1366,1804,1825]" captionStart="FIGURES 27–29" captionStartId="6.[151,264,1972,1994]" captionTargetBox="[189,1394,1000,1934]" captionTargetId="figure-386@6.[177,1414,985,1955]" captionTargetPageId="6" captionText="FIGURES 27–29. Aedeagus of Lesteva binotata Reitter (Kandavan, Iran): 27—parameral view, 28—lateral view, 29—parameral surface of median lobe of aedeagus. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951574" httpUri="https://zenodo.org/record/4951574/files/figure.png" pageId="16" pageNumber="138">Figs. 27–29</figureCitation>
). Forebody brown to black; each elytron often with distinct yellow brown to red brown elongate macula in anterior third (
<figureCitation id="13292A59411EE206FAFE7A01F44CF8FE" captionStart="FIGURES 12–20" captionStartId="4.[151,264,1936,1958]" captionTargetBox="[379,1272,205,1902]" captionTargetId="figure-54@4.[300,1292,195,1916]" captionTargetPageId="4" captionText="FIGURES 12–20. Elytron of Lesteva binotata Reitter (12—Kandavan (Iran), 13—Tbilisi (Georgia), 14—Qohrud (Iran), 15—Kandavan (Iran), 16—Gukan (Iran), 7—Semnan (Iran), 18—Sertavul-Pass (Turkey), 19—Elmadi (Turkey)) and L. schawalleri sp.n. (Fig. 20). Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951568" httpUri="https://zenodo.org/record/4951568/files/figure.png" pageId="16" pageNumber="138">Fig. 12–19</figureCitation>
); mouthparts and tarsi yellow. Antenna as in
<figureCitation id="13292A59411EE206FD5B7A6EF666F8FE" box="[720,778,1862,1883]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 5</figureCitation>
. Body length:
<quantity id="4CEA9B39411EE206FC1C7A6EF161F8FE" box="[919,1037,1862,1883]" metricMagnitude="-3" metricUnit="m" metricValue="4.6" metricValueMax="5.5" metricValueMin="3.7" pageId="16" pageNumber="138" unit="mm" value="4.6" valueMax="5.5" valueMin="3.7">3.7–5.5 mm</quantity>
.............
<taxonomicName id="4C124D5F411EE206FB2E7A6EF0F0F8FE" ID-CoL="6PR3G" authority="Reitter" authorityName="Reitter" authorityYear="1901" box="[1189,1436,1861,1883]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="species" species="binotata">
<emphasis id="B966EACE411EE206FB2E7A6EF1DBF8FE" box="[1189,1207,1862,1883]" italics="true" pageId="16" pageNumber="138">L</emphasis>
. (s.str.)
<emphasis id="B966EACE411EE206FA8F7A6DF038F8FE" box="[1284,1364,1861,1883]" italics="true" pageId="16" pageNumber="138">binotata</emphasis>
Reitter
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</paragraph>
<paragraph id="8BAD36DC411EE206FF1C7A4CF0F0F874" blockId="16.[151,1437,1158,2030]" pageId="16" pageNumber="138">
<keyLead id="30E395E9411EE206FF1C7A4CF0F0F874" pageId="16" pageNumber="138">
- Apex of aedeagus nipple-shaped in parameral view; parameres slightly longer than apex of aedeagus (
<figureCitation id="13292A59411EE206FB4A7A4CF042F8DC" box="[1217,1326,1892,1913]" captionStart="FIGURES 42–44" captionStartId="12.[151,264,1969,1991]" captionTargetBox="[171,1402,887,1913]" captionTargetId="figure-329@12.[152,1437,877,1934]" captionTargetPageId="12" captionText="FIGURES 42–44. Aedeagus of Lesteva maculipennis Luze (Yeniköy, Turkey): 42—parameral view, 43—lateral view, 44—parameral surface of median lobe of aedeagus. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4951586" httpUri="https://zenodo.org/record/4951586/files/figure.png" pageId="16" pageNumber="138">Fig. 42–44</figureCitation>
). Forebody and abdomen brown to black; each elytron without or with distinct to indistinct yellow to red-brown macula in anterior third (Fig. 21–26); ocelli, antennae and legs brown; mouthparts, sometimes base of antennomeres I−III and tarsi yellow. Antenna as in
<figureCitation id="13292A59411EE206FF727A94F45FF874" box="[249,307,1980,2001]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 8</figureCitation>
. Body length: 4.0–
<quantity id="4CEA9B39411EE206FE607A94F72FF874" box="[491,579,1980,2001]" metricMagnitude="-3" metricUnit="m" metricValue="4.95" pageId="16" pageNumber="138" unit="mm" value="4.95">4.95 mm</quantity>
................................................
<taxonomicName id="4C124D5F411EE206FB057A94F0F0F874" ID-CoL="6Q33D" authority="Luze" authorityName="Luze" authorityYear="1903" box="[1166,1436,1979,2001]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="16" pageNumber="138" phylum="Arthropoda" rank="species" species="maculipennis">
<emphasis id="B966EACE411EE206FB057A94F1F7F874" box="[1166,1179,1980,2001]" italics="true" pageId="16" pageNumber="138">L</emphasis>
. (s.str)
<emphasis id="B966EACE411EE206FB6D7A93F00AF874" box="[1254,1382,1979,2001]" italics="true" pageId="16" pageNumber="138">maculipennis</emphasis>
Luze
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7 Antennomeres IV−X significantly shorter (
<figureCitation id="13292A59411EE206FD027AF1F7AAF84B" box="[649,710,2009,2030]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="16" pageNumber="138">Fig. 6</figureCitation>
). Apex of aedeagus wider, parameres longer than apex of aedeagus (
<figureCitation id="13292A59411EE207FAFE7AF1F44EFF09" captionStart="FIGURES 30–32" captionStartId="8.[151,264,1662,1684]" captionTargetBox="[164,1410,519,1628]" captionTargetId="figure-209@8.[152,1437,515,1648]" captionTargetPageId="8" captionText="FIGURES 30–32. Aedeagus of Lesteva frischi sp.n.: 30—parameral view, 31—lateral view, 32—parameral surface of median lobe of aedeagus. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4951578" httpUri="https://zenodo.org/record/4951578/files/figure.png" lastPageId="17" lastPageNumber="139" pageId="16" pageNumber="138">Fig. 30−32</figureCitation>
). Head with moderately flat frons and vertex, 1.6−1.8 times as wide as long. Eyes more than two times as long as temples. Forebody, abdomen, femora and antennomeres II and III black; tibiae, antennomeres III–XI, ocelli, mouthparts and apical margin of tergite VIII brown, base of antennomeres I–III and tarsi yellowish brown. Body length:
<quantity id="4CEA9B39411FE207FB067DFAF06FFF42" box="[1165,1283,210,231]" metricMagnitude="-3" metricUnit="m" metricValue="4.199999999999999" metricValueMax="4.3" metricValueMin="4.1" pageId="17" pageNumber="139" unit="mm" value="4.199999999999999" valueMax="4.3" valueMin="4.1">4.1–4.3 mm</quantity>
.................................................................................................
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<emphasis id="B966EACE411FE207FB427DD8F1B6FEA0" box="[1225,1242,240,261]" italics="true" pageId="17" pageNumber="139">L</emphasis>
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<emphasis id="B966EACE411FE207FAA37DC7F009FEA0" box="[1320,1381,239,261]" italics="true" pageId="17" pageNumber="139">frischi</emphasis>
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<paragraph id="8BAD36DC411FE207FF1C7C25F0F0FEDF" blockId="17.[151,1436,151,378]" pageId="17" pageNumber="139">
<keyLead id="30E395E9411FE207FF1C7C25F0F0FEDF" pageId="17" pageNumber="139">
- Antennomeres IV−X elongated (
<figureCitation id="13292A59411FE207FDA97C25F706FE87" box="[546,618,269,290]" captionStart="FIGURES 4–11" captionStartId="3.[151,264,1687,1709]" captionTargetBox="[195,1356,578,1614]" captionTargetId="figure-211@3.[184,1394,565,1633]" captionTargetPageId="3" captionText="FIGURES 4–11. Antennae of Lesteva: 4—L. antennata sp.n., 5—L. binotata Reitter, 6—L. frischi sp.n., 7—L. longoelytrata longoelytrata (Goeze), 8—L. maculipennis Luze, 9—L. morvani Jarrige, 10—L. schawalleri sp.n., 11—L. punctata Erichson. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4951562" httpUri="https://zenodo.org/record/4951562/files/figure.png" pageId="17" pageNumber="139">Fig. 10</figureCitation>
). Apex of aedeagus narrower, parameres shorter (
<figureCitation id="13292A59411FE207FBCC7C25F1DAFE87" box="[1095,1206,269,290]" captionStart="FIGURES 45–47" captionStartId="13.[151,264,1398,1420]" captionTargetBox="[166,1410,214,1340]" captionTargetId="figure-293@13.[149,1434,201,1373]" captionTargetPageId="13" captionText="FIGURES 45–47. Aedeagus of Lesteva schawalleri sp.n.: 45—parameral view, 46—lateral view, 47—parameral surface of median lobe of aedeagus. Scale bar: 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4951592" httpUri="https://zenodo.org/record/4951592/files/figure.png" pageId="17" pageNumber="139">Fig. 45−47</figureCitation>
). Head with moderately convex frons and vertex, 1.1 times as wide as long. Eyes approximately 3 times as long as temples. Head, pronotum and antennae black, elytra dark brown with indistinct yellow-brown macula in anterior third of each elytron; abdomen and legs brown; ocelli, mouthparts and base of antennomeres II−III red-brown. Body length:
<quantity id="4CEA9B39411FE207FC4B7C4DF159FEDF" box="[960,1077,357,378]" metricMagnitude="-3" metricUnit="m" metricValue="4.4" metricValueMax="4.7" metricValueMin="4.1" pageId="17" pageNumber="139" unit="mm" value="4.4" valueMax="4.7" valueMin="4.1">4.1–4.7 mm</quantity>
..........
<emphasis id="B966EACE411FE207FB387C4DF1ACFEDF" box="[1203,1216,357,378]" italics="true" pageId="17" pageNumber="139">L</emphasis>
.
<taxonomicName id="4C124D5F411FE207FB477C4DF00AFEDF" ID-CoL="6PR2Z" authority="Shavrin, 2014" authorityName="Shavrin" authorityYear="2014" box="[1228,1382,356,378]" class="Insecta" family="Staphylinidae" genus="Lesteva" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="139" phylum="Arthropoda" rank="species" species="schawalleri" status="sp. nov." subGenus="Lestevidia">
(
<emphasis id="B966EACE411FE207FB5F7C4DF18DFEDF" box="[1236,1249,357,378]" italics="true" pageId="17" pageNumber="139">L</emphasis>
.)
<emphasis id="B966EACE411FE207FB7D7C4CF00AFEDF" box="[1270,1382,356,378]" italics="true" pageId="17" pageNumber="139">schawalleri</emphasis>
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