455 lines
56 KiB
XML
455 lines
56 KiB
XML
<document id="4CD9AC3CF503C88C329B9913D1A318DC" ID-DOI="10.5281/zenodo.277567" ID-GBIF-Dataset="8b3c1d83-f60c-4704-9a6d-ea9b7af4d28d" ID-ISSN="1175-5326" ID-Zenodo-Dep="277567" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460315008605" checkinUser="plazi" docAuthor="Korsós, Zoltán, Nakamura, Yasuyuki & Tanabe, Tsutomu" docDate="2011" docId="03BB8784C926783FFF77C5F6D2A5FA19" docLanguage="en" docName="zt02877p068.pdf" docOrigin="Zootaxa 2877" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Riukiaria maculata Korsós, Nakamura & Tanabe, 2011, sp. n." docType="treatment" docVersion="11" lastPageNumber="62" masterDocId="FF82FFFCC9257838FFE0C52BD366FFFE" masterDocTitle="Two new millipede species of the genus Riukiaria (Diplopoda, Polydesmida, Xystodesmidae) endemic to the Ryukyu Archipelago, Japan" masterLastPageNumber="68" masterPageNumber="55" pageNumber="58" updateTime="1698254750703" updateUser="plazi">
|
||
<mods:mods id="4BB5E420412BCC2957E8D64D57E05DCE" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="4D3DF86079B1CECF8A8A268A7A9DFEB5">
|
||
<mods:title id="824F21216A619EFC4E8BCB0F1EAD8C28">Two new millipede species of the genus Riukiaria (Diplopoda, Polydesmida, Xystodesmidae) endemic to the Ryukyu Archipelago, Japan</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="CDC1059D1DEFF20A1E0C958F92C24F87" type="personal">
|
||
<mods:role id="066D3E59F0FC93AB75B612E0680C7030">
|
||
<mods:roleTerm id="1AA02ECECE5806F9F729B5617DC51711">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="037F2A1E75A434179FAE59CDC965B77A">Korsós, Zoltán</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="F045F69CE8BC7773DEA265611491C289" type="personal">
|
||
<mods:role id="7263AC447F9F2A158C50E893410AB0A2">
|
||
<mods:roleTerm id="BCF59A6BADAD849A5F38A5CAA2317AC6">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="3B0F5944367790B782FF97BCA2D953B1">Nakamura, Yasuyuki</mods:namePart>
|
||
</mods:name>
|
||
<mods:name id="D9C86BFF6B7DB772617F977A3D7BAF1F" type="personal">
|
||
<mods:role id="641CE0A40F8E5C6D83AB0E46C41E6BA4">
|
||
<mods:roleTerm id="37FA2AECA7B0E0DBAD4F33B7146541E1">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="6D1D0FB054B25B17FBDD34CF9BA87219">Tanabe, Tsutomu</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource id="3F8AD6CACBE549E9DDBFC4FD7E10076B">text</mods:typeOfResource>
|
||
<mods:relatedItem id="30C92166A8978CEE2CEB063FCB09EE21" type="host">
|
||
<mods:titleInfo id="DC35B32A1B35F5AD94BB69CAE88C88F4">
|
||
<mods:title id="EC0034B5EE54D75AB03026FC4616672E">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="F5BF7653F4F81551FB2E2A75F7493BF1">
|
||
<mods:date id="469E01F638C752ABC65FEB9E0D913D32">2011</mods:date>
|
||
<mods:detail id="D090DBBDAA150FA8DABA6B0267429F1A" type="volume">
|
||
<mods:number id="7B2128ADA6F4B3AE5EE8B82C3210F6D8">2877</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="13AB135669AEBD19DE08C2028DD7EE1F" unit="page">
|
||
<mods:start id="FA2BAE8F525692320475E0D2940ED6E1">55</mods:start>
|
||
<mods:end id="70EFC6870A55B8EFC4AF7D6F875F5CF4">68</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification id="26CF88C9B4268D681B74752C163BC9B0">journal article</mods:classification>
|
||
<mods:identifier id="2CB9AC00B4A2E90F97AB5E916A6C0BCB" type="DOI">10.5281/zenodo.277567</mods:identifier>
|
||
<mods:identifier id="08B9488F26908F2B1FBB9F0536F4FBEA" type="GBIF-Dataset">8b3c1d83-f60c-4704-9a6d-ea9b7af4d28d</mods:identifier>
|
||
<mods:identifier id="01CCE348B4509D112B6555E414C124AE" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="112243E866E6AF2BCE3D7DA345FCAB45" type="Zenodo-Dep">277567</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03BB8784C926783FFF77C5F6D2A5FA19" ID-DOI="http://doi.org/10.5281/zenodo.5678521" ID-GBIF-Taxon="119401250" ID-Zenodo-Dep="5678521" LSID="urn:lsid:plazi:treatment:03BB8784C926783FFF77C5F6D2A5FA19" httpUri="http://treatment.plazi.org/id/03BB8784C926783FFF77C5F6D2A5FA19" lastPageId="7" lastPageNumber="62" pageId="3" pageNumber="58">
|
||
<subSubSection id="C3086519C926783BFF77C5F6D223FEE9" pageId="3" pageNumber="58" type="nomenclature">
|
||
<paragraph id="8BAD3692C926783BFF77C5F6D2ADFF09" blockId="3.[151,459,221,279]" box="[151,459,221,247]" pageId="3" pageNumber="58">
|
||
<heading id="D0E581FEC926783BFF77C5F6D2ADFF09" bold="true" box="[151,459,221,247]" fontSize="11" level="1" pageId="3" pageNumber="58" reason="1">
|
||
<emphasis id="B966EA80C926783BFF77C5F6D2ADFF09" bold="true" box="[151,459,221,247]" pageId="3" pageNumber="58">
|
||
<taxonomicName id="4C124D11C926783BFF77C5F6D2E4FF09" ID-CoL="4T6XQ" box="[151,386,221,247]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C926783BFF77C5F6D2E4FF09" bold="true" box="[151,386,221,247]" italics="true" pageId="3" pageNumber="58">Riukiaria maculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC926783BFE69C5F6D2ADFF09" box="[393,459,221,247]" pageId="3" pageNumber="58" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C926783BFF77C5D4D223FEE9" blockId="3.[151,459,221,279]" box="[151,325,255,279]" pageId="3" pageNumber="58">
|
||
<figureCitation id="13292A17C926783BFF77C5D4D39DFEE9" box="[151,251,255,279]" captionStart="FIGURES 2 – 3" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[273,1322,204,1900]" captionTargetId="figure@4.[260,1334,193,1912]" captionTargetPageId="4" captionText="FIGURES 2 – 3. Riukiaria maculata sp. n., holotype male. 2 = Live coloration, Tane-ga-shima; 3 = Live specimen in UV-light. FIGURES 4 – 6. Riukiaria mundyi sp. n., paratype specimens. 4 = Live coloration, Yonaguni-jima; 5 – 6 = The same specimens in ambient flashlight, and in UV-light." httpUri="https://zenodo.org/record/277569/files/figure.png" pageId="3" pageNumber="58">Figs 2–3</figureCitation>
|
||
, 7–13.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086519C926783BFF77C46CD3BEFDC9" pageId="3" pageNumber="58" type="materials_examined">
|
||
<paragraph id="8BAD3692C926783BFF77C46CD388FE59" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">
|
||
<typeStatus id="54A98830C926783BFF77C46CD399FEA1" box="[151,255,327,351]" pageId="3" pageNumber="58" type="holotype">Holotype</typeStatus>
|
||
male (NSMT-My 377)—
|
||
<collectingCountry id="F3057602C926783BFDD3C46CD113FEA1" box="[563,629,327,351]" name="Japan" pageId="3" pageNumber="58">Japan</collectingCountry>
|
||
, Northern Ryukyus, Osumi Group, Tane-ga-shima Island, Nakatane Town,
|
||
<taxonomicName id="4C124D11C926783BFF04C446D215FE7A" box="[228,371,365,388]" class="Insecta" family="Noctuidae" genus="Cryptomeria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA80C926783BFF04C446D215FE7A" box="[228,371,365,388]" italics="true" pageId="3" pageNumber="58">Cryptomeria</emphasis>
|
||
</taxonomicName>
|
||
mixed forest close to the airport,
|
||
<quantity id="4CEA9B77C926783BFD0FC447D050FE7A" box="[751,822,364,388]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="58" unit="m" value="260.0">260 m</quantity>
|
||
alt.,
|
||
<geoCoordinate id="EE265055C926783BFC8FC447D08FFE7A" box="[879,1001,364,388]" direction="north" orientation="latitude" pageId="3" pageNumber="58" precision="5" value="30.6401">N30.6401°</geoCoordinate>
|
||
<geoCoordinate id="EE265055C926783BFC10C447D71FFE7A" box="[1008,1145,364,388]" direction="east" orientation="longitude" pageId="3" pageNumber="58" precision="5" value="130.9797">E130.9797°</geoCoordinate>
|
||
,
|
||
<date id="FFAC1052C926783BFB64C446D66CFE7A" box="[1156,1290,364,388]" pageId="3" pageNumber="58" value="2010-07-07">7 July 2010</date>
|
||
, leg. R. & Z. Korsós.
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C926783BFF25C49ED72CFE32" blockId="3.[151,1437,327,2008]" box="[197,1098,436,461]" pageId="3" pageNumber="58">
|
||
<typeStatus id="54A98830C926783BFF25C49ED250FE33" box="[197,310,437,461]" pageId="3" pageNumber="58" type="paratype">Paratypes</typeStatus>
|
||
:
|
||
<specimenCount id="9D14FD1BC926783BFEA0C49FD2C9FE32" box="[320,431,436,460]" pageId="3" pageNumber="58" type="female">3 females</specimenCount>
|
||
, 3 juvs (NSMT-My 378,
|
||
<collectionCode id="ED03AE57C926783BFD29C49FD046FE32" LSID="urn:lsid:biocol.org:col:34991" box="[713,800,436,460]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="3" pageNumber="58">HNHM</collectionCode>
|
||
)—Same locality and date.
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C926783BFF27C4FCD3BEFDC9" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">
|
||
<specimenCount id="9D14FD1BC926783BFF27C4FCD252FE11" box="[199,308,471,495]" pageId="3" pageNumber="58" type="female">3 females</specimenCount>
|
||
(
|
||
<collectionCode id="ED03AE57C926783BFEA5C4FCD2FEFE11" box="[325,408,471,495]" pageId="3" pageNumber="58">RUMF</collectionCode>
|
||
)—
|
||
<collectingCountry id="F3057602C926783BFE5CC4FCD298FE11" box="[444,510,471,495]" name="Japan" pageId="3" pageNumber="58">Japan</collectingCountry>
|
||
, Northern Ryukyus, Osumi Group, Tane-ga-shima Island, Nakatane Town,
|
||
<taxonomicName id="4C124D11C926783BFABBC4F3D388FDEA" class="Insecta" family="Noctuidae" genus="Cryptomeria" kingdom="Animalia" order="Lepidoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA80C926783BFABBC4F3D388FDEA" italics="true" pageId="3" pageNumber="58">Cryptomeria</emphasis>
|
||
</taxonomicName>
|
||
forest close to the airport,
|
||
<quantity id="4CEA9B77C926783BFDFAC4D7D107FDEA" box="[538,609,508,532]" metricMagnitude="2" metricUnit="m" metricValue="2.6" pageId="3" pageNumber="58" unit="m" value="260.0">260 m</quantity>
|
||
alt.,
|
||
<geoCoordinate id="EE265055C926783BFD79C4D7D075FDEA" box="[665,787,508,532]" direction="north" orientation="latitude" pageId="3" pageNumber="58" precision="5" value="30.6401">N30.6401°</geoCoordinate>
|
||
<geoCoordinate id="EE265055C926783BFCFAC4D7D0C4FDEA" box="[794,930,508,532]" direction="east" orientation="longitude" pageId="3" pageNumber="58" precision="5" value="130.9797">E130.9797°</geoCoordinate>
|
||
,
|
||
<date id="FFAC1052C926783BFC4DC4D7D70AFDEA" box="[941,1132,508,532]" pageId="3" pageNumber="58" value="2009-10-12">12 October 2009</date>
|
||
, leg. Z. Korsós & Y. Nakamura.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086519C926783BFF25C768D7E3FD12" pageId="3" pageNumber="58" type="diagnosis">
|
||
<paragraph id="8BAD3692C926783BFF25C768D7E3FD12" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">
|
||
<emphasis id="B966EA80C926783BFF25C768D259FDA2" bold="true" box="[197,319,579,604]" pageId="3" pageNumber="58">Diagnosis.</emphasis>
|
||
A species of the genus
|
||
<taxonomicName id="4C124D11C926783BFDAAC76FD1D2FDA2" box="[586,692,580,604]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="3" pageNumber="58" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA80C926783BFDAAC76FD1D2FDA2" box="[586,692,580,604]" italics="true" pageId="3" pageNumber="58">Riukiaria</emphasis>
|
||
</taxonomicName>
|
||
as defined by
|
||
<bibRefCitation id="EF834B63C926783BFCB7C76FD77CFDA2" author="Shinohara" box="[855,1050,580,604]" pageId="3" pageNumber="58" refString="Shinohara, K. (1977) Revaluation on Riukiaria (Diplopoda). Acta Arachnologica, 27 (special number), 115 - 119." type="journal article" year="1977">Shinohara (1977)</bibRefCitation>
|
||
and
|
||
<bibRefCitation id="EF834B63C926783BFBB1C76FD6FAFDA2" author="Tanabe" box="[1105,1436,580,604]" pageId="3" pageNumber="58" refString="Tanabe, T. & Shinohara, K. (1996) Revision of the millipede genus Xystodesmus, with reference to the status of the tribe Xystodesmini (Diplopoda: Xystodesmidae). Journal of Natural History, 30, 1459 - 1494." type="journal article" year="1996">Tanabe and Shinohara (1996)</bibRefCitation>
|
||
with the simple, forceps-like male gonopod conformation. It differs from all congeners by its coloration (a pair of distinct dark, brownish-black patches on each segment, including collum), by its exclusive occurrence on a single island (Tane-ga-shima), and in details of gonopod morphology: a small but definite triangular process ventrally on coxa (
|
||
<figureCitation id="13292A17C926783BFF3AC7FFD233FD12" box="[218,341,724,748]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1945,1968]" captionTargetBox="[170,1402,362,1920]" captionTargetId="figure@6.[151,1436,357,1924]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Riukiaria maculata sp. n. 10 – 12 = Left gonopod of holotype male, lateral, mesal, and apical (ventral) views, respectively; 13 = Left cyphopod of paratype female, posterior view. Scales 0.5 mm. c = coxa, tp = triangular process, v = valve, r = receptacle, op = operculum, other abbreviations as in Fig. 1." httpUri="https://zenodo.org/record/277570/files/figure.png" pageId="3" pageNumber="58">Figs 10-11</figureCitation>
|
||
,
|
||
<emphasis id="B966EA80C926783BFE81C7FED211FD12" box="[353,375,725,748]" italics="true" pageId="3" pageNumber="58">tp</emphasis>
|
||
), and by shape and length of both prefemoral process and acropodite.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086519C926783BFF27C7D3D71BFCF1" box="[199,1149,759,785]" pageId="3" pageNumber="58" type="etymology">
|
||
<paragraph id="8BAD3692C926783BFF27C7D3D71BFCF1" blockId="3.[151,1437,327,2008]" box="[199,1149,759,785]" pageId="3" pageNumber="58">
|
||
<emphasis id="B966EA80C926783BFF27C7D3D228FCEE" bold="true" box="[199,334,760,784]" pageId="3" pageNumber="58">Etymology.</emphasis>
|
||
Named after the color pattern, unusual in the genus (adjective, feminin).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086519C926783FFF27C630D2BAFEC1" lastPageId="7" lastPageNumber="62" pageId="3" pageNumber="58" type="description">
|
||
<paragraph id="8BAD3692C926783BFF27C630D074FC61" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">
|
||
<emphasis id="B966EA80C926783BFF27C630D23FFCCA" bold="true" box="[199,345,795,820]" pageId="3" pageNumber="58">Description.</emphasis>
|
||
Measurements: Medium sized species in comparison to other members of the genus. Length of
|
||
<typeStatus id="54A98830C926783BFF77C614D39FFCA9" box="[151,249,831,855]" pageId="3" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
male
|
||
<quantity id="4CEA9B77C926783BFEDFC614D2F7FCA6" box="[319,401,831,856]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="3" pageNumber="58" unit="mm" value="40.0">40 mm</quantity>
|
||
, midbody paranotal width
|
||
<quantity id="4CEA9B77C926783BFD5FC614D07FFCA6" box="[703,793,831,856]" metricMagnitude="-3" metricUnit="m" metricValue="7.5" pageId="3" pageNumber="58" unit="mm" value="7.5">7.5 mm</quantity>
|
||
, metatergal length
|
||
<quantity id="4CEA9B77C926783BFC0FC614D72EFCA6" box="[1007,1096,831,856]" metricMagnitude="-3" metricUnit="m" metricValue="1.9" pageId="3" pageNumber="58" unit="mm" value="1.9">1.9 mm</quantity>
|
||
, collum width
|
||
<quantity id="4CEA9B77C926783BFB11C614D62CFCA6" box="[1265,1354,831,856]" metricMagnitude="-3" metricUnit="m" metricValue="6.2" pageId="3" pageNumber="58" unit="mm" value="6.2">6.2 mm</quantity>
|
||
, length
|
||
<quantity id="4CEA9B77C926783BFF77C64FD394FC82" box="[151,242,868,892]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="3" pageNumber="58" unit="mm" value="2.7">2.7 mm</quantity>
|
||
. Adult female body length
|
||
<quantity id="4CEA9B77C926783BFDCEC64FD1CBFC82" box="[558,685,868,892]" metricMagnitude="-2" metricUnit="m" metricValue="4.15" metricValueMax="4.3" metricValueMin="4.0" pageId="3" pageNumber="58" unit="mm" value="41.5" valueMax="43.0" valueMin="40.0">40–43 mm</quantity>
|
||
, midbody paranotal width
|
||
<quantity id="4CEA9B77C926783BFC01C64FD70BFC82" box="[993,1133,868,892]" metricMagnitude="-3" metricUnit="m" metricValue="8.1" metricValueMax="8.5" metricValueMin="7.7" pageId="3" pageNumber="58" unit="mm" value="8.1" valueMax="8.5" valueMin="7.7">7.7–8.5 mm</quantity>
|
||
, metatergal length
|
||
<quantity id="4CEA9B77C926783BFAA8C64FD3A3FC5E" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="3" pageNumber="58" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
|
||
, collum width
|
||
<quantity id="4CEA9B77C926783BFE8CC6ACD293FC5E" box="[364,501,903,928]" metricMagnitude="-3" metricUnit="m" metricValue="6.55" metricValueMax="6.7" metricValueMin="6.4" pageId="3" pageNumber="58" unit="mm" value="6.55" valueMax="6.7" valueMin="6.4">6.4–6.7 mm</quantity>
|
||
, length
|
||
<quantity id="4CEA9B77C926783BFDADC6ACD1D9FC5E" box="[589,703,903,928]" metricMagnitude="-3" metricUnit="m" metricValue="2.95" metricValueMax="3.0" metricValueMin="2.9" pageId="3" pageNumber="58" unit="mm" value="2.95" valueMax="3.0" valueMin="2.9">2.9–3 mm</quantity>
|
||
(n= 3).
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C926783BFF27C687D0A2FB3E" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">
|
||
Color in life (
|
||
<figureCitation id="13292A17C926783BFE85C687D2CAFC3A" box="[357,428,940,964]" captionStart="FIGURES 2 – 3" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[273,1322,204,1900]" captionTargetId="figure@4.[260,1334,193,1912]" captionTargetPageId="4" captionText="FIGURES 2 – 3. Riukiaria maculata sp. n., holotype male. 2 = Live coloration, Tane-ga-shima; 3 = Live specimen in UV-light. FIGURES 4 – 6. Riukiaria mundyi sp. n., paratype specimens. 4 = Live coloration, Yonaguni-jima; 5 – 6 = The same specimens in ambient flashlight, and in UV-light." httpUri="https://zenodo.org/record/277569/files/figure.png" pageId="3" pageNumber="58">Fig. 2</figureCitation>
|
||
): Same coloration in both sexes, prozona, metazona with yellow background, metaterga laterally with a pair of distinct, oval, brownish-black patches, occupying about 1/4th of total paranotal width on both sides. Paranota yellow with slightly darker, orange margins. Head dark greyish-brown, antennae, legs, and underside pale yellow or whitish. Collum with broad yellow anterior and lateral margins, the two dark spots closer to each other than on following metaterga. Last segment with epiproct uniformly yellow. Dark patches also distinct in UV light (
|
||
<figureCitation id="13292A17C926783BFEC6C14BD20FFB86" box="[294,361,1120,1144]" captionStart="FIGURES 2 – 3" captionStartId="4.[151,264,1934,1957]" captionTargetBox="[273,1322,204,1900]" captionTargetId="figure@4.[260,1334,193,1912]" captionTargetPageId="4" captionText="FIGURES 2 – 3. Riukiaria maculata sp. n., holotype male. 2 = Live coloration, Tane-ga-shima; 3 = Live specimen in UV-light. FIGURES 4 – 6. Riukiaria mundyi sp. n., paratype specimens. 4 = Live coloration, Yonaguni-jima; 5 – 6 = The same specimens in ambient flashlight, and in UV-light." httpUri="https://zenodo.org/record/277569/files/figure.png" pageId="3" pageNumber="58">Fig. 3</figureCitation>
|
||
), their fluorescence much less intense. Preserved specimens (70% ethanol) after 10 months still have the typical pattern, although the patches are somewhat faded. Juvenile specimens (with body length of ca.
|
||
<quantity id="4CEA9B77C926783BFA60C1AFD3A2FB3E" metricMagnitude="-2" metricUnit="m" metricValue="1.5" pageId="3" pageNumber="58" unit="mm" value="15.0">15 mm</quantity>
|
||
) uniformly greyish-yellow, and do not show the characteristic spots.
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C926783BFF27C1E7D796FAAE" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">Head smooth, epicranial suture distinct, 2+2 frontal setae, 1+1 between antennal sockets, 1+1 above clypeus, and 2 dense rows at its margin and on labrum. Antennae straight, first article globose, 2nd about twice as long as first, articles 3–6 subequal in length, slightly longer than 2nd, 7th smallest, shorter than wide. Anterior part of gnathochilarial stipites and lamellae linguales densely hairy, mentum with a distinct, median hair-field.</paragraph>
|
||
<paragraph id="8BAD3692C926783BFF27C077D12AF946" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">Collum convex, smooth, shiny, lateral and posterior margins with very weak ridge, lateral corners triangular, pointed caudad. Pro- and metaterga (Fig. 7) smooth without traces of tubercles or punctuation, but from 5th onwards with definite posterior marginal ridge. Posteriolateral edge of paranota 2–3 triangular, of 4th and onwards pointed caudad. Pores on paranota 5,7,9,10,12,13,15,16,17, and 18, in median depression in lateral view. Body sides between segments 5-16 parallel, segments 17–19 strongly tapering, posteriolateral projections becoming more pointed. Epiproct in dorsal view triangular, its straight shape only broken by 3+3 lateral and 1+1 apical knobs (Fig. 8), in lateral view protruding over paraprocts (anal valves), parallel-sided, straight. Setation: 1+1 anterio-laterally on knobs, 4+4 medially on lateral sides, 2 of them on knobs, and 2+2 apically, not on knobs. Paraprocts marginate with 2+2 setae, one pair marginally, the other pair posteriorly on sides of paraprocts, hypoproct wide subtrapezoid, with 1+1 setae on knobs.</paragraph>
|
||
<paragraph id="8BAD3692C926783BFF27C3EFD6F0F892" blockId="3.[151,1437,327,2008]" pageId="3" pageNumber="58">
|
||
Midbody legs well separated, in male closer to each other (
|
||
<quantity id="4CEA9B77C926783BFC9CC3EFD0BCF922" box="[892,986,1732,1756]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="58" unit="mm" value="1.8">1.8 mm</quantity>
|
||
), in females separated by
|
||
<quantity id="4CEA9B77C926783BFAECC3EFD6FEF922" box="[1292,1432,1732,1756]" metricMagnitude="-3" metricUnit="m" metricValue="2.45" metricValueMax="2.5" metricValueMin="2.4" pageId="3" pageNumber="58" unit="mm" value="2.45" valueMax="2.5" valueMin="2.4">2.4–2.5 mm</quantity>
|
||
, sterna smooth. Postgonopodal legs (in both sexes) with well-developed ventral spine on prefemur, increasingly stronger towards body end, coxa and prefemur subequal in length, femur long and slender, slightly clavate, about 2.5x as long as prefemur, postfemur and tibia subequal in length, postfemur thicker, tibia more slender, its length about 1/3rd of femur, tarsus even more slender, about 1.5x as long as tibia, claw (ca.
|
||
<quantity id="4CEA9B77C926783BFBA9C27FD7F9F892" box="[1097,1183,1876,1900]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="3" pageNumber="58" unit="mm" value="0.5">0.5 mm</quantity>
|
||
long) slightly curved.
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C926783DFF27C253D7A9F926" blockId="3.[151,1437,327,2008]" lastBlockId="5.[151,1436,1694,1752]" lastPageId="5" lastPageNumber="60" pageId="3" pageNumber="58">
|
||
<emphasis id="B966EA80C926783BFF27C253D2FBF86E" box="[199,413,1912,1936]" italics="true" pageId="3" pageNumber="58">Sexual characters:</emphasis>
|
||
Coxa of 2nd legpair (Fig. 9) of
|
||
<typeStatus id="54A98830C926783BFCC1C253D0E5F86E" box="[801,899,1912,1936]" pageId="3" pageNumber="58" type="holotype">holotype</typeStatus>
|
||
male with well-developed median projections about half as long as length of coxa, apically rounded without membraneous tubulae, with several short setae, 2+2 macrosetae sitting proximally from j oint of prefemur (1 anteriorly, 1 posteriorly), and 1+1 small ones on lateral
|
||
<emphasis id="B966EA80C920783DFF77C38BD25AF948" bold="true" box="[151,316,1696,1719]" pageId="5" pageNumber="60">FIGURES 7–9.</emphasis>
|
||
<taxonomicName id="4C124D11C920783DFEA3C38BD16EF948" box="[323,520,1696,1718]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="5" pageNumber="60" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C920783DFEA3C38BD16EF948" box="[323,520,1696,1718]" italics="true" pageId="5" pageNumber="60">Riukiaria maculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC920783DFDF0C38AD12DF948" box="[528,587,1697,1718]" pageId="5" pageNumber="60" rank="species">
|
||
<emphasis id="B966EA80C920783DFDF0C38AD12DF948" bold="true" box="[528,587,1697,1718]" pageId="5" pageNumber="60">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
<typeStatus id="54A98830C920783DFDB3C38BD1CDF948" box="[595,683,1696,1718]" pageId="5" pageNumber="60" type="holotype">holotype</typeStatus>
|
||
male. 7= Anterior body part, dorsal view; 8= Epiproct and 19th segment, dorsal view; 9= Sternum, coxa, and prefemur of 2nd legpair, posterior view. Scales
|
||
<quantity id="4CEA9B77C920783DFC39C3E9D775F929" box="[985,1043,1730,1751]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="5" pageNumber="60" unit="mm" value="1.0">1 mm</quantity>
|
||
(7,8),
|
||
<quantity id="4CEA9B77C920783DFBB7C3EAD7C2F929" box="[1111,1188,1729,1751]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="5" pageNumber="60" unit="mm" value="0.5">0.5 mm</quantity>
|
||
(9).
|
||
</paragraph>
|
||
<caption id="DF6D661AC921783CFF77C2A5D17DF81F" httpUri="https://zenodo.org/record/277569/files/figure.png" pageId="4" pageNumber="59" targetBox="[273,1322,204,1900]" targetPageId="4">
|
||
<paragraph id="8BAD3692C921783CFF77C2A5D17DF81F" blockId="4.[151,1437,1934,2017]" pageId="4" pageNumber="59">
|
||
<emphasis id="B966EA80C921783CFF77C2A5D25DF85D" bold="true" box="[151,315,1934,1957]" pageId="4" pageNumber="59">FIGURES 2–3.</emphasis>
|
||
<taxonomicName id="4C124D11C921783CFEA2C2A5D161F85A" box="[322,519,1934,1956]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C921783CFEA2C2A5D161F85A" box="[322,519,1934,1956]" italics="true" pageId="4" pageNumber="59">Riukiaria maculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC921783CFDEEC2A4D12EF85A" box="[526,584,1935,1956]" pageId="4" pageNumber="59" rank="species">
|
||
<emphasis id="B966EA80C921783CFDEEC2A4D12EF85A" bold="true" box="[526,584,1935,1956]" pageId="4" pageNumber="59">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, holotype male. 2= Live coloration, Tane-ga-shima; 3= Live specimen in UV-light.
|
||
<emphasis id="B966EA80C921783CFF77C287D25DF83C" bold="true" box="[151,315,1964,1987]" pageId="4" pageNumber="59">FIGURES 4–6.</emphasis>
|
||
<taxonomicName id="4C124D11C921783CFEA2C286D294F83D" box="[322,498,1965,1987]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="mundyi" status="sp. nov.">
|
||
<emphasis id="B966EA80C921783CFEA2C286D294F83D" box="[322,498,1965,1987]" italics="true" pageId="4" pageNumber="59">Riukiaria mundyi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC921783CFE1AC286D152F83C" box="[506,564,1965,1986]" pageId="4" pageNumber="59" rank="species">
|
||
<emphasis id="B966EA80C921783CFE1AC286D152F83C" bold="true" box="[506,564,1965,1986]" pageId="4" pageNumber="59">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, paratype specimens. 4= Live coloration, Yonaguni-jima; 5–6 = The same specimens in ambient flashlight, and in UV-light.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD3692C920783EFF77C22FD6ECFEC1" blockId="5.[151,1436,1796,2036]" lastBlockId="6.[151,1437,151,320]" lastPageId="6" lastPageNumber="61" pageId="5" pageNumber="60">
|
||
sides of coxa. Sternum on segment 4 (legpair 3) narrow, without lobes, on segment 5 (legpairs 4–5) and 6 (legpairs 6–7) gradually wider and smoother, bridging the
|
||
<quantity id="4CEA9B77C920783DFD5EC203D072F8BE" box="[702,788,1832,1856]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="5" pageNumber="60" unit="mm" value="1.8">1.8 mm</quantity>
|
||
intercoxal distance on postgonopodal legs. No other sternal or leg modifications could be observed. Male gonopodal aperture on 7th segment wide, elliptical, about twice as wide as long, gonopods in situ usually deeply embedded, with acropodites crossing each other. Coxa (
|
||
<figureCitation id="13292A17C920783DFAD8C25BD3D2F852" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1945,1968]" captionTargetBox="[170,1402,362,1920]" captionTargetId="figure@6.[151,1436,357,1924]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Riukiaria maculata sp. n. 10 – 12 = Left gonopod of holotype male, lateral, mesal, and apical (ventral) views, respectively; 13 = Left cyphopod of paratype female, posterior view. Scales 0.5 mm. c = coxa, tp = triangular process, v = valve, r = receptacle, op = operculum, other abbreviations as in Fig. 1." httpUri="https://zenodo.org/record/277570/files/figure.png" pageId="5" pageNumber="60">Figs 10– 11</figureCitation>
|
||
,
|
||
<emphasis id="B966EA80C920783DFF5FC2BED3ADF852" box="[191,203,1941,1964]" italics="true" pageId="5" pageNumber="60">c</emphasis>
|
||
) stout, approximately as long as wide, without proximal apophysis but with small apophyseal macroseta (
|
||
<emphasis id="B966EA80C920783DFA8BC2BED6EAF852" box="[1387,1420,1941,1964]" italics="true" pageId="5" pageNumber="60">cm</emphasis>
|
||
), and distally (ventrally) with small but definite, triangular projection (
|
||
<emphasis id="B966EA80C920783DFC48C292D0D8F82E" box="[936,958,1977,2000]" italics="true" pageId="5" pageNumber="60">tp</emphasis>
|
||
). Cannula normal, situated on mesal side. Telopodite has two simple processes (
|
||
<figureCitation id="13292A17C920783DFDA9C2F7D1ABF80A" box="[585,717,2012,2036]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1945,1968]" captionTargetBox="[170,1402,362,1920]" captionTargetId="figure@6.[151,1436,357,1924]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Riukiaria maculata sp. n. 10 – 12 = Left gonopod of holotype male, lateral, mesal, and apical (ventral) views, respectively; 13 = Left cyphopod of paratype female, posterior view. Scales 0.5 mm. c = coxa, tp = triangular process, v = valve, r = receptacle, op = operculum, other abbreviations as in Fig. 1." httpUri="https://zenodo.org/record/277570/files/figure.png" pageId="5" pageNumber="60">Figs 10–12</figureCitation>
|
||
), a shorter, more slender prefemoral process (
|
||
<emphasis id="B966EA80C920783DFB3EC2F7D664F80A" box="[1246,1282,2012,2036]" italics="true" pageId="5" pageNumber="60">pfp</emphasis>
|
||
), and a long, curved, leaflike acropodite (solenomere,
|
||
<emphasis id="B966EA80C923783EFD84C5B2D109FF4E" box="[612,623,153,176]" italics="true" pageId="6" pageNumber="61">s</emphasis>
|
||
), delimitation of which from the densely setose prefemur is hardly visible. The two processes form the the simple, forceps-like gonopodal apparatus typical for
|
||
<taxonomicName id="4C124D11C923783EFB9EC597D257FF09" authority="Tanabe & Shinohara 1996" authorityName="Tanabe & Shinohara" authorityYear="1996" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA80C923783EFB9EC597D78EFF2A" box="[1150,1256,188,212]" italics="true" pageId="6" pageNumber="61">Riukiaria</emphasis>
|
||
(
|
||
<bibRefCitation id="EF834B63C923783EFB16C597D24EFF09" author="Tanabe" pageId="6" pageNumber="61" refString="Tanabe, T. & Shinohara, K. (1996) Revision of the millipede genus Xystodesmus, with reference to the status of the tribe Xystodesmini (Diplopoda: Xystodesmidae). Journal of Natural History, 30, 1459 - 1494." type="journal article" year="1996">Tanabe & Shinohara 1996</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
. Prefemoral process slender, triangular (
|
||
<figureCitation id="13292A17C923783EFCE1C5F4D030FF09" box="[769,854,223,247]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1945,1968]" captionTargetBox="[170,1402,362,1920]" captionTargetId="figure@6.[151,1436,357,1924]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Riukiaria maculata sp. n. 10 – 12 = Left gonopod of holotype male, lateral, mesal, and apical (ventral) views, respectively; 13 = Left cyphopod of paratype female, posterior view. Scales 0.5 mm. c = coxa, tp = triangular process, v = valve, r = receptacle, op = operculum, other abbreviations as in Fig. 1." httpUri="https://zenodo.org/record/277570/files/figure.png" pageId="6" pageNumber="61">Fig. 12</figureCitation>
|
||
,
|
||
<emphasis id="B966EA80C923783EFC84C5CBD0EEFF06" box="[868,904,224,248]" italics="true" pageId="6" pageNumber="61">pfp</emphasis>
|
||
), about 3/4th of length of acropodite, with no setae, hairs, or processes. Tip of acropodite (
|
||
<figureCitation id="13292A17C923783EFD43C42FD19FFEE2" box="[675,761,260,284]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1945,1968]" captionTargetBox="[170,1402,362,1920]" captionTargetId="figure@6.[151,1436,357,1924]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Riukiaria maculata sp. n. 10 – 12 = Left gonopod of holotype male, lateral, mesal, and apical (ventral) views, respectively; 13 = Left cyphopod of paratype female, posterior view. Scales 0.5 mm. c = coxa, tp = triangular process, v = valve, r = receptacle, op = operculum, other abbreviations as in Fig. 1." httpUri="https://zenodo.org/record/277570/files/figure.png" pageId="6" pageNumber="61">Fig. 12</figureCitation>
|
||
, s) bending backwards toward prefemoral process, almost touching it, its sides gradually tapering, prostatic groove running along mesal side, and ending on the pointed tip.
|
||
</paragraph>
|
||
<caption id="DF6D661AC923783EFF77C2B2D068F80D" httpUri="https://zenodo.org/record/277570/files/figure.png" pageId="6" pageNumber="61" targetBox="[170,1402,362,1920]" targetPageId="6">
|
||
<paragraph id="8BAD3692C923783EFF77C2B2D068F80D" blockId="6.[151,1436,1945,2036]" pageId="6" pageNumber="61">
|
||
<emphasis id="B966EA80C923783EFF77C2B2D235F851" bold="true" box="[151,339,1945,1968]" pageId="6" pageNumber="61">FIGURES 10–13.</emphasis>
|
||
<taxonomicName id="4C124D11C923783EFEB8C2B2D17AF851" box="[344,540,1945,1967]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="6" pageNumber="61" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C923783EFEB8C2B2D17AF851" box="[344,540,1945,1967]" italics="true" pageId="6" pageNumber="61">Riukiaria maculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC923783EFDC2C2B1D13DF851" box="[546,603,1946,1967]" pageId="6" pageNumber="61" rank="species">
|
||
<emphasis id="B966EA80C923783EFDC2C2B1D13DF851" bold="true" box="[546,603,1946,1967]" pageId="6" pageNumber="61">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
10–12 = Left gonopod of holotype male, lateral, mesal, and apical (ventral) views, respectively; 13= Left cyphopod of paratype female, posterior view. Scales 0.5 mm. c= coxa, tp= triangular process, v= valve, r= receptacle, op= operculum, other abbreviations as in Fig. 1.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BAD3692C922783FFF27C5BCD2BAFEC1" blockId="7.[151,1437,151,1511]" pageId="7" pageNumber="62">
|
||
Female cyphopods (
|
||
<figureCitation id="13292A17C922783FFE49C5BCD29FFF51" box="[425,505,151,175]" captionStart="FIGURES 10 – 13" captionStartId="6.[151,264,1945,1968]" captionTargetBox="[170,1402,362,1920]" captionTargetId="figure@6.[151,1436,357,1924]" captionTargetPageId="6" captionText="FIGURES 10 – 13. Riukiaria maculata sp. n. 10 – 12 = Left gonopod of holotype male, lateral, mesal, and apical (ventral) views, respectively; 13 = Left cyphopod of paratype female, posterior view. Scales 0.5 mm. c = coxa, tp = triangular process, v = valve, r = receptacle, op = operculum, other abbreviations as in Fig. 1." httpUri="https://zenodo.org/record/277570/files/figure.png" pageId="7" pageNumber="62">Fig. 13</figureCitation>
|
||
) embedded closely behind 2nd legpair, in large, ∞-shaped aperture, valves (
|
||
<emphasis id="B966EA80C922783FFAAFC5B2D63DFF4E" box="[1359,1371,153,176]" italics="true" pageId="7" pageNumber="62">v</emphasis>
|
||
) subrectangular, almost twice as wide as high, densely setose, with basal bumps of each seta giving a rough appearance to its surface and margins. Lateral operculum (
|
||
<emphasis id="B966EA80C922783FFD40C5CAD1DAFF06" box="[672,700,225,248]" italics="true" pageId="7" pageNumber="62">op</emphasis>
|
||
) large, thick, almost as high as valves, with dense setation, receptacles (
|
||
<emphasis id="B966EA80C922783FFF08C42ED395FEE2" box="[232,243,261,284]" italics="true" pageId="7" pageNumber="62">r</emphasis>
|
||
) embracing valves both anteriorly and posteriorly, subrectangular, with several series of short hairs along ventral, serrated-like margin.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086519C922783FFF27C460D1D3FE31" pageId="7" pageNumber="62" type="distribution">
|
||
<paragraph id="8BAD3692C922783FFF27C460D1D3FE31" blockId="7.[151,1437,151,1511]" pageId="7" pageNumber="62">
|
||
<emphasis id="B966EA80C922783FFF27C460D206FE9A" bold="true" box="[199,352,331,356]" pageId="7" pageNumber="62">Distribution.</emphasis>
|
||
Known only from Tane-ga-shima Island, northern Ryukyus (Osumi Island Group), Kagoshima Prefecture,
|
||
<collectingCountry id="F3057602C922783FFEF8C45BD23AFE79" box="[280,348,368,391]" name="Japan" pageId="7" pageNumber="62">Japan</collectingCountry>
|
||
. Specimens were found in the single locality of a mixed forest of deciduous trees, possibly an old
|
||
<taxonomicName id="4C124D11C922783FFF77C4BED2F4FE52" box="[151,402,405,428]" class="Insecta" family="Noctuidae" genus="Cryptomeria" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="B966EA80C922783FFF77C4BED2F4FE52" box="[151,402,405,428]" italics="true" pageId="7" pageNumber="62">Cryptomeria japonica</emphasis>
|
||
</taxonomicName>
|
||
plantation. Search in other similar habitats on the island (about 445 square kilometers of total surface area) produced no more specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3086519C922783FFF27C4F7D2A5FA19" pageId="7" pageNumber="62" type="discussion">
|
||
<paragraph id="8BAD3692C922783FFF27C4F7D07FFCC4" blockId="7.[151,1437,151,1511]" pageId="7" pageNumber="62">
|
||
<emphasis id="B966EA80C922783FFF27C4F7D25DFE0A" bold="true" box="[199,315,476,500]" pageId="7" pageNumber="62">Remarks.</emphasis>
|
||
Tane-ga-shima island, the locality of the new species’ population, is a member of the Osumi Islands, Northern Ryukyus, about
|
||
<quantity id="4CEA9B77C922783FFDFAC4D4D104FDE9" box="[538,610,511,535]" metricMagnitude="4" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="62" unit="km" value="50.0">50 km</quantity>
|
||
southeast of Kyushu. It is an elongate island
|
||
<quantity id="4CEA9B77C922783FFB89C4D4D7D7FDE9" box="[1129,1201,511,535]" metricMagnitude="4" metricUnit="m" metricValue="5.7" pageId="7" pageNumber="62" unit="km" value="57.0">57 km</quantity>
|
||
from north to south, and about
|
||
<quantity id="4CEA9B77C922783FFEEAC70FD20BFDC2" box="[266,365,548,572]" metricMagnitude="3" metricUnit="m" metricValue="8.5" metricValueMax="12.0" metricValueMin="5.0" pageId="7" pageNumber="62" unit="km" value="8.5" valueMax="12.0" valueMin="5.0">5–12 km</quantity>
|
||
from east to west, with the highest elevation
|
||
<specimenCount id="9D14FD1BC922783FFC86C70FD0D2FDC2" box="[870,948,548,572]" pageId="7" pageNumber="62" type="generic">
|
||
<quantity id="4CEA9B77C922783FFC86C70FD0C9FDC2" box="[870,943,548,572]" metricMagnitude="2" metricUnit="m" metricValue="2.82" pageId="7" pageNumber="62" unit="m" value="282.0">282 m</quantity>
|
||
.
|
||
</specimenCount>
|
||
The nearest island is the small, uninhabited Mage-jima, with an area of only
|
||
<quantity id="4CEA9B77C922783FFDEBC761D122FD9C" box="[523,580,586,610]" metricMagnitude="3" metricUnit="m" metricValue="8.0" pageId="7" pageNumber="62" unit="km" value="8.0">8 km</quantity>
|
||
2, and with no worth-while habitat for native millipedes. Yaku-shima island, on the other hand, the largest member of the Osumi Group, is about
|
||
<quantity id="4CEA9B77C922783FFC6DC744D0B2FD79" box="[909,980,623,647]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="7" pageNumber="62" unit="km" value="30.0">30 km</quantity>
|
||
to the west, and has a notable vegetation and fauna. Its high mountains (up to
|
||
<quantity id="4CEA9B77C922783FFDA4C7B9D1FDFD54" box="[580,667,658,682]" metricMagnitude="3" metricUnit="m" metricValue="1.936" pageId="7" pageNumber="62" unit="m" value="1936.0">1936 m</quantity>
|
||
a.s.l.) are mainly covered by ancient Japanese cedar (
|
||
<taxonomicName id="4C124D11C922783FFAEDC7B8D39FFD31" class="Insecta" family="Noctuidae" genus="Cryptomeria" kingdom="Animalia" order="Lepidoptera" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="japonica">
|
||
<emphasis id="B966EA80C922783FFAEDC7B8D39FFD31" italics="true" pageId="7" pageNumber="62">Cryptomeria japonica</emphasis>
|
||
</taxonomicName>
|
||
) forest, and provide an optimal habitat for many endemics. Two
|
||
<taxonomicName id="4C124D11C922783FFC08C79CD737FD31" box="[1000,1105,695,719]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA80C922783FFC08C79CD737FD31" box="[1000,1105,695,719]" italics="true" pageId="7" pageNumber="62">Riukiaria</emphasis>
|
||
</taxonomicName>
|
||
species have been described from Yaku-shima (
|
||
<taxonomicName id="4C124D11C922783FFE8EC7F7D2B3FD0D" box="[366,469,731,755]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="B966EA80C922783FFE8EC7F7D2B3FD0D" box="[366,469,731,755]" italics="true" pageId="7" pageNumber="62">R. puella</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D11C922783FFDEFC7F7D111FD0D" box="[527,631,731,755]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="jamila">
|
||
<emphasis id="B966EA80C922783FFDEFC7F7D111FD0D" box="[527,631,731,755]" italics="true" pageId="7" pageNumber="62">R. jamila</emphasis>
|
||
</taxonomicName>
|
||
), but they both differ considerably in size (
|
||
<quantity id="4CEA9B77C922783FFB8BC7F1D78FFD0C" box="[1131,1257,730,754]" metricMagnitude="-2" metricUnit="m" metricValue="3.4" metricValueMax="3.6" metricValueMin="3.2" pageId="7" pageNumber="62" unit="mm" value="34.0" valueMax="36.0" valueMin="32.0">32–36 mm</quantity>
|
||
) and coloration from
|
||
<taxonomicName id="4C124D11C922783FFF35C62BD23BFCE9" box="[213,349,767,791]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C922783FFF35C62BD23BFCE9" box="[213,349,767,791]" italics="true" pageId="7" pageNumber="62">R. maculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC922783FFE85C7D4D2C3FCE9" box="[357,421,767,791]" pageId="7" pageNumber="62" rank="species">
|
||
<emphasis id="B966EA80C922783FFE85C7D4D2C3FCE9" bold="true" box="[357,421,767,791]" pageId="7" pageNumber="62">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
:
|
||
<taxonomicName id="4C124D11C922783FFE54C62BD17CFCE9" box="[436,538,767,791]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="puella">
|
||
<emphasis id="B966EA80C922783FFE54C62BD17CFCE9" box="[436,538,767,791]" italics="true" pageId="7" pageNumber="62">R. puella</emphasis>
|
||
</taxonomicName>
|
||
is almost entirely yellow, whereas tergites of
|
||
<taxonomicName id="4C124D11C922783FFBC2C62BD7ECFCE9" box="[1058,1162,767,791]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="jamila">
|
||
<emphasis id="B966EA80C922783FFBC2C62BD7ECFCE9" box="[1058,1162,767,791]" italics="true" pageId="7" pageNumber="62">R. jamila</emphasis>
|
||
</taxonomicName>
|
||
are gray with yellowish white paranota. (For gonopod differences see Diagnosis.)
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C922783FFF27C663D216FB81" blockId="7.[151,1437,151,1511]" pageId="7" pageNumber="62">
|
||
In the southern part of Kyushu, one of the four main islands of
|
||
<collectingCountry id="F3057602C922783FFC47C663D08CFCA1" box="[935,1002,840,863]" name="Japan" pageId="7" pageNumber="62">Japan</collectingCountry>
|
||
, three
|
||
<taxonomicName id="4C124D11C922783FFBD8C66CD7C4FCA1" box="[1080,1186,839,863]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA80C922783FFBD8C66CD7C4FCA1" box="[1080,1186,839,863]" italics="true" pageId="7" pageNumber="62">Riukiaria</emphasis>
|
||
</taxonomicName>
|
||
species are known to occur:
|
||
<taxonomicName id="4C124D11C922783FFF04C647D23DFC7D" box="[228,347,876,899]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cornuta">
|
||
<emphasis id="B966EA80C922783FFF04C647D23DFC7D" box="[228,347,876,899]" italics="true" pageId="7" pageNumber="62">R. cornuta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C124D11C922783FFE8BC647D16FFC7D" box="[363,521,875,899]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="anachoreta">
|
||
<emphasis id="B966EA80C922783FFE8BC647D16FFC7D" box="[363,521,875,899]" italics="true" pageId="7" pageNumber="62">R. anachoreta</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C124D11C922783FFDA8C647D061FC7D" box="[584,775,875,899]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="semicircularis">
|
||
<emphasis id="B966EA80C922783FFDA8C647D061FC7D" box="[584,775,875,899]" italics="true" pageId="7" pageNumber="62">R. semicircularis</emphasis>
|
||
</taxonomicName>
|
||
. Whereas their size is similar (
|
||
<taxonomicName id="4C124D11C922783FFB89C647D661FC7D" box="[1129,1287,875,899]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="anachoreta">
|
||
<emphasis id="B966EA80C922783FFB89C647D661FC7D" box="[1129,1287,875,899]" italics="true" pageId="7" pageNumber="62">R. anachoreta</emphasis>
|
||
</taxonomicName>
|
||
, body length
|
||
<quantity id="4CEA9B77C922783FFF77C6A4D272FC59" box="[151,276,911,935]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.5" metricValueMin="3.9" pageId="7" pageNumber="62" unit="mm" value="42.0" valueMax="45.0" valueMin="39.0">39–45 mm</quantity>
|
||
,
|
||
<taxonomicName id="4C124D11C922783FFEC1C6BBD287FC59" box="[289,481,911,935]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="semicircularis">
|
||
<emphasis id="B966EA80C922783FFEC1C6BBD287FC59" box="[289,481,911,935]" italics="true" pageId="7" pageNumber="62">R. semicircularis</emphasis>
|
||
</taxonomicName>
|
||
<quantity id="4CEA9B77C922783FFE0AC6A4D15CFC59" box="[490,570,911,935]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" pageId="7" pageNumber="62" unit="mm" value="40.0">40 mm</quantity>
|
||
) to, or larger (
|
||
<taxonomicName id="4C124D11C922783FFD05C6BBD03BFC59" box="[741,861,912,935]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cornuta">
|
||
<emphasis id="B966EA80C922783FFD05C6BBD03BFC59" box="[741,861,912,935]" italics="true" pageId="7" pageNumber="62">R. cornuta</emphasis>
|
||
</taxonomicName>
|
||
,
|
||
<quantity id="4CEA9B77C922783FFC8DC6A4D0A6FC59" box="[877,960,911,935]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" pageId="7" pageNumber="62" unit="mm" value="55.0">55 mm</quantity>
|
||
) than
|
||
<taxonomicName id="4C124D11C922783FFBE9C6BBD7F4FC59" box="[1033,1170,911,935]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C922783FFBE9C6BBD7F4FC59" box="[1033,1170,911,935]" italics="true" pageId="7" pageNumber="62">R. maculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC922783FFB7BC6A4D7BBFC59" box="[1179,1245,911,935]" pageId="7" pageNumber="62" rank="species">
|
||
<emphasis id="B966EA80C922783FFB7BC6A4D7BBFC59" bold="true" box="[1179,1245,911,935]" pageId="7" pageNumber="62">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
, all three have a generally greenish or brownish gray color, with a little yellow or whitish tint on the paranota in
|
||
<taxonomicName id="4C124D11C922783FFB34C69FD62DFC35" box="[1236,1355,948,971]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cornuta">
|
||
<emphasis id="B966EA80C922783FFB34C69FD62DFC35" box="[1236,1355,948,971]" italics="true" pageId="7" pageNumber="62">R. cornuta</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C124D11C922783FFA64C69FD250FC11" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="semicircularis">
|
||
<emphasis id="B966EA80C922783FFA64C69FD250FC11" italics="true" pageId="7" pageNumber="62">R. semicircularis</emphasis>
|
||
</taxonomicName>
|
||
. Moreover, metatergites of
|
||
<taxonomicName id="4C124D11C922783FFD97C6F3D07EFC11" box="[631,792,983,1007]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="anachoreta">
|
||
<emphasis id="B966EA80C922783FFD97C6F3D07EFC11" box="[631,792,983,1007]" italics="true" pageId="7" pageNumber="62">R. anachoreta</emphasis>
|
||
</taxonomicName>
|
||
show 3 transverse rows of conspicuous tubercles. Male gonopods also differ in the length and shape of the acropodite and prefemoral process, the former in case of
|
||
<taxonomicName id="4C124D11C922783FFAAFC6D7D3AFFBC9" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="cornuta">
|
||
<emphasis id="B966EA80C922783FFAAFC6D7D3AFFBC9" italics="true" pageId="7" pageNumber="62">R. cornuta</emphasis>
|
||
</taxonomicName>
|
||
bearing a small mesal tooth. The prefemoral process of
|
||
<taxonomicName id="4C124D11C922783FFCA2C10BD766FBC9" box="[834,1024,1055,1079]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="semicircularis">
|
||
<emphasis id="B966EA80C922783FFCA2C10BD766FBC9" box="[834,1024,1055,1079]" italics="true" pageId="7" pageNumber="62">R. semicircularis</emphasis>
|
||
</taxonomicName>
|
||
is long and slender, undulated like a flagellum, and only a little shorter than the acropodite, the tip of which bends backwards to form an almost completely closed oval.
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C922783FFF27C1A1D38DFA84" blockId="7.[151,1437,151,1511]" pageId="7" pageNumber="62">
|
||
Based only on superficially similar coloration, we also might to compare
|
||
<taxonomicName id="4C124D11C922783FFBEAC1A7D7F4FB5D" box="[1034,1170,1163,1187]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C922783FFBEAC1A7D7F4FB5D" box="[1034,1170,1163,1187]" italics="true" pageId="7" pageNumber="62">R. maculata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B966EA80C922783FFB7BC1A0D7BDFB5D" bold="true" box="[1179,1243,1163,1187]" pageId="7" pageNumber="62">
|
||
<taxonomicNameLabel id="A25557FBC922783FFB7BC1A0D7BDFB5D" box="[1179,1243,1163,1187]" pageId="7" pageNumber="62" rank="species">sp. n.</taxonomicNameLabel>
|
||
</emphasis>
|
||
with
|
||
<taxonomicName id="4C124D11C922783FFAFDC1A7D6FAFB5D" box="[1309,1436,1163,1187]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="falcifera">
|
||
<emphasis id="B966EA80C922783FFAFDC1A7D6FAFB5D" box="[1309,1436,1163,1187]" italics="true" pageId="7" pageNumber="62">R. falcifera</emphasis>
|
||
</taxonomicName>
|
||
from Okinawa-jima island. The two locations are more than
|
||
<quantity id="4CEA9B77C922783FFCA7C184D0FAFB39" box="[839,924,1199,1223]" metricMagnitude="5" metricUnit="m" metricValue="5.0" pageId="7" pageNumber="62" unit="km" value="500.0">500 km</quantity>
|
||
apart by sea, and
|
||
<taxonomicName id="4C124D11C922783FFB8BC19BD78CFB39" box="[1131,1258,1199,1223]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="falcifera">
|
||
<emphasis id="B966EA80C922783FFB8BC19BD78CFB39" box="[1131,1258,1199,1223]" italics="true" pageId="7" pageNumber="62">R. falcifera</emphasis>
|
||
</taxonomicName>
|
||
is considerably larger (
|
||
<quantity id="4CEA9B77C922783FFF08C1F9D201FB14" box="[232,359,1234,1258]" metricMagnitude="-2" metricUnit="m" metricValue="5.9" metricValueMax="6.5" metricValueMin="5.3" pageId="7" pageNumber="62" unit="mm" value="59.0" valueMax="65.0" valueMin="53.0">53–65 mm</quantity>
|
||
), but its coloration also differs from the new species, the dark spots never being so strong, and not of the same size. In
|
||
<taxonomicName id="4C124D11C922783FFE47C1D3D149FAF1" box="[423,559,1271,1295]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="maculata">
|
||
<emphasis id="B966EA80C922783FFE47C1D3D149FAF1" box="[423,559,1271,1295]" italics="true" pageId="7" pageNumber="62">R. maculata</emphasis>
|
||
</taxonomicName>
|
||
, the spots are always widely separated by the yellow median part, and all are the same size on each segment, giving really a 'spotted' appearance to the animal. In
|
||
<taxonomicName id="4C124D11C922783FFBA7C037D7A3FACD" box="[1095,1221,1307,1331]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="falcifera">
|
||
<emphasis id="B966EA80C922783FFBA7C037D7A3FACD" box="[1095,1221,1307,1331]" italics="true" pageId="7" pageNumber="62">R. falcifera</emphasis>
|
||
</taxonomicName>
|
||
, on the other hand, the spots are blurred, sometimes missing or fused together, and change in their size and shape along the body of the animal.
|
||
</paragraph>
|
||
<paragraph id="8BAD3692C922783FFF27C0ACD2A5FA19" blockId="7.[151,1437,151,1511]" pageId="7" pageNumber="62">
|
||
Since
|
||
<taxonomicName id="4C124D11C922783FFEEDC0A3D2F0FA61" box="[269,406,1415,1439]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="species" species="maculata" status="sp. nov.">
|
||
<emphasis id="B966EA80C922783FFEEDC0A3D2F0FA61" box="[269,406,1415,1439]" italics="true" pageId="7" pageNumber="62">R. maculata</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A25557FBC922783FFE7FC0ACD287FA61" box="[415,481,1415,1439]" pageId="7" pageNumber="62" rank="species">
|
||
<emphasis id="B966EA80C922783FFE7FC0ACD287FA61" bold="true" box="[415,481,1415,1439]" pageId="7" pageNumber="62">sp. n.</emphasis>
|
||
</taxonomicNameLabel>
|
||
is only known from a single population, and because its attractive, colorful appearance we recommend the new species for full legal protection. The new species could well be one of the rarest
|
||
<taxonomicName id="4C124D11C922783FFF77C0E4D267FA19" box="[151,257,1487,1511]" class="Diplopoda" family="Xystodesmidae" genus="Riukiaria" kingdom="Animalia" order="Polydesmida" pageId="7" pageNumber="62" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B966EA80C922783FFF77C0E4D267FA19" box="[151,257,1487,1511]" italics="true" pageId="7" pageNumber="62">Riukiaria</emphasis>
|
||
</taxonomicName>
|
||
species in
|
||
<collectingCountry id="F3057602C922783FFE9CC0FBD2D9FA19" box="[380,447,1488,1511]" name="Japan" pageId="7" pageNumber="62">Japan</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |