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<document id="171E841CF51FF90EAF7A463FB3D8EEE9" ID-DOI="10.11646/phytotaxa.614.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="8389307" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1695979127173" checkinUser="plazi" docAuthor="Henderson, Andrew" docDate="2023" docId="03B387DAFFB81F27FF50F99FFA8E8A5E" docLanguage="en" docName="phytotaxa.614.1.1.pdf" docOrigin="Phytotaxa 614 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.614.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.3:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="3" docTitle="Coccothrinax pumila Borhidi &amp; Hernandez" docType="treatment" docVersion="2" lastPageNumber="73" masterDocId="FF8AFFA2FFFF1F6FFFD8FFB9FFF9881B" masterDocTitle="A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)" masterLastPageNumber="115" masterPageNumber="1" pageNumber="72" updateTime="1695980191673" updateUser="ExternalLinkService">
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<mods:title id="11E3442E498B06E2766A7A1F4A520724">A revision of Coccothrinax, Hemithrinax, Leucothrinax, Thrinax, and Zombia (Arecaceae)</mods:title>
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<subSubSection id="C3006547FFB81F28FF50F99FFD548E92" pageId="71" pageNumber="72" type="nomenclature">
<paragraph id="8BA536CCFFB81F28FF50F99FFD548E92" blockId="71.[136,1451,1573,1673]" pageId="71" pageNumber="72">
<emphasis id="B96EEADEFFB81F28FF50F99FFBC18E5B" bold="true" box="[136,1080,1573,1601]" pageId="71" pageNumber="72">
1.30.
<taxonomicName id="4C1A4D4FFFB81F28FF11F99FFD2A8E5B" ID-CoL="WLZB" authority="Borhidi &amp; Hernandez" authorityName="Borhidi &amp; Hernandez" box="[201,723,1573,1601]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="71" pageNumber="72" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B96EEADEFFB81F28FF11F99FFE388E5A" bold="true" box="[201,449,1574,1601]" italics="true" pageId="71" pageNumber="72">Coccothrinax pumila</emphasis>
Borhidi &amp; Hernández
</taxonomicName>
(19931994 publ. 1995: 195).
</emphasis>
<materialsCitation id="3B723C91FFB81F28FB9AF99EFD548E92" collectingDate="1994-03" collectionCode="J, HAC" collectorName="de Hatibonico &amp; de la Base Naval" country="Cuba" location="Guantanamo" pageId="71" pageNumber="72" specimenCount="2" stateProvince="Guantanamo" typeStatus="lectotype">
<typeStatus id="54A1886EFFB81F28FB9AF99EFB4B8E5A" box="[1090,1202,1575,1601]" pageId="71" pageNumber="72">Lectotype</typeStatus>
(designated by
<bibRefCitation id="EF8B4B3DFFB81F28FAB2F99EFF388E7E" author="Moya, C." pageId="71" pageNumber="72" pagination="1 - 27" refId="ref63288" refString="Moya, C. (2021) An annotated checklist of Cuban palms 6. Coccothrinax, Pt. 2. 1972 - 1995. Nomenclature, typification, and distribution. PalmArbor 3: 1 - 27." type="journal article" year="2021">Moya 2021</bibRefCitation>
):—
<collectingCountry id="F30D765CFFB81F28FF34F9F2FEC68E7E" box="[236,319,1611,1637]" name="Cuba" pageId="71" pageNumber="72">CUBA</collectingCountry>
. Prov.
<collectingRegion id="49DEF82EFFB81F28FE49F9F2FDDA8E7E" box="[401,547,1611,1637]" country="Cuba" name="Guantanamo" pageId="71" pageNumber="72">Guantánamo</collectingRegion>
, Morrillo Chico, area
<collectorName id="26EF531AFFB81F28FCF0F9F3FC358E7F" box="[808,972,1610,1636]" pageId="71" pageNumber="72">de Hatibonico</collectorName>
, oeste
<collectorName id="26EF531AFFB81F28FBC7F9F3FB1F8E7F" box="[1055,1254,1610,1637]" pageId="71" pageNumber="72">de la Base Naval</collectorName>
de
<collectingRegion id="49DEF82EFFB81F28FACDF9F2FA5E8E7E" box="[1301,1447,1611,1637]" country="Cuba" name="Guantanamo" pageId="71" pageNumber="72">Guantánamo</collectingRegion>
,
<date id="FFA4100CFFB81F28FF50F9D7FEEA8E92" box="[136,275,1646,1673]" pageId="71" pageNumber="72" value="1994-03">
<collectingDate id="EFE0E9E4FFB81F28FF50F9D7FEEA8E92" box="[136,275,1646,1673]" pageId="71" pageNumber="72" value="1994-03">March 1994</collectingDate>
</date>
,
<emphasis id="B96EEADEFFB81F28FEC6F9D6FE188E93" box="[286,481,1646,1672]" italics="true" pageId="71" pageNumber="72">
<collectionCode id="ED0BAE09FFB81F28FEC6F9D6FED78E93" box="[286,302,1647,1672]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="71" pageNumber="72" type="Herbarium">J</collectionCode>
. Hernández s.n.
</emphasis>
(
<typeStatus id="54A1886EFFB81F28FE28F9D7FDA08E93" box="[496,601,1646,1672]" pageId="71" pageNumber="72">lectotype</typeStatus>
<collectionCode id="ED0BAE09FFB81F28FD87F9D6FD588E92" box="[607,673,1647,1673]" country="Cuba" httpUri="http://grbio.org/cool/cenz-te88" name="Instituto de Ecología y Sistemática" pageId="71" pageNumber="72" type="Herbarium">HAC</collectionCode>
!)
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFB81F27FF50F90FFF0B893F" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72" type="description">
<paragraph id="8BA536CCFFB81F27FF50F90FFF0B893F" blockId="71.[136,1452,1718,2069]" lastBlockId="72.[136,1452,158,581]" lastPageId="72" lastPageNumber="73" pageId="71" pageNumber="72">
<emphasis id="B96EEADEFFB81F28FF50F90FFF298ECB" bold="true" box="[136,208,1718,1744]" pageId="71" pageNumber="72">Stems</emphasis>
length not recorded, 7.0 cm diameter, clustered.
<emphasis id="B96EEADEFFB81F28FCC8F90EFC9B8ECA" bold="true" box="[784,866,1719,1745]" pageId="71" pageNumber="72">Leaves</emphasis>
not recorded; leaf sheath fibers
<quantity id="4CE29B29FFB81F28FB38F90EFAC08ECA" box="[1248,1337,1719,1745]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="71" pageNumber="72" unit="mm" value="1.8">1.8 mm</quantity>
diameter, stout, woody, loosely woven, ± joined or briefly free at the apices; petioles
<quantity id="4CE29B29FFB81F28FC2DF963FBB78EEE" box="[1013,1102,1754,1781]" metricMagnitude="-3" metricUnit="m" metricValue="7.6" pageId="71" pageNumber="72" unit="mm" value="7.6">7.6 mm</quantity>
diameter just below the apex; palmans
<quantity id="4CE29B29FFB81F28FF36F947FEBC8F02" box="[238,325,1790,1817]" metricMagnitude="-3" metricUnit="m" metricValue="4.5" pageId="71" pageNumber="72" unit="mm" value="4.5">4.5 mm</quantity>
long, relatively short, with the adaxial veins prominent and terminating in a slight raised ridge and distinct pulvinus; leaf blades not wedge-shaped; segments 22 per leaf, the middle ones 38.0 cm long and
<quantity id="4CE29B29FFB81F28FAC3F89AFA9F8F26" box="[1307,1382,1827,1853]" metricMagnitude="-2" metricUnit="m" metricValue="2.4" pageId="71" pageNumber="72" unit="cm" value="2.4">2.4 cm</quantity>
wide; segments not pendulous at the apices, giving the leaf a flat appearance; middle leaf segments relatively short and broad, abruptly narrowed (shoulder) toward the apex, otherwise parallel-sided, often strongly folded, stiff and leathery, the apices briefly splitting; middle leaf segment apices attenuate; leaf segments with a deciduous, thin layer of wax adaxially, densely indumentose abaxially, with irregularly shaped, persistent, interlocking, fimbriate hairs, each one with a rounded, raised, light green to greenish-brown center, without transverse veinlets.
<emphasis id="B96EEADEFFB81F28FBB9F86FFAF38FEB" bold="true" box="[1121,1290,2006,2032]" pageId="71" pageNumber="72">Inflorescences</emphasis>
erect amongst the leaves, with few partial inflorescences at apex of inflorescence; rachis bracts not recorded; partial inflorescences 2; proximalmost rachillae straight,
<quantity id="4CE29B29FFB71F27FE2EFF27FDBA88A2" box="[502,579,158,185]" metricMagnitude="-2" metricUnit="m" metricValue="3.5" pageId="72" pageNumber="73" unit="cm" value="3.5">3.5 cm</quantity>
long and
<quantity id="4CE29B29FFB71F27FD6CFF27FCF288A2" box="[692,779,158,185]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" pageId="72" pageNumber="73" unit="mm" value="0.8">0.8 mm</quantity>
diameter in fruit; rachillae not recorded at or near anthesis; stamens 9; fruit pedicels
<quantity id="4CE29B29FFB71F27FE70FF7BFDF988C6" box="[424,512,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" pageId="72" pageNumber="73" unit="mm" value="2.5">2.5 mm</quantity>
long;
<emphasis id="B96EEADEFFB71F27FD93FF7BFD7588C7" bold="true" box="[587,652,194,220]" pageId="72" pageNumber="73">fruits</emphasis>
<quantity id="4CE29B29FFB71F27FD4EFF7BFD1788C6" box="[662,750,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="72" pageNumber="73" unit="mm" value="8.5">8.5 mm</quantity>
long and
<quantity id="4CE29B29FFB71F27FCBBFF7BFC4288C6" box="[867,955,194,221]" metricMagnitude="-3" metricUnit="m" metricValue="8.5" pageId="72" pageNumber="73" unit="mm" value="8.5">8.5 mm</quantity>
diameter, color not recorded; fruit surfaces smooth or sometimes with projecting fibers; seed surfaces deeply lobed, the lobes running from base of seeds almost to apices.
</paragraph>
</subSubSection>
<subSubSection id="C3006547FFB71F27FF65FE97FA8E8A5E" pageId="72" pageNumber="73" type="distribution">
<paragraph id="8BA536CCFFB71F27FF65FE97FE758976" blockId="72.[136,1452,158,581]" pageId="72" pageNumber="73">
<emphasis id="B96EEADEFFB71F27FF65FE97FDFC8953" bold="true" box="[189,517,302,328]" pageId="72" pageNumber="73">Distribution and habitat:—</emphasis>
<collectingCountry id="F30D765CFFB71F27FDDDFE97FDB98953" box="[517,576,302,328]" name="Cuba" pageId="72" pageNumber="73">Cuba</collectingCountry>
(
<collectingRegion id="49DEF82EFFB71F27FD96FE97FD188953" box="[590,737,302,328]" country="Cuba" name="Guantanamo" pageId="72" pageNumber="73">Guantánamo</collectingRegion>
) (
<figureCitation id="13212A49FFB71F27FD2FFE97FCB18953" box="[759,840,302,328]" captionStart="FIGURE 16" captionStartId="71.[136,229,1506,1528]" captionTargetBox="[155,1429,198,1477]" captionTargetId="figure-15@71.[150,1434,193,1482]" captionTargetPageId="71" captionText="FIGURE 16. Distribution of Coccothrinax pseudorigida, C. pumila, C. rigida, and C. salvatoris." figureDoi="http://doi.org/10.5281/zenodo.8389405" httpUri="https://zenodo.org/record/8389405/files/figure.png" pageId="72" pageNumber="73">Fig. 16</figureCitation>
) on coastal savannas or hills on limestone rocks at low elevations near the sea.
</paragraph>
<paragraph id="8BA536CCFFB71F27FF65FECFFA8E8A5E" blockId="72.[136,1452,158,581]" pageId="72" pageNumber="73">
<emphasis id="B96EEADEFFB71F27FF65FECFFE56898B" bold="true" box="[189,431,374,400]" pageId="72" pageNumber="73">Taxonomic notes:—</emphasis>
As a preliminary species,
<taxonomicName id="4C1A4D4FFFB71F27FD39FECFFC2B898B" box="[737,978,374,400]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B96EEADEFFB71F27FD39FECFFC2B898B" box="[737,978,374,400]" italics="true" pageId="72" pageNumber="73">Coccothrinax pumila</emphasis>
</taxonomicName>
has a unique combination of qualitative character states and is recognized as a phylogenetic species. Only
<specimenCount id="9D1CFD45FFB71F27FCA4FE22FBEF89AF" box="[892,1046,410,437]" pageId="72" pageNumber="73" type="generic">one specimen</specimenCount>
has been examined and the species remains poorly known. In the protologue,
<taxonomicName id="4C1A4D4FFFB71F27FD8AFE06FD4689C3" box="[594,703,446,473]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B96EEADEFFB71F27FD8AFE06FD4689C3" box="[594,703,446,473]" italics="true" pageId="72" pageNumber="73">C. pumila</emphasis>
</taxonomicName>
is said to belong to section
<taxonomicName id="4C1A4D4FFFB71F27FC3DFE07FB7389C3" box="[997,1162,446,472]" pageId="72" pageNumber="73" rank="section" section="Longispadicae">
<emphasis id="B96EEADEFFB71F27FC3DFE07FB7389C3" box="[997,1162,446,472]" italics="true" pageId="72" pageNumber="73">Longispadicae</emphasis>
</taxonomicName>
subsection
<taxonomicName id="4C1A4D4FFFB71F27FAD2FE06FA5589C3" box="[1290,1452,447,472]" pageId="72" pageNumber="73" rank="subSection" subSection="Pauciramosae">
<emphasis id="B96EEADEFFB71F27FAD2FE06FA5589C3" box="[1290,1452,447,472]" italics="true" pageId="72" pageNumber="73">Pauciramosae</emphasis>
</taxonomicName>
and to be similar to
<taxonomicName id="4C1A4D4FFFB71F27FEBDFE5AFDE389E7" box="[357,538,482,509]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="pseudorigida">
<emphasis id="B96EEADEFFB71F27FEBDFE5AFDE389E7" box="[357,538,482,509]" italics="true" pageId="72" pageNumber="73">C. pseudorigida</emphasis>
</taxonomicName>
. It does share several character states with that species. An image in
<bibRefCitation id="EF8B4B3DFFB71F27FAC1FE5AFA5289E7" author="Moya, C." box="[1305,1451,482,509]" pageId="72" pageNumber="73" pagination="1 - 27" refId="ref63288" refString="Moya, C. (2021) An annotated checklist of Cuban palms 6. Coccothrinax, Pt. 2. 1972 - 1995. Nomenclature, typification, and distribution. PalmArbor 3: 1 - 27." type="journal article" year="2021">Moya (2021)</bibRefCitation>
of a plant from the
<typeStatus id="54A1886EFFB71F27FEB4FDBEFE658A3A" box="[364,412,519,545]" pageId="72" pageNumber="73">type</typeStatus>
locality of
<taxonomicName id="4C1A4D4FFFB71F27FDFCFDBEFD6C8A3B" box="[548,661,518,545]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B96EEADEFFB71F27FDFCFDBEFD6C8A3B" box="[548,661,518,545]" italics="true" pageId="72" pageNumber="73">C. pumila</emphasis>
</taxonomicName>
shows stems covered with a skirt of persistent dead leaves (as in
<taxonomicName id="4C1A4D4FFFB71F27FA4AFDBEFEE58A5F" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="pseudorigida">
<emphasis id="B96EEADEFFB71F27FA4AFDBEFEE58A5F" italics="true" pageId="72" pageNumber="73">C. pseudorigida</emphasis>
</taxonomicName>
). However, the images of
<taxonomicName id="4C1A4D4FFFB71F27FD9DFD92FD4A8A5F" box="[581,691,554,581]" class="Liliopsida" family="Arecaceae" genus="Coccothrinax" kingdom="Plantae" order="Arecales" pageId="72" pageNumber="73" phylum="Tracheophyta" rank="species" species="pumila">
<emphasis id="B96EEADEFFB71F27FD9DFD92FD4A8A5F" box="[581,691,554,581]" italics="true" pageId="72" pageNumber="73">C. pumila</emphasis>
</taxonomicName>
in
<bibRefCitation id="EF8B4B3DFFB71F27FD0FFD93FC9B8A5F" author="Craft, P." box="[727,866,554,580]" pageId="72" pageNumber="73" refId="ref61560" refString="Craft, P. (2017) The Palms of Cuba. Palm Nut Pages, Florida, 232 pp." type="book" year="2017">Craft (2017)</bibRefCitation>
do not show skirts of dead leaves on the stems.
</paragraph>
</subSubSection>
</treatment>
</document>