289 lines
36 KiB
XML
289 lines
36 KiB
XML
<document id="953364B168BA349E699D16F8DDABD896" ID-CLB-Dataset="30030" ID-DOI="10.11646/zootaxa.4422.4.3" ID-GBIF-Dataset="d2c5d8c4-d2e2-44a7-add4-3a94812d2581" ID-ISSN="1175-5326" ID-Zenodo-Dep="1253219" ID-ZooBank="45E16185-7EE6-4768-88DF-6ACA1D29DCE4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1527266432226" checkinUser="plazi" docAuthor="Yang, Ho Jin, Seo, Ji Eun & Gordon, Dennis P." docDate="2018" docId="03B387C39C79FFB176DCC4E25572FF71" docLanguage="en" docName="zootaxa.4422.4.3.pdf.imf" docOrigin="Zootaxa 4422 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Torquatella imperforata Yang & Seo & Gordon 2018, n. sp." docType="treatment" docVersion="7" lastPageNumber="510" masterDocId="FF8AFFBB9C69FFA0764BC508565FFFC1" masterDocTitle="Sixteen new generic records of Korean Bryozoa from southern coastal waters and Jeju Island, East China Sea: evidence of tropical affinities" masterLastPageNumber="518" masterPageNumber="493" pageNumber="509" updateTime="1698545350699" updateUser="ExternalLinkService">
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<mods:title id="309853A02F12134EF5AB5664F21BB2BC">Sixteen new generic records of Korean Bryozoa from southern coastal waters and Jeju Island, East China Sea: evidence of tropical affinities</mods:title>
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<mods:namePart id="D3A25013A6E7F1E785153323CA2959E7">Yang, Ho Jin</mods:namePart>
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<mods:namePart id="E3783FA25CE8D8A1A2C1AD4E1B656B5F">Seo, Ji Eun</mods:namePart>
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<mods:namePart id="2B16AAA50C99B13C78C700497D0E32A9">Gordon, Dennis P.</mods:namePart>
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<mods:title id="2D71229A1A7546A8E75236CF4C9C1085">Zootaxa</mods:title>
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<mods:date id="B5F67B5142FABF40EFB28F7FD7B5DE05">2018</mods:date>
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<mods:number id="F6744F49CCB8041262B248AB10F64349">4422</mods:number>
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<treatment id="03B387C39C79FFB176DCC4E25572FF71" ID-DOI="http://doi.org/10.5281/zenodo.5964321" ID-GBIF-Taxon="144128889" ID-Zenodo-Dep="5964321" LSID="urn:lsid:plazi:treatment:03B387C39C79FFB176DCC4E25572FF71" httpUri="http://treatment.plazi.org/id/03B387C39C79FFB176DCC4E25572FF71" lastPageId="17" lastPageNumber="510" pageId="16" pageNumber="509">
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<subSubSection id="C300655E9C79FFB076DCC4E25778FDE7" pageId="16" pageNumber="509" type="nomenclature">
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<paragraph id="8BA536D59C79FFB076DCC4E2545DFDC5" blockId="16.[151,514,490,550]" box="[151,514,490,516]" pageId="16" pageNumber="509">
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<heading id="D0ED81B99C79FFB076DCC4E2545DFDC5" bold="true" box="[151,514,490,516]" fontSize="11" level="1" pageId="16" pageNumber="509" reason="1">
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<emphasis id="B96EEAC79C79FFB076DCC4E2545DFDC5" bold="true" box="[151,514,490,516]" pageId="16" pageNumber="509">
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<taxonomicName id="4C1A4D569C79FFB076DCC4E257E7FDC5" authority="Yang & Seo & Gordon, 2018" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[151,440,490,516]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="imperforata" status="sp. nov.">
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<emphasis id="B96EEAC79C79FFB076DCC4E257E7FDC5" bold="true" box="[151,440,490,516]" italics="true" pageId="16" pageNumber="509">Torquatella imperforata</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A25D57BC9C79FFB077F4C4E2545DFDC5" box="[447,514,490,516]" pageId="16" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BA536D59C79FFB076DCC7055778FDE7" blockId="16.[151,514,490,550]" box="[151,295,525,550]" pageId="16" pageNumber="509">
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(
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<figureCitation id="13212A509C79FFB076D4C7055778FDE7" box="[159,295,525,550]" captionStart="FIGURE 28" captionStartId="15.[151,250,1539,1561]" captionTargetBox="[250,1339,293,1521]" captionTargetId="figure@15.[235,1351,281,1530]" captionTargetPageId="15" captionText="FIGURE 28. Hippaliosina setiformis Tilbrook, 2006, Munseom Island, showing, at centre, a female zooid (♀) with enlarged orifice and ooecium, surrounded by autozooids with two types of avicularia. FIGURES 29–31. Torquatella imperforata n. sp., holotype, Munseom Island. 29, ovicellate and non-ovicellate zooids from colony margin to colony center; 30, autozooid at colony margin and ooecium of proximal zooid; 31, two orifices, each with an intraperistomial avicularium. FIGURE 32. Stephanollona boreopacifica n. sp., holotype, Munseom Island, showing a six-spined autozooidal orifice (right), ovicellate orifices and two types of adventitious avicularia. Scale bars: 28, 500 µm; 29, 400 µm; 30, 31, 100 µm; 32, 300 µm." httpUri="https://zenodo.org/record/1253233/files/figure.png" pageId="16" pageNumber="509">Figs 29–31)</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C300655E9C79FFB076DCC75D54AEFDAF" box="[151,753,597,622]" pageId="16" pageNumber="509" type="etymology">
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<paragraph id="8BA536D59C79FFB076DCC75D54AEFDAF" blockId="16.[151,1437,597,2026]" box="[151,753,597,622]" pageId="16" pageNumber="509">
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<emphasis id="B96EEAC79C79FFB076DCC75D5741FDAF" bold="true" box="[151,286,597,622]" pageId="16" pageNumber="509">Etymology.</emphasis>
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Alluding to the non-foraminate ooecium.
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</paragraph>
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</subSubSection>
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<subSubSection id="C300655E9C79FFB0768CC7715245FD77" pageId="16" pageNumber="509" type="materials_examined">
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<paragraph id="8BA536D59C79FFB0768CC7715245FD77" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">
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<materialsCitation id="3B723C889C79FFB0768CC7715245FD77" ID-GBIF-Occurrence="1845321963" collectingDate="2016-12-24" collectorName="Ho Jin Yang." location="Seogwipo" municipality="Munseom Island" pageId="16" pageNumber="509" specimenCount="1" stateProvince="Jeju" typeStatus="holotype">
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<emphasis id="B96EEAC79C79FFB0768CC77157EBFD53" bold="true" box="[199,436,633,658]" pageId="16" pageNumber="509">Material examined.</emphasis>
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<emphasis id="B96EEAC79C79FFB077F4C7715470FD53" box="[447,559,633,658]" italics="true" pageId="16" pageNumber="509">
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<typeStatus id="54A188779C79FFB077F4C7715477FD53" box="[447,552,633,658]" pageId="16" pageNumber="509" type="holotype">Holotype</typeStatus>
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:
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</emphasis>
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NIBRIV0000805892, unique colony found at
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<collectingMunicipality id="6BC1ACAF9C79FFB07218C7725349FD53" box="[1107,1302,633,658]" pageId="16" pageNumber="509">Munseom Island</collectingMunicipality>
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,
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<location id="8EC5600E9C79FFB0736EC77153C7FD53" LSID="urn:lsid:plazi:treatment:03B387C39C79FFB176DCC4E25572FF71:8EC5600E9C79FFB0736EC77153C7FD53" box="[1317,1432,633,658]" municipality="Munseom Island" name="Seogwipo" pageId="16" pageNumber="509" stateProvince="Jeju">Seogwipo</location>
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,
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<collectingRegion id="49DEF8379C79FFB076DCC795569BFD77" box="[151,196,669,694]" country="South Korea" name="Jeju" pageId="16" pageNumber="509">Jeju</collectingRegion>
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Island,
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<date id="FFA410159C79FFB07756C79657AAFD77" box="[285,501,669,694]" pageId="16" pageNumber="509" value="2016-12-24">
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<collectingDate id="EFE0E9FD9C79FFB07756C79657AAFD77" box="[285,501,669,694]" pageId="16" pageNumber="509" value="2016-12-24">
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24 December
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<quantity id="4CE29B309C79FFB077F1C79557AAFD77" box="[442,501,669,694]" metricMagnitude="3" metricUnit="m" metricValue="2.0162" pageId="16" pageNumber="509" unit="m" value="2016.2">2016</quantity>
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</collectingDate>
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</date>
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, 20 m, on oyster shell, collected by
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<collectorName id="26EF53039C79FFB075C3C7965245FD77" box="[904,1050,669,694]" pageId="16" pageNumber="509">Ho Jin Yang.</collectorName>
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</materialsCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="C300655E9C79FFB0768CC7C9522FFB13" pageId="16" pageNumber="509" type="description">
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<paragraph id="8BA536D59C79FFB0768CC7C955C0FCAB" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">
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<emphasis id="B96EEAC79C79FFB0768CC7C95706FD1B" bold="true" box="[199,345,705,730]" pageId="16" pageNumber="509">Description.</emphasis>
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Colony encrusting, multiserial, unilaminar, small,
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<quantity id="4CE29B309C79FFB075FAC7CA5253FD18" box="[945,1036,706,730]" metricMagnitude="-3" metricUnit="m" metricValue="4.7" pageId="16" pageNumber="509" unit="mm" value="4.7">4.7 mm</quantity>
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across. Autozooids with irregular indistinct boundaries except at margin, frontal shield porcellanous, surface topography irregular, sparsely porous (fewer than a dozen pores). Orifice a little wider than long, anter proportionately large, delimited from broad shallow poster (sinus) by small triangular condyles in proximolateral corners. Peristome well developed, forming thick crescentic ridge around proximal half of orifice. No oral spines.
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</paragraph>
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<paragraph id="8BA536D59C79FFB0768CC67D521DFC17" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">Avicularia not in every zooid, adventitious, set singly on one side of inner face of peristome. Rostrum and opesia roundly triangular, edge of latter at midline of peristome; face of avicularium concave, curving obliquely laterally within curvature of peristome, rostral rim slightly projecting, crossbar thin.</paragraph>
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<paragraph id="8BA536D59C79FFB0768CC6EA54E4FB83" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">Most zooids in colony ovicellate. Ooecium recumbent on distal zooid, subglobular or with slightly flattened apex, opening above zooidal operculum, skeletal surface smooth, non-foraminate, proximal margin becoming elevated and continuous with orificial peristome.</paragraph>
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<paragraph id="8BA536D59C79FFB0768CC1455724FB48" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">Interzooidal communications via basal pore-chambers, a large distolateral pair and smaller distal pair. Ancestrula not seen.</paragraph>
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<paragraph id="8BA536D59C79FFB0768CC19D522FFB13" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">
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<emphasis id="B96EEAC79C79FFB0768CC19D5722FB6F" bold="true" box="[199,381,1173,1198]" pageId="16" pageNumber="509">Measurements.</emphasis>
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ZL 129–250 (179) µm; ZW 90–159 (124) µm; OrL 32–51 (37) µm; OrW 38–47 (42) µm; AvL 21–28 (25) µm; AvW 11–14 (12) µm; OoL 53–82 (69) µm; OoW 87–112 (98) µm.
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</paragraph>
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</subSubSection>
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<subSubSection id="C300655E9C79FFB0768CC1D55267F967" pageId="16" pageNumber="509" type="discussion">
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<paragraph id="8BA536D59C79FFB0768CC1D55267F967" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">
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<emphasis id="B96EEAC79C79FFB0768CC1D55764FB37" bold="true" box="[199,315,1245,1270]" pageId="16" pageNumber="509">Remarks.</emphasis>
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The present species raises the question of relationship between
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<taxonomicName id="4C1A4D569C79FFB07272C1D552E5FB37" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[1081,1210,1245,1270]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
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<emphasis id="B96EEAC79C79FFB07272C1D552E5FB37" box="[1081,1210,1245,1270]" italics="true" pageId="16" pageNumber="509">Torquatella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1A4D569C79FFB072B1C1D5571DFADB" authority="Tilbrook, 2006" authorityName="Tilbrook" authorityYear="2006" class="Gymnolaemata" family="Celleporidae" genus="Predanophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
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<emphasis id="B96EEAC79C79FFB072B1C1D553C3FB37" box="[1274,1436,1245,1270]" italics="true" pageId="16" pageNumber="509">Predanophora</emphasis>
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<bibRefCitation id="EF8B4B249C79FFB076DCC009571DFADB" box="[151,322,1281,1306]" pageId="16" pageNumber="509" refId="ref16335" refString="Tilbrook, K. J. (2006) Cheilostomatous Bryozoa from the Solomon Islands. Santa Barbara Museum of Natural History Monographs 4, Studies in Biodiversity, 3, 1 - 385." type="journal article">Tilbrook, 2006</bibRefCitation>
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</taxonomicName>
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. As was pointed out by
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<bibRefCitation id="EF8B4B249C79FFB07418C0095556FADB" box="[595,777,1281,1306]" pageId="16" pageNumber="509" refId="ref16335" refString="Tilbrook, K. J. (2006) Cheilostomatous Bryozoa from the Solomon Islands. Santa Barbara Museum of Natural History Monographs 4, Studies in Biodiversity, 3, 1 - 385." type="journal article">Tilbrook (2006)</bibRefCitation>
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, the peristomial orifice, which incorporates a more-or-less transversely orientated avicularium on its inner face, is “identical” in the two genera, but he segregated
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<taxonomicName id="4C1A4D569C79FFB076DCC0415765FAA3" authorityName="Tilbrook" authorityYear="2006" box="[151,314,1353,1378]" class="Gymnolaemata" family="Celleporidae" genus="Predanophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
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<emphasis id="B96EEAC79C79FFB076DCC0415765FAA3" box="[151,314,1353,1378]" italics="true" pageId="16" pageNumber="509">Predanophora</emphasis>
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</taxonomicName>
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on the grounds that the ooecium is bifenestrate, whereas in
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<taxonomicName id="4C1A4D569C79FFB075A3C0415236FAA3" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[1000,1129,1353,1378]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
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<emphasis id="B96EEAC79C79FFB075A3C0415236FAA3" box="[1000,1129,1353,1378]" italics="true" pageId="16" pageNumber="509">Torquatella</emphasis>
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</taxonomicName>
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it is unifenestrate. Further, he created a new family,
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<taxonomicName id="4C1A4D569C79FFB07788C0655438FA47" authorityName="Tilbrook" authorityYear="2006" box="[451,615,1389,1414]" class="Gymnolaemata" family="Torquatellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="family">Torquatellidae</taxonomicName>
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, monogeneric for
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<taxonomicName id="4C1A4D569C79FFB07574C065559FFA47" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[831,960,1389,1414]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
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<emphasis id="B96EEAC79C79FFB07574C065559FFA47" box="[831,960,1389,1414]" italics="true" pageId="16" pageNumber="509">Torquatella</emphasis>
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</taxonomicName>
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, on the grounds that its frontal shield is pseudoporous, an alien character in
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<taxonomicName id="4C1A4D569C79FFB0747CC0995495FA6B" authorityName="Johnston" authorityYear="1838" box="[567,714,1425,1450]" class="Gymnolaemata" family="Celleporidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="family">Celleporidae</taxonomicName>
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. We would argue that the similarities between
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<taxonomicName id="4C1A4D569C79FFB072A2C0995335FA6B" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[1257,1386,1425,1450]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
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<emphasis id="B96EEAC79C79FFB072A2C0995335FA6B" box="[1257,1386,1425,1450]" italics="true" pageId="16" pageNumber="509">Torquatella</emphasis>
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</taxonomicName>
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and
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<taxonomicName id="4C1A4D569C79FFB076DCC0BD5766FA0F" authorityName="Tilbrook" authorityYear="2006" box="[151,313,1461,1486]" class="Gymnolaemata" family="Celleporidae" genus="Predanophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B96EEAC79C79FFB076DCC0BD5766FA0F" box="[151,313,1461,1486]" italics="true" pageId="16" pageNumber="509">Predanophora</emphasis>
|
||
</taxonomicName>
|
||
are greater than the differences. Hence, we unite the two genera and abandon
|
||
<taxonomicName id="4C1A4D569C79FFB072B8C0BD53C7FA0F" authorityName="Tilbrook" authorityYear="2006" box="[1267,1432,1461,1486]" class="Gymnolaemata" family="Torquatellidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="family">Torquatellidae</taxonomicName>
|
||
. Pseudopores are in fact known in
|
||
<taxonomicName id="4C1A4D569C79FFB07460C0D154E2FA33" authorityName="Johnston" authorityYear="1838" box="[555,701,1497,1522]" class="Gymnolaemata" family="Celleporidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="family">Celleporidae</taxonomicName>
|
||
, providing the basis for the genus
|
||
<emphasis id="B96EEAC79C79FFB07212C0D152BDFA33" box="[1113,1250,1497,1522]" italics="true" pageId="16" pageNumber="509">Portalesella</emphasis>
|
||
<bibRefCitation id="EF8B4B249C79FFB072A6C0D153C7FA33" box="[1261,1432,1497,1522]" pageId="16" pageNumber="509" refId="ref16533" refString="Winston, J. E. (2005) Re-description and revision of Smitt's " Floridan Bryozoa " in the collection of the Museum of Comparative Zoology, Harvard University. Virginia Museum of Natural History Memoir, 7, 1 - 147." type="journal article">Winston, 2005</bibRefCitation>
|
||
, which, as
|
||
<bibRefCitation id="EF8B4B249C79FFB07742C0F55415F9D7" box="[265,586,1533,1558]" pageId="16" pageNumber="509" refId="ref11722" refString="Berning, B. & Kuklinski, P. (2008) North-east Atlantic and Mediterranean species of the genus Buffonellaria (Bryozoa, Cheilostomata): implications for biodiversity and biogeography. Zoological Journal of the Linnean Society, 152, 537 - 566. https: // doi. org / 10.1111 / j. 1096 - 3642.2007.00379. x" type="journal article">Berning & Kuklinski (2008)</bibRefCitation>
|
||
pointed out, is so close to
|
||
<taxonomicName id="4C1A4D569C79FFB07531C0F55250F9D7" box="[890,1039,1533,1558]" class="Gymnolaemata" family="Celleporidae" genus="Buffonellaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B96EEAC79C79FFB07531C0F55250F9D7" box="[890,1039,1533,1558]" italics="true" pageId="16" pageNumber="509">Buffonellaria</emphasis>
|
||
</taxonomicName>
|
||
, with some partially pseudoporous species [including
|
||
<taxonomicName id="4C1A4D569C79FFB07724C3295532F9FB" authority="Seo & Gong, 2006" authorityName="Seo & Gong" authorityYear="2006" box="[367,877,1569,1594]" class="Gymnolaemata" family="Celleporidae" genus="Buffonellaria" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="acutrostris">
|
||
<emphasis id="B96EEAC79C79FFB07724C32954D8F9F8" box="[367,647,1569,1594]" italics="true" pageId="16" pageNumber="509">Buffonellaria acutrostris</emphasis>
|
||
<bibRefCitation id="EF8B4B249C79FFB074D9C32A5532F9FB" box="[658,877,1569,1594]" pageId="16" pageNumber="509" refId="ref15654" refString="Seo, J. E. & Gong, Y. H. (2006) A new species and two new records of Cheilostomata (Bryozoa) from Korea. Korean Journal of Systematic Zoology, 22, 13 - 16." type="journal article">Seo & Gong, 2006</bibRefCitation>
|
||
</taxonomicName>
|
||
from
|
||
<collectingRegion id="49DEF8379C79FFB075F3C32955BAF9FB" box="[952,997,1569,1594]" country="South Korea" name="Jeju" pageId="16" pageNumber="509">Jeju</collectingRegion>
|
||
Island], that there is little to separate them. In parallel, these authors also noted that pseudopores may be an inadequate basis for separating
|
||
<taxonomicName id="4C1A4D569C79FFB076DCC3615765F943" authorityName="Tilbrook" authorityYear="2006" box="[151,314,1641,1666]" class="Gymnolaemata" family="Celleporidae" genus="Predanophora" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B96EEAC79C79FFB076DCC3615765F943" box="[151,314,1641,1666]" italics="true" pageId="16" pageNumber="509">Predanophora</emphasis>
|
||
</taxonomicName>
|
||
from
|
||
<taxonomicName id="4C1A4D569C79FFB077C7C3615455F943" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[396,522,1641,1666]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="genus">
|
||
<emphasis id="B96EEAC79C79FFB077C7C3615455F943" box="[396,522,1641,1666]" italics="true" pageId="16" pageNumber="509">Torquatella</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
<bibRefCitation id="EF8B4B249C79FFB0746AC3615490F943" box="[545,719,1641,1666]" pageId="16" pageNumber="509" refId="ref13209" refString="Gordon, D. P. (2000) Towards a phylogeny of cheilostomes-morphological models of frontal wall / shield evolution. In: Herrera Cubilla, A. & Jackson, J. B. C. (Eds.), Proceedings of the 11 th International Bryozoology Association Conference. Smithsonian Tropical Research Institution, Panama City, pp. 17 - 37." type="book chapter">Gordon (2000)</bibRefCitation>
|
||
hypothesized that pseudopores are a derived character that originated independently from branching areolar-septular pores in several families.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300655E9C79FFB0768CC3B9537EF82B" pageId="16" pageNumber="509" type="reference_group">
|
||
<paragraph id="8BA536D59C79FFB0768CC3B9528DF8BC" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">
|
||
<taxonomicName id="4C1A4D569C79FFB0768CC3B9578AF90B" authority="Yang & Seo & Gordon, 2018" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[199,469,1713,1738]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="imperforata" status="sp. nov.">
|
||
<emphasis id="B96EEAC79C79FFB0768CC3B9578AF90B" box="[199,469,1713,1738]" italics="true" pageId="16" pageNumber="509">Torquatella imperforata</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B96EEAC79C79FFB07796C3B95442F90B" bold="true" box="[477,541,1713,1738]" pageId="16" pageNumber="509">
|
||
<taxonomicNameLabel id="A25D57BC9C79FFB07796C3B95442F90B" box="[477,541,1713,1738]" pageId="16" pageNumber="509" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
resembles
|
||
<taxonomicName id="4C1A4D569C79FFB074D7C3B9520BF90B" authority="Scholz, 1991" authorityName="Scholz" authorityYear="1991" box="[668,1108,1713,1738]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="duolamellata">
|
||
<emphasis id="B96EEAC79C79FFB074D7C3B955E6F90B" box="[668,953,1713,1738]" italics="true" pageId="16" pageNumber="509">Torquatella duolamellata</emphasis>
|
||
<bibRefCitation id="EF8B4B249C79FFB0758BC3B9520BF90B" box="[960,1108,1713,1738]" pageId="16" pageNumber="509" refId="ref15439" refString="Scholz, J. (1991) Die Bryozoenfauna der philippinischen Riffregion Cebu. Mitteilungen aus dem Geologisch- Palaontologischen Institut der Universitat Hamburg, 71, 253 - 403." type="journal article">
|
||
Scholz,
|
||
<quantity id="4CE29B309C79FFB07257C3BA520BF90B" box="[1052,1108,1714,1738]" metricMagnitude="1" metricUnit="m" metricValue="5.05714" pageId="16" pageNumber="509" unit="in" value="1991.0">1991</quantity>
|
||
</bibRefCitation>
|
||
</taxonomicName>
|
||
in having some pseudopores but differs in having a non-fenestrate ooecium. Likewise,
|
||
<taxonomicName id="4C1A4D569C79FFB0750FC3DE55B5F92F" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[836,1002,1750,1774]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="imperforata">
|
||
<emphasis id="B96EEAC79C79FFB0750FC3DE55B5F92F" box="[836,1002,1750,1774]" italics="true" pageId="16" pageNumber="509">T. imperforata</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C1A4D569C79FFB072C6C3DD571BF8D3" authority="(Harmer, 1957)" baseAuthorityName="Harmer" baseAuthorityYear="1957" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="longiuscula" status="comb. nov.">
|
||
<emphasis id="B96EEAC79C79FFB072C6C3DD53C3F92F" box="[1165,1436,1749,1774]" italics="true" pageId="16" pageNumber="509">Torquatella longiuscula</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B249C79FFB076D5C3F25764F8D3" box="[158,315,1785,1810]" pageId="16" pageNumber="509" refId="ref13424" refString="Harmer, S. F. (1957) The Polyzoa of the Siboga Expedition. Part 4. Cheilostomata Ascophora. II. Siboga-Expeditie, 28 d, 641 - 1147, pls. 42 - 74." type="journal article">Harmer, 1957</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B96EEAC79C79FFB07701C3F157EFF8D3" bold="true" box="[330,432,1785,1810]" pageId="16" pageNumber="509">
|
||
<taxonomicNameLabel id="A25D57BC9C79FFB07701C3F157EFF8D3" box="[330,432,1785,1810]" pageId="16" pageNumber="509">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
and
|
||
<taxonomicName id="4C1A4D569C79FFB077ADC3F155C7F8D3" authority="(Tilbrook, 2006)" baseAuthorityName="Tilbrook" baseAuthorityYear="2006" box="[486,920,1785,1810]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="ensenada" status="comb. nov.">
|
||
<emphasis id="B96EEAC79C79FFB077ADC3F15488F8D3" box="[486,727,1785,1810]" italics="true" pageId="16" pageNumber="509">Torquatella ensenada</emphasis>
|
||
(
|
||
<bibRefCitation id="EF8B4B249C79FFB074ADC3F155CFF8D3" box="[742,912,1785,1810]" pageId="16" pageNumber="509" refId="ref16335" refString="Tilbrook, K. J. (2006) Cheilostomatous Bryozoa from the Solomon Islands. Santa Barbara Museum of Natural History Monographs 4, Studies in Biodiversity, 3, 1 - 385." type="journal article">Tilbrook, 2006</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
<emphasis id="B96EEAC79C79FFB075D5C3F1525BF8D3" bold="true" box="[926,1028,1785,1810]" pageId="16" pageNumber="509">
|
||
<taxonomicNameLabel id="A25D57BC9C79FFB075D5C3F1525BF8D3" box="[926,1028,1785,1810]" pageId="16" pageNumber="509">n. comb.</taxonomicNameLabel>
|
||
</emphasis>
|
||
in lacking paired lateral windows in the ooecium.
|
||
<taxonomicName id="4C1A4D569C79FFB07773C2155417F8F7" authorityName="Yang & Seo & Gordon" authorityYear="2018" baseAuthorityName="Yang & Seo & Gordon" baseAuthorityYear="2018" box="[312,584,1821,1846]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="longiuscula">
|
||
<emphasis id="B96EEAC79C79FFB07773C2155417F8F7" box="[312,584,1821,1846]" italics="true" pageId="16" pageNumber="509">Torquatella longiuscula</emphasis>
|
||
</taxonomicName>
|
||
also has a thinner peristome and
|
||
<taxonomicName id="4C1A4D569C79FFB075ABC2165235F8F7" authorityName="Yang & Seo & Gordon" authorityYear="2018" baseAuthorityName="Yang & Seo & Gordon" baseAuthorityYear="2018" box="[992,1130,1821,1846]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="ensenada">
|
||
<emphasis id="B96EEAC79C79FFB075ABC2165235F8F7" box="[992,1130,1821,1846]" italics="true" pageId="16" pageNumber="509">T. ensenada</emphasis>
|
||
</taxonomicName>
|
||
has a smaller peristomial avicularium and additional, larger frontally adventitious avicularia. An ancestrula was not seen in the
|
||
<typeStatus id="54A188779C79FFB07371C24953C3F89B" box="[1338,1436,1857,1882]" pageId="16" pageNumber="509" type="holotype">holotype</typeStatus>
|
||
colony of
|
||
<taxonomicName id="4C1A4D569C79FFB07743C26E57F5F8BF" authorityName="Yang & Seo & Gordon" authorityYear="2018" box="[264,426,1894,1918]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="imperforata">
|
||
<emphasis id="B96EEAC79C79FFB07743C26E57F5F8BF" box="[264,426,1894,1918]" italics="true" pageId="16" pageNumber="509">T. imperforata</emphasis>
|
||
</taxonomicName>
|
||
but
|
||
<bibRefCitation id="EF8B4B249C79FFB07790C26D54D1F8BF" box="[475,654,1893,1918]" pageId="16" pageNumber="509" refId="ref16335" refString="Tilbrook, K. J. (2006) Cheilostomatous Bryozoa from the Solomon Islands. Santa Barbara Museum of Natural History Monographs 4, Studies in Biodiversity, 3, 1 - 385." type="journal article">Tilbrook (2006)</bibRefCitation>
|
||
noted that it was small and tatiform in
|
||
<taxonomicName id="4C1A4D569C79FFB0720EC26E5294F8BF" authorityName="Yang & Seo & Gordon" authorityYear="2018" baseAuthorityName="Yang & Seo & Gordon" baseAuthorityYear="2018" box="[1093,1227,1893,1918]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="ensenada">
|
||
<emphasis id="B96EEAC79C79FFB0720EC26E5294F8BF" box="[1093,1227,1893,1918]" italics="true" pageId="16" pageNumber="509">T. ensenada</emphasis>
|
||
</taxonomicName>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8BA536D59C79FFB0768CC281537EF82B" blockId="16.[151,1437,597,2026]" pageId="16" pageNumber="509">
|
||
<collectingRegion id="49DEF8379C79FFB0768CC28156ABF863" box="[199,244,1929,1954]" country="South Korea" name="Jeju" pageId="16" pageNumber="509">Jeju</collectingRegion>
|
||
Island represents the northernmost distribution of this tropical to subtropical genus.
|
||
<taxonomicName id="4C1A4D569C79FFB07350C2815774F807" authorityName="Scholz" authorityYear="1991" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="duolamellata">
|
||
<emphasis id="B96EEAC79C79FFB07350C2815774F807" italics="true" pageId="16" pageNumber="509">Torquatella duolamellata</emphasis>
|
||
</taxonomicName>
|
||
ranges from the
|
||
<collectingCountry id="F30D76459C79FFB077A2C2A55423F807" box="[489,636,1965,1990]" name="Cook Islands" pageId="16" pageNumber="509">Cook Islands</collectingCountry>
|
||
and
|
||
<collectingCountry id="F30D76459C79FFB074F8C2A65550F804" box="[691,783,1966,1989]" name="Vanuatu" pageId="16" pageNumber="509">Vanuatu</collectingCountry>
|
||
through the
|
||
<collectingCountry id="F30D76459C79FFB075D5C2A55206F807" box="[926,1113,1965,1990]" name="Solomon Islands" pageId="16" pageNumber="509">Solomon Islands</collectingCountry>
|
||
to the central
|
||
<collectingCountry id="F30D76459C79FFB072B1C2A55323F807" box="[1274,1404,1965,1990]" name="Philippines" pageId="16" pageNumber="509">Philippines</collectingCountry>
|
||
;
|
||
<taxonomicName id="4C1A4D569C79FFB073CCC2A6575EF82B" authorityName="Yang & Seo & Gordon" authorityYear="2018" baseAuthorityName="Yang & Seo & Gordon" baseAuthorityYear="2018" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="ensenada">
|
||
<emphasis id="B96EEAC79C79FFB073CCC2A6575EF82B" italics="true" pageId="16" pageNumber="509">T. ensenada</emphasis>
|
||
</taxonomicName>
|
||
is found in the
|
||
<collectingCountry id="F30D76459C79FFB077E4C2D95432F82B" box="[431,621,2001,2026]" name="Solomon Islands" pageId="16" pageNumber="509">Solomon Islands</collectingCountry>
|
||
, and
|
||
<taxonomicName id="4C1A4D569C79FFB074E3C2DA5518F82B" authorityName="Yang & Seo & Gordon" authorityYear="2018" baseAuthorityName="Yang & Seo & Gordon" baseAuthorityYear="2018" box="[680,839,2001,2026]" class="Gymnolaemata" family="Torquatellidae" genus="Torquatella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Cheilostomatida" pageId="16" pageNumber="509" phylum="Bryozoa" rank="species" species="longiuscula">
|
||
<emphasis id="B96EEAC79C79FFB074E3C2DA5518F82B" box="[680,839,2001,2026]" italics="true" pageId="16" pageNumber="509">T. longiuscula</emphasis>
|
||
</taxonomicName>
|
||
from the Red Sea and possibly
|
||
<collectingCountry id="F30D76459C79FFB072E7C2D95342F82B" box="[1196,1309,2001,2026]" name="Sri Lanka" pageId="16" pageNumber="509">Sri Lanka</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C300655E9C78FFB1768CC59F5572FF71" box="[199,813,151,176]" pageId="17" pageNumber="510" type="distribution">
|
||
<paragraph id="8BA536D59C78FFB1768CC59F5572FF71" blockId="17.[199,813,151,176]" box="[199,813,151,176]" pageId="17" pageNumber="510">
|
||
<emphasis id="B96EEAC79C78FFB1768CC59F573FFF71" bold="true" box="[199,352,151,176]" pageId="17" pageNumber="510">Distribution.</emphasis>
|
||
<collectingRegion id="49DEF8379C78FFB1772CC59F57CBFF71" box="[359,404,151,176]" country="South Korea" name="Jeju" pageId="17" pageNumber="510">Jeju</collectingRegion>
|
||
Island, southern
|
||
<collectingCountry id="F30D76459C78FFB1741EC59054C3FF6E" box="[597,668,152,175]" name="South Korea" pageId="17" pageNumber="510">Korea</collectingCountry>
|
||
,
|
||
<quantity id="4CE29B309C78FFB174ECC59054BFFF6E" box="[679,736,152,176]" metricMagnitude="1" metricUnit="m" metricValue="2.0" pageId="17" pageNumber="510" unit="m" value="20.0">20 m</quantity>
|
||
depth.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |