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<document id="6A73EE5517F31301A986A0CC1A9CF162" ID-DOI="10.11646/zootaxa.4810.1.12" ID-ISSN="1175-5326" ID-Zenodo-Dep="3937036" ID-ZooBank="424415DC-807E-493A-8883-903D2ADE9427" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1594292134250" checkinUser="plazi" docAuthor="Zhu, Wenhui, Yao, Zhiyuan, Zheng, Guo &amp; Li, Shuqiang" docDate="2020" docId="039FEF343D0FFFCAFF3CF3749471FD28" docLanguage="en" docName="zootaxa.4810.1.12.pdf" docOrigin="Zootaxa 4810 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Belisana guilin Zhu &amp; Yao &amp; Zheng &amp; Li 2020, sp. nov." docType="treatment" docVersion="3" lastPageNumber="179" masterDocId="FFA6974C3D0EFFCEFFABF735972AFFDB" masterDocTitle="Six new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from southern China" masterLastPageNumber="197" masterPageNumber="175" pageNumber="176" updateTime="1700277344912" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="3FD8B0DAB431470DF27D4A04D5591420">Six new species of the spider genus Belisana Thorell, 1898 (Araneae: Pholcidae) from southern China</mods:title>
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<mods:namePart id="F05474337E305699F6ABF8204658286F">Zhu, Wenhui</mods:namePart>
<mods:affiliation id="F7335B1456C7389BFBB5AA7CA86EBDCC">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhuwh 1016 @ outlook. com; https: // orcid. org / 0000 - 0001 - 5650 - 1670</mods:affiliation>
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<mods:namePart id="AAB022209346425AD8A83FEE5296AB84">Yao, Zhiyuan</mods:namePart>
<mods:affiliation id="11E338CE20CC7B5DFA79063DC8BA34F7">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; yaozy @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 1631 - 0949</mods:affiliation>
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<mods:namePart id="BFE6836540942F1F49D4DC8FAC4858D2">Zheng, Guo</mods:namePart>
<mods:affiliation id="5477947F02FBF14F23CA31B77933D296">College of Life Science, Shenyang Normal University, Shenyang 110034, Liaoning, China &amp; zhengguo @ synu. edu. cn; https: // orcid. org / 0000 - 0002 - 9735 - 9700</mods:affiliation>
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<mods:namePart id="514F86A9576F8F193C0BF53E851F3275">Li, Shuqiang</mods:namePart>
<mods:affiliation id="BC5944FD039FA200E76252883883C272">Institute of Zoology, Chinese Academy of Sciences, Beijing 100101, China &amp; lisq @ ioz. ac. cn; https: // orcid. org / 0000 - 0002 - 3290 - 5416</mods:affiliation>
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<mods:title id="53A30AFF1735323CC7F203BA5AAB9E9C">Zootaxa</mods:title>
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<mods:date id="54993DD6B602307AE0E82372B51CE9AF">2020</mods:date>
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<mods:number id="A6229152270109C9540F2A4A0CF39593">4810</mods:number>
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<emphasis id="B94282303D0FFFCFFF3CF3749507FB80" bold="true" box="[151,557,1089,1116]" pageId="1" pageNumber="176">
<taxonomicName id="4C3625A13D0FFFCFFF3CF37496E3FB80" ID-CoL="93VJD" authority="Yao &amp; Li" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[151,457,1089,1116]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="176" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B94282303D0FFFCFFF3CF374967AFB87" bold="true" box="[151,336,1089,1116]" italics="true" pageId="1" pageNumber="176">Belisana guilin</emphasis>
Yao &amp; Li
</taxonomicName>
<taxonomicNameLabel id="A2713F4B3D0FFFCFFE7BF3749507FB80" box="[464,557,1089,1115]" pageId="1" pageNumber="176" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
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<subSubSection id="C32C0DA93D0FFFCFFF3CF350969DFBA4" box="[151,439,1125,1151]" pageId="1" pageNumber="176" type="description">
<paragraph id="8B895E223D0FFFCFFF3CF350969DFBA4" blockId="1.[151,557,1089,1151]" box="[151,439,1125,1151]" pageId="1" pageNumber="176">
<figureCitation id="130D42A73D0FFFCFFF3CF35097D0FBA4" box="[151,250,1125,1151]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStartId-0="2.[151,250,1893,1918]" captionStartId-1="3.[151,250,1910,1935]" captionTargetBox-0="[220,1367,179,1865]" captionTargetBox-1="[214,1373,181,1885]" captionTargetId-0="figure@2.[220,1367,179,1865]" captionTargetId-1="figure@3.[214,1373,181,1885]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." captionText-1="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi-0="http://doi.org/10.5281/zenodo.3937038" figureDoi-1="http://doi.org/10.5281/zenodo.3937040" httpUri-0="https://zenodo.org/record/3937038/files/figure.png" httpUri-1="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Figs 12</figureCitation>
,
<figureCitation id="130D42A73D0FFFCFFEADF350960AFBA4" box="[262,288,1125,1151]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="1" pageNumber="176">13</figureCitation>
AB, 14AB
</paragraph>
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<subSubSection id="C32C0DA93D0FFFCFFF3CF39896BFFAE8" pageId="1" pageNumber="176" type="materials_examined">
<paragraph id="8B895E223D0FFFCFFF3CF39896BFFAE8" blockId="1.[151,1437,1196,1979]" pageId="1" pageNumber="176">
<emphasis id="B94282303D0FFFCFFF3CF3989662FB1C" bold="true" box="[151,328,1196,1223]" pageId="1" pageNumber="176">Type material.</emphasis>
<materialsCitation id="3B5E547F3D0FFFCFFEFAF398952CFAD4" collectingDate="2012-12-06" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zhao" country="China" county="Guilin" elevation="144" latitude="25.215557" location="Unnamed cave" longLatPrecision="1" longitude="110.154976" municipality="Lingui District" pageId="1" pageNumber="176" specimenCount="1" stateProvince="Guangxi" typeStatus="holotype">
<emphasis id="B94282303D0FFFCFFEFAF39896E2FB1C" bold="true" box="[337,456,1197,1223]" pageId="1" pageNumber="176">
<typeStatus id="548DE0803D0FFFCFFEFAF39896E8FB1C" box="[337,450,1197,1223]" pageId="1" pageNumber="176" type="holotype">Holotype</typeStatus>
:
</emphasis>
Male (
<collectionCode id="ED27C6E73D0FFFCFFDB7F3989545FB1C" box="[540,623,1197,1223]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="176">IZCAS</collectionCode>
Ar39846),
<location id="8EE908F93D0FFFCFFD5EF39894B4FB1C" LSID="urn:lsid:plazi:treatment:039FEF343D0FFFCAFF3CF3749471FD28:8EE908F93D0FFFCFFD5EF39894B4FB1C" box="[757,926,1197,1223]" country="China" county="Guilin" latitude="25.215557" longLatPrecision="1" longitude="110.154976" municipality="Lingui District" name="Unnamed cave" pageId="1" pageNumber="176" stateProvince="Guangxi">Unnamed cave</location>
(
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,
<geoCoordinate id="EE0238E53D0FFFCFFBC4F3989202FB1C" box="[1135,1320,1197,1223]" degrees="110" direction="east" minutes="09" orientation="longitude" pageId="1" pageNumber="176" precision="1" seconds="17.90" value="110.154976">110°0917.90”E</geoCoordinate>
,
<elevation id="001BB9113D0FFFCFFA9DF39897CBFB30" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="1" pageNumber="176" unit="m" value="144.0">
elevation
<quantity id="4CCEF3C73D0FFFCFFF3CF3E497CBFB30" box="[151,225,1233,1259]" metricMagnitude="2" metricUnit="m" metricValue="1.44" pageId="1" pageNumber="176" unit="m" value="144.0">144 m</quantity>
</elevation>
), S306
<location id="8EE908F93D0FFFCFFE9DF3E496C4FB30" LSID="urn:lsid:plazi:treatment:039FEF343D0FFFCAFF3CF3749471FD28:8EE908F93D0FFFCFFE9DF3E496C4FB30" box="[310,494,1233,1259]" country="China" county="Guilin" latitude="25.215557" longLatPrecision="1" longitude="110.154976" municipality="Lingui District" name="Provincial Road" pageId="1" pageNumber="176" stateProvince="Guangxi">Provincial Road</location>
, 30 meters east of
<location id="8EE908F93D0FFFCFFD15F3E4941EFB30" LSID="urn:lsid:plazi:treatment:039FEF343D0FFFCAFF3CF3749471FD28:8EE908F93D0FFFCFFD15F3E4941EFB30" box="[702,820,1233,1259]" country="China" county="Guilin" latitude="25.215557" longLatPrecision="1" longitude="110.154976" municipality="Lingui District" name="Feihulinju" pageId="1" pageNumber="176" stateProvince="Guangxi">Feihulinju</location>
,
<collectingMunicipality id="6BEDC4583D0FFFCFFC94F3E494C2FB30" box="[831,1000,1233,1259]" pageId="1" pageNumber="176">Lingui District</collectingMunicipality>
,
<collectingCounty id="62E826AE3D0FFFCFFC58F3E49317FB30" box="[1011,1085,1233,1259]" pageId="1" pageNumber="176">Guilin</collectingCounty>
,
<emphasis id="B94282303D0FFFCFFBE3F3E59385FB30" bold="true" box="[1096,1199,1232,1259]" pageId="1" pageNumber="176">
<collectingRegion id="49F290C03D0FFFCFFBE3F3E59385FB30" box="[1096,1199,1232,1259]" country="China" name="Guangxi" pageId="1" pageNumber="176">Guangxi</collectingRegion>
</emphasis>
,
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<collectingCountry id="F3211EB23D0FFFCFFB16F3E5922CFB30" box="[1213,1286,1232,1259]" name="China" pageId="1" pageNumber="176">China</collectingCountry>
</emphasis>
,
<date id="FF8878E23D0FFFCFFABFF3E497F8FAD4" pageId="1" pageNumber="176" value="2012-12-06">
<collectingDate id="EFCC810A3D0FFFCFFABFF3E497F8FAD4" pageId="1" pageNumber="176" value="2012-12-06">6 December 2012</collectingDate>
</date>
,
<collectorName id="26C33BF43D0FFFCFFF4BF3C09616FAD4" box="[224,316,1269,1295]" pageId="1" pageNumber="176">Z. Chen</collectorName>
and
<collectorName id="26C33BF43D0FFFCFFEDCF3C096F9FAD4" box="[375,467,1269,1295]" pageId="1" pageNumber="176">Z. Zhao</collectorName>
leg.
</materialsCitation>
<materialsCitation id="3B5E547F3D0FFFCFFDA5F3C094DBFAD4" box="[526,1009,1269,1295]" collectionCode="IZCAS" country="China" pageId="1" pageNumber="176" specimenCount="1" specimenCount-male="1" stateProvince="Guangxi" typeStatus="paratype">
<emphasis id="B94282303D0FFFCFFDA5F3C095BAFAD4" bold="true" box="[526,656,1269,1295]" pageId="1" pageNumber="176">
<typeStatus id="548DE0803D0FFFCFFDA5F3C095A0FAD4" box="[526,650,1269,1295]" pageId="1" pageNumber="176" type="paratype">Paratypes</typeStatus>
:
</emphasis>
<specimenCount id="9D3095AB3D0FFFCFFD32F3C095CCFAD4" box="[665,742,1269,1295]" pageId="1" pageNumber="176" type="male">1 male</specimenCount>
(
<collectionCode id="ED27C6E73D0FFFCFFD53F3C09461FAD4" box="[760,843,1269,1295]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="176">IZCAS</collectionCode>
Ar39847) and
</materialsCitation>
<materialsCitation id="3B5E547F3D0FFFCFFC51F3C096B8FAE8" collectingDate="2012-12-06" collectionCode="IZCAS" collectorName="Z. Chen &amp; Z. Zhao" country="China" county="Guilin" elevation="144" latitude="25.215557" location="Unnamed cave" longLatPrecision="1" longitude="110.154976" municipality="Lingui District" pageId="1" pageNumber="176" specimenCount="2" specimenCount-female="2" stateProvince="Guangxi" typeStatus="paratype">
<specimenCount id="9D3095AB3D0FFFCFFC51F3C09343FAD4" box="[1018,1129,1269,1295]" pageId="1" pageNumber="176" type="female">2 females</specimenCount>
(
<collectionCode id="ED27C6E73D0FFFCFFBD1F3C093E7FAD4" box="[1146,1229,1269,1295]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="1" pageNumber="176">IZCAS</collectionCode>
Ar3984839849), same data as holotype
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D0FFFCFFF6CF2089356FA8C" box="[199,1148,1341,1367]" pageId="1" pageNumber="176" type="etymology">
<paragraph id="8B895E223D0FFFCFFF6CF2089356FA8C" blockId="1.[151,1437,1196,1979]" box="[199,1148,1341,1367]" pageId="1" pageNumber="176">
<emphasis id="B94282303D0FFFCFFF6CF2089664FA8C" bold="true" box="[199,334,1341,1367]" pageId="1" pageNumber="176">Etymology.</emphasis>
The specific name refers to the
<typeStatus id="548DE0803D0FFFCFFD1EF20895CFFA8C" box="[693,741,1341,1367]" pageId="1" pageNumber="176">type</typeStatus>
locality and is a noun in apposition.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D0FFFCFFF6CF25594F4F940" pageId="1" pageNumber="176" type="diagnosis">
<paragraph id="8B895E223D0FFFCFFF6CF25594F4F940" blockId="1.[151,1437,1196,1979]" pageId="1" pageNumber="176">
<emphasis id="B94282303D0FFFCFFF6CF2559668FAA0" bold="true" box="[199,322,1376,1403]" pageId="1" pageNumber="176">Diagnosis.</emphasis>
This species resembles
<taxonomicName id="4C3625A13D0FFFCFFDE5F25794E1FAA0" authority="Zhang &amp; Peng, 2011" authorityName="Zhang &amp; Peng" authorityYear="2011" box="[590,971,1377,1403]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="176" phylum="Arthropoda" rank="species" species="longinqua">
<emphasis id="B94282303D0FFFCFFDE5F25795F4FAA0" box="[590,734,1377,1403]" italics="true" pageId="1" pageNumber="176">B. longinqua</emphasis>
<bibRefCitation id="EFA723D33D0FFFCFFD4FF25494E1FAA0" author="Zhang, F. &amp; Peng, Y." box="[740,971,1377,1403]" pageId="1" pageNumber="176" pagination="51 - 68" refId="ref9108" refString="Zhang, F. &amp; Peng, Y. (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa, 2989 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1.2" type="journal article" year="2011">Zhang &amp; Peng, 2011</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFA723D33D0FFFCFFBADF2549238FAA0" author="Zhang, F. &amp; Peng, Y." box="[1030,1298,1376,1403]" pageId="1" pageNumber="176" pagination="51 - 68" refId="ref9108" refString="Zhang, F. &amp; Peng, Y. (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa, 2989 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1.2" type="journal article" year="2011">Zhang &amp; Peng 2011: 58</bibRefCitation>
, fig. 5, male only) with similar male chelicerae (
<figureCitation id="130D42A73D0FFFCFFD82F2B095ABFA44" box="[553,641,1413,1439]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Fig. 2D</figureCitation>
) and bulbal apophyses (
<figureCitation id="130D42A73D0FFFCFFC3FF2B094C0FA44" box="[916,1002,1413,1439]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Fig. 2C</figureCitation>
) and resembles
<taxonomicName id="4C3625A13D0FFFCFFB08F2B3963FFA18" authority="Zhang &amp; Peng, 2011" authorityName="Zhang &amp; Peng" authorityYear="2011" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="176" phylum="Arthropoda" rank="species" species="colubrina">
<emphasis id="B94282303D0FFFCFFB08F2B3921AFA44" box="[1187,1328,1413,1439]" italics="true" pageId="1" pageNumber="176">B. colubrina</emphasis>
<bibRefCitation id="EFA723D33D0FFFCFFA9CF2B0963FFA18" author="Zhang, F. &amp; Peng, Y." pageId="1" pageNumber="176" pagination="51 - 68" refId="ref9108" refString="Zhang, F. &amp; Peng, Y. (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa, 2989 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1.2" type="journal article" year="2011">Zhang &amp; Peng, 2011</bibRefCitation>
</taxonomicName>
(see
<bibRefCitation id="EFA723D33D0FFFCFFEFBF29C954AFA18" author="Zhang, F. &amp; Peng, Y." box="[336,608,1449,1475]" pageId="1" pageNumber="176" pagination="51 - 68" refId="ref9108" refString="Zhang, F. &amp; Peng, Y. (2011) Eleven new species of the genus Belisana Thorell (Araneae: Pholcidae) from South China. Zootaxa, 2989 (1), 51 - 68. https: // doi. org / 10.11646 / zootaxa. 2989.1.2" type="journal article" year="2011">Zhang &amp; Peng 2011: 64</bibRefCitation>
, fig. 9) with similar bulbal apophyses (
<figureCitation id="130D42A73D0FFFCFFBB3F29C9344FA18" box="[1048,1134,1449,1475]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Fig. 2C</figureCitation>
) and external female genitalia (
<figureCitation id="130D42A73D0FFFCFFF73F2F89612FA3C" box="[216,312,1485,1511]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Figs 2A</figureCitation>
,
<figureCitation id="130D42A73D0FFFCFFEEDF2F89653FA3C" box="[326,377,1485,1511]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="1" pageNumber="176">14A</figureCitation>
), but can be distinguished by presence of large, disto-prolateral membranous lamella (arrow
<quantity id="4CCEF3C73D0FFFCFFF3CF2C497E9F9D0" box="[151,195,1521,1547]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="1" pageNumber="176" unit="in" value="1.0">1 in</quantity>
<figureCitation id="130D42A73D0FFFCFFF61F2C4960EF9D0" box="[202,292,1521,1547]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="1" pageNumber="176">Figs 1C</figureCitation>
,
<figureCitation id="130D42A73D0FFFCFFE99F2C49649F9D0" box="[306,355,1521,1547]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="1" pageNumber="176">13A</figureCitation>
) and protruding subdisto-dorsal sclerite (arrow
<quantity id="4CCEF3C73D0FFFCFFCD4F2C49481F9D0" box="[895,939,1521,1547]" metricMagnitude="-2" metricUnit="m" metricValue="5.08" pageId="1" pageNumber="176" unit="in" value="2.0">2 in</quantity>
<figureCitation id="130D42A73D0FFFCFFC19F2C49326F9D0" box="[946,1036,1521,1547]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="1" pageNumber="176">Figs 1C</figureCitation>
,
<figureCitation id="130D42A73D0FFFCFFBB1F2C49361F9D0" box="[1050,1099,1521,1547]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="1" pageNumber="176">13A</figureCitation>
) on procursus, by short retrolatero-ventral apophysis of male pedipalpal trochanter (
<figureCitation id="130D42A73D0FFFCFFCA6F120944FF9F4" box="[781,869,1557,1583]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="1" pageNumber="176">Fig. 1B</figureCitation>
), and by absence of marks on carapace (
<figureCitation id="130D42A73D0FFFCFFA9DF12092A5F9F4" box="[1334,1423,1557,1583]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Fig. 2E</figureCitation>
); also distinguished from
<taxonomicName id="4C3625A13D0FFFCFFE0DF10F9512F988" authorityName="Zhang &amp; Peng" authorityYear="2011" box="[422,568,1593,1619]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="176" phylum="Arthropoda" rank="species" species="longinqua">
<emphasis id="B94282303D0FFFCFFE0DF10F9512F988" box="[422,568,1593,1619]" italics="true" pageId="1" pageNumber="176">B. longinqua</emphasis>
</taxonomicName>
by presence of small proximo-dorsal hump on male pedipalpal femur (arrow in
<figureCitation id="130D42A73D0FFFCFFF1FF1689622F9AC" box="[180,264,1629,1655]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="1" pageNumber="176">Fig. 1B</figureCitation>
) and from
<taxonomicName id="4C3625A13D0FFFCFFE2DF16B9538F9AC" authorityName="Zhang &amp; Peng" authorityYear="2011" box="[390,530,1629,1655]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="1" pageNumber="176" phylum="Arthropoda" rank="species" species="colubrina">
<emphasis id="B94282303D0FFFCFFE2DF16B9538F9AC" box="[390,530,1629,1655]" italics="true" pageId="1" pageNumber="176">B. colubrina</emphasis>
</taxonomicName>
by male cheliceral distal apophyses directed toward downward (arrow da in
<figureCitation id="130D42A73D0FFFCFFADBF1689790F940" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Fig. 2D</figureCitation>
) and vulval pore plates not strongly curved (arrow pp in
<figureCitation id="130D42A73D0FFFCFFC92F1B494BEF940" box="[825,916,1665,1691]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Figs 2B</figureCitation>
,
<figureCitation id="130D42A73D0FFFCFFC34F1B494FBF940" box="[927,977,1665,1691]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="1" pageNumber="176">14B</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D0FFFCAFF6CF19193DAFD14" lastPageId="4" lastPageNumber="179" pageId="1" pageNumber="176" type="description">
<paragraph id="8B895E223D0FFFCAFF6CF191969AFE50" blockId="1.[151,1437,1196,1979]" lastBlockId="4.[151,1437,153,755]" lastPageId="4" lastPageNumber="179" pageId="1" pageNumber="176">
<emphasis id="B94282303D0FFFCFFF6CF19196B5F964" bold="true" box="[199,415,1700,1727]" pageId="1" pageNumber="176">Description. Male</emphasis>
(
<emphasis id="B94282303D0FFFCFFE19F1909533F964" bold="true" box="[434,537,1701,1727]" pageId="1" pageNumber="176">
<typeStatus id="548DE0803D0FFFCFFE19F1909533F964" box="[434,537,1701,1727]" pageId="1" pageNumber="176" type="holotype">holotype</typeStatus>
</emphasis>
, IZCAS Ar39846): Total length 1.68 (1.78 with clypeus), carapace 0.74 long, 0.68 wide, opisthosoma 0.94 long, 0.90 wide. Leg I: 16.84 (4.10 + 0.43 + 4.62 + 6.41 + 1.28), leg II: 10.89 (3.01 + 0.38 + 2.69 + 4.17 + 0.64), leg III: 7.44 (2.12 + 0.26 + 1.86 + 2.49 + 0.71), leg IV: 10.07 (3.27 + 0.32 + 2.44 + 3.27 + 0.77); tibia I L/d: 74. Distance PME-PME 0.09, diameter PME 0.06, distance PME-ALE 0.03, AME absent. Sternum slightly wider than long (0.60/0.50). Habitus as in
<figureCitation id="130D42A73D0FFFCFFC9DF0009451F894" box="[822,891,1845,1871]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Figs 2</figureCitation>
EF. Carapace and sternum pale, without marks. Legs pale, without darker rings. Opisthosoma yellowish white, without spots. Ocular area not elevated. Thoracic furrow absent. Clypeus unmodified. Chelicerae (
<figureCitation id="130D42A73D0FFFCFFD10F0489438F84C" box="[699,786,1917,1943]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="1" pageNumber="176">Fig. 2D</figureCitation>
) with pair of small proximo-lateral apophyses and pair of long, curved distal apophyses (distance between tips of distal apophyses: 0.31). Pedipalps as in
<figureCitation id="130D42A73D0FFFCFFB70F0949234F860" box="[1243,1310,1953,1979]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="1" pageNumber="176">Figs 1</figureCitation>
AB; coxa unmodified; trochanter with short retrolatero-ventral apophysis; femur with small proximo-dorsal hump (arrow in
<figureCitation id="130D42A73D0AFFCAFF3CF78897C6FF0C" box="[151,236,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="4" pageNumber="179">Fig. 1B</figureCitation>
); procursus (
<figureCitation id="130D42A73D0AFFCAFED5F7889695FF0C" box="[382,447,189,215]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="4" pageNumber="179">Figs 1</figureCitation>
AD, 13AB) simple proximally but complex distally, with large, disto-prolateral membranous lamella (arrow
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<figureCitation id="130D42A73D0AFFCAFE7AF7D49500FF20" box="[465,554,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="4" pageNumber="179">Figs 1C</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFD9DF7D49543FF20" box="[566,617,225,251]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13A</figureCitation>
), small subdisto-dorsal sclerite (arrow
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<figureCitation id="130D42A73D0AFFCAFBE0F7D4938EFF20" box="[1099,1188,225,251]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="4" pageNumber="179">Figs 1C</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFB1BF7D493C9FF20" box="[1200,1251,225,251]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13A</figureCitation>
), large subdistoprolateral sclerite (arrow
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<figureCitation id="130D42A73D0AFFCAFE4EF6309514FEC4" box="[485,574,261,287]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="4" pageNumber="179">Figs 1C</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFDE1F6309557FEC4" box="[586,637,261,287]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13A</figureCitation>
), curved distal spine and large retrolateral membranous flap (arrow f in
<figureCitation id="130D42A73D0AFFCAFF3CF61C97D9FE98" box="[151,243,297,323]" captionStart="FIGURE 1" captionStartId="2.[151,250,1893,1918]" captionTargetBox="[220,1367,179,1865]" captionTargetId="figure@2.[220,1367,179,1865]" captionTargetPageId="2" captionText="FIGURE 1. Belisana guilin sp. nov., holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD)." figureDoi="http://doi.org/10.5281/zenodo.3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="4" pageNumber="179">Figs 1D</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFEABF61C9618FE98" box="[256,306,297,323]" captionStart="FIGURE 13" captionStartId="18.[151,250,1945,1970]" captionTargetBox="[241,1355,190,1911]" captionTargetId="figure@18.[225,1362,181,1919]" captionTargetPageId="18" captionText="FIGURE 13. Drawings of male left procursus in prolateral and retrolateral views (arrows point at same structures as photos of each species, respectively). AB. B. guilin sp. nov.; CD. B. maoer sp. nov.; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov.; IJ. B. xuanguan sp. nov.; KL. B. yuexiu sp. nov. f = flap. Scale bars: 0.05." figureDoi="http://doi.org/10.5281/zenodo.3937062" httpUri="https://zenodo.org/record/3937062/files/figure.png" pageId="4" pageNumber="179">13B</figureCitation>
); bulb (
<figureCitation id="130D42A73D0AFFCAFE21F61C96CAFE98" box="[394,480,297,323]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="4" pageNumber="179">Fig. 2C</figureCitation>
) with long hooked apophysis and simple embolus. Retrolateral trichobothria of tibia I at 10% proximally; legs with short vertical setae on metatarsi, without spines and curved setae; tarsus I with 24 distinct pseudosegments.
</paragraph>
<caption id="DF490EAA3D0CFFCCFF3CF05092BCF831" ID-DOI="http://doi.org/10.5281/zenodo.3937038" ID-Zenodo-Dep="3937038" httpUri="https://zenodo.org/record/3937038/files/figure.png" pageId="2" pageNumber="177" startId="2.[151,250,1893,1918]" targetBox="[220,1367,179,1865]" targetPageId="2">
<paragraph id="8B895E223D0CFFCCFF3CF05092BCF831" blockId="2.[151,1437,1893,2026]" pageId="2" pageNumber="177">
<emphasis id="B94282303D0CFFCCFF3CF050963EF8A5" bold="true" box="[151,276,1893,1918]" pageId="2" pageNumber="177">FIGURE 1.</emphasis>
<taxonomicName id="4C3625A13D0CFFCCFEB0F053969CF8A5" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[283,438,1894,1918]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="2" pageNumber="177" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B94282303D0CFFCCFEB0F053969CF8A5" box="[283,438,1894,1918]" italics="true" pageId="2" pageNumber="177">Belisana guilin</emphasis>
</taxonomicName>
<emphasis id="B94282303D0CFFCCFE16F0539525F8A5" bold="true" box="[445,527,1894,1918]" pageId="2" pageNumber="177">
<taxonomicNameLabel id="A2713F4B3D0CFFCCFE16F0539525F8A5" box="[445,527,1894,1918]" pageId="2" pageNumber="177" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male. AB. Pedipalp (A. Prolateral view; B. Retrolateral view, arrow points at proximo-dorsal hump on pedipalpal femur); CD. Distal part of procursus (C. Prolateral view, arrow 1 points at disto-prolateral membranous lamella, arrow 2 points at small subdisto-dorsal sclerite, arrow 3 points at large subdisto-prolateral sclerite; D. Retrolateral view). b = bulb, ba = bulbal apophysis, e = embolus, f = flap, pr = procursus. Scale bars: 0.05 (AB), 0.02 (CD).
</paragraph>
</caption>
<caption id="DF490EAA3D0DFFCDFF3CF043936FF820" ID-DOI="http://doi.org/10.5281/zenodo.3937040" ID-Zenodo-Dep="3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="3" pageNumber="178" startId="3.[151,250,1910,1935]" targetBox="[214,1373,181,1885]" targetPageId="3">
<paragraph id="8B895E223D0DFFCDFF3CF043936FF820" blockId="3.[151,1436,1910,2043]" pageId="3" pageNumber="178">
<emphasis id="B94282303D0DFFCDFF3CF043963FF854" bold="true" box="[151,277,1910,1935]" pageId="3" pageNumber="178">FIGURE 2.</emphasis>
<taxonomicName id="4C3625A13D0DFFCDFEB5F0429691F854" authority="Zhu &amp; Yao &amp; Zheng &amp; Li, 2020" authorityName="Zhu &amp; Yao &amp; Zheng &amp; Li" authorityYear="2020" box="[286,443,1911,1935]" class="Arachnida" family="Pholcidae" genus="Belisana" kingdom="Animalia" order="Araneae" pageId="3" pageNumber="178" phylum="Arthropoda" rank="species" species="guilin" status="sp. nov.">
<emphasis id="B94282303D0DFFCDFEB5F0429691F854" box="[286,443,1911,1935]" italics="true" pageId="3" pageNumber="178">Belisana guilin</emphasis>
</taxonomicName>
<emphasis id="B94282303D0DFFCDFE6FF042953DF854" bold="true" box="[452,535,1911,1935]" pageId="3" pageNumber="178">
<taxonomicNameLabel id="A2713F4B3D0DFFCDFE6FF042953DF854" box="[452,535,1911,1935]" pageId="3" pageNumber="178" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH).
</paragraph>
</caption>
<paragraph id="8B895E223D0AFFCAFF6CF6A093A6FDB8" blockId="4.[151,1437,153,755]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF6A09634FE74" bold="true" box="[199,286,405,431]" pageId="4" pageNumber="179">Female</emphasis>
(IZCAS Ar39848): Similar to male, habitus as in
<figureCitation id="130D42A73D0AFFCAFCCBF6A0948DFE74" box="[864,935,405,431]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="4" pageNumber="179">Figs 2</figureCitation>
GH. Total length 1.50 (1.66 with clypeus), carapace 0.56 long, 0.63 wide, opisthosoma 0.94 long, 0.94 wide. Tibia I: 2.88; tibia I L/d: 58. Distance PME-PME 0.08, diameter PME 0.04, distance PME-ALE 0.03, AME absent. Sternum approximately as wide as long (0.47). External female genitalia (
<figureCitation id="130D42A73D0AFFCAFE17F5349532FDC0" box="[444,536,513,539]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="4" pageNumber="179">Figs 2A</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFD89F534957EFDC0" box="[546,596,513,539]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="4" pageNumber="179">14A</figureCitation>
) simple and flat, with pair of frontal “epigynal” pockets postero-laterally on scletotized plate (arrow ep in
<figureCitation id="130D42A73D0AFFCAFE48F5109502FDE4" box="[483,552,549,575]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="4" pageNumber="179">Figs 2</figureCitation>
AB, 14AB), 0.28 apart. Vulva (
<figureCitation id="130D42A73D0AFFCAFC09F51094D4FDE4" box="[930,1022,549,575]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="4" pageNumber="179">Figs 2B</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFBA1F5109313FDE4" box="[1034,1081,549,575]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="4" pageNumber="179">14B</figureCitation>
) with roof-shaped anterior arch and pair of long, narrow pore plates (arrow pp in
<figureCitation id="130D42A73D0AFFCAFD6BF57C9431FDB8" box="[704,795,585,611]" captionStart="FIGURE 2" captionStartId="3.[151,250,1910,1935]" captionTargetBox="[214,1373,181,1885]" captionTargetId="figure@3.[214,1373,181,1885]" captionTargetPageId="3" captionText="FIGURE 2. Belisana guilin sp. nov., holotype male (CF) and paratype female (AB, GH). A. External female genitalia, ventral view; B. Vulva, dorsal view; C. Bulb, prolateral view; D. Chelicerae, frontal view; EH. Habitus (E, G. Dorsal view; F. Lateral view; H. Ventral view). b = bulb, ba = bulbal apophysis, da = distal apophysis, e = embolus, ep = frontal “epigynal” pocket, pa = proximo-lateral apophysis, pp = pore plate. Scale bars: 0.05 (AD), 0.30 (EH)." figureDoi="http://doi.org/10.5281/zenodo.3937040" httpUri="https://zenodo.org/record/3937040/files/figure.png" pageId="4" pageNumber="179">Figs 2B</figureCitation>
,
<figureCitation id="130D42A73D0AFFCAFC8DF57C9472FDB8" box="[806,856,585,611]" captionStart="FIGURE 14" captionStartId="19.[151,250,1945,1970]" captionTargetBox="[202,1387,196,1913]" captionTargetId="figure@19.[193,1395,181,1919]" captionTargetPageId="19" captionText="FIGURE 14. Drawings of female genitalia in ventral and dorsal views. AB. B. guilin sp. nov.; CD. B. maoer sp. nov., arrow points at strongly protruding median ridge of anterior arch; EF. B. naling sp. nov.; GH. B. xiyuan sp. nov. ep = frontal “epigynal” pocket, pp = pore plate. Scale bars: 0.10." figureDoi="http://doi.org/10.5281/zenodo.3937064" httpUri="https://zenodo.org/record/3937064/files/figure.png" pageId="4" pageNumber="179">14B</figureCitation>
), without serrated sclerites.
</paragraph>
<paragraph id="8B895E223D0AFFCAFF6CF5599661FD70" blockId="4.[151,1437,153,755]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF559966BFD5C" bold="true" box="[199,321,620,647]" pageId="4" pageNumber="179">Variation:</emphasis>
<materialsCitation id="3B5E547F3D0AFFCAFEE7F5589662FD70" collectionCode="IZCAS" collectorName="Tibia I" pageId="4" pageNumber="179" specimenCount="2" typeStatus="paratype">
<collectorName id="26C33BF43D0AFFCAFEE7F55896B1FD5C" box="[332,411,621,647]" pageId="4" pageNumber="179">Tibia I</collectorName>
in male
<typeStatus id="548DE0803D0AFFCAFDA7F5589546FD5C" box="[524,620,621,647]" pageId="4" pageNumber="179" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED27C6E73D0AFFCAFD2BF55895F9FD5C" box="[640,723,621,647]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="179">IZCAS</collectionCode>
Ar39847): 3.98.
<collectorName id="26C33BF43D0AFFCAFC0AF55894DAFD5C" box="[929,1008,621,647]" pageId="4" pageNumber="179">Tibia I</collectorName>
in another female
<typeStatus id="548DE0803D0AFFCAFB7DF558921CFD5C" box="[1238,1334,621,647]" pageId="4" pageNumber="179" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED27C6E73D0AFFCAFAE1F55892B7FD5C" box="[1354,1437,621,647]" country="China" httpUri="http://grbio.org/cool/crjh-4ne1" name="Institute of Zoology, Chinese Academy of Sciences" pageId="4" pageNumber="179">IZCAS</collectionCode>
Ar39849): 2.88
</materialsCitation>
.
</paragraph>
<paragraph id="8B895E223D0AFFCAFF6CF58093DAFD14" blockId="4.[151,1437,153,755]" box="[199,1264,692,719]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF58096AFFD14" bold="true" box="[199,389,692,719]" pageId="4" pageNumber="179">Natural history.</emphasis>
The species was found underside of rocks in the aphotic zone inside the cave.
</paragraph>
</subSubSection>
<subSubSection id="C32C0DA93D0AFFCAFF6CF5ED9471FD28" box="[199,859,728,755]" pageId="4" pageNumber="179" type="distribution">
<paragraph id="8B895E223D0AFFCAFF6CF5ED9471FD28" blockId="4.[151,1437,153,755]" box="[199,859,728,755]" pageId="4" pageNumber="179">
<emphasis id="B94282303D0AFFCAFF6CF5ED964BFD28" bold="true" box="[199,353,728,755]" pageId="4" pageNumber="179">Distribution.</emphasis>
<collectingCountry id="F3211EB23D0AFFCAFECCF5EC9680FD28" box="[359,426,729,755]" name="China" pageId="4" pageNumber="179">China</collectingCountry>
(
<collectingRegion id="49F290C03D0AFFCAFE12F5EC9534FD28" box="[441,542,729,755]" country="China" name="Guangxi" pageId="4" pageNumber="179">Guangxi</collectingRegion>
,
<typeStatus id="548DE0803D0AFFCAFD82F5EC9573FD28" box="[553,601,729,755]" pageId="4" pageNumber="179">type</typeStatus>
locality only;
<figureCitation id="130D42A73D0AFFCAFD57F5EC9464FD28" box="[764,846,728,755]" captionStart="FIGURE 15" captionStartId="20.[151,250,1181,1206]" captionTargetBox="[151,1436,181,1156]" captionTargetId="figure@20.[151,1436,181,1156]" captionTargetPageId="20" captionText="FIGURE 15. Distribution records of six new species of the genus Belisana from southern China. 1. B. guilin sp. nov.; 2. B. maoer sp. nov.; 3. B. naling sp. nov.; 4. B. xiyuan sp. nov.; 5. B. xuanguan sp. nov.; 6. B. yuexiu sp. nov." figureDoi="http://doi.org/10.5281/zenodo.3937066" httpUri="https://zenodo.org/record/3937066/files/figure.png" pageId="4" pageNumber="179">Fig. 15</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>