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<document id="49CC5435D245D3DBD137DC74E77BDA04" ID-CLB-Dataset="7956" ID-DOI="10.11646/zootaxa.4878.1.11" ID-GBIF-Dataset="f6a0a87c-cb08-43d3-a804-cd6e6d44f95f" ID-ISSN="1175-5326" ID-Zenodo-Dep="4424692" ID-ZooBank="3F87D306-1CDD-48A3-8236-5F0F809F26ED" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1610034191052" checkinUser="plazi" docAuthor="Chen, Xin-Yu &amp; Zhang, Hua-Chuan" docDate="2020" docId="039F87CBF43AFFFDFF4AFD26FE6A29C2" docLanguage="en" docName="zootaxa.4878.1.11.pdf" docOrigin="Zootaxa 4878 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Adamas hukawngensis Chen &amp; Zhang 2020, sp. nov." docType="treatment" docVersion="7" lastPageNumber="196" masterDocId="FFA6FFB3F43BFFFCFFDDFF95FFC82E1F" masterDocTitle="A new fossil record of Lymexylidae (Insecta: Coleoptera) from mid-Cretaceous amber of northern Myanmar" masterLastPageNumber="200" masterPageNumber="195" pageNumber="196" updateTime="1698923531004" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="FC229999041EB9C1FBF12263075078C3">A new fossil record of Lymexylidae (Insecta: Coleoptera) from mid-Cretaceous amber of northern Myanmar</mods:title>
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<mods:namePart id="941D7627C11F87A5119E4D5FAC2CD760">Chen, Xin-Yu</mods:namePart>
<mods:affiliation id="9663A4758F4B0E59ECEC4B7975F259E3">Cores and Samples Center of Land &amp; Resources (CSCLR), Sanhe, 065201, Hebei, China</mods:affiliation>
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<mods:namePart id="D81063551B013A570F07248B9494FDA0">Zhang, Hua-Chuan</mods:namePart>
<mods:nameIdentifier id="80F2E6814C5AE865B42E0A6F8BD0C32E" type="ORCID">0000-0002-6870-5395</mods:nameIdentifier>
<mods:affiliation id="00B37FEA1E70B83927EC4EE6990A1EA7">Cores and Samples Center of Land &amp; Resources (CSCLR), Sanhe, 065201, Hebei, China &amp; 250247883 @ qq. com; https: // orcid. org / 0000 - 0002 - 6870 - 5395</mods:affiliation>
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<mods:title id="4F13BF1C6E2827407FE095F33A2F4CA1">Zootaxa</mods:title>
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<mods:date id="73B9F8F3BD6239DCA081B23C586B14CD">2020</mods:date>
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<treatment id="039F87CBF43AFFFDFF4AFD26FE6A29C2" ID-DOI="http://doi.org/10.5281/zenodo.4570463" ID-GBIF-Taxon="179983497" ID-Zenodo-Dep="4570463" LSID="urn:lsid:plazi:treatment:039F87CBF43AFFFDFF4AFD26FE6A29C2" httpUri="http://treatment.plazi.org/id/039F87CBF43AFFFDFF4AFD26FE6A29C2" lastPageNumber="196" pageId="1" pageNumber="196">
<subSubSection id="C32C6556F43AFFFDFF4AFD26FDE02CD2" box="[151,552,691,718]" pageId="1" pageNumber="196" type="nomenclature">
<paragraph id="8B8936DDF43AFFFDFF4AFD26FDE02CD2" blockId="1.[151,552,691,753]" box="[151,552,691,718]" pageId="1" pageNumber="196">
<heading id="D0C181B1F43AFFFDFF4AFD26FDE02CD2" bold="true" box="[151,552,691,718]" fontSize="11" level="1" pageId="1" pageNumber="196" reason="1">
<emphasis id="B942EACFF43AFFFDFF4AFD26FDE02CD2" bold="true" box="[151,552,691,718]" pageId="1" pageNumber="196">
<taxonomicName id="4C364D5EF43AFFFDFF70FD26FE0B2CD1" authority="Chen &amp; Zhang, 2020" authorityName="Chen &amp; Zhang" authorityYear="2020" box="[173,451,691,718]" family="Lymexylidae" genus="Adamas" higherTaxonomySource="CoL" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Chordata" rank="species" species="hukawngensis" status="sp. nov.">
<emphasis id="B942EACFF43AFFFDFF70FD26FE0B2CD1" bold="true" box="[173,451,691,718]" italics="true" pageId="1" pageNumber="196">Adamas hukawngensis</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A27157B4F43AFFFDFE17FD26FDE02CD2" box="[458,552,691,717]" pageId="1" pageNumber="196" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="C32C6556F43AFFFDFF4AFD42FEC32CEE" box="[151,267,727,753]" pageId="1" pageNumber="196" type="description">
<paragraph id="8B8936DDF43AFFFDFF4AFD42FEC32CEE" blockId="1.[151,552,691,753]" box="[151,267,727,753]" pageId="1" pageNumber="196">
(
<figureCitation id="130D2A58F43AFFFDFF42FD42FECA2CEE" box="[159,258,727,753]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,250,918,942]" captionStartId-1="3.[151,250,1805,1829]" captionStartId-2="4.[151,250,1810,1834]" captionTargetBox-0="[245,1342,181,895]" captionTargetBox-1="[290,1282,181,1780]" captionTargetBox-2="[380,1231,188,1781]" captionTargetId-0="figure-21@2.[245,1342,181,897]" captionTargetId-1="figure-19@3.[290,1298,181,1781]" captionTargetId-2="figure-21@4.[348,1239,179,1785]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Adamas hukawngensis, holotype (CSCLRCAB100180). A. habitus, dorsal view. B. habitus, ventral view. Scale bars: 1.0 mm." captionText-1="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." captionText-2="FIGURE 3. Adamas hukawngensis, holotype (CSCLRCAB100180). A. Metendosternite. B. Image of metathoracic wing. C. Drawing of metathoracic wing. D. Image of male genitalia, dorsal view. E. Drawing of male genitalia, dorsal view. F. Image of male genitalia, lateral view. G. Drawing of male genitalia, lateral view. Abbreviations: ll = lateral lobes; ml = median lobe. Scale bars: AC = 1.0 mm; DG = 0.2 mm." figureDoi-0="http://doi.org/10.5281/zenodo.4424694" figureDoi-1="http://doi.org/10.5281/zenodo.4424698" figureDoi-2="http://doi.org/10.5281/zenodo.4424700" httpUri-0="https://zenodo.org/record/4424694/files/figure.png" httpUri-1="https://zenodo.org/record/4424698/files/figure.png" httpUri-2="https://zenodo.org/record/4424700/files/figure.png" pageId="1" pageNumber="196">Figs 13</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32C6556F43AFFFDFF4AFC8AFDFA2D42" pageId="1" pageNumber="196" type="etymology">
<paragraph id="8B8936DDF43AFFFDFF4AFC8AFDFA2D42" blockId="1.[151,1437,799,2013]" pageId="1" pageNumber="196">
<emphasis id="B942EACFF43AFFFDFF4AFC8AFED72D26" bold="true" box="[151,287,799,825]" pageId="1" pageNumber="196">Etymology.</emphasis>
The specific epithet is derived from the Hukawng Valley in northern
<collectingCountry id="F321764DF43AFFFDFBFFFCB5FB472D25" box="[1058,1167,800,826]" name="Myanmar" pageId="1" pageNumber="196">Myanmar</collectingCountry>
and is a reference to the location discovered of the specimen.
</paragraph>
</subSubSection>
<subSubSection id="C32C6556F43AFFFDFF1AFCF2FBCC2D9E" box="[199,1028,871,898]" pageId="1" pageNumber="196" type="materials_examined">
<paragraph id="8B8936DDF43AFFFDFF1AFCF2FBCC2D9E" blockId="1.[151,1437,799,2013]" box="[199,1028,871,898]" pageId="1" pageNumber="196">
<emphasis id="B942EACFF43AFFFDFF1AFCF2FE212D9E" bold="true" box="[199,489,871,897]" pageId="1" pageNumber="196">
Type material.
<typeStatus id="548D887FF43AFFFDFEA1FCF2FE212D9E" box="[380,489,871,897]" pageId="1" pageNumber="196" type="holotype">Holotype</typeStatus>
</emphasis>
: male, in Burmese amber, CSCLRCAB100180.
</paragraph>
</subSubSection>
<subSubSection id="C32C6556F43AFFFDFF1AFC1EFDCF2DB9" box="[199,519,907,934]" pageId="1" pageNumber="196" type="diagnosis">
<paragraph id="8B8936DDF43AFFFDFF1AFC1EFDCF2DB9" blockId="1.[151,1437,799,2013]" box="[199,519,907,934]" pageId="1" pageNumber="196">
<emphasis id="B942EACFF43AFFFDFF1AFC1EFE8A2DBA" bold="true" box="[199,322,907,933]" pageId="1" pageNumber="196">Diagnosis.</emphasis>
As for the genus.
</paragraph>
</subSubSection>
<subSubSection id="C32C6556F43AFFFDFF1AFC3AFDB12886" pageId="1" pageNumber="196" type="description">
<paragraph id="8B8936DDF43AFFFDFF1AFC3AFDB12886" blockId="1.[151,1437,799,2013]" pageId="1" pageNumber="196">
<emphasis id="B942EACFF43AFFFDFF1AFC3AFE912DD6" bold="true" box="[199,345,943,969]" pageId="1" pageNumber="196">Description.</emphasis>
Male. Body length from anterior margin of frons to abdominal terminal 6.0 mm; elongate, nearly parallel-sided, moderately dorsoventrally flattened (
<figureCitation id="130D2A58F43AFFFDFD07FC46FCD32DF2" box="[730,795,979,1005]" captionStart="FIGURE 1" captionStartId="2.[151,250,918,942]" captionTargetBox="[245,1342,181,895]" captionTargetId="figure-21@2.[245,1342,181,897]" captionTargetPageId="2" captionText="FIGURE 1. Adamas hukawngensis, holotype (CSCLRCAB100180). A. habitus, dorsal view. B. habitus, ventral view. Scale bars: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424694" httpUri="https://zenodo.org/record/4424694/files/figure.png" pageId="1" pageNumber="196">Fig. 1</figureCitation>
); lightly sclerotized; clothed in fine setae; legs long, thin. Head (
<figureCitation id="130D2A58F43AFFFDFF3EFC62FEEF2A0E" box="[227,295,1015,1041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2</figureCitation>
AB): ovate, length about 1.5X width, narrowed behind eyes, forming &quot;neck&quot;; eyes (
<figureCitation id="130D2A58F43AFFFDFB29FC62FAF02A0E" box="[1268,1336,1015,1041]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2</figureCitation>
AB; e), entire and large, separated by less than single eye width, with stout intrafacetal setae; epicranial pit absent; antennae 11-segmented, with flagellum pectinate (
<figureCitation id="130D2A58F43AFFFDFDBBFBAAFD622A46" box="[614,682,1087,1113]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2</figureCitation>
CD), rami of segments 3-11 approximatly
<quantity id="4CCE9B38F43AFFFDFB43FBAAFACF2A45" box="[1182,1287,1087,1114]" metricMagnitude="-4" metricUnit="m" metricValue="6.8999999999999995" pageId="1" pageNumber="196" unit="mm" value="0.69">0.69 mm</quantity>
; scapus (
<figureCitation id="130D2A58F43AFFFDFAB2FBAAFF1D2A61" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2D; I</figureCitation>
) short, 3/4 as long as pedicellus (
<figureCitation id="130D2A58F43AFFFDFD80FBF6FD712A62" box="[605,697,1123,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2D</figureCitation>
; II); antennal segments IIIXI (
<figureCitation id="130D2A58F43AFFFDFBF7FBF6FB4D2A62" box="[1066,1157,1123,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2D</figureCitation>
; IIIXI) relatively long, almost equal in length; maxillary palp organ (
<figureCitation id="130D2A58F43AFFFDFD76FB12FCCB2ABE" box="[683,771,1159,1185]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2F</figureCitation>
; mp) present, multi-branched, with two primary branches and parallel secondary branches forming a basketlike structure; maxillary palpus 4-segmented, segment IV (
<figureCitation id="130D2A58F43AFFFDFAB2FB3EFF1B2AF5" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2F; p</figureCitation>
IV) elongate, with numerous apical sensilla. Thorax: pronotum length
<quantity id="4CCE9B38F43AFFFDFC20FB5AFBAD2AF5" box="[1021,1125,1231,1258]" metricMagnitude="-4" metricUnit="m" metricValue="9.5" pageId="1" pageNumber="196" unit="mm" value="0.95">0.95 mm</quantity>
, width
<quantity id="4CCE9B38F43AFFFDFB67FB5AFAEB2AF5" box="[1210,1315,1231,1258]" metricMagnitude="-4" metricUnit="m" metricValue="5.6" pageId="1" pageNumber="196" unit="mm" value="0.56">0.56 mm</quantity>
; pronotum slightly narrower than head, with mesal longitudinal depression, posterior and anterior widths equivalent (i.e., not constricted); scutellum (
<figureCitation id="130D2A58F43AFFFDFE7AFA82FDD02B2D" box="[423,536,1303,1330]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2H; s</figureCitation>
) lozenge-shaped, with longitudinal median groove. Legs: long, thin, with dense shallow punctures and short, fine setae; portion of prothoracic leg lost, tarsal formula?-5-5, tibial spur formula?-0- 0. Each elytron (
<figureCitation id="130D2A58F43AFFFDFE8CFACAFE5B2B66" box="[337,403,1375,1401]" captionStart="FIGURE 2" captionStartId="3.[151,250,1805,1829]" captionTargetBox="[290,1282,181,1780]" captionTargetId="figure-19@3.[290,1298,181,1781]" captionTargetPageId="3" captionText="FIGURE 2. Adamas hukawngensis, holotype (CSCLRCAB100180). A. dorsal image of head. B. Drawing of dorsal head. C. Image of antenna. D. Drawing of antenna. E. Image of maxillary palp organ. F. Drawing of maxillary palp organ. G. Image of elytron and scutellum. H. Drawing of elytron and scutellum. Abbreviations: 1º = primary branch; 2 º = secondary branch; I, II, III-XI = scapus, pedicellus, antennal segments IIIXI; e = eyes; mp = maxillary palporgan; p IV = palpomere IV; S = scutellum. Scale bars: AB = 0.1 mm; CH = 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4424698" httpUri="https://zenodo.org/record/4424698/files/figure.png" pageId="1" pageNumber="196">Fig. 2</figureCitation>
GH) but slightly shortened, not covering abdomen; lightly sclerotized, with dense, shallow punctures and long, fine setae. Each metathoracic wing (
<figureCitation id="130D2A58F43AFFFDFCCCFA16FC9A2B82" box="[785,850,1411,1437]" captionStart="FIGURE 3" captionStartId="4.[151,250,1810,1834]" captionTargetBox="[380,1231,188,1781]" captionTargetId="figure-21@4.[348,1239,179,1785]" captionTargetPageId="4" captionText="FIGURE 3. Adamas hukawngensis, holotype (CSCLRCAB100180). A. Metendosternite. B. Image of metathoracic wing. C. Drawing of metathoracic wing. D. Image of male genitalia, dorsal view. E. Drawing of male genitalia, dorsal view. F. Image of male genitalia, lateral view. G. Drawing of male genitalia, lateral view. Abbreviations: ll = lateral lobes; ml = median lobe. Scale bars: AC = 1.0 mm; DG = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4424700" httpUri="https://zenodo.org/record/4424700/files/figure.png" pageId="1" pageNumber="196">Fig. 3</figureCitation>
BC): large, membranous, not covered by elytron at rest, folded longitudinally only; venation only partially observed; C+Sc+R as well as r-m crossvein fragmentary, radius (R) fused with C + Sc at about middle; radial cell and radial crossvein absent. Metendosternite (
<figureCitation id="130D2A58F43AFFFDFB33FA5EFA802BFA" box="[1262,1352,1483,1509]" captionStart="FIGURE 3" captionStartId="4.[151,250,1810,1834]" captionTargetBox="[380,1231,188,1781]" captionTargetId="figure-21@4.[348,1239,179,1785]" captionTargetPageId="4" captionText="FIGURE 3. Adamas hukawngensis, holotype (CSCLRCAB100180). A. Metendosternite. B. Image of metathoracic wing. C. Drawing of metathoracic wing. D. Image of male genitalia, dorsal view. E. Drawing of male genitalia, dorsal view. F. Image of male genitalia, lateral view. G. Drawing of male genitalia, lateral view. Abbreviations: ll = lateral lobes; ml = median lobe. Scale bars: AC = 1.0 mm; DG = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4424700" httpUri="https://zenodo.org/record/4424700/files/figure.png" pageId="1" pageNumber="196">Fig. 3A</figureCitation>
): furcal stalk long, parallel sided. Abdomen: flat in form, with seventh sternite visible; tergites and sternites wider than long; segments IV-VII similar in form, VII smaller than VI; VII forms circumgenitalic capsule; length
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excluding genitalia, width at most
<quantity id="4CCE9B38F43AFFFDFE75F9A2FDC5284D" box="[424,525,1591,1618]" metricMagnitude="-4" metricUnit="m" metricValue="8.299999999999999" pageId="1" pageNumber="196" unit="mm" value="0.83">0.83 mm</quantity>
on IV segment. Male genitalia: lateral lobes (
<figureCitation id="130D2A58F43AFFFDFBCEF9A2FB7A284E" box="[1043,1202,1591,1617]" captionStart="FIGURE 3" captionStartId="4.[151,250,1810,1834]" captionTargetBox="[380,1231,188,1781]" captionTargetId="figure-21@4.[348,1239,179,1785]" captionTargetPageId="4" captionText="FIGURE 3. Adamas hukawngensis, holotype (CSCLRCAB100180). A. Metendosternite. B. Image of metathoracic wing. C. Drawing of metathoracic wing. D. Image of male genitalia, dorsal view. E. Drawing of male genitalia, dorsal view. F. Image of male genitalia, lateral view. G. Drawing of male genitalia, lateral view. Abbreviations: ll = lateral lobes; ml = median lobe. Scale bars: AC = 1.0 mm; DG = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4424700" httpUri="https://zenodo.org/record/4424700/files/figure.png" pageId="1" pageNumber="196">Fig. 3E &amp; 3G</figureCitation>
; ll) symmetrical; abdominal segment IX of male symmetrical; abdominal sternite VIII of male without large setose pads; median lobe (
<figureCitation id="130D2A58F43AFFFDFF7DF9EAFF312886" box="[160,249,1663,1689]" captionStart="FIGURE 3" captionStartId="4.[151,250,1810,1834]" captionTargetBox="[380,1231,188,1781]" captionTargetId="figure-21@4.[348,1239,179,1785]" captionTargetPageId="4" captionText="FIGURE 3. Adamas hukawngensis, holotype (CSCLRCAB100180). A. Metendosternite. B. Image of metathoracic wing. C. Drawing of metathoracic wing. D. Image of male genitalia, dorsal view. E. Drawing of male genitalia, dorsal view. F. Image of male genitalia, lateral view. G. Drawing of male genitalia, lateral view. Abbreviations: ll = lateral lobes; ml = median lobe. Scale bars: AC = 1.0 mm; DG = 0.2 mm." figureDoi="http://doi.org/10.5281/zenodo.4424700" httpUri="https://zenodo.org/record/4424700/files/figure.png" pageId="1" pageNumber="196">Fig. 3G</figureCitation>
; ml) large relative to lateral lobes.
</paragraph>
</subSubSection>
<subSubSection id="C32C6556F43AFFFDFF1AF936FE6A29C2" pageId="1" pageNumber="196" type="discussion">
<paragraph id="8B8936DDF43AFFFDFF1AF936FADF29A6" blockId="1.[151,1437,799,2013]" pageId="1" pageNumber="196">
<emphasis id="B942EACFF43AFFFDFF1AF936FEFC28A2" bold="true" box="[199,308,1699,1725]" pageId="1" pageNumber="196">Remarks</emphasis>
. The new species is distinguished from all recent and extinct genera of
<taxonomicName id="4C364D5EF43AFFFDFB8BF936FB2028A2" authorityName="Fleming" authorityYear="1821" box="[1110,1256,1699,1725]" class="Insecta" family="Lymexylidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="family">Lymexylidae</taxonomicName>
by the lozengeshaped scutellum with a longitudinal median groove (that of
<taxonomicName id="4C364D5EF43AFFFDFC97F95DFA9128FE" authority="(Wolf-Schwenninger, 2011)" baseAuthorityName="Wolf-Schwenninger" baseAuthorityYear="2011" box="[842,1369,1735,1762]" class="Insecta" family="Lymexylidae" genus="Cratoatractocerus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B942EACFF43AFFFDFC97F95DFBD128FD" box="[842,1049,1736,1762]" italics="true" pageId="1" pageNumber="196">Cratoatractocerus</emphasis>
(
<bibRefCitation id="EFA74B2CF43AFFFDFBF5F952FA9A28FE" author="Wolf-Schwenninger, K." box="[1064,1362,1735,1761]" pageId="1" pageNumber="196" pagination="205 - 212" refId="ref3149" refString="Wolf-Schwenninger, K. (2011) The oldest fossil record of Lymexylidae (Insecta: Coleoptera) from the Lower Cretaceous Crato formation of Brazil. Insect Systematics &amp; Evolution, 42 (2), 205 - 212. https: // doi. org / 10.1163 / 187631211 X 578352" type="journal article" year="2011">Wolf-Schwenninger, 2011</bibRefCitation>
)
</taxonomicName>
bears two longitudinal ridges plus a slightly grooved meson). The following characters are of diagnostic value: lengthened longitudinal oval-shaped head; pectinate antennae the same as “flabellate” morphs in
<taxonomicName id="4C364D5EF43AFFFDFB5DF89AFEA92952" authority="(Schneider, 1791)" baseAuthorityName="Schneider" baseAuthorityYear="1791" class="Insecta" family="Lymexylidae" genus="Hylecoetus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="species" species="flabellicornis">
<emphasis id="B942EACFF43AFFFDFB5DF89AFA542936" box="[1152,1436,1807,1833]" italics="true" pageId="1" pageNumber="196">Hylecoetus flabellicornis</emphasis>
(
<bibRefCitation id="EFA74B2CF43AFFFDFF42F8A6FE912952" author="Schneider, D. H." box="[159,345,1843,1869]" pageId="1" pageNumber="196" pagination="109" refId="ref2955" refString="Schneider, D. H. (1791) Anmerkung des Herausgebers. Neues Magazin fur die Liebhaber der Entomologie, 1, 109." type="journal article" year="1791">Schneider, 1791</bibRefCitation>
)
</taxonomicName>
but unlike all other lymexylids; huge eyes that occupy vast parts of the head surface and are strongly approximate dorsally (eyes large, ovate, conspicuous, occupying much of both the dorsal and ventral surfaces of the head in
<taxonomicName id="4C364D5EF43AFFFDFEABF8EEFDCF298A" box="[374,519,1915,1941]" class="Insecta" family="Lymexylidae" genus="Atractocerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B942EACFF43AFFFDFEABF8EEFDCF298A" box="[374,519,1915,1941]" italics="true" pageId="1" pageNumber="196">Atractocerus</emphasis>
</taxonomicName>
); each elytron slightly shortened, not covering all the parts of the abdomen (elytra very short, exposing most abdominal terga in
<taxonomicName id="4C364D5EF43AFFFDFD45F80AFCE129A6" box="[664,809,1951,1977]" class="Insecta" family="Lymexylidae" genus="Atractocerus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="196" phylum="Arthropoda" rank="genus">
<emphasis id="B942EACFF43AFFFDFD45F80AFCE129A6" box="[664,809,1951,1977]" italics="true" pageId="1" pageNumber="196">Atractocerus</emphasis>
</taxonomicName>
); large median lobe relative to lateral lobes.
</paragraph>
<paragraph id="8B8936DDF43AFFFDFF1AF856FE6A29C2" blockId="1.[151,1437,799,2013]" box="[199,418,1987,2013]" pageId="1" pageNumber="196">
<emphasis id="B942EACFF43AFFFDFF1AF856FEED29C2" bold="true" box="[199,293,1987,2013]" pageId="1" pageNumber="196">Female.</emphasis>
Unknown.
</paragraph>
</subSubSection>
</treatment>
</document>