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<document id="CDC8370F7CE2A09E28812AFEA67AFEA1" ID-DOI="10.11646/zootaxa.5389.2.3" ID-ISSN="1175-5326" ID-Zenodo-Dep="10406905" ID-ZooBank="F3FAC345-624C-4BE1-912F-E56EEB8EBE3B" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1703001578362" checkinUser="plazi" docAuthor="Zámbó, András, Kovács, Szilvia &amp; Kondorosy, Előd" docDate="2023" docId="03D287D0FFBFFFE8EF8DFF35036C823B" docLanguage="en" docName="zootaxa.5389.2.3.pdf" docOrigin="Zootaxa 5389 (2)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5389.2.3/52510" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Orbellis madagascariensis Zámbó &amp; Kovács &amp; Kondorosy 2023, comb. nov." docType="treatment" docVersion="2" lastPageNumber="200" masterDocId="FFEBFFA8FFB8FFEFEF1AFFA2016E8041" masterDocTitle="Revision of the genus Orbellis Distant, 1913 (Heteroptera: Rhyparochromidae)" masterLastPageNumber="212" masterPageNumber="193" pageNumber="200" updateTime="1703002295063" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="C7B1B84BD2BBBE93158C8D7F050F70EE">Revision of the genus Orbellis Distant, 1913 (Heteroptera: Rhyparochromidae)</mods:title>
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<mods:name id="2D0EC76BF0B0BE52993EC017A153BEBB" type="personal">
<mods:role id="2AA75A72F56F41295D098636FE4F1B1D">
<mods:roleTerm id="54CAABDBF70AE2DF9CD872CCA7DEB13E">Author</mods:roleTerm>
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<mods:namePart id="A28623539CB7CCC18A134653E690EFD4">Zámbó, András</mods:namePart>
<mods:affiliation id="EF5D491A150E50FA023CE9D9A9855069">Festetics Doctoral School, Hungarian University of Agriculture and Life Sciences, Georgikon Campus, str. Deák F. 16, Keszthely, H- 8360 Hungary</mods:affiliation>
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<mods:namePart id="CD20B9C5038594D8974CCBDE7BE323E2">Kovács, Szilvia</mods:namePart>
<mods:affiliation id="63DE766569A02AC3EE1BEE18A8D74A38">Department of Conservation Biology, Hungarian University of Agriculture and Life Sciences, Georgikon Campus, str. Deák F. 16, Keszthely, H- 8360 Hungary</mods:affiliation>
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</mods:role>
<mods:namePart id="8E1F5541D352BA5BF5ED1BA5AACF484D">Kondorosy, Előd</mods:namePart>
<mods:affiliation id="8EF120C3A4C986A0FEC68A1A0A9B2390">Department of Conservation Biology, Hungarian University of Agriculture and Life Sciences, Georgikon Campus, str. Deák F. 16, Keszthely, H- 8360 Hungary</mods:affiliation>
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<mods:title id="C18274E7BADCFEF9D50DC973FFD1B1E7">Zootaxa</mods:title>
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<mods:part id="422DA97A35C7A205186AE0B366052B48">
<mods:date id="788195ECCD53A3DBFC01DF73D77FFF8B">2023</mods:date>
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<mods:number id="E6D459C8974A1F186423DB2743DB5916">2023-12-19</mods:number>
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<mods:number id="D16EAC9D027E1531C5FE63EDFCD599FA">5389</mods:number>
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<mods:number id="009EEE9C1AE4391421BCE3671E076ABA">2</mods:number>
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<mods:identifier id="59CBF973720B25BEA1A18F05D4236FAC" type="DOI">10.11646/zootaxa.5389.2.3</mods:identifier>
<mods:identifier id="75CB671DEE20190C7943EDDC2F270B14" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="161B6E2D779D55FD39DFE708A3D5BC9A" type="Zenodo-Dep">10406905</mods:identifier>
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<treatment id="03D287D0FFBFFFE8EF8DFF35036C823B" LSID="urn:lsid:plazi:treatment:03D287D0FFBFFFE8EF8DFF35036C823B" httpUri="http://treatment.plazi.org/id/03D287D0FFBFFFE8EF8DFF35036C823B" lastPageNumber="200" pageId="7" pageNumber="200">
<subSubSection id="C361654DFFBFFFE8EF8DFF3502B480F0" box="[151,986,151,178]" pageId="7" pageNumber="200" type="nomenclature">
<paragraph id="8BC436C6FFBFFFE8EF8DFF3502B480F0" blockId="7.[151,1117,151,249]" box="[151,986,151,178]" pageId="7" pageNumber="200">
<heading id="D08C81AAFFBFFFE8EF8DFF3502B480F0" bold="true" box="[151,986,151,178]" fontSize="11" level="1" pageId="7" pageNumber="200" reason="1">
<emphasis id="B90FEAD4FFBFFFE8EF8DFF3502B480F0" bold="true" box="[151,986,151,178]" pageId="7" pageNumber="200">
<taxonomicName id="4C7B4D45FFBFFFE8EF8DFF35022880F0" authority="(Slater and O'Donnell, 1999)" authorityName="Zámbó &amp; Kovács &amp; Kondorosy" authorityYear="2023" baseAuthorityName="Slater and O'Donnell" baseAuthorityYear="1999" box="[151,838,151,178]" class="Insecta" family="Rhyparochromidae" genus="Orbellis" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="200" phylum="Arthropoda" rank="species" species="madagascariensis" status="comb. nov.">
<emphasis id="B90FEAD4FFBFFFE8EF8DFF3500B880F3" bold="true" box="[151,470,151,178]" italics="true" pageId="7" pageNumber="200">Orbellis madagascariensis</emphasis>
(
<bibRefCitation id="EFEA4B37FFBFFFE8EEFFFF35025380F0" author="Slater, J. A. &amp; O'Donnell, J. E." box="[485,829,151,177]" pageId="7" pageNumber="200" pagination="256 - 267" refId="ref19021" refString="Slater, J. A. &amp; O'Donnell, J. E. (1999) New species of Lethaeini from Madagascar (Heteroptera: Lygaeoidea: Rhyparochromidae). Journal of the New York Entomological Society, 107, 256 - 267." type="journal article" year="1999">Slater and O’Donnell, 1999</bibRefCitation>
)
</taxonomicName>
,
<taxonomicNameLabel id="A23C57AFFFBFFFE8EC49FF3502B480F0" box="[851,986,151,177]" pageId="7" pageNumber="200" rank="species">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C361654DFFBFFFE8EF8DFF19053380B8" pageId="7" pageNumber="200" type="description">
<paragraph id="8BC436C6FFBFFFE8EF8DFF1900248094" blockId="7.[151,1117,151,249]" box="[151,330,187,213]" pageId="7" pageNumber="200">
(
<figureCitation id="13402A43FFBFFFE8EF85FF1901818094" box="[159,239,187,213]" captionStart="FIGURES 1–11" captionStartId="6.[151,264,1313,1337]" captionTargetBox="[159,1433,352,1282]" captionTargetId="figure-91@6.[151,1436,345,1290]" captionTargetPageId="6" captionText="FIGURES 1–11. Dorsal views of Orbellis species: 1. O. typicus Distant, 1913 lectotype; 2. O. polhemi Slater &amp; O’Donnell, 1999; 3. O. maculosus Slater &amp; O’Donnell, 1999 holotype; 4. O. madagascariensis Slater &amp; O’Donnell, 1999; 5. O. calcaratus Zámbó &amp; Kondorosy, sp. nov. holotype; 6. O. dentifemoralis Zámbó &amp; Kondorosy, sp. nov. paratype; 7. O. odonnellae Zámbó &amp; Kondorosy, sp. nov. holotype; 8. O. flavescens Zámbó &amp; Kondorosy, sp. nov. holotype; 9. O. guilberti Zámbó &amp; Kondorosy, sp. nov. holotype; 10. O. nigriapicalis Zámbó &amp; Kondorosy, sp. nov. holotype; 11. O. longirostris Zámbó &amp; Kondorosy, sp. nov. holotype. (Fig. 1 by E. Kondorosy, Fig. 3 E. Guilbert, Figs. 2, 4–11 A. Zámbó)." figureDoi="http://doi.org/10.5281/zenodo.10406908" httpUri="https://zenodo.org/record/10406908/files/figure.png" pageId="7" pageNumber="200">Figs. 4</figureCitation>
,
<figureCitation id="13402A43FFBFFFE8EFE1FF1900778094" box="[251,281,187,213]" captionStart="FIGURES 12–22" captionStartId="13.[151,264,1709,1733]" captionTargetBox="[151,1429,390,1677]" captionTargetId="figure-90@13.[151,1436,381,1685]" captionTargetPageId="13" captionText="FIGURES 12–22. Male genitalia of Orbellis species: 12–15 pygophore (dorsal view); 16–22 left paramere (inner view): 16. O. madagascariensis; 12, 17. O. calcaratus; 13, 18. O. dentifemoralis; 14, 21. O. guilberti; 15, 22. O. nigriapicalis; 20. O. flavescens. (Figs. 12–15, 18 by Norbert Völgyi, 16–17, 19–22 Szilvia Kovács)." figureDoi="http://doi.org/10.5281/zenodo.10406912" httpUri="https://zenodo.org/record/10406912/files/figure.png" pageId="7" pageNumber="200">16</figureCitation>
,
<figureCitation id="13402A43FFBFFFE8EE3FFF19002C8094" box="[293,322,187,213]" captionStart="FIGURES 26–28" captionStartId="15.[151,264,1151,1175]" captionTargetBox="[161,1426,571,1118]" captionTargetId="figure-103@15.[151,1436,560,1127]" captionTargetPageId="15" captionText="FIGURES 26–28. Distribution of Orbellis species in Madagascar: 26. previously described Orbellis species (solid symbol: literature data; empty symbol: new data; no new data available for O. maculosus); 27. O. calcaratus and O. dentifemoralis; 28. other new Orbellis species." figureDoi="http://doi.org/10.5281/zenodo.10406916" httpUri="https://zenodo.org/record/10406916/files/figure.png" pageId="7" pageNumber="200">26</figureCitation>
)
</paragraph>
<paragraph id="8BC436C6FFBFFFE8EF8DFF7D053380B8" blockId="7.[151,1117,151,249]" box="[151,1117,223,249]" pageId="7" pageNumber="200">(http://lygaeoidea.speciesfile.org/Common/basic/Taxa.aspx?TaxonNameID=1236380)</paragraph>
</subSubSection>
<subSubSection id="C361654DFFBFFFE8EF8DFEBC04438177" box="[151,1325,286,310]" pageId="7" pageNumber="200" type="reference_group">
<paragraph id="8BC436C6FFBFFFE8EF8DFEBC04438177" blockId="7.[151,1325,286,310]" box="[151,1325,286,310]" pageId="7" pageNumber="200">
<treatmentCitationGroup id="AB6B11E8FFBFFFE8EF8DFEBC04438177" box="[151,1325,286,310]" pageId="7" pageNumber="200">
<treatmentCitation id="0ADA10D7FFBFFFE8EF8DFEBC02388177" author="Slater, J. A. &amp; O'Donnell, J. E." box="[151,854,286,310]" page="263" pageId="7" pageNumber="200" year="1999">
<taxonomicName id="4C7B4D45FFBFFFE8EF8DFEBC02388177" ID-CoL="46KS4" authority="Slater and O'Donnell, 1999: 261 - 263" authorityName="Slater and O'Donnell" authorityPageNumber="261 - 263" authorityYear="1999" box="[151,854,286,310]" class="Insecta" family="Rhyparochromidae" genus="Neolethaeus" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="200" phylum="Arthropoda" rank="species" species="madagascariensis">
<emphasis id="B90FEAD4FFBFFFE8EF8DFEBC00BC8177" box="[151,466,286,310]" italics="true" pageId="7" pageNumber="200">Neolethaeus madagascariensis</emphasis>
<bibRefCitation id="EFEA4B37FFBFFFE8EEC2FEBC02388177" author="Slater, J. A. &amp; O'Donnell, J. E." box="[472,854,286,310]" pageId="7" pageNumber="200" pagination="256 - 267" refId="ref19021" refString="Slater, J. A. &amp; O'Donnell, J. E. (1999) New species of Lethaeini from Madagascar (Heteroptera: Lygaeoidea: Rhyparochromidae). Journal of the New York Entomological Society, 107, 256 - 267." type="journal article" year="1999">Slater and O’Donnell, 1999: 261–263</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(original description, figure of male genitalia)
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C361654DFFBFFFE8EF8DFEC7031C824E" pageId="7" pageNumber="200" type="materials_examined">
<paragraph id="8BC436C6FFBFFFE8EF8DFEC7023481E2" blockId="7.[151,1437,356,635]" pageId="7" pageNumber="200">
<emphasis id="B90FEAD4FFBFFFE8EF8DFEC700A9813F" bold="true" box="[151,455,356,383]" pageId="7" pageNumber="200">Type specimen examined.</emphasis>
<materialsCitation id="3B133C9BFFBFFFE8EED7FEC7023481E2" collectionCode="MNHN" country="Madagascar" county="Antsingy Forest" location="Central" municipality="Melaky Region" pageId="7" pageNumber="200" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<typeStatus id="54C08864FFBFFFE8EED7FEC7030B813E" box="[461,613,357,383]" pageId="7" pageNumber="200" type="holotype">HOLOTYPE</typeStatus>
: label picture not taken; see data in original description (
<specimenCount id="9D7DFD4FFFBFFFE8EBCDFEC70580813E" box="[1239,1262,357,383]" count="1" pageId="7" pageNumber="200" type="male">♂</specimenCount>
,
<collectionCode id="ED6AAE03FFBFFFE8EBE3FEC7043A813E" box="[1273,1364,357,383]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" lsid="urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="7" pageNumber="200">MNHN</collectionCode>
). Type locality.
<location id="8EA4601DFFBFFFE8EFEDFE2A002781E3" LSID="urn:lsid:plazi:treatment:03D287D0FFBFFFE8EF8DFF35036C823B:8EA4601DFFBFFFE8EFEDFE2A002781E3" box="[247,329,392,418]" country="Madagascar" county="Antsingy Forest" municipality="Melaky Region" name="Central" pageId="7" pageNumber="200">Central</location>
<collectingCountry id="F36C7656FFBFFFE8EE4BFE2A00B481E3" box="[337,474,392,418]" name="Madagascar" pageId="7" pageNumber="200">Madagascar</collectingCountry>
,
<collectingMunicipality id="6BA0ACBCFFBFFFE8EEFFFE2A03FB81E3" box="[485,661,392,418]" pageId="7" pageNumber="200">Melaky Region</collectingMunicipality>
,
<collectingCounty id="62A54E4AFFBFFFE8ED85FE2A023881E2" box="[671,854,392,419]" pageId="7" pageNumber="200">Antsingy Forest</collectingCounty>
.
</materialsCitation>
</paragraph>
<paragraph id="8BC436C6FFBFFFE8EFDDFE0E031C824E" blockId="7.[151,1437,356,635]" pageId="7" pageNumber="200">
<emphasis id="B90FEAD4FFBFFFE8EFDDFE0E03608187" bold="true" box="[199,526,428,454]" pageId="7" pageNumber="200">Other specimens examined.</emphasis>
<materialsCitation id="3B133C9BFFBFFFE8ED0EFE0E01AE81AA" collectionCode="EHIA" collectorName="V. Dolin" country="Madagascar" location="Madagascar" pageId="7" pageNumber="200" specimenCount="1" specimenCount-male="1">
<collectingCountry id="F36C7656FFBFFFE8ED0EFE0E03F58187" box="[532,667,428,454]" name="Madagascar" pageId="7" pageNumber="200">Madagascar</collectingCountry>
/ Ankarafantsika / prot. area,
<date id="FFC51006FFBFFFE8ECF8FE0E051B8186" box="[994,1141,428,455]" pageId="7" pageNumber="200" value="2001-12-10">10.xii. / 2001</date>
<collectorName id="268E5310FFBFFFE8EB61FE0F05B68187" box="[1147,1240,428,455]" pageId="7" pageNumber="200">V. Dolin</collectorName>
lgt. (
<specimenCount id="9D7DFD4FFFBFFFE8EA0EFE0F04538186" box="[1300,1341,429,455]" count="1" pageId="7" pageNumber="200" type="male">1 ♂</specimenCount>
,
<collectionCode id="ED6AAE03FFBFFFE8EA52FE0F04FE8186" box="[1352,1424,429,455]" pageId="7" pageNumber="200">EHIA</collectionCode>
); SW
</materialsCitation>
<materialsCitation id="3B133C9BFFBFFFE8EFDDFE720300824E" collectionCode="MMBC" collectorName="Zombitse N. P. &amp; M. Tryzna" country="Madagascar" elevation="816" latitude="-22.886389" location="Ambakintany forest" longLatPrecision="21" longitude="44.691944" pageId="7" pageNumber="200" specimenCount="18" specimenCount-female="14" specimenCount-male="4">
<collectingCountry id="F36C7656FFBFFFE8EFDDFE72002081AB" box="[199,334,464,490]" name="Madagascar" pageId="7" pageNumber="200">Madagascar</collectingCountry>
2013 /
<collectorName id="268E5310FFBFFFE8EEB8FE72032681AA" box="[418,584,464,491]" pageId="7" pageNumber="200">Zombitse N. P.</collectorName>
(main entr.) /
<location id="8EA4601DFFBFFFE8EDF9FE7202AB81AB" LSID="urn:lsid:plazi:treatment:03D287D0FFBFFFE8EF8DFF35036C823B:8EA4601DFFBFFFE8EDF9FE7202AB81AB" box="[739,965,464,490]" country="Madagascar" latitude="-22.886389" longLatPrecision="21" longitude="44.691944" name="Ambakintany forest" pageId="7" pageNumber="200">Ambakintany forest</location>
,
<quantity id="4C839B23FFBFFFE8ECD5FE72056081AB" box="[975,1038,464,490]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="7" pageNumber="200" unit="m" value="816.0">
<elevation id="0056D1F5FFBFFFE8ECD5FE72056081AB" box="[975,1038,464,490]" metricMagnitude="2" metricUnit="m" metricValue="8.16" pageId="7" pageNumber="200" unit="m" value="816.0">816m</elevation>
</quantity>
/
<geoCoordinate id="EE4F5001FFBFFFE8EB39FE7205C881AA" box="[1059,1190,464,491]" degrees="22" direction="south" minutes="53" orientation="latitude" pageId="7" pageNumber="200" precision="15" seconds="11" value="-22.886389">S22°53’11”</geoCoordinate>
<geoCoordinate id="EE4F5001FFBFFFE8EBB7FE73045D81AA" box="[1197,1331,465,491]" degrees="44" direction="east" minutes="41" orientation="longitude" pageId="7" pageNumber="200" precision="15" seconds="31" value="44.691944">E44°41’31”</geoCoordinate>
/ 24.–27. i.
<collectorName id="268E5310FFBFFFE8EFB7FE57004F824E" box="[173,289,501,527]" pageId="7" pageNumber="200">M. Trýzna</collectorName>
leg. (
<specimenCount id="9D7DFD4FFFBFFFE8EE7BFE5700CD824E" box="[353,419,501,527]" count="4" pageId="7" pageNumber="200" type="male">4 ♂♂</specimenCount>
,
<specimenCount id="9D7DFD4FFFBFFFE8EEB5FE570093824E" box="[431,509,501,527]" count="14" pageId="7" pageNumber="200" type="female">14 ♀♀</specimenCount>
,
<collectionCode id="ED6AAE03FFBFFFE8ED10FE570308824E" box="[522,614,501,527]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="7" pageNumber="200">MMBC</collectionCode>
)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C361654DFFBFFFE8EFDDFDBB036C823B" pageId="7" pageNumber="200" type="discussion">
<paragraph id="8BC436C6FFBFFFE8EFDDFDBB036C823B" blockId="7.[151,1437,356,635]" pageId="7" pageNumber="200">
<emphasis id="B90FEAD4FFBFFFE8EFDDFDBB005A8272" bold="true" box="[199,308,537,563]" pageId="7" pageNumber="200">Remarks</emphasis>
. Paramere (
<figureCitation id="13402A43FFBFFFE8EEADFDBA03698273" box="[439,519,536,562]" captionStart="FIGURES 12–22" captionStartId="13.[151,264,1709,1733]" captionTargetBox="[151,1429,390,1677]" captionTargetId="figure-90@13.[151,1436,381,1685]" captionTargetPageId="13" captionText="FIGURES 12–22. Male genitalia of Orbellis species: 12–15 pygophore (dorsal view); 16–22 left paramere (inner view): 16. O. madagascariensis; 12, 17. O. calcaratus; 13, 18. O. dentifemoralis; 14, 21. O. guilberti; 15, 22. O. nigriapicalis; 20. O. flavescens. (Figs. 12–15, 18 by Norbert Völgyi, 16–17, 19–22 Szilvia Kovács)." figureDoi="http://doi.org/10.5281/zenodo.10406912" httpUri="https://zenodo.org/record/10406912/files/figure.png" pageId="7" pageNumber="200">Fig. 16</figureCitation>
) is subtrapezoidal, outer projection rounded basally, continuing to apex in a straight line, inner projection short, very widely rounded, inner margin between projection and base straight, nearly parallel with apical half of outer margin.
</paragraph>
</subSubSection>
</treatment>
</document>
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