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<document id="428774AC69ED9D3809D59424B30E7F94" ID-ISSN="1175­5334" IM.materialsCitations_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629589942712" checkinUser="felipe" docAuthor="Vaz-De-Mello, Fernando Z." docDate="2008" docId="03D287CEFFABAF3AD1DBD9C3FE1A1571" docLanguage="en" docName="zt01955p075.pdf" docOrigin="Zootaxa 1955" docStyle="DocumentStyle:C4791ACFF01C5C140D2C1A203DB3C9B0.4:Zootaxa.2007-2008.monograph" docStyleId="C4791ACFF01C5C140D2C1A203DB3C9B0" docStyleName="Zootaxa.2007-2008.monograph" docStyleVersion="4" docTitle="Leotrichillum Vaz-De-Mello 2008, new genus" docType="treatment" docVersion="2" lastPageNumber="33" masterDocId="FFEBFFB6FFB4AF1AD14CDF74FFA61358" masterDocTitle="Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus" masterLastPageNumber="75" masterPageNumber="1" pageNumber="32" updateTime="1699081972826" updateUser="plazi" zenodo-license-document="CLOSED">
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<mods:title id="FD259F08761F99DCF3F9755126F0A988">Synopsis of the new subtribe Scatimina (Coleoptera: Scarabaeidae: Scarabaeinae: Ateuchini), with descriptions of twelve new genera and review of Genieridium, new genus</mods:title>
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<mods:namePart id="CA57BB43515C33D3BA9A0C56D2CD1761">Vaz-De-Mello, Fernando Z.</mods:namePart>
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<mods:date id="743E63985068B56DB962095C609ACD60">2008</mods:date>
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<treatment id="03D287CEFFABAF3AD1DBD9C3FE1A1571" LSID="urn:lsid:plazi:treatment:03D287CEFFABAF3AD1DBD9C3FE1A1571" httpUri="http://treatment.plazi.org/id/03D287CEFFABAF3AD1DBD9C3FE1A1571" lastPageId="32" lastPageNumber="33" pageId="31" pageNumber="32">
<subSubSection id="C3616553FFABAF05D1DBD9C3FE441589" box="[151,482,1719,1745]" pageId="31" pageNumber="32" type="nomenclature">
<paragraph id="8BC436D8FFABAF05D1DBD9C3FE441589" blockId="31.[151,482,1719,1745]" box="[151,482,1719,1745]" pageId="31" pageNumber="32">
<heading id="D08C81B4FFABAF05D1DBD9C3FE441589" bold="true" box="[151,482,1719,1745]" fontSize="11" level="1" pageId="31" pageNumber="32" reason="1">
<emphasis id="B90FEACAFFABAF05D1DBD9C3FE441589" bold="true" box="[151,482,1719,1745]" pageId="31" pageNumber="32">
7.
<taxonomicName id="4C7B4D5BFFABAF05D1F8D9C3FEFC1589" ID-CoL="5CCP" authority="Vaz-De-Mello, 2008" authorityName="Vaz-De-Mello" authorityYear="2008" box="[180,346,1719,1745]" class="Insecta" family="Scarabaeidae" genus="Leotrichillum" kingdom="Animalia" order="Coleoptera" pageId="31" pageNumber="32" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90FEACAFFABAF05D1F8D9C3FEFC1589" bold="true" box="[180,346,1719,1745]" italics="true" pageId="31" pageNumber="32">Leotrichillum</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A23C57B1FFABAF05D02ED9C3FE441589" box="[354,482,1719,1745]" pageId="31" pageNumber="32" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3616553FFABAF05D1DBD876FC5C14BC" pageId="31" pageNumber="32" type="diagnosis">
<paragraph id="8BC436D8FFABAF05D1DBD876FC5C14BC" blockId="31.[151,1436,1794,2020]" pageId="31" pageNumber="32">
<emphasis id="B90FEACAFFABAF05D1DBD876FEBF1444" bold="true" box="[151,281,1794,1820]" pageId="31" pageNumber="32">Diagnosis:</emphasis>
Small (
<quantity id="4C839B3DFFABAF05D036D876FDB61444" box="[378,528,1794,1820]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="31" pageNumber="32" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.22.8 mm</quantity>
), oval, and elongated body (
<figureCitation id="13402A5DFFABAF05D225D876FC651444" box="[873,963,1794,1820]" captionStart="FIGURES 2330" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 2330. Dorsal views. 23. Leotrichillum louzadaorum; 2425: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="31" pageNumber="32">Fig. 23</figureCitation>
); color tan to brown; clypeus with two teeth and evenly rounded laterally (
<figureCitation id="13402A5DFFABAF05D371D85EFD30141C" box="[573,662,1834,1860]" captionStart="FIGURES 7178" captionStartId="63.[151,269,1933,1957]" captionTargetBox="[221,1365,194,1906]" captionTargetId="figure-52@63.[221,1365,194,1910]" captionTargetPageId="63" captionText="FIGURES 7178. 71. Genieridium zanunciorum, parameres; 7273: Leotrichillum louzadaorum: 72. head; 73. parameres; 7476: Martinezidium: 7475: M. galileoae: 74. head; 75. parameres; 76. M. maya, head; 7778: Nunoidium argentinum: 77. head; 78. parameres." pageId="31" pageNumber="32">Fig. 72</figureCitation>
). Pronotum separated of hypomeron by sharp carina, hypomeron with sharp lateral longitudinal carina posteriorly. Elytral interstriae with uniseriate setose punctures; pseudoepipleura glabrous and gradually narrowed posteriorly. Mesofemora and metafemora oval; mesotibiae and metatibiae apical width of tibia much longer than third of tibial length. Metatarsi with basal tarsomere slightly longer than second. Parameres with an apicolateral invagination (
<figureCitation id="13402A5DFFABAF05D2D9D8BEFC4B14BC" box="[917,1005,1994,2020]" captionStart="FIGURES 7178" captionStartId="63.[151,269,1933,1957]" captionTargetBox="[221,1365,194,1906]" captionTargetId="figure-52@63.[221,1365,194,1910]" captionTargetPageId="63" captionText="FIGURES 7178. 71. Genieridium zanunciorum, parameres; 7273: Leotrichillum louzadaorum: 72. head; 73. parameres; 7476: Martinezidium: 7475: M. galileoae: 74. head; 75. parameres; 76. M. maya, head; 7778: Nunoidium argentinum: 77. head; 78. parameres." pageId="31" pageNumber="32">Fig. 73</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF94AF3AD18ADFE3FC48108A" pageId="32" pageNumber="33" type="description">
<paragraph id="8BC436D8FF94AF3AD18ADFE3FB3C106A" blockId="32.[151,1436,151,1577]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18ADFE3FEC613E9" bold="true" box="[198,352,151,177]" pageId="32" pageNumber="33">Description:</emphasis>
Length
<quantity id="4C839B3DFF94AF3AD08BDFECFDFC13EA" box="[455,602,152,178]" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="2.8" metricValueMin="2.2" pageId="32" pageNumber="33" unit="mm" value="2.5" valueMax="2.8" valueMin="2.2">2.22.8 mm</quantity>
, body oval, elongated, convex (
<figureCitation id="13402A5DFF94AF3AD297DFECFB9313EA" box="[987,1077,152,178]" captionStart="FIGURES 2330" captionStartId="57.[151,269,1934,1958]" captionTargetBox="[222,1365,194,1910]" captionTargetId="figure-54@57.[222,1365,194,1911]" captionTargetPageId="57" captionText="FIGURES 2330. Dorsal views. 23. Leotrichillum louzadaorum; 2425: Martinezidium: 24. M. fulgens; 25. M. maya; 26. Nunoidium argentinum; 27. Onoreidium cristatum; 28. Onoreidium bottimeri; 29. Pedaridium hirsutum; 30. Pereiraidium almeidai, ♂." pageId="32" pageNumber="33">Fig. 23</figureCitation>
); tan to brown, shiny, lacking metallic sheen, dorsal setae slightly claviform. Clypeus with two short triangular teeth, separated by wide Vshaped emargination, evenly curved laterally, continuous with genal margin. Clypeofrontal and clypeogenal sutures indistinct. Eyes, in dorsal view, rounded, each about as wide as seven-tenths of its length; interocular width five to six times eye width (
<figureCitation id="13402A5DFF94AF3AD308DE4CFD06120A" box="[580,672,312,338]" captionStart="FIGURES 7178" captionStartId="63.[151,269,1933,1957]" captionTargetBox="[221,1365,194,1906]" captionTargetId="figure-52@63.[221,1365,194,1910]" captionTargetPageId="63" captionText="FIGURES 7178. 71. Genieridium zanunciorum, parameres; 7273: Leotrichillum louzadaorum: 72. head; 73. parameres; 7476: Martinezidium: 7475: M. galileoae: 74. head; 75. parameres; 76. M. maya, head; 7778: Nunoidium argentinum: 77. head; 78. parameres." pageId="32" pageNumber="33">Fig. 72</figureCitation>
). Pronotum lacking without anterior or posterior beads; lateral pronotal callosity indicated by indistinct dark spot; disc covered by simple setose punctures, separated from hypomeron by very weak carina. Hypomeron with strong longitudinal lateral carina; mesoepimera with weak anterior transverse carina, stronger mesally; metasternal disc covered by simple punctures. Elytral interstriae with one row of setose punctures, flat. Elytral striae not distinctly widened or deeper on declivity. Pseudoepipleura glabrous, gradually narrowed to apex, with longitudinal fold near metacoxa, causing anterior part to be almost vertical in position, and posterior part almost horizontal. Protibiae with three strong teeth distributed along apical three-fifths of lateral margin, which is denticulate from base to basal tooth, with ventral scale-like setae on teeth. Mesofemora and metafemora oval; mesotibiae and metatibiae apically wider than third of tibial length. Pygidium nearly vertical. Parameres shorter than half length of phallobase, flattened and apically rounded, laterally incised (
<figureCitation id="13402A5DFF94AF3AD0A8DDBCFDE511BA" box="[484,579,712,738]" captionStart="FIGURES 7178" captionStartId="63.[151,269,1933,1957]" captionTargetBox="[221,1365,194,1906]" captionTargetId="figure-52@63.[221,1365,194,1910]" captionTargetPageId="63" captionText="FIGURES 7178. 71. Genieridium zanunciorum, parameres; 7273: Leotrichillum louzadaorum: 72. head; 73. parameres; 7476: Martinezidium: 7475: M. galileoae: 74. head; 75. parameres; 76. M. maya, head; 7778: Nunoidium argentinum: 77. head; 78. parameres." pageId="32" pageNumber="33">Fig. 73</figureCitation>
). Pseudoflagellum helicoidal and thick, with flat elongated auxiliary lamella. Coxites large, oval and symmetrical. Spermatheca simply C-shaped, gradually narrowed apically, bulbous at base, with duct insertion at ventral part of the base of the spermathecal body.
</paragraph>
<paragraph id="8BC436D8FF94AF3AD18ADC4BFC48108A" blockId="32.[151,1436,151,1577]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18ADC4BFE651001" bold="true" box="[198,451,831,857]" pageId="32" pageNumber="33">Sexual dimorphism:</emphasis>
Males with proclaws bent in the middle, protarsi with apical tarsomere modified to receive claws; mesotarsi and metatarsi with claws bigger and stronger than in females. All tibiae with apicomesal tooth in males, mesotibial and metatibial teeth smaller than protibial one. Metasternum slightly concave in males, flat in females; pygidium slightly longer in males than females.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF94AF3AD18ADCABFC16177A" pageId="32" pageNumber="33" type="type_taxon">
<paragraph id="8BC436D8FF94AF3AD18ADCABFC16177A" blockId="32.[151,1436,151,1577]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18ADCABFEC910A1" bold="true" box="[198,367,991,1017]" pageId="32" pageNumber="33">
<typeStatus id="54C0887AFF94AF3AD18ADCABFEA510A1" box="[198,259,991,1017]" pageId="32" pageNumber="33">Type</typeStatus>
species:
</emphasis>
<taxonomicName id="4C7B4D5BFF94AF3AD037DC94FD0A10A1" ID-CoL="75WS2" authorityName="Vaz-de-Mello &amp; Canhedo" authorityYear="1998" box="[379,684,992,1017]" class="Insecta" family="Scarabaeidae" genus="Pedaridium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="louzadaorum">
<emphasis id="B90FEACAFF94AF3AD037DC94FD0A10A1" box="[379,684,992,1017]" italics="true" pageId="32" pageNumber="33">Pedaridium louzadaorum</emphasis>
</taxonomicName>
<bibRefCitation id="EFEA4B29FF94AF3AD3F4DC94FB9B10A2" author="Vaz-de-Mello, F. Z. &amp; Canhedo, V. L." box="[696,1085,992,1018]" pageId="32" pageNumber="33" pagination="95 - 100" refId="ref51217" refString="Vaz-de-Mello, F. Z. &amp; Canhedo, V. L. (1998) Duas novas especies de Pedaridium (Coleoptera, Scarabaeidae, Coprinae, Ateuchini). Iheringia, Serie Zoologia, 84, 95 - 100." type="journal article" year="1998">Vaz-de-Mello &amp; Canhedo, 1998</bibRefCitation>
(monotypy) =
<taxonomicName id="4C7B4D5BFF94AF3AD5B0DC94FE971779" ID-CoL="3T7D9" baseAuthorityName="Vaz-de-Mello &amp; Canhedo" baseAuthorityYear="1998" class="Insecta" family="Scarabaeidae" genus="Leotrichillum" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="louzadaorum">
<emphasis id="B90FEACAFF94AF3AD5B0DC94FE971779" italics="true" pageId="32" pageNumber="33">Leotrichillum louzadaorum</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFEA4B29FF94AF3AD00CDB7CFD18177A" author="Vaz-de-Mello, F. Z. &amp; Canhedo, V. L." box="[320,702,1032,1058]" pageId="32" pageNumber="33" pagination="95 - 100" refId="ref51217" refString="Vaz-de-Mello, F. Z. &amp; Canhedo, V. L. (1998) Duas novas especies de Pedaridium (Coleoptera, Scarabaeidae, Coprinae, Ateuchini). Iheringia, Serie Zoologia, 84, 95 - 100." type="journal article" year="1998">Vaz-de-Mello &amp; Canhedo, 1998</bibRefCitation>
),
<emphasis id="B90FEACAFF94AF3AD39FDB73FC0E1779" bold="true" box="[723,936,1031,1057]" pageId="32" pageNumber="33">new combination</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF94AF3AD18ADB5BFBAA1712" box="[198,1036,1071,1098]" pageId="32" pageNumber="33" type="etymology">
<paragraph id="8BC436D8FF94AF3AD18ADB5BFBAA1712" blockId="32.[151,1436,151,1577]" box="[198,1036,1071,1098]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18ADB5BFEF11711" bold="true" box="[198,343,1071,1097]" pageId="32" pageNumber="33">Etymology:</emphasis>
After my son, Léo Falqueto Vaz de Mello. Gender neutral.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF94AF3AD18ADB23FC0217C2" pageId="32" pageNumber="33" type="distribution">
<paragraph id="8BC436D8FF94AF3AD18ADB23FC0217C2" blockId="32.[151,1436,151,1577]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18ADB23FECF1729" bold="true" box="[198,361,1111,1137]" pageId="32" pageNumber="33">Distribution:</emphasis>
Northern
<collectingCountry id="F36C7648FF94AF3AD0AFDB2CFDFE172A" box="[483,600,1112,1138]" name="Argentina" pageId="32" pageNumber="33">Argentina</collectingCountry>
and southern
<collectingCountry id="F36C7648FF94AF3AD24CDB2CFCC8172A" box="[768,878,1112,1138]" name="Paraguay" pageId="32" pageNumber="33">Paraguay</collectingCountry>
, central and northeastern
<collectingCountry id="F36C7648FF94AF3AD5D2DB2CFB4F172A" box="[1182,1257,1112,1138]" name="Brazil" pageId="32" pageNumber="33">Brazil</collectingCountry>
, occupying the Chaco, Caatinga and Cerrado provinces of the Chacoan subregion.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF94AF3AD18ADBD3FB5B155A" pageId="32" pageNumber="33" type="discussion">
<paragraph id="8BC436D8FF94AF3AD18ADBD3FBC61682" blockId="32.[151,1436,151,1577]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18ADBD3FEE71799" bold="true" box="[198,321,1191,1217]" pageId="32" pageNumber="33">Remarks:</emphasis>
The pygidial sulcus of
<taxonomicName id="4C7B4D5BFF94AF3AD315DBDCFCB41799" baseAuthorityName="Vaz-de-Mello &amp; Canhedo" baseAuthorityYear="1998" box="[601,786,1192,1217]" class="Insecta" family="Scarabaeidae" genus="Leotrichillum" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="species" species="louzadaorum">
<emphasis id="B90FEACAFF94AF3AD315DBDCFCB41799" box="[601,786,1192,1217]" italics="true" pageId="32" pageNumber="33">L. louzadaorum</emphasis>
</taxonomicName>
is, in fact, not divided as described originally (Vaz-de- Mello &amp; Canhedo 1998); it is entire. The relationship of
<taxonomicName id="4C7B4D5BFF94AF3AD272DBA4FC7817B1" authorityName="Vaz-De-Mello" authorityYear="2008" box="[830,990,1232,1257]" class="Insecta" family="Scarabaeidae" genus="Leotrichillum" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF94AF3AD272DBA4FC7817B1" box="[830,990,1232,1257]" italics="true" pageId="32" pageNumber="33">Leotrichillum</emphasis>
</taxonomicName>
to other genera is uncertain, as it has several characters relating it both to
<taxonomicName id="4C7B4D5BFF94AF3AD301DB8CFD471649" authorityName="Vaz-De-Mello" authorityYear="2008" box="[589,737,1272,1297]" class="Insecta" family="Scarabaeidae" genus="Genieridium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF94AF3AD301DB8CFD471649" box="[589,737,1272,1297]" italics="true" pageId="32" pageNumber="33">Genieridium</emphasis>
</taxonomicName>
(e.g. oval and elongated body, presence of mesotibial and metatibial mesoapical teeth in males, slightly modified mesotibial and metatibial male claws, weak pronotalhypomeral carina) and to
<taxonomicName id="4C7B4D5BFF94AF3AD088DA3CFDD11639" authorityName="Vaz-De-Mello" authorityYear="2008" box="[452,631,1352,1377]" class="Insecta" family="Scarabaeidae" genus="Degallieridium" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF94AF3AD088DA3CFDD11639" box="[452,631,1352,1377]" italics="true" pageId="32" pageNumber="33">Degallieridium</emphasis>
</taxonomicName>
(e.g. very small size, color tan to brown with darkened pronotal callosities, claviform dorsal setae, longitudinally bent pseudoepipleura, large eyes). Unambiguous synapomorphies for
<taxonomicName id="4C7B4D5BFF94AF3AD05CDAECFE1616E9" authorityName="Vaz-De-Mello" authorityYear="2008" box="[272,432,1432,1457]" class="Insecta" family="Scarabaeidae" genus="Leotrichillum" kingdom="Animalia" order="Coleoptera" pageId="32" pageNumber="33" phylum="Arthropoda" rank="genus">
<emphasis id="B90FEACAFF94AF3AD05CDAECFE1616E9" box="[272,432,1432,1457]" italics="true" pageId="32" pageNumber="33">Leotrichillum</emphasis>
</taxonomicName>
are the laterally incised parameres and the ventral position of spermathecal duct insertion, apart of others that depend on the resolution of its phylogenetic position.
</paragraph>
<paragraph id="8BC436D8FF94AF3AD18ADA93FB5B155A" blockId="32.[151,1436,151,1577]" box="[198,1277,1511,1538]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18ADA93FEC91559" bold="true" box="[198,367,1511,1537]" pageId="32" pageNumber="33">Composition:</emphasis>
Besides the
<typeStatus id="54C0887AFF94AF3AD348DA9CFD90155A" box="[516,566,1512,1538]" pageId="32" pageNumber="33">type</typeStatus>
species, the genus contains at least one undescribed species.
</paragraph>
</subSubSection>
<subSubSection id="C3616553FF94AF3AD18AD97BFE1A1571" box="[198,444,1551,1577]" pageId="32" pageNumber="33" type="materials_examined">
<paragraph id="8BC436D8FF94AF3AD18AD97BFE1A1571" blockId="32.[151,1436,151,1577]" box="[198,444,1551,1577]" pageId="32" pageNumber="33">
<emphasis id="B90FEACAFF94AF3AD18AD97BFE1A1571" bold="true" box="[198,444,1551,1577]" pageId="32" pageNumber="33">Material examined:</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>