241 lines
27 KiB
XML
241 lines
27 KiB
XML
<document id="D1059E6A14A30AA535AC5301959576B7" ID-DOI="10.5281/zenodo.282248" ID-GBIF-Dataset="e68171f3-6fa8-4644-b5b4-1741a65298ba" ID-ISSN="1175-5326" ID-Zenodo-Dep="282248" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460523931932" checkinUser="plazi" docAuthor="Yan, Zhuo, Bai, Ming & Ren, Dong" docDate="2012" docId="03D287A9EC2DFFD98DADFAFC1E77FEAA" docLanguage="en" docName="zt03478p204.pdf" docOrigin="Zootaxa 3478" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Pulcherhybosorus Yan , Bai" docType="treatment" docVersion="7" lastPageNumber="203" masterDocId="FFEBFFD1EC2DFFDB8D3AFFAD1B68FFB6" masterDocTitle="A new fossil Hybosoridae (Coleoptera: Scarabaeoidea) from the Yixian Formation of China" masterLastPageNumber="204" masterPageNumber="201" pageNumber="201" updateTime="1698314873610" updateUser="plazi">
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<mods:mods id="547E8A15E6E18CB23229FF5A0CCF1B96" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="58BFB8430A8FFC5221DE29188A8F5B8B">
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<mods:title id="B706272EDBE18935F992104C06F9DBB4">A new fossil Hybosoridae (Coleoptera: Scarabaeoidea) from the Yixian Formation of China</mods:title>
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</mods:titleInfo>
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<mods:name id="03425F97BB141ADC4FB77E15EA9E5EB7" type="personal">
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<mods:roleTerm id="329DA277A3E904C0194B90EE8ED34F8B">Author</mods:roleTerm>
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<mods:namePart id="0FC6573DFC658D5E28D8FDD37BC5BA42">Yan, Zhuo</mods:namePart>
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<mods:roleTerm id="5660DE513F7E8B8BB30BE531B02C2286">Author</mods:roleTerm>
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<mods:namePart id="D32AC029B111DCF338A189AFFC00E81C">Bai, Ming</mods:namePart>
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</mods:name>
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<mods:name id="16CF2C4FB6E3218208FCBA8ABE1ADA24" type="personal">
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<mods:roleTerm id="211E459F4C27B2C8A16C060B07CFBF29">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="0EA8A057624508A1614A4FB50E036804">Ren, Dong</mods:namePart>
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<mods:typeOfResource id="E03EF46D650C956393B2DFFB7134F837">text</mods:typeOfResource>
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<mods:title id="DAEEB56FCD920BC13866DB5512B90477">Zootaxa</mods:title>
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<mods:part id="E2ED3CCC8505C08875C179E68C6779CD">
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<mods:date id="A0B52C45D2F8EC3E3D6949A84D1476E5">2012</mods:date>
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<mods:detail id="129709F8F9431A686A94F4A4651FC455" type="volume">
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<mods:number id="F1FB81759A6454B64E7AEBCA3AE53C2D">3478</mods:number>
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<mods:classification id="5A3D80D48BABA65731B2B94FE96F979A">journal article</mods:classification>
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<mods:identifier id="2C46922623B1A6A775B3DB2E1CE60AED" type="DOI">10.5281/zenodo.282248</mods:identifier>
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<mods:identifier id="919278FD9596AF387607AE3309825C1A" type="ISSN">1175-5326</mods:identifier>
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<treatment id="03D287A9EC2DFFD98DADFAFC1E77FEAA" ID-DOI="http://doi.org/10.5281/zenodo.6171920" ID-GBIF-Taxon="119526050" ID-Zenodo-Dep="6171920" LSID="urn:lsid:plazi:treatment:03D287A9EC2DFFD98DADFAFC1E77FEAA" httpUri="http://treatment.plazi.org/id/03D287A9EC2DFFD98DADFAFC1E77FEAA" lastPageId="2" lastPageNumber="203" pageId="0" pageNumber="201">
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<subSubSection id="C3616534EC2DFFDB8DADFAFC1A63FA3D" pageId="0" pageNumber="201" type="nomenclature">
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<paragraph id="8BC436BFEC2DFFDB8DADFAFC19C8FADF" blockId="0.[151,672,1361,1419]" box="[151,672,1361,1385]" pageId="0" pageNumber="201">
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<heading id="D08C81D3EC2DFFDB8DADFAFC19C8FADF" bold="true" box="[151,672,1361,1385]" fontSize="10" level="2" pageId="0" pageNumber="201" reason="2">
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<emphasis id="B90FEAADEC2DFFDB8DADFAFC19C8FADF" bold="true" box="[151,672,1361,1385]" pageId="0" pageNumber="201">
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<taxonomicName id="4C7B4D3CEC2DFFDB8DADFAFC1AABFADF" ID-CoL="75CN" authority="Yan , Bai" authorityName="Yan , Bai" box="[151,451,1361,1385]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="201" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAADEC2DFFDB8DADFAFC1A37FADF" bold="true" box="[151,351,1361,1385]" italics="true" pageId="0" pageNumber="201">Pulcherhybosorus</emphasis>
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Yan, Bai
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</taxonomicName>
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, & Ren, new genus
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC436BFEC2DFFDB8DADFAD91A63FA3D" blockId="0.[151,672,1361,1419]" box="[151,267,1396,1419]" pageId="0" pageNumber="201">
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(
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<figureCitation id="13402A3AEC2DFFDB8DA5FAD91A6BFA3D" box="[159,259,1396,1419]" captionStart-0="FIGURES 1 – 2" captionStart-1="FIGURES 3 – 4" captionStartId-0="1.[151,257,699,720]" captionStartId-1="1.[151,257,1537,1558]" captionTargetBox-0="[151,1436,182,675]" captionTargetBox-1="[167,1417,1000,1516]" captionTargetId-0="figure@1.[151,1436,182,675]" captionTargetId-1="figure@1.[151,1436,986,1533]" captionTargetPageId-0="1" captionTargetPageId-1="1" captionText-0="FIGURES 1 – 2. Pulcherhybosorus tridentatus photographs of the holotype. 1 — CNU-COL-NN 2011002 P (dorsal view of body), 2 — CNU-COL-NN 2011002 C (ventral view of body)." captionText-1="FIGURES 3 – 4. Pulcherhybosorus tridentatus line drawings of the holotype. 3 — dorsal view of body; 4 — ventral view of body." httpUri-0="https://zenodo.org/record/282249/files/figure.png" httpUri-1="https://zenodo.org/record/282250/files/figure.png" pageId="0" pageNumber="201">Figs. 1–4</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616534EC2DFFDB8DADFA1718C3FA67" box="[151,939,1466,1489]" pageId="0" pageNumber="201" type="materials_examined">
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<paragraph id="8BC436BFEC2DFFDB8DADFA1718C3FA67" blockId="0.[151,939,1466,1489]" box="[151,939,1466,1489]" pageId="0" pageNumber="201">
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<typeStatus id="54C0881DEC2DFFDB8DADFA171BA4FA67" box="[151,204,1466,1489]" pageId="0" pageNumber="201">Type</typeStatus>
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species:
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<taxonomicName id="4C7B4D3CEC2DFFDB8C17FA171901FA67" box="[301,617,1466,1489]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="201" phylum="Arthropoda" rank="species" species="tridentatus" status="sp. nov.">
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<emphasis id="B90FEAADEC2DFFDB8C17FA171901FA67" box="[301,617,1466,1489]" italics="true" pageId="0" pageNumber="201">Pulcherhybosorus tridentatus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A23C57D6EC2DFFDB8F4AFA17199BFA67" box="[624,755,1466,1489]" pageId="0" pageNumber="201" rank="species">new species</taxonomicNameLabel>
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, here designated.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616534EC2DFFDB8DADFA511F78F968" pageId="0" pageNumber="201" type="diagnosis">
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<paragraph id="8BC436BFEC2DFFDB8DADFA511F78F968" blockId="0.[151,1437,1532,2027]" pageId="0" pageNumber="201">
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<emphasis id="B90FEAADEC2DFFDB8DADFA511A64F9A2" bold="true" box="[151,268,1532,1556]" pageId="0" pageNumber="201">Diagnosis.</emphasis>
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Elongate oval. Mandibles and labrum prominent, clearly visible in dorsal view of head, labrum exposed beyond apex of clypeus, mandibles exposed beyond apex of clypeus (fig. 5), anterior margin of clypeus moderately emarginate. Antennae with a 3-antennomere lamellate club, club compact (
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<figureCitation id="13402A3AEC2DFFDB8EFBF9921F6AF9E0" box="[961,1026,1599,1622]" captionStart="FIGURES 5 – 8" captionStartId="2.[151,257,1297,1318]" captionTargetBox="[151,1435,328,1283]" captionTargetId="figure@2.[151,1436,328,1284]" captionTargetPageId="2" captionText="FIGURES 5 – 8. Pulcherhybosorus tridentatus magnified photographs of some details, 5 — labrum and mandibles; 6 — two spurs on the end of mesotibia; 7 — hind wing; 8 — antennae." httpUri="https://zenodo.org/record/282251/files/figure.png" pageId="0" pageNumber="201">Fig. 8</figureCitation>
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). Eyes large. Pronotum slightly wider than elytra, nearly trapezoidal, with widest part of pronotum at base. Elytra convex, striae well defined and without tubercles; hind wings well developed, the MP3 vein present and MP4 vein absent. Abdomen with five visible sternites. Protibia with three teeth on outer margin, mesotibia and metatibia without a transverse carina, two spurs on the end of mesotibia subequal in length, and two symmetrical claws on the end of mesotarsus.
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</paragraph>
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</subSubSection>
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<subSubSection id="C3616534EC2DFFD98DFDF9451A9CFF41" lastPageId="2" lastPageNumber="203" pageId="0" pageNumber="201" type="discussion">
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<paragraph id="8BC436BFEC2DFFDB8DFDF9451965F85D" blockId="0.[151,1437,1532,2027]" pageId="0" pageNumber="201">
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<emphasis id="B90FEAADEC2DFFDB8DFDF9451A5DF8B6" bold="true" box="[199,309,1768,1792]" pageId="0" pageNumber="201">Remarks.</emphasis>
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The new genus is tentatively assigned to the family
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<taxonomicName id="4C7B4D3CEC2DFFDB8E48F9451892F949" box="[882,1018,1768,1791]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="201" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
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based on the following characters: (1) antennae with a 3-antennomere lamellate club, club compact; (2) eyes developed; (3) labrum and mandibles produced beyond apex of clypeus; (4) pronotum trapezoidal, convex, base wider than elytral base; (5) elytra convex, striae well defined, Wings well developed, the MP3 vein present; (6) legs with anterior coxae conical and mesocoxae contiguous; (7) protibia with three teeth; (8) mesotibia and metatibia slightly dilated at the apex, mesotibia with 2 subequal spurs; (9) mesotibia and metatibia without a transverse carina; (10) claws equal in size, simple; (11) abdomen with five visible sternites.
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<taxonomicName id="4C7B4D3CEC2DFFDB8C3BF81E1AABF87C" box="[257,451,1971,1994]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="0" pageNumber="201" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAADEC2DFFDB8C3BF81E1AABF87C" box="[257,451,1971,1994]" italics="true" pageId="0" pageNumber="201">Pulcherhybosorus</emphasis>
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</taxonomicName>
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is not placed in a subfamily because of the lack of some key visible characters and the unusual combination of characters.
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</paragraph>
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<caption id="DF046637EC2CFFDA8DADFD1619FFFD58" httpUri="https://zenodo.org/record/282249/files/figure.png" pageId="1" pageNumber="202" targetBox="[151,1436,182,675]" targetPageId="1">
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<paragraph id="8BC436BFEC2CFFDA8DADFD1619FFFD58" blockId="1.[151,1436,699,750]" pageId="1" pageNumber="202">
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<emphasis id="B90FEAADEC2CFFDA8DADFD161A5EFD66" bold="true" box="[151,310,699,720]" pageId="1" pageNumber="202">FIGURES 1–2.</emphasis>
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<taxonomicName id="4C7B4D3CEC2CFFDA8C7AFD111908FD67" box="[320,608,700,721]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="species" species="tridentatus">
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<emphasis id="B90FEAADEC2CFFDA8C7AFD111908FD67" box="[320,608,700,721]" italics="true" pageId="1" pageNumber="202">Pulcherhybosorus tridentatus</emphasis>
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</taxonomicName>
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photographs of the holotype. 1—CNU-COL-NN2011002P (dorsal view of body), 2—CNU-COL-NN2011002C (ventral view of body).
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</paragraph>
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</caption>
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<paragraph id="8BC436BFEC2CFFDA8DFDFCB819F5FC0A" blockId="1.[151,1436,789,956]" pageId="1" pageNumber="202">
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Beetle fossils with hind wings not covered by the elytra are extremely rare.
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<taxonomicName id="4C7B4D3CEC2CFFDA8ED5FCB81E43FC9A" box="[1007,1323,789,812]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="species" species="tridentatus">
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<emphasis id="B90FEAADEC2CFFDA8ED5FCB81E43FC9A" box="[1007,1323,789,812]" italics="true" pageId="1" pageNumber="202">Pulcherhybosorus tridentatus</emphasis>
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</taxonomicName>
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is the first
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<taxonomicName id="4C7B4D3CEC2CFFDA8DADFC951A77FCF9" box="[151,287,824,847]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
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fossil species with the hind wing visible and preserved. The hind wings are well preserved and well developed showing the
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<collectingCountry id="F36C762FEC2CFFDA8CA1FCF71AD3FCC7" box="[411,443,858,881]" name="Poland" pageId="1" pageNumber="202">RP</collectingCountry>
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, RA3+4, MP1+2, MP3,
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<collectingCountry id="F36C762FEC2CFFDA8FA5FCF719A8FCC7" box="[671,704,858,881]" name="Poland" pageId="1" pageNumber="202">RP</collectingCountry>
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3+4,
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<collectingCountry id="F36C762FEC2CFFDA8FDCFCF7186FFCC7" box="[742,775,858,881]" name="Poland" pageId="1" pageNumber="202">RP</collectingCountry>
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2 and RA4+
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<collectingCountry id="F36C762FEC2CFFDA8EBBFCF718CAFCC7" box="[897,930,858,881]" name="Poland" pageId="1" pageNumber="202">RP</collectingCountry>
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1 veins. The presence of the MP3 vein and the absence of the MP4 vein are different character states from extant
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<taxonomicName id="4C7B4D3CEC2CFFDA8EB6FC2D1F70FC21" box="[908,1048,896,919]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
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. This fossil provides an important evidence for the early evolution of
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<taxonomicName id="4C7B4D3CEC2CFFDA8F34FC0819F2FC0A" box="[526,666,933,956]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
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.
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</paragraph>
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<caption id="DF046637EC2CFFDA8DADF9AC1E36F9A0" box="[151,1374,1537,1559]" httpUri="https://zenodo.org/record/282250/files/figure.png" pageId="1" pageNumber="202" targetBox="[167,1417,1000,1516]" targetPageId="1">
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<paragraph id="8BC436BFEC2CFFDA8DADF9AC1E36F9A0" blockId="1.[151,1374,1537,1559]" box="[151,1374,1537,1559]" pageId="1" pageNumber="202">
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<emphasis id="B90FEAADEC2CFFDA8DADF9AC1A59F9A0" bold="true" box="[151,305,1537,1558]" pageId="1" pageNumber="202">FIGURES 3–4.</emphasis>
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<taxonomicName id="4C7B4D3CEC2CFFDA8C0DF9AF193BF9A1" box="[311,595,1538,1559]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="species" species="tridentatus">
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<emphasis id="B90FEAADEC2CFFDA8C0DF9AF193BF9A1" box="[311,595,1538,1559]" italics="true" pageId="1" pageNumber="202">Pulcherhybosorus tridentatus</emphasis>
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</taxonomicName>
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line drawings of the holotype. 3—dorsal view of body; 4—ventral view of body.
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</paragraph>
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</caption>
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<paragraph id="8BC436BFEC2CFFD98DFDF9EC1A9CFF41" blockId="1.[151,1436,1601,2006]" lastBlockId="2.[151,1436,152,284]" lastPageId="2" lastPageNumber="203" pageId="1" pageNumber="202">
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<emphasis id="B90FEAADEC2CFFDA8DFDF9EC1A33F9EF" bold="true" box="[199,347,1601,1625]" pageId="1" pageNumber="202">Comparison.</emphasis>
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According to the
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<taxonomicName id="4C7B4D3CEC2CFFDA8F1BF9EF19C1F9EF" box="[545,681,1602,1625]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
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fossil record, there are nine genera described from the Mesozoic Era that warrant comparison:
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<taxonomicName id="4C7B4D3CEC2CFFDA8C83F9C91992F9CD" authority="Nikolajev, 1996" authorityName="Nikolajev" authorityYear="1996" box="[441,762,1636,1659]" class="Insecta" family="Hybosoridae" genus="Cretanaides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAADEC2CFFDA8C83F9C91952F9CD" box="[441,570,1636,1659]" italics="true" pageId="1" pageNumber="202">Cretanaides</emphasis>
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Nikolajev, 1996
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</taxonomicName>
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;
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<taxonomicName id="4C7B4D3CEC2CFFDA8E30F9C91F1BF9CD" authority="Nikolajev, 1999" authorityName="Nikolajev" authorityYear="1999" box="[778,1139,1636,1659]" class="Insecta" family="Hybosoridae" genus="Cretohybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAADEC2CFFDA8E30F9C918DBF9CD" box="[778,947,1636,1659]" italics="true" pageId="1" pageNumber="202">Cretohybosorus</emphasis>
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Nikolajev, 1999
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</taxonomicName>
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;
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<taxonomicName id="4C7B4D3CEC2CFFDA89B9F9C91BB8F928" authority="Nikolajev, 2005" authorityName="Nikolajev" authorityYear="2005" class="Insecta" family="Hybosoridae" genus="Jurahybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
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<emphasis id="B90FEAADEC2CFFDA89B9F9C91E49F9CD" box="[1155,1313,1636,1659]" italics="true" pageId="1" pageNumber="202">Jurahybosorus</emphasis>
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Nikolajev, 2005
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</taxonomicName>
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;
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8DE7F92A1967F928" authority="Nikolajev, 2006" authorityName="Nikolajev" authorityYear="2006" box="[221,527,1671,1694]" class="Insecta" family="Hybosoridae" genus="Leptosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8DE7F92A1A3BF928" box="[221,339,1671,1694]" italics="true" pageId="1" pageNumber="202">Leptosorus</emphasis>
|
||
Nikolajev, 2006
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8F27F92A1F2DF928" authority="Kirejtshuk, Azar" authorityName="Kirejtshuk, Azar" authorityYear="2011" box="[541,1093,1671,1694]" class="Insecta" family="Hybosoridae" genus="Libanochrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8F27F92A19CCF928" box="[541,676,1671,1694]" italics="true" pageId="1" pageNumber="202">Libanochrus</emphasis>
|
||
<bibRefCitation id="EFEA4B4EEC2CFFDA8F94F92A1F2DF928" author="Kirejtshuk" box="[686,1093,1671,1694]" pageId="1" pageNumber="202" refString="Kirejtshuk, A. G., Azar, D. & Montreuil, O. (2011) First Mesozoic representative of the subfamily Liparochrinae (Coleoptera: Hybosoridae) from the Lower Cretaceous Lebanese amber. Zoosystematica Rossica, 20, 62 - 70." type="journal article" year="2011">Kirejtshuk, Azar, & Montreuil, 2011</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8969F92A1AD0F977" authority="Nikolajev, Wang, Liu" authorityName="Nikolajev, Wang, Liu" authorityYear="2010" class="Insecta" family="Hybosoridae" genus="Mesoceratocanthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8969F92A1E4CF928" box="[1107,1316,1671,1694]" italics="true" pageId="1" pageNumber="202">Mesoceratocanthus</emphasis>
|
||
<bibRefCitation id="EFEA4B4EEC2CFFDA8817F92A1AD0F977" author="Nikolajev" pageId="1" pageNumber="202" refString="Nikolajev, G. V., Wang, B., Liu, Y. & Zhang, H. C. (2010) First record of Mesozoic Ceratocanthinae (Coleoptera: Hybosoridae). Acta Palaeontologica Sinica, 49, 443 - 447." type="journal article" year="2010">Nikolajev, Wang, Liu, & Zhang, 2010</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8CF8F9071860F977" authority="Nikolajev, 2007" authorityName="Nikolajev" authorityYear="2007" box="[450,776,1706,1729]" class="Insecta" family="Hybosoridae" genus="Mimaphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8CF8F907193DF977" box="[450,597,1706,1729]" italics="true" pageId="1" pageNumber="202">Mimaphodius</emphasis>
|
||
<bibRefCitation id="EFEA4B4EEC2CFFDA8F61F9071860F977" author="Nikolajev" box="[603,776,1706,1729]" pageId="1" pageNumber="202" refString="Nikolajev, G. V. (2007) Novoe podsemeystvo plastinchatousih zhukov semeystva Hybosoridae (Coleoptera, Scaraboidea) iz Mezozoya Azii. Izvestiya Natsionalnoi Akademii Nauk Respubliki Kazakhstan Seriya Biologicheskaya i Meditsinskaya, 8 (260), 47 - 48." type="journal article" year="2007">Nikolajev, 2007</bibRefCitation>
|
||
</taxonomicName>
|
||
;
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8E29F9071F2FF977" authority="Nikolajev, 2010" authorityName="Nikolajev" authorityYear="2010" box="[787,1095,1706,1729]" class="Insecta" family="Hybosoridae" genus="Protanaides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8E29F90718FDF977" box="[787,917,1706,1729]" italics="true" pageId="1" pageNumber="202">Protanaides</emphasis>
|
||
Nikolajev, 2010
|
||
</taxonomicName>
|
||
; and
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8944F9071BB8F955" authority="Nikolajev, 2010" authorityName="Nikolajev" authorityYear="2010" class="Insecta" family="Hybosoridae" genus="Protohybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8944F9071E40F977" box="[1150,1320,1706,1729]" italics="true" pageId="1" pageNumber="202">Protohybosorus</emphasis>
|
||
Nikolajev, 2010
|
||
</taxonomicName>
|
||
. The new genus is readily distinguished from the genera
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8E16F96118BDF955" box="[812,981,1740,1763]" class="Insecta" family="Hybosoridae" genus="Cretohybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8E16F96118BDF955" box="[812,981,1740,1763]" italics="true" pageId="1" pageNumber="202">Cretohybosorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8932F9611FCEF955" box="[1032,1190,1740,1763]" class="Insecta" family="Hybosoridae" genus="Jurahybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8932F9611FCEF955" box="[1032,1190,1740,1763]" italics="true" pageId="1" pageNumber="202">Jurahybosorus</emphasis>
|
||
</taxonomicName>
|
||
by the large body size, the mandibles visible in dorsal view, and the long and narrow shape of mandibles.
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8912F9421F82F8B0" box="[1064,1258,1775,1798]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8912F9421F82F8B0" box="[1064,1258,1775,1798]" italics="true" pageId="1" pageNumber="202">Pulcherhybosorus</emphasis>
|
||
</taxonomicName>
|
||
is distinguished from the genus
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8C05F8BF1ADDF89F" box="[319,437,1810,1833]" class="Insecta" family="Hybosoridae" genus="Leptosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8C05F8BF1ADDF89F" box="[319,437,1810,1833]" italics="true" pageId="1" pageNumber="202">Leptosorus</emphasis>
|
||
</taxonomicName>
|
||
by the protibia dilated apically and with three teeth on outer margin and by the two spurs on the end of the mesotibia subequal in length. The abdomen with five visible sternites is a reliable diagnostic character that differentiates the new genus from the genus
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8F99F8FA1824F8D8" box="[675,844,1879,1902]" class="Insecta" family="Hybosoridae" genus="Protohybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8F99F8FA1824F8D8" box="[675,844,1879,1902]" italics="true" pageId="1" pageNumber="202">Protohybosorus</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8EB4F8FA1F37F8D8" box="[910,1119,1879,1902]" class="Insecta" family="Hybosoridae" genus="Mesoceratocanthus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8EB4F8FA1F37F8D8" box="[910,1119,1879,1902]" italics="true" pageId="1" pageNumber="202">Mesoceratocanthus</emphasis>
|
||
</taxonomicName>
|
||
. The new genus is readily distinguished from the genus
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8CDAF8D71909F827" box="[480,609,1914,1937]" class="Insecta" family="Hybosoridae" genus="Protanaides" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8CDAF8D71909F827" box="[480,609,1914,1937]" italics="true" pageId="1" pageNumber="202">Protanaides</emphasis>
|
||
</taxonomicName>
|
||
by absence of a transverse carina on the mesotibia and metatibia, and the protibia with three teeth on the outer margin; from
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8F88F8311851F805" box="[690,825,1948,1971]" class="Insecta" family="Hybosoridae" genus="Libanochrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8F88F8311851F805" box="[690,825,1948,1971]" italics="true" pageId="1" pageNumber="202">Libanochrus</emphasis>
|
||
</taxonomicName>
|
||
by the antennae with a 3-antennomere lamellate club and exposed mandibles; from
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8C97F8121928F860" box="[429,576,1983,2006]" class="Insecta" family="Hybosoridae" genus="Mimaphodius" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8C97F8121928F860" box="[429,576,1983,2006]" italics="true" pageId="1" pageNumber="202">Mimaphodius</emphasis>
|
||
</taxonomicName>
|
||
by labrum and mandibles visible in dorsal view.
|
||
<taxonomicName id="4C7B4D3CEC2CFFDA8969F8121E7DF860" box="[1107,1301,1983,2006]" class="Insecta" family="Hybosoridae" genus="Pulcherhybosorus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="202" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2CFFDA8969F8121E7DF860" box="[1107,1301,1983,2006]" italics="true" pageId="1" pageNumber="202">Pulcherhybosorus</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="4C7B4D3CEC2FFFD98DADFF351A70FF19" box="[151,280,152,175]" class="Insecta" family="Hybosoridae" genus="Cretanaides" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B90FEAADEC2FFFD98DADFF351A70FF19" box="[151,280,152,175]" italics="true" pageId="2" pageNumber="203">Cretanaides</emphasis>
|
||
</taxonomicName>
|
||
, known from a single elytron impression, by the body size and the absence of tubercles on the elytra. The new genus is distinguished from the extant genera of
|
||
<taxonomicName id="4C7B4D3CEC2FFFD98FECFF171836FF67" box="[726,862,186,209]" class="Insecta" family="Hybosoridae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="203" phylum="Arthropoda" rank="family">Hybosoridae</taxonomicName>
|
||
by the large body size, the presence of the MP3 vein, and the absence of the MP4 vein.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3616534EC2FFFD98DFDFEA91E77FEAA" box="[199,1311,260,284]" pageId="2" pageNumber="203" type="etymology">
|
||
<paragraph id="8BC436BFEC2FFFD98DFDFEA91E77FEAA" blockId="2.[151,1436,152,284]" box="[199,1311,260,284]" pageId="2" pageNumber="203">
|
||
<emphasis id="B90FEAADEC2FFFD98DFDFEA91A2FFEAA" bold="true" box="[199,327,260,284]" pageId="2" pageNumber="203">Etymology.</emphasis>
|
||
The Latin ‘
|
||
<emphasis id="B90FEAADEC2FFFD98CFFFEA81977FEAA" box="[453,543,261,284]" italics="true" pageId="2" pageNumber="203">pulcher-</emphasis>
|
||
’ means ‘beautiful’ in reference to this beautifully well-preserved fossil.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |