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<document id="CF837B8FA78A9CD614F2A514ADA39B19" ID-DOI="10.11646/zootaxa.3670.2.9" ID-GBIF-Dataset="492d3ca4-105c-4c92-8a7d-6cb180d00b08" ID-ISSN="1175-5326" ID-Zenodo-Dep="223576" ID-ZooBank="33F668E4-95BC-4203-B892-80365B4F3B89" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460633857078" checkinUser="plazi" docAuthor="Filippini, Valentina, Micó, Estefania &amp; Galante, Eduardo" docDate="2013" docId="03D28781FFF5FFDB5AA663BCFC26FD33" docLanguage="en" docName="zt03670p273.pdf" docOrigin="Zootaxa 3670 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala eucoma Bates 1888" docType="treatment" docVersion="8" lastPageNumber="264" masterDocId="FFEBFFF9FFF7FFD25A31622DFFA8FFB7" masterDocTitle="Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)" masterLastPageNumber="273" masterPageNumber="255" pageNumber="257" updateTime="1698334918277" updateUser="plazi">
<mods:mods id="DCC1B8EC7C3B3941C6E3E6D62100C32F" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="E74BB5255B23A9B3ADB2403B0D92AA10">
<mods:title id="0FDA4DDB2010C6BB8D2EB4A34BD8D2E0">Redescription of Anomala eucoma Bates, 1888 and a description of three new species from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae)</mods:title>
</mods:titleInfo>
<mods:name id="47450B23E291B68B813643AFE4DD8FD9" type="personal">
<mods:role id="199AAD32361BAA2C7AE4B7DE7F73A7D2">
<mods:roleTerm id="14814678AFDFB732127376D5B8DC3070">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A9CBDDDFA35A164FB406B193C506226E">Filippini, Valentina</mods:namePart>
</mods:name>
<mods:name id="52F445D75724DCC41FD15C8B046FBE40" type="personal">
<mods:role id="BBC0DD63F695292D0B58A3D234ED9D8A">
<mods:roleTerm id="A33FC065B038A640EECC23CC81F29C40">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="F3E884C1C42A3860AEAD0B1C00059A56">Micó, Estefania</mods:namePart>
</mods:name>
<mods:name id="4C89022BC0DAA526DF396EB60E8E755A" type="personal">
<mods:role id="D53FCAB1CFCC0649D45A854EB22C7F6C">
<mods:roleTerm id="C4BCEA4EBF4DE3798E15339B39322761">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="6B8B3C1E693DE120D6AB4F1262FF7C04">Galante, Eduardo</mods:namePart>
</mods:name>
<mods:typeOfResource id="F40F240632660C1F94EE4119D24FB2C3">text</mods:typeOfResource>
<mods:relatedItem id="4776C3DBA0F3ECEC6A7C40D8C9C6E04C" type="host">
<mods:titleInfo id="B0E1D360740FB6B132E32E2A113B9FCB">
<mods:title id="E96F1AAB39B7195CDDDC344C69987BE3">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="30F12EC01B4177A46C2CBF48244F8A90">
<mods:date id="F871787F7C36848A8BC5FB534AB2D37C">2013</mods:date>
<mods:detail id="6CE1BE26126C95D96D45098340B8C68C" type="volume">
<mods:number id="E18A600551884B9AF1ED5193F5FC3E9F">3670</mods:number>
</mods:detail>
<mods:detail id="B31439395F5786953EA3DCAE86F94ACC" type="issue">
<mods:number id="BD469D6F49A50A4415A6716FA7D03E93">2</mods:number>
</mods:detail>
<mods:extent id="E6D73D16E17690D915FF3D81512857CF" unit="page">
<mods:start id="97E5D18572D98A3D24D5359AEEE59F68">255</mods:start>
<mods:end id="28ED52E58BAF3920D236F8C4BFD48800">273</mods:end>
</mods:extent>
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<mods:classification id="796A21FE271E63897FF9F01D2C793429">journal article</mods:classification>
<mods:identifier id="E65E002BDE93567D6EC098849AAA3BDB" type="DOI">10.11646/zootaxa.3670.2.9</mods:identifier>
<mods:identifier id="CA79E81134778EA89A4E5FEEAAA83F30" type="GBIF-Dataset">492d3ca4-105c-4c92-8a7d-6cb180d00b08</mods:identifier>
<mods:identifier id="8AA3D9E6A813F2227BB24441D9CA92B1" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="F30E7659053629B65CF86E003E01D79E" type="Zenodo-Dep">223576</mods:identifier>
<mods:identifier id="5229D36F4D9F524CE75F55266CC96114" type="ZooBank">33F668E4-95BC-4203-B892-80365B4F3B89</mods:identifier>
</mods:mods>
<treatment id="03D28781FFF5FFDB5AA663BCFC26FD33" ID-DOI="http://doi.org/10.5281/zenodo.6159031" ID-GBIF-Taxon="119570135" ID-Zenodo-Dep="6159031" LSID="urn:lsid:plazi:treatment:03D28781FFF5FFDB5AA663BCFC26FD33" httpUri="http://treatment.plazi.org/id/03D28781FFF5FFDB5AA663BCFC26FD33" lastPageId="9" lastPageNumber="264" pageId="2" pageNumber="257">
<subSubSection id="C361651CFFF5FFD05AA663BCFF43FE7B" pageId="2" pageNumber="257" type="nomenclature">
<paragraph id="8BC43697FFF5FFD05AA663BCFDA8FE1D" blockId="2.[151,512,400,460]" box="[151,512,400,427]" pageId="2" pageNumber="257">
<heading id="D08C81FBFFF5FFD05AA663BCFDA8FE1D" bold="true" box="[151,512,400,427]" fontSize="11" level="1" pageId="2" pageNumber="257" reason="1">
<taxonomicName id="4C7B4D14FFF5FFD05AA663BCFDA8FE1D" authority="Bates, 1888" authorityName="Bates" authorityYear="1888" box="[151,512,400,427]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF5FFD05AA663BCFDA8FE1D" bold="true" box="[151,512,400,427]" pageId="2" pageNumber="257">
<emphasis id="B90FEA85FFF5FFD05AA663BCFEC3FE1C" bold="true" box="[151,363,401,427]" italics="true" pageId="2" pageNumber="257">Anomala eucoma</emphasis>
<bibRefCitation id="EFEA4B66FFF5FFD05B4263BDFDA8FE1D" author="Bates" box="[371,512,400,426]" pageId="2" pageNumber="257" refString="Bates, H. W. (1888) Insecta. Coleoptera. Pectinicornia and Lamellicornia. Volume 2, Part 2. In: Godman &amp; Salvin (Eds.), Biologia Centrali - Americana, R. H. Porter, London, United Kingdom, 432 pp." type="book" year="1888">Bates, 1888</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8BC43697FFF5FFD05AA66399FF43FE7B" blockId="2.[151,512,400,460]" box="[151,235,436,460]" pageId="2" pageNumber="257">
(
<figureCitation id="13402A12FFF5FFD05AAE6399FF4BFE7B" box="[159,227,436,460]" captionStart="FIGURES 2 – 5" captionStartId="4.[151,269,1929,1953]" captionTargetBox="[193,1406,206,1890]" captionTargetId="figure@4.[184,1420,194,1919]" captionTargetPageId="4" captionText="FIGURES 2 – 5. Habitus images. 2: Anomala eucoma (paralectotype, Las Mercedes, Guatemala). 3: A. flavaeucoma (female paratype, Estación Hitoy Cerere, Costa Rica). 4. A. megaparamera (holotype). 5: A. pseudoeucoma (holotype). Scale = 5 mm." httpUri="https://zenodo.org/record/223578/files/figure.png" pageId="2" pageNumber="257">Fig. 2</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFF5FFD15AA663D6FAA8FB73" lastPageId="3" lastPageNumber="258" pageId="2" pageNumber="257" type="materials_examined">
<paragraph id="8BC43697FFF5FFD05AA663D6FD09FC83" blockId="2.[151,1437,507,2008]" pageId="2" pageNumber="257">
<emphasis id="B90FEA85FFF5FFD05AA663D6FE2FFDA3" bold="true" box="[151,391,507,532]" pageId="2" pageNumber="257">Material examined:</emphasis>
<typeStatus id="54C08835FFF5FFD05BA063D0FDA9FDA2" box="[401,513,509,533]" pageId="2" pageNumber="257" type="lectotype">Lectotype</typeStatus>
(here designated): 1ɞ &quot;Syn-type /
<typeStatus id="54C08835FFF5FFD0599163D0FC70FDA2" box="[928,984,509,533]" pageId="2" pageNumber="257">Type</typeStatus>
/ Las Mercedes,
<quantity id="4C839B72FFF5FFD05E9963D1FB57FDA3" box="[1192,1279,508,532]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="2" pageNumber="257" unit="ft" value="3000.0">3000 ft</quantity>
. Champion. /
<taxonomicName id="4C7B4D14FFF5FFD05AA66032FE37FD80" authority="Bates" authorityName="Bates" box="[151,415,543,567]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="eucoma">Anomala eucoma Bates</taxonomicName>
(handwritten) / B.C.A., Col., II(2)
<taxonomicName id="4C7B4D14FFF5FFD059176032FC27FD80" box="[806,911,543,567]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
&quot; (
<collectionCode id="ED6AAE52FFF5FFD059966032FBA9FD80" box="[935,1025,543,567]" pageId="2" pageNumber="257">BMNH</collectionCode>
).
<typeStatus id="54C08835FFF5FFD05E226032FB13FD80" box="[1043,1211,543,567]" pageId="2" pageNumber="257" type="paralectotype">Paralectotypes</typeStatus>
: 1ɞ &quot;Las Mercedes,
<quantity id="4C839B72FFF5FFD05AA66069FF46FDEB" box="[151,238,580,604]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="2" pageNumber="257" unit="ft" value="3000.0">3000 ft</quantity>
. Champion. /
<taxonomicName id="4C7B4D14FFF5FFD05BA26069FD0BFDEB" authority="Bates" authorityName="Bates" box="[403,675,580,604]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="eucoma">Anomala eucoma Bates</taxonomicName>
(handwritten) / H.W. Bates Biol. Cent. Amer.&quot; (
<collectionCode id="ED6AAE52FFF5FFD05EE96069FA9CFDEB" LSID="urn:lsid:biocol.org:col:34988" box="[1240,1332,580,604]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="257">MNHN</collectionCode>
). ɞ &quot;Las Mercedes,
<quantity id="4C839B72FFF5FFD05B20604AFECDFDC8" box="[273,357,615,639]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="2" pageNumber="257" unit="ft" value="3000.0">3000 ft</quantity>
. Champion. /
<taxonomicName id="4C7B4D14FFF5FFD058316045FD34FDC8" authority="Bates" authorityName="Bates" box="[512,668,616,639]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="eucoma">eucoma Bates</taxonomicName>
(handwritten) / H.W. Bates Biol. Cent. Amer.&quot; (
<collectionCode id="ED6AAE52FFF5FFD05E86604AFABBFDC8" LSID="urn:lsid:biocol.org:col:34988" box="[1207,1299,615,639]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="2" pageNumber="257">MNHN</collectionCode>
). 1ɞ &quot;SYN-
<collectionCode id="ED6AAE52FFF5FFD05AA660A0FF75FD13" box="[151,221,653,676]" pageId="2" pageNumber="257">
<typeStatus id="54C08835FFF5FFD05AA660A0FF75FD13" box="[151,221,653,676]" pageId="2" pageNumber="257">TYPE</typeStatus>
</collectionCode>
/
<collectingCountry id="F36C7607FFF5FFD05AC260A1FED8FD13" box="[243,368,652,676]" name="Costa Rica" pageId="2" pageNumber="257">Costa Rica</collectingCountry>
. V.P. (handwritten) / Locality doubtful (handwritten) / B. C. A., Col. II (2)
<taxonomicName id="4C7B4D14FFF5FFD05E8460A1FAB6FD13" box="[1205,1310,652,676]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
&quot; (
<collectionCode id="ED6AAE52FFF5FFD05F0760A1FA38FD13" box="[1334,1424,652,676]" pageId="2" pageNumber="257">BMNH</collectionCode>
); 1ɞ &quot;SYN-TYPE /
<collectingCountry id="F36C7607FFF5FFD05B566082FE49FD70" box="[359,481,687,711]" name="Costa Rica" pageId="2" pageNumber="257">Costa Rica</collectingCountry>
/ Salle Coll. / 1195 /
<taxonomicName id="4C7B4D14FFF5FFD058FD6082FC7CFD70" authority="Bates" authorityName="Bates" box="[716,980,687,711]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="eucoma">Anomala eucoma Bates</taxonomicName>
(handwritten) / Isonychus sp. apud Sallé (handwritten) / B. C. A., Col. II (2)
<taxonomicName id="4C7B4D14FFF5FFD0580B60F9FD0CFD5B" box="[570,676,724,748]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
&quot; (
<collectionCode id="ED6AAE52FFF5FFD0588C60F9FCBFFD5B" box="[701,791,724,748]" pageId="2" pageNumber="257">BMNH</collectionCode>
). 1ɞ &quot;Pantaleon,
<quantity id="4C839B72FFF5FFD059D260F9FB91FD5B" box="[995,1081,724,748]" metricMagnitude="2" metricUnit="m" metricValue="5.1816" pageId="2" pageNumber="257" unit="ft" value="1700.0">1700 ft</quantity>
. Champion /
<taxonomicName id="4C7B4D14FFF5FFD05EE460F9FA34FD5B" box="[1237,1436,724,748]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="eucoma">Anomala eucoma</taxonomicName>
(handwritten) / B. C. A., Col. II (2)
<taxonomicName id="4C7B4D14FFF5FFD0585D60DAFD71FCB8" box="[620,729,759,783]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
&quot;. 1Ƥ &quot;San Isidro,
<quantity id="4C839B72FFF5FFD059F560DAFB88FCB8" box="[964,1056,759,783]" metricMagnitude="2" metricUnit="m" metricValue="4.8768" pageId="2" pageNumber="257" unit="ft" value="1600.0">1600 ft</quantity>
. Champion /
<taxonomicName id="4C7B4D14FFF5FFD05EFE60DAFA34FCB8" box="[1231,1436,759,783]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="species" species="eucoma">Anomala eucoma</taxonomicName>
(handwritten) / B. C. A., Col. II (2)
<taxonomicName id="4C7B4D14FFF5FFD058196131FD3DFC83" box="[552,661,796,820]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
&quot;.
</paragraph>
<paragraph id="8BC43697FFF5FFD15AF66112FAA8FB73" blockId="2.[151,1437,507,2008]" lastBlockId="3.[151,1436,151,1832]" lastPageId="3" lastPageNumber="258" pageId="2" pageNumber="257">
<emphasis id="B90FEA85FFF5FFD05AF66112FE68FCEF" bold="true" box="[199,448,831,856]" pageId="2" pageNumber="257">Additional material:</emphasis>
1ɞ &quot;
<collectionCode id="ED6AAE52FFF5FFD058336112FDCFFCE0" box="[514,615,831,855]" pageId="2" pageNumber="257">MUSEO</collectionCode>
<collectingCountry id="F36C7607FFF5FFD05843616DFD3FFCE0" box="[626,663,832,855]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="257">DE</collectingCountry>
INSECTOS UNIVERSIDAD
<collectingCountry id="F36C7607FFF5FFD059CF616DFB8BFCE0" box="[1022,1059,832,855]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="257">DE</collectingCountry>
<collectingCountry id="F36C7607FFF5FFD05E1E6112FB74FCE0" box="[1071,1244,831,855]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
,
<collectingCountry id="F36C7607FFF5FFD05EDA6112FA30FCE0" box="[1259,1432,831,855]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
,
<collectionCode id="ED6AAE52FFF5FFD05AA66149FF4DFCCB" box="[151,229,868,892]" pageId="2" pageNumber="257">PROV</collectionCode>
. (printed) San José U.C.R.
<date id="FFC51057FFF5FFD058286149FD26FCCB" box="[537,654,868,892]" pageId="2" pageNumber="257" value="1964-11-12">12–XI–64</date>
. G. Fuentes (handwritten) /
<taxonomicName id="4C7B4D14FFF5FFD059FB6149FB99FCCB" box="[970,1073,868,892]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
? id: Sullivan 86&quot; (
<collectionCode id="ED6AAE52FFF5FFD05F3D6149FACDFCCB" box="[1292,1381,868,892]" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="2" pageNumber="257">MUCR</collectionCode>
); 1ɞ &quot;
<collectionCode id="ED6AAE52FFF5FFD05AAE61AAFEACFC28" box="[159,260,903,927]" pageId="2" pageNumber="257">MUSEO</collectionCode>
<collectingCountry id="F36C7607FFF5FFD05B3C61A5FE9AFC28" box="[269,306,904,927]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="257">DE</collectingCountry>
INSECTOS UNIVERSIDAD
<collectingCountry id="F36C7607FFF5FFD058A061A5FD1EFC28" box="[657,694,904,927]" name="South Georgia and the South Sandwich Islands" pageId="2" pageNumber="257">DE</collectingCountry>
<collectingCountry id="F36C7607FFF5FFD0588E61AAFCC1FC28" box="[703,873,903,927]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
,
<collectingCountry id="F36C7607FFF5FFD0594761AAFB88FC28" box="[886,1056,903,927]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
,
<collectionCode id="ED6AAE52FFF5FFD05E1C61AAFBD4FC28" box="[1069,1148,903,927]" pageId="2" pageNumber="257">PROV</collectionCode>
. (printed) San José Univ. de
<collectingCountry id="F36C7607FFF5FFD05A8A6181FE90FC73" box="[187,312,940,964]" name="Costa Rica" pageId="2" pageNumber="257">Costa Rica</collectingCountry>
<collectingCountry id="F36C7607FFF5FFD05B726181FE13FC73" box="[323,443,940,964]" name="Saint Pierre and Miquelon" pageId="2" pageNumber="257">San Pedro</collectingCountry>
, San José. Alt.
<quantity id="4C839B72FFF5FFD058456181FD67FC73" box="[628,719,940,964]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="2" pageNumber="257" unit="m" value="1200.0">1200 m</quantity>
.
<date id="FFC51057FFF5FFD058EC6181FCC7FC73" box="[733,879,940,964]" pageId="2" pageNumber="257" value="1968-10-10">10–oct 1968</date>
. P.L. Kazan (handwritten) /
<taxonomicName id="4C7B4D14FFF5FFD05E8D6181FA8AFC73" box="[1212,1314,940,964]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="257" phylum="Arthropoda" rank="genus">Anomala</taxonomicName>
sp. Id: D. Corde 1994&quot; (
<collectionCode id="ED6AAE52FFF5FFD05B0F61E2FE3FFC50" box="[318,407,975,999]" httpUri="http://grbio.org/cool/ww9x-1v7b" name="Museo de Insectos" pageId="2" pageNumber="257">MUCR</collectionCode>
); 1ɞ &quot;Amubri, A.C. Amistad, Prov. Limon,
<collectingCountry id="F36C7607FFF5FFD0599D61E2FBF2FC50" box="[940,1114,975,999]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF5FFD05E5B61E2FB07FC50" box="[1130,1199,975,999]" pageId="2" pageNumber="257" type="generic">
<quantity id="4C839B72FFF5FFD05E5B61E2FB03FC50" box="[1130,1195,975,999]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="257" unit="m" value="70.0">70 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF5FFD05E8B61E2FAD9FC50" box="[1210,1393,975,999]" pageId="2" pageNumber="257" value="1994-04-04" valueMax="1994-04-30" valueMin="1994-04-04">4–30 Abr 1994</date>
, G. Gallardo, L S 385500_578100 # 2822 /
<collectingCountry id="F36C7607FFF5FFD058B561D9FC98FBBB" box="[644,816,1012,1036]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI001785409&quot;; 1Ƥ &quot;Amubri, Prov. Limón,
<collectingCountry id="F36C7607FFF5FFD05AA6663AFEE7FB98" box="[151,335,1047,1071]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF5FFD05B59663AFE09FB98" box="[360,417,1047,1071]" pageId="2" pageNumber="257" type="generic">
<quantity id="4C839B72FFF5FFD05B59663AFE36FB98" box="[360,414,1047,1071]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="257" unit="m" value="70.0">70m</quantity>
.
</specimenCount>
<date id="FFC51057FFF5FFD05B87663AFD20FB98" box="[438,648,1047,1071]" pageId="2" pageNumber="257" value="1995-06-01" valueMax="1995-06-22" valueMin="1995-06-01">1–22 JUN 1995</date>
. G. Gallardo, L S 385000 578100 #5333 /
<collectingCountry id="F36C7607FFF5FFD05ED9663AFA34FB98" box="[1256,1436,1047,1071]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI002235510&quot;; 1ɞ &quot;Amubri, Talamanca., A.C. Amistad, Prov. Limón,
<collectingCountry id="F36C7607FFF5FFD05E186611FB7DFBE3" box="[1065,1237,1084,1108]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF5FFD05ED06611FA83FBE3" box="[1249,1323,1084,1108]" pageId="2" pageNumber="257" type="generic">
<quantity id="4C839B72FFF5FFD05ED06611FAB4FBE3" box="[1249,1308,1084,1108]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="257" unit="m" value="70.0">70 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF5FFD05F056611FF7AFBC0" pageId="2" pageNumber="257" value="1994-06-06" valueMax="1994-06-28" valueMin="1994-06-06">6–28 Jun 1994</date>
, G. Gallardo, L N 385000_578100 #3006 /
<collectingCountry id="F36C7607FFF5FFD058DC6672FC31FBC0" box="[749,921,1119,1143]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI001875336&quot;; 1Ƥ &quot;Amubri, A.C. Amistad, Prov. Limon,
<collectingCountry id="F36C7607FFF5FFD05B8066A9FDC9FB2B" box="[433,609,1156,1180]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF5FFD0584066A9FD1FFB2B" box="[625,695,1156,1180]" pageId="2" pageNumber="257" type="generic">
<quantity id="4C839B72FFF5FFD0584066A9FD1BFB2B" box="[625,691,1156,1180]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="257" unit="m" value="70.0">70 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF5FFD058F566A9FC2CFB2B" box="[708,900,1156,1181]" pageId="2" pageNumber="257" value="1994-05-02" valueMax="1994-05-31" valueMin="1994-05-02">2–31 May 1994</date>
, G. Gallardo, L N 385000_578100 # 2928 /
<collectionCode id="ED6AAE52FFF5FFD05AA6668AFF4EFB08" box="[151,230,1191,1215]" pageId="2" pageNumber="257">CEUA</collectionCode>
00105270&quot;; 1ɞ &quot;Amubri, Prov. Limón,
<collectingCountry id="F36C7607FFF5FFD05886668AFCCDFB08" box="[695,869,1191,1215]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF5FFD05942668AFC04FB08" box="[883,940,1191,1215]" pageId="2" pageNumber="257" type="generic">
<quantity id="4C839B72FFF5FFD05942668AFC01FB08" box="[883,937,1191,1215]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="257" unit="m" value="70.0">70m</quantity>
.
</specimenCount>
<date id="FFC51057FFF5FFD05986668AFBC1FB08" box="[951,1129,1191,1215]" pageId="2" pageNumber="257" value="1994-10-01" valueMax="1994-10-22" valueMin="1994-10-01">1–22 Oct 1994</date>
. G. Gallardo, L S 385500 578000 #3266 /
<collectingCountry id="F36C7607FFF5FFD05B7F66E1FE5BFB53" box="[334,499,1228,1252]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI002008852&quot;; 1ɞ &quot;Amubri, Prov. Limón,
<collectingCountry id="F36C7607FFF5FFD05E7466E1FB47FB53" box="[1093,1263,1228,1252]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF5FFD05EC866E1FA9AFB53" box="[1273,1330,1228,1252]" pageId="2" pageNumber="257" type="generic">
<quantity id="4C839B72FFF5FFD05EC866E1FA87FB53" box="[1273,1327,1228,1252]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="257" unit="m" value="70.0">70m</quantity>
.
</specimenCount>
<date id="FFC51057FFF5FFD05F0866E1FF7BFAB0" pageId="2" pageNumber="257" value="1994-09-03" valueMax="1994-09-09" valueMin="1994-09-03">3–9 SET 1994</date>
. G.M. Gallardo, L S 385500 578000 #3201 /
<collectingCountry id="F36C7607FFF5FFD058C766C2FC08FAB0" box="[758,928,1263,1287]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI002018611&quot;; 1ɞ &quot;Amubri, A.C. Amistad, Prov. Limon,
<collectingCountry id="F36C7607FFF5FFD05B826739FDCBFA9B" box="[435,611,1300,1324]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF5FFD058456739FD13FA9B" box="[628,699,1300,1324]" pageId="2" pageNumber="257" type="generic">
<quantity id="4C839B72FFF5FFD058456739FD1EFA9B" box="[628,694,1300,1324]" metricMagnitude="1" metricUnit="m" metricValue="7.0" pageId="2" pageNumber="257" unit="m" value="70.0">70 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF5FFD058F96739FC2BFA9B" box="[712,899,1300,1324]" pageId="2" pageNumber="257" value="1993-12-04" valueMax="1993-12-21" valueMin="1993-12-04">4–21 Dec 1993</date>
, G. Gallardo, L S 385500_578100 # 2480 /
<collectionCode id="ED6AAE52FFF5FFD05AA6671AFF4FFAF8" box="[151,231,1335,1359]" pageId="2" pageNumber="257">CEUA</collectionCode>
00105271&quot;; 1ɞ &quot;Est. Sirena, P. N. Corcovado,
<quantity id="4C839B72FFF5FFD058C0671AFCF5FAF8" box="[753,861,1335,1359]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0 – 100m</quantity>
, Prov. Punt.,
<collectingCountry id="F36C7607FFF5FFD059C2671AFB35FAF8" box="[1011,1181,1335,1359]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
. N. Obando,
<date id="FFC51057FFF5FFD05F1E6715FA30FAF8" box="[1327,1432,1335,1359]" pageId="2" pageNumber="257" value="1990-06">Jun 1990</date>
, L- S
<date id="FFC51057FFF5FFD05AFF6771FE8AFAC3" box="[206,290,1372,1396]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD05BA86771FD97FAC3" box="[409,575,1372,1396]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000644524&quot;; 1ɞ &quot;Est Sirena, Corcovado N. P., Puntarenas Prov.
<collectingCountry id="F36C7607FFF5FFD05AA66752FEEBFA20" box="[151,323,1407,1431]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
,
<quantity id="4C839B72FFF5FFD05B616752FE06FA20" box="[336,430,1407,1431]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
,
<date id="FFC51057FFF5FFD05B8C67ADFD85FA20" box="[445,557,1407,1431]" pageId="2" pageNumber="257" value="1990-01">Jan. 1990</date>
, G. Fonseca, L_S_
<date id="FFC51057FFF5FFD059386752FCF6FA20" box="[777,862,1407,1431]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD059ED6752FB2DFA20" box="[988,1157,1407,1431]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000198374&quot;; 1ɞ &quot;Sirena, Corcovado N. P. Puntarenas Province
<collectingCountry id="F36C7607FFF5FFD058CC6789FC05FA0B" box="[765,941,1444,1468]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
.
<quantity id="4C839B72FFF5FFD0598F6789FB90FA0B" box="[958,1080,1444,1468]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
0 –
<specimenCount id="9D7DFD1EFFF5FFD059C46789FB90FA0B" box="[1013,1080,1444,1468]" pageId="2" pageNumber="257" type="generic">100m</specimenCount>
</quantity>
. G. Fonseca,
<date id="FFC51057FFF5FFD05ED36789FAFBFA0B" box="[1250,1363,1444,1468]" pageId="2" pageNumber="257" value="1989-12">Dic 1989</date>
, L- S
<date id="FFC51057FFF5FFD05AA667EAFF43FA68" box="[151,235,1479,1503]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD05B5267EAFDA1FA68" box="[355,521,1479,1503]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000203726&quot;; 1ɞ &quot;Est. Sirena, P. N. Corcovado,
<quantity id="4C839B72FFF5FFD05E9C67EAFAA3FA68" box="[1197,1291,1479,1503]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, Prov. Punt.,
<collectingCountry id="F36C7607FFF5FFD05AA667C1FEE8F9B3" box="[151,320,1516,1540]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
F. Quesada,
<date id="FFC51057FFF5FFD05BEF67C0FDE3F9B3" box="[478,587,1516,1540]" pageId="2" pageNumber="257" value="1990-06">Jun 1990</date>
, L- S
<date id="FFC51057FFF5FFD058A867C1FD45F9B3" box="[665,749,1516,1540]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD0595D67C1FBBDF9B3" box="[876,1045,1516,1540]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000376353&quot;; 1ɞ &quot;Est. Sirena, P. N. Corcovado,
<quantity id="4C839B72FFF5FFD05BF36422FD83F990" box="[450,555,1551,1575]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0 –100m</quantity>
, Prov. Punt.,
<collectingCountry id="F36C7607FFF5FFD058FD6422FCD0F990" box="[716,888,1551,1575]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
, G. Fonseca,
<date id="FFC51057FFF5FFD05E286422FB2EF990" box="[1049,1158,1551,1575]" pageId="2" pageNumber="257" value="1989-10">Oct 1989</date>
, L- S
<date id="FFC51057FFF5FFD05EE56422FA80F990" box="[1236,1320,1551,1575]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD05AA66419FE95F9FB" box="[151,317,1588,1612]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000446957&quot;; 1ɞ &quot;Est. Sirena,
<quantity id="4C839B72FFF5FFD059296419FCDFF9FB" box="[792,887,1588,1612]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, P. N. Corcovado, Prov. Puntarenas,
<collectingCountry id="F36C7607FFF5FFD05F2B6419FA30F9FB" box="[1306,1432,1588,1612]" name="Costa Rica" pageId="2" pageNumber="257">Costa Rica</collectingCountry>
, G. Rodriguez,
<date id="FFC51057FFF5FFD05B70647AFE19F9D8" box="[321,433,1623,1647]" pageId="2" pageNumber="257" value="1992-04">Abr 1992</date>
, L- S
<date id="FFC51057FFF5FFD05BCA647AFDE7F9D8" box="[507,591,1623,1647]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectionCode id="ED6AAE52FFF5FFD058FD647AFCB3F9D8" box="[716,795,1623,1647]" pageId="2" pageNumber="257">CEUA</collectionCode>
00105265&quot;; 1Ƥ &quot;Est. Sirena,
<quantity id="4C839B72FFF5FFD05E58647AFB6FF9D8" box="[1129,1223,1623,1647]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, P. N. Corcovado, Prov. Punt.,
<collectingCountry id="F36C7607FFF5FFD05B136451FE63F923" box="[290,459,1660,1684]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
, G. Fonseca,
<date id="FFC51057FFF5FFD058526450FD79F923" box="[611,721,1660,1684]" pageId="2" pageNumber="257" value="1992-01">Ene 1992</date>
, L- S
<date id="FFC51057FFF5FFD059246451FCC1F923" box="[789,873,1660,1684]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD059D06451FB2FF923" box="[993,1159,1660,1684]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000506778&quot;; 1ɞ &quot;Est. Sirena, P. N. Corcovado,
<quantity id="4C839B72FFF5FFD0586A64B2FD12F900" box="[603,698,1695,1719]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, Prov. Punt.,
<collectingCountry id="F36C7607FFF5FFD0594164B2FB8CF900" box="[880,1060,1695,1719]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
. N. Obando,
<date id="FFC51057FFF5FFD05EE6648DFAE2F900" box="[1239,1354,1695,1719]" pageId="2" pageNumber="257" value="1990-06">Jun 1990</date>
, L- S
<date id="FFC51057FFF5FFD05AA664E9FF43F96B" box="[151,235,1732,1756]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD05B5E64E9FDB4F96B" box="[367,540,1732,1756]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000644521&quot;; 1Ƥ &quot;Est. Sirena,
<quantity id="4C839B72FFF5FFD05E2564E9FBDAF96B" box="[1044,1138,1732,1756]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, P. N. Corcovado, Prov. Puntarenas,
<collectingCountry id="F36C7607FFF5FFD05B2E64CAFE34F948" box="[287,412,1767,1791]" name="Costa Rica" pageId="2" pageNumber="257">Costa Rica</collectingCountry>
,
<date id="FFC51057FFF5FFD05B9664C5FDA7F948" box="[423,527,1767,1791]" pageId="2" pageNumber="257" value="1992-06">Jun 1992</date>
, G. Fonseca, L -S
<date id="FFC51057FFF5FFD058EC64CAFC99F948" box="[733,817,1767,1791]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectionCode id="ED6AAE52FFF5FFD0599964CAFC5FF948" box="[936,1015,1767,1791]" pageId="2" pageNumber="257">CEUA</collectionCode>
00105264&quot;; 1ɞ &quot;Est. Sirena,
<quantity id="4C839B72FFF5FFD05F0864CAFA3FF948" box="[1337,1431,1767,1791]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">0–100m</quantity>
, P. N. Corcovado, Prov. Puntarenas,
<collectingCountry id="F36C7607FFF5FFD0580E6521FD17F893" box="[575,703,1804,1828]" name="Costa Rica" pageId="2" pageNumber="257">Costa Rica</collectingCountry>
,
<date id="FFC51057FFF5FFD058FE6520FC93F893" box="[719,827,1804,1828]" pageId="2" pageNumber="257" value="1992-06">Jun 1992</date>
, G. Fonseca, L -S
<date id="FFC51057FFF5FFD05E2E6521FBDBF893" box="[1055,1139,1804,1828]" pageId="2" pageNumber="257" value="00-05-27">270500</date>
_508300 /
<collectingCountry id="F36C7607FFF5FFD05EC36521FA34F893" box="[1266,1436,1804,1828]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
INBIOCRI000714400&quot;; 1Ƥ &quot;
<collectingCountry id="F36C7607FFF5FFD05BD76502FD3FF8F0" box="[486,663,1839,1863]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
. Prov. Puntarenas. P.N. Corcovado, Est. Sirena.
<quantity id="4C839B72FFF5FFD05EE26502FA9BF8F0" box="[1235,1331,1839,1863]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
0–
<specimenCount id="9D7DFD1EFFF5FFD05EC06502FA9BF8F0" box="[1265,1331,1839,1863]" pageId="2" pageNumber="257" type="generic">100m</specimenCount>
</quantity>
.
<date id="FFC51057FFF5FFD05F706502FF7BF8DB" pageId="2" pageNumber="257" value="2004-02-21">21 FEB 2004</date>
. E. Holzer. C. Libre. L_S_
<date id="FFC51057FFF5FFD058266579FDC4F8DB" box="[535,620,1876,1900]" pageId="2" pageNumber="257" value="50-08-27">270850</date>
_509176 #92946 / INB0004129881 INBIOCRI
<collectingCountry id="F36C7607FFF5FFD05E936579FAE5F8DB" box="[1186,1357,1876,1900]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
&quot;; 1Ƥ &quot;
<collectingCountry id="F36C7607FFF5FFD05AAE655AFEF9F838" box="[159,337,1911,1935]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
. Prov. Puntarenas. P.N. Corcovado, Est. Sirena.
<quantity id="4C839B72FFF5FFD059A1655AFC47F838" box="[912,1007,1911,1935]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
0–
<specimenCount id="9D7DFD1EFFF5FFD0599F655AFC47F838" box="[942,1007,1911,1935]" pageId="2" pageNumber="257" type="generic">100m</specimenCount>
</quantity>
.
<date id="FFC51057FFF5FFD059CE655AFB09F838" box="[1023,1185,1911,1935]" pageId="2" pageNumber="257" value="2004-02-21">21 FEB 2004</date>
. E. Holzer. C. Libre. L_S_
<date id="FFC51057FFF5FFD05AE065B1FE8EF803" box="[209,294,1948,1972]" pageId="2" pageNumber="257" value="50-08-27">270850</date>
_509176 #92946 / INB0004129887 INBIOCRI
<collectingCountry id="F36C7607FFF5FFD0594D65B1FB9CF803" box="[892,1076,1948,1972]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
&quot;; 1ɞ &quot;
<collectingCountry id="F36C7607FFF5FFD05EA365B1FAE2F803" box="[1170,1354,1948,1972]" name="Costa Rica" pageId="2" pageNumber="257">COSTA RICA</collectingCountry>
. Prov. Puntarenas. P.N. Corcovado, Est. Sirena.
<quantity id="4C839B72FFF5FFD058B665EDFD4EF86F" box="[647,742,1984,2008]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="2" pageNumber="257" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
0–
<specimenCount id="9D7DFD1EFFF5FFD0589465EDFD4EF86F" box="[677,742,1984,2008]" pageId="2" pageNumber="257" type="generic">100m</specimenCount>
</quantity>
.
<date id="FFC51057FFF5FFD058C665EDFC34F86F" box="[759,924,1984,2008]" pageId="2" pageNumber="257" value="2004-02-21">21 FEB 2004</date>
. E. Holzer. C. Libre. L_S_
<date id="FFC51057FFF5FFD05ED565EDFA91F86F" box="[1252,1337,1984,2008]" pageId="2" pageNumber="257" value="50-08-27">270850</date>
_509176 #92946 / INB0004129886 INBIOCRI
<collectingCountry id="F36C7607FFF4FFD1586262BAFCABFF18" box="[595,771,151,175]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
&quot;; 1ɞ &quot;
<collectingCountry id="F36C7607FFF4FFD1597E62BAFC56FF18" box="[847,1022,151,175]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
. Prov. Puntarenas. P.N. Corcovado, Est. Sirena.
<quantity id="4C839B72FFF4FFD15B1E6291FE27FF63" box="[303,399,188,212]" metricMagnitude="1" metricUnit="m" metricValue="5.0" metricValueMax="10.0" metricValueMin="0.0" pageId="3" pageNumber="258" unit="m" value="50.0" valueMax="100.0" valueMin="0.0">
0–
<specimenCount id="9D7DFD1EFFF4FFD15B7D6291FE27FF63" box="[332,399,188,212]" pageId="3" pageNumber="258" type="generic">100m</specimenCount>
</quantity>
.
<date id="FFC51057FFF4FFD15B906291FDE3FF63" box="[417,587,188,212]" pageId="3" pageNumber="258" value="2004-02-21">21 FEB 2004</date>
. E. Holzer. C. Libre. L_S_
<date id="FFC51057FFF4FFD159AF6291FC5BFF63" box="[926,1011,188,212]" pageId="3" pageNumber="258" value="50-08-27">270850</date>
_509176 #92946 / INB0004129883 INBIOCRI
<collectingCountry id="F36C7607FFF4FFD15B2F62F2FE66FF40" box="[286,462,223,247]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
&quot;; 1 ɞ &quot;Est. Palo Verde,
<quantity id="4C839B72FFF4FFD158C362F2FC8EFF40" box="[754,806,223,247]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="258" unit="m" value="10.0">10m</quantity>
, Ref. Nac. Fauna Silv. R. L. Rodriguez, Prov. Guan.,
<collectingCountry id="F36C7607FFF4FFD15AA66329FEEAFEAB" box="[151,322,260,284]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
, D. Acevedo,
<date id="FFC51057FFF4FFD15BD96328FDFAFEAB" box="[488,594,260,284]" pageId="3" pageNumber="258" value="1991-06">Jun 1991</date>
, L- N 259000_388400 /
<collectingCountry id="F36C7607FFF4FFD159406329FBB2FEAB" box="[881,1050,260,284]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI000652257&quot;; 1ɞ &quot;Est. Palo Verde,
<quantity id="4C839B72FFF4FFD15B10630AFEFEFE88" box="[289,342,295,319]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="3" pageNumber="258" unit="m" value="10.0">10m</quantity>
, Ref. Nac. Fauna Silv. R. L. Rodriguez, Prov. Guan.,
<collectingCountry id="F36C7607FFF4FFD1599C630AFBFEFE88" box="[941,1110,295,319]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
, D. Acevedo,
<date id="FFC51057FFF4FFD15EC26305FAF3FE88" box="[1267,1371,295,319]" pageId="3" pageNumber="258" value="1991-06">Jun 1991</date>
, L- N 259000_388400 /
<collectingCountry id="F36C7607FFF4FFD15B596361FDB9FED3" box="[360,529,332,356]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI000652258&quot;; 1ɞ &quot;Est. Las Pailas,
<quantity id="4C839B72FFF4FFD15E146361FBCFFED3" box="[1061,1127,332,356]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800m</quantity>
, P. N. Rincon de la Vieja, Prov. Guanacaste,
<collectingCountry id="F36C7607FFF4FFD15BB36342FDA4FE30" box="[386,524,367,391]" name="Costa Rica" pageId="3" pageNumber="258">Costa Rica</collectingCountry>
,
<date id="FFC51057FFF4FFD158156342FCA9FE30" box="[548,769,367,391]" pageId="3" pageNumber="258" value="1992-07-01" valueMax="1992-07-22" valueMin="1992-07-01">1 a 22 jul 1992</date>
, D. Garcia, L- N 306300_388600 /
<collectingCountry id="F36C7607FFF4FFD15EDB6342FA34FE30" box="[1258,1436,367,391]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI000713719&quot;; 1ɞ &quot;Fca. Jenny,
<quantity id="4C839B72FFF4FFD1585963B9FD18FE1B" box="[616,688,404,428]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="258" unit="km" value="30.0">30 km</quantity>
N de
<collectingCountry id="F36C7607FFF4FFD158C563B9FCE0FE1B" box="[756,840,404,428]" name="Liberia" pageId="3" pageNumber="258">Liberia</collectingCountry>
, P. N. Guanacaste, Prov. Guanacaste,
<collectingCountry id="F36C7607FFF4FFD15EC663B9FADCFE1B" box="[1271,1396,404,428]" name="Costa Rica" pageId="3" pageNumber="258">Costa Rica</collectingCountry>
,
<date id="FFC51057FFF4FFD15FB163B9FECEFE78" pageId="3" pageNumber="258" value="1992-08-20" valueMax="1992-09-12" valueMin="1992-08-20">20 ago a 12 set 1992</date>
, E. Araya, L N 316200_364400 /
<collectingCountry id="F36C7607FFF4FFD158DF639AFC3EFE78" box="[750,918,439,463]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI000849675&quot;; 1ɞ &quot;Est. Las Pailas,
<quantity id="4C839B72FFF4FFD15AA663F1FF4FFE43" box="[151,231,476,500]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800 m</quantity>
, P.N. Rincon de la Vieja, Prov. Guan.
<collectingCountry id="F36C7607FFF4FFD158E563F1FC2DFE43" box="[724,901,476,500]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<date id="FFC51057FFF4FFD159A763F1FB34FE43" box="[918,1180,476,500]" pageId="3" pageNumber="258" value="1993-05-15" valueMax="1993-06-11" valueMin="1993-05-15">15 May–11 Jun 1993</date>
, K. E. Taylor, L- N 306300_388600 /
<collectionCode id="ED6AAE52FFF4FFD15B5E63D2FE16FDA0" box="[367,446,511,535]" pageId="3" pageNumber="258">CEUA</collectionCode>
00105267&quot;; 1ɞ &quot;Finca Jenny,
<quantity id="4C839B72FFF4FFD1591663D2FCDCFDA0" box="[807,884,511,535]" metricMagnitude="4" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="258" unit="km" value="30.0">30 km</quantity>
N de
<collectingCountry id="F36C7607FFF4FFD159FB63D2FBB6FDA0" box="[970,1054,511,535]" name="Liberia" pageId="3" pageNumber="258">Liberia</collectingCountry>
, P.N. Guanacaste, Prov. Guana,
<collectingCountry id="F36C7607FFF4FFD15AA66009FEFBFD8B" box="[151,339,548,572]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF4FFD15B5F6009FE1EFD8B" box="[366,438,548,572]" pageId="3" pageNumber="258" type="generic">
<quantity id="4C839B72FFF4FFD15B5F6009FE19FD8B" box="[366,433,548,572]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="258" unit="m" value="300.0">300m</quantity>
.
</specimenCount>
<date id="FFC51057FFF4FFD15BFF6008FDF9FD8B" box="[462,593,548,572]" pageId="3" pageNumber="258" value="1991-03">Mar 1991</date>
. R. Espinoza, L N 316200_364400 # 1678 /
<collectingCountry id="F36C7607FFF4FFD15ED56009FA34FD8B" box="[1252,1436,548,572]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001693034&quot;; 1ɞ &quot;Fca. Jenny,
<quantity id="4C839B72FFF4FFD15840606AFD1BFDE8" box="[625,691,583,607]" metricMagnitude="2" metricUnit="m" metricValue="3.0" pageId="3" pageNumber="258" unit="m" value="300.0">300m</quantity>
, 31 Km N.
<collectingCountry id="F36C7607FFF4FFD1590E606AFC3BFDE8" box="[831,915,583,607]" name="Liberia" pageId="3" pageNumber="258">Liberia</collectingCountry>
, Guanacaste Prov.
<collectingCountry id="F36C7607FFF4FFD15E40606AFAB5FDE8" box="[1137,1309,583,607]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
,
<date id="FFC51057FFF4FFD15F1B6065FA34FDE8" box="[1322,1436,583,607]" pageId="3" pageNumber="258" value="1988-11">Nov 1988</date>
GNP Biodiversity Survey
<geoCoordinate id="EE4F5050FFF4FFD15BF06041FDECFD33" box="[449,580,620,644]" direction="west" orientation="longitude" pageId="3" pageNumber="258" precision="15" value="-85.574165">W85 34 27</geoCoordinate>
&quot;,
<geoCoordinate id="EE4F5050FFF4FFD1586B6041FD7DFD33" box="[602,725,620,644]" direction="north" orientation="latitude" pageId="3" pageNumber="258" precision="15" value="10.865278">N10 51 55</geoCoordinate>
&quot; /
<collectingCountry id="F36C7607FFF4FFD158C66041FC35FD33" box="[759,925,620,644]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI002517326&quot;; 1ɞ &quot;Estac. Maritza,
<quantity id="4C839B72FFF4FFD15AA660A2FF77FD10" box="[151,223,655,679]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="258" unit="m" value="600.0">600 m</quantity>
, W side Volcan Orosi Guanac. Pr.
<collectingCountry id="F36C7607FFF4FFD1585560A2FCA5FD10" box="[612,781,655,679]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<date id="FFC51057FFF4FFD1592660BDFC24FD10" box="[791,908,655,679]" pageId="3" pageNumber="258" value="1988-06">June 1988</date>
. Janzen &amp; Hallwachs
<geoCoordinate id="EE4F5050FFF4FFD15EB360A2FB4CFD10" box="[1154,1252,655,679]" direction="west" orientation="longitude" pageId="3" pageNumber="258" precision="925" value="-85.48333">W85 29`</geoCoordinate>
37&quot;.
<geoCoordinate id="EE4F5050FFF4FFD15F2460A2FADCFD10" box="[1301,1396,655,679]" direction="north" orientation="latitude" pageId="3" pageNumber="258" precision="925" value="10.95">N10 57`</geoCoordinate>
39&quot; /
<collectingCountry id="F36C7607FFF4FFD15A996099FEF8FD7B" box="[168,336,692,716]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI002517343&quot;; 1ɞ &quot;Estac. Maritza,
<quantity id="4C839B72FFF4FFD1596D6099FC0FFD7B" box="[860,935,692,716]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="3" pageNumber="258" unit="m" value="600.0">600 m</quantity>
, W side Volcan Orosi Guanac. Pr.
<collectingCountry id="F36C7607FFF4FFD15F716099FF76FD58" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<date id="FFC51057FFF4FFD15ADA60F5FECCFD58" box="[235,356,727,751]" pageId="3" pageNumber="258" value="1988-06">June 1988</date>
. Janzen &amp; Hallwachs
<geoCoordinate id="EE4F5050FFF4FFD1585B60FAFD79FD58" box="[618,721,727,751]" direction="west" orientation="longitude" pageId="3" pageNumber="258" precision="925" value="-85.48333">W85 29`</geoCoordinate>
37&quot;.
<geoCoordinate id="EE4F5050FFF4FFD1593760FAFCC1FD58" box="[774,873,727,751]" direction="north" orientation="latitude" pageId="3" pageNumber="258" precision="925" value="10.95">N10 57`</geoCoordinate>
39&quot; /
<collectingCountry id="F36C7607FFF4FFD1599C60FAFBFFFD58" box="[941,1111,727,751]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI002517348&quot;; 1Ƥ &quot;Tierras Morenas,
<quantity id="4C839B72FFF4FFD15B5D60D1FE07FCA3" box="[364,431,764,788]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="258" unit="m" value="700.0">700m</quantity>
, Prov. Guan.,
<collectingCountry id="F36C7607FFF4FFD1586560D1FD57FCA3" box="[596,767,764,788]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<date id="FFC51057FFF4FFD1593A60D0FC2AFCA3" box="[779,898,764,789]" pageId="3" pageNumber="258" value="1993-05">May 1993</date>
. G. Rodriguez. L-N-283950, 424500 /
<collectingCountry id="F36C7607FFF4FFD15F7160D1FF72FC80" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001180822&quot;; 1Ƥ &quot;Tierras Morenas,
<quantity id="4C839B72FFF4FFD159156132FCCEFC80" box="[804,870,799,823]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="258" unit="m" value="700.0">700m</quantity>
, Prov. Guan.,
<collectingCountry id="F36C7607FFF4FFD15E156132FB70FC80" box="[1060,1240,799,823]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<date id="FFC51057FFF4FFD15EDD610DFAC4FC80" box="[1260,1388,799,824]" pageId="3" pageNumber="258" value="1993-05">May 1993</date>
. G. Rodriguez. L-N-283950, 424500 /
<collectingCountry id="F36C7607FFF4FFD158136169FD61FCEB" box="[546,713,836,860]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001180834&quot;; 1ɞ &quot;Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana.
<collectingCountry id="F36C7607FFF4FFD1587D614AFD5FFCC8" box="[588,759,871,895]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF4FFD15933614AFCF9FCC8" box="[770,849,871,895]" pageId="3" pageNumber="258" type="generic">
<quantity id="4C839B72FFF4FFD15933614AFCE5FCC8" box="[770,845,871,895]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF4FFD15968614AFBB8FCC8" box="[857,1040,871,896]" pageId="3" pageNumber="258" value="1994-05-08" valueMax="1994-05-26" valueMin="1994-05-08">8–26 May 1994</date>
. K.E. Taylor, L N 306300_388600 #2912 /
<collectingCountry id="F36C7607FFF4FFD15AC661A1FE08FC13" box="[247,416,908,932]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001861399&quot;; 1ɞ &quot;Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana.
<collectingCountry id="F36C7607FFF4FFD15B046182FE48FC70" box="[309,480,943,967]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF4FFD15BDD6182FD94FC70" box="[492,572,943,967]" pageId="3" pageNumber="258" type="generic">
<quantity id="4C839B72FFF4FFD15BDD6182FD90FC70" box="[492,568,943,967]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF4FFD158756182FD54FC70" box="[580,764,943,968]" pageId="3" pageNumber="258" value="1994-05-08" valueMax="1994-05-26" valueMin="1994-05-08">8–26 May 1994</date>
. K.E. Taylor, L N 306300_388600 #2912 /
<collectingCountry id="F36C7607FFF4FFD15EC56182FA34FC70" box="[1268,1436,943,967]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001861400&quot;; 1Ƥ &quot;Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana.
<collectingCountry id="F36C7607FFF4FFD15F7161F9FF76FBB8" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF4FFD15AD961DAFE9FFBB8" box="[232,311,1015,1039]" pageId="3" pageNumber="258" type="generic">
<quantity id="4C839B72FFF4FFD15AD961DAFE9BFBB8" box="[232,307,1015,1039]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF4FFD15B0F61DAFE5CFBB8" box="[318,500,1015,1040]" pageId="3" pageNumber="258" value="1994-05-08" valueMax="1994-05-26" valueMin="1994-05-08">8–26 May 1994</date>
. K.E. Taylor, L N 306300_388600 #2912 /
<collectingCountry id="F36C7607FFF4FFD159D061DAFB2FFBB8" box="[993,1159,1015,1039]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001861401&quot;; 1Ƥ &quot;Est. Las Pailas, P.N. Rincón de la Vieja, A.C. Guanacaste, Prov. Guana.
<collectingCountry id="F36C7607FFF4FFD15E266631FB6AFB83" box="[1047,1218,1052,1076]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF4FFD15EFF6631FAB7FB83" box="[1230,1311,1052,1076]" pageId="3" pageNumber="258" type="generic">
<quantity id="4C839B72FFF4FFD15EFF6631FAB3FB83" box="[1230,1307,1052,1076]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF4FFD15F166631FF7BFBE0" pageId="3" pageNumber="258" value="1994-05-07" valueMax="1994-05-26" valueMin="1994-05-07">7–26 May 1994</date>
. D. García, L N 306300_388600 #2910 /
<collectingCountry id="F36C7607FFF4FFD158856612FCF3FBE0" box="[692,859,1087,1111]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001879319&quot;; 1Ƥ &quot;Est. Las Pailas, P.N. Rincón de la Vieja, Prov. Guanacaste,
<collectingCountry id="F36C7607FFF4FFD1587C6649FD63FBCB" box="[589,715,1124,1148]" name="Costa Rica" pageId="3" pageNumber="258">Costa Rica</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF4FFD158E76649FC8DFBCB" box="[726,805,1124,1148]" pageId="3" pageNumber="258" type="generic">
<quantity id="4C839B72FFF4FFD158E76649FC89FBCB" box="[726,801,1124,1148]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF4FFD1591C6649FBAEFBCB" box="[813,1030,1124,1148]" pageId="3" pageNumber="258" value="1993-06-19" valueMax="1993-07-01" valueMin="1993-06-19">19 Jun– 1 Jul 1993</date>
. D. G. García, L N 306300_388600 #2189 /
<collectingCountry id="F36C7607FFF4FFD15ACF66AAFE01FB28" box="[254,425,1159,1183]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
INBIOCRI001967801&quot;; 1Ƥ &quot;Est. Las Pailas, P.N. Rincón de la Vieja, Prov. Guana.
<collectingCountry id="F36C7607FFF4FFD15AA66681FEE9FB73" box="[151,321,1196,1220]" name="Costa Rica" pageId="3" pageNumber="258">COSTA RICA</collectingCountry>
.
<specimenCount id="9D7DFD1EFFF4FFD15B7A6681FE31FB73" box="[331,409,1196,1220]" pageId="3" pageNumber="258" type="generic">
<quantity id="4C839B72FFF4FFD15B7A6681FE3DFB73" box="[331,405,1196,1220]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="258" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
<date id="FFC51057FFF4FFD15B916681FDF7FB73" box="[416,607,1196,1220]" pageId="3" pageNumber="258" value="1993-08-16" valueMax="1993-08-24" valueMin="1993-08-16">16–24 Ago 1993</date>
. D. García, L N 306300_388600 #2268 /
<collectionCode id="ED6AAE52FFF4FFD15E016681FBD7FB73" box="[1072,1151,1196,1220]" pageId="3" pageNumber="258">CEUA</collectionCode>
00105266&quot;.
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFF4FFDA5AF666E2FE97FCA3" lastPageId="8" lastPageNumber="263" pageId="3" pageNumber="258" type="description">
<paragraph id="8BC43697FFF4FFD15AF666E2FBD4FA98" blockId="3.[151,1436,151,1832]" pageId="3" pageNumber="258">
<emphasis id="B90FEA85FFF4FFD15AF666E2FEF1FB5F" bold="true" box="[199,345,1231,1256]" pageId="3" pageNumber="258">Description.</emphasis>
Male. Body shape oval. Length
<quantity id="4C839B72FFF4FFD158EB66E2FCFBFB50" box="[730,851,1231,1255]" metricMagnitude="-2" metricUnit="m" metricValue="1.215" pageId="3" pageNumber="258" unit="mm" value="12.15">12.15 mm</quantity>
. Width
<quantity id="4C839B72FFF4FFD1598066E2FBB4FB50" box="[945,1052,1231,1255]" metricMagnitude="-3" metricUnit="m" metricValue="6.9" pageId="3" pageNumber="258" unit="mm" value="6.9">6.90 mm</quantity>
. Head, pronotum, scutellum, and pygidium dark brown; legs and underside reddish brown; elytra dark brown with two slightly lighter horizontal bands (along anterior margin and on elytral disc). Surface covered with fine, blond setae.
</paragraph>
<paragraph id="8BC43697FFF4FFD15AF66711FEA6FA08" blockId="3.[151,1436,151,1832]" pageId="3" pageNumber="258">
Clypeus trapezoidal, densely punctate-reticulate; ratio width/length 1.91. Anterior angles rounded. Frons densely punctate, flat, with apparent boundary with vertex (due to an abrupt change in inclination of the two surfaces) (
<figureCitation id="13402A12FFF4FFD15B3D67A9FEE7FA2B" box="[268,335,1412,1436]" captionStart="FIGURE 6" captionStartId="5.[151,255,561,585]" captionTargetBox="[362,1227,193,533]" captionTargetId="figure@5.[340,1248,176,556]" captionTargetPageId="5" captionText="FIGURE 6. Head of a: Anomala eucoma (paralectotype, Las Mercedes, Guatemala) and b: A. eucoma (paralectotype, precise locality unknown, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223579/files/figure.png" pageId="3" pageNumber="258">Fig. 6</figureCitation>
a). Ocular canthum long and thin, with rounded apex. Interocular ratio (interocular width/width of eye): 3.15.
</paragraph>
<paragraph id="8BC43697FFF4FFD15AF667E0FD29F99B" blockId="3.[151,1436,151,1832]" pageId="3" pageNumber="258">Pronotum trapezoidal, width 1.57 times length. Lateral margins forming an obtuse angle at 1/3 of pronotum length. Anterior angles right and sharp, posterior angles obtuse and rounded. Basal margin thin and complete, sinuate. Entire surface with deep punctures.</paragraph>
<paragraph id="8BC43697FFF4FFD15AF6641AFF56F9C3" blockId="3.[151,1436,151,1832]" pageId="3" pageNumber="258">Scutellum sub-pentagonal in shape, with rounded sides and acute apex, ratio width/length 1.26; sparsely punctate.</paragraph>
<paragraph id="8BC43697FFF4FFD15AF66452FE7CF957" blockId="3.[151,1436,151,1832]" pageId="3" pageNumber="258">Elytra with defined costae and interstices with sparse, large punctures, somewhat obliterated by the numerous and smaller setae-bearing punctures. Lateral margin convex, larger at base, disappearing on apex. Marginal membrane almost complete.</paragraph>
<paragraph id="8BC43697FFF4FFD15AF664C1FE44F89F" blockId="3.[151,1436,151,1832]" pageId="3" pageNumber="258">Pygidium finely granulate, with longer setae at hind margins. Triangular in shape. Slightly convex in lateral view. Ratio width/length 1.58.</paragraph>
<caption id="DF04661FFFF3FFD65AA665A4FE9AF856" httpUri="https://zenodo.org/record/223578/files/figure.png" pageId="4" pageNumber="259" targetBox="[193,1406,206,1890]" targetPageId="4">
<paragraph id="8BC43697FFF3FFD65AA665A4FE9AF856" blockId="4.[151,1437,1929,2017]" pageId="4" pageNumber="259">
<emphasis id="B90FEA85FFF3FFD65AA665A4FEEEF816" bold="true" box="[151,326,1929,1953]" pageId="4" pageNumber="259">FIGURES 2–5.</emphasis>
Habitus images. 2:
<taxonomicName id="4C7B4D14FFF3FFD6581B65A7FD40F816" box="[554,744,1930,1953]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF3FFD6581B65A7FD40F816" box="[554,744,1930,1953]" italics="true" pageId="4" pageNumber="259">Anomala eucoma</emphasis>
</taxonomicName>
(paralectotype, Las Mercedes, Guatemala). 3:
<taxonomicName id="4C7B4D14FFF3FFD65EC365A7FA34F816" box="[1266,1436,1930,1953]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="flavaeucoma">
<emphasis id="B90FEA85FFF3FFD65EC365A7FA34F816" box="[1266,1436,1930,1953]" italics="true" pageId="4" pageNumber="259">A. flavaeucoma</emphasis>
</taxonomicName>
(female paratype, Estación Hitoy Cerere, Costa Rica). 4.
<taxonomicName id="4C7B4D14FFF3FFD658CE6587FC69F876" box="[767,961,1962,1985]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="megaparamera">
<emphasis id="B90FEA85FFF3FFD658CE6587FC69F876" box="[767,961,1962,1985]" italics="true" pageId="4" pageNumber="259">A. megaparamera</emphasis>
</taxonomicName>
(holotype). 5:
<taxonomicName id="4C7B4D14FFF3FFD65E536587FAB7F876" box="[1122,1311,1962,1985]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="259" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFF3FFD65E536587FAB7F876" box="[1122,1311,1962,1985]" italics="true" pageId="4" pageNumber="259">A. pseudoeucoma</emphasis>
</taxonomicName>
(holotype). Scale = 5 mm.
</paragraph>
</caption>
<caption id="DF04661FFFF2FFD75AA6601CFD7EFDDF" httpUri="https://zenodo.org/record/223579/files/figure.png" pageId="5" pageNumber="260" targetBox="[362,1227,193,533]" targetPageId="5">
<paragraph id="8BC43697FFF2FFD75AA6601CFD7EFDDF" blockId="5.[151,1436,561,616]" pageId="5" pageNumber="260">
<emphasis id="B90FEA85FFF2FFD75AA6601CFEB2FDFE" bold="true" box="[151,282,561,585]" pageId="5" pageNumber="260">FIGURE 6.</emphasis>
Head of a:
<taxonomicName id="4C7B4D14FFF2FFD75BAE601CFDF3FDFF" box="[415,603,561,584]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF2FFD75BAE601CFDF3FDFF" box="[415,603,561,584]" italics="true" pageId="5" pageNumber="260">Anomala eucoma</emphasis>
</taxonomicName>
(paralectotype, Las Mercedes, Guatemala) and b:
<taxonomicName id="4C7B4D14FFF2FFD75EB3601CFB5CFDFF" box="[1154,1268,561,584]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF2FFD75EB3601CFB5CFDFF" box="[1154,1268,561,584]" italics="true" pageId="5" pageNumber="260">A. eucoma</emphasis>
</taxonomicName>
(paralectotype, precise locality unknown, Costa Rica). Scale = 1 mm.
</paragraph>
</caption>
<caption id="DF04661FFFF2FFD75AA666A9FB17FB6B" httpUri="https://zenodo.org/record/223580/files/figure.png" pageId="5" pageNumber="260" targetBox="[174,1395,662,1128]" targetPageId="5">
<paragraph id="8BC43697FFF2FFD75AA666A9FB17FB6B" blockId="5.[151,1436,1156,1244]" pageId="5" pageNumber="260">
<emphasis id="B90FEA85FFF2FFD75AA666A9FEF9FB2B" bold="true" box="[151,337,1156,1180]" pageId="5" pageNumber="260">FIGURES 7–10.</emphasis>
Shape of protibia in 7:
<taxonomicName id="4C7B4D14FFF2FFD7586466A8FCB9FB2B" box="[597,785,1157,1180]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF2FFD7586466A8FCB9FB2B" box="[597,785,1157,1180]" italics="true" pageId="5" pageNumber="260">Anomala eucoma</emphasis>
</taxonomicName>
(paralectotype, Las Mercedes, Guatemala); 8:
<taxonomicName id="4C7B4D14FFF2FFD75F3F66A8FA34FB2B" box="[1294,1436,1157,1180]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="flavacoma">
<emphasis id="B90FEA85FFF2FFD75F3F66A8FA34FB2B" box="[1294,1436,1157,1180]" italics="true" pageId="5" pageNumber="260">A. flavacoma</emphasis>
</taxonomicName>
(3 holotype, Ƥ Hitoy Cerere, Costa Rica); 9:
<taxonomicName id="4C7B4D14FFF2FFD758B16688FCE9FB0B" box="[640,833,1189,1212]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="megaparamera">
<emphasis id="B90FEA85FFF2FFD758B16688FCE9FB0B" box="[640,833,1189,1212]" italics="true" pageId="5" pageNumber="260">A. megaparamera</emphasis>
</taxonomicName>
(3 holotype, Ƥ paratype, Cuatro Esquinas, Costa Rica); 10:
<taxonomicName id="4C7B4D14FFF2FFD75A8E66E8FED3FB6B" box="[191,379,1221,1244]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="260" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFF2FFD75A8E66E8FED3FB6B" box="[191,379,1221,1244]" italics="true" pageId="5" pageNumber="260">A. pseudoeucoma</emphasis>
</taxonomicName>
(3 holotype, Ƥ paratype, Reserva de San Ramón, Costa Rica). Scale = 1 mm.
</paragraph>
</caption>
<paragraph id="8BC43697FFF2FFD75AF66724FAAFFAF2" blockId="5.[151,1437,1289,2033]" pageId="5" pageNumber="260">
Metasternal disc sulcated, with setigerous punctures on the entire surface. Space between the mesocoxae narrow, slightly convex, width
<quantity id="4C839B72FFF2FFD75BC46700FDF4FAF2" box="[501,604,1325,1349]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="5" pageNumber="260" unit="mm" value="0.25">0.25 mm</quantity>
. Mesometasternal suture well defined, at base of mesocoxae.
</paragraph>
<paragraph id="8BC43697FFF2FFD75AF6677CFE82FA06" blockId="5.[151,1437,1289,2033]" pageId="5" pageNumber="260">Abdominal sternites with 5–8 irregular transverse rows of setae, concentrated on bottom on sternites 2–4. Surface with elongate punctures that coalesce at sides. Male last sternite strigate, with apical margin well defined and rounded.</paragraph>
<paragraph id="8BC43697FFF2FFD75AF66790FA92F9AA" blockId="5.[151,1437,1289,2033]" pageId="5" pageNumber="260">
Protibia with 2 teeth (
<figureCitation id="13402A12FFF2FFD75B8F6790FDACFA62" box="[446,516,1469,1493]" captionStart="FIGURES 7 – 10" captionStartId="5.[151,269,1156,1180]" captionTargetBox="[174,1395,662,1128]" captionTargetId="figure@5.[151,1436,658,1144]" captionTargetPageId="5" captionText="FIGURES 7 – 10. Shape of protibia in 7: Anomala eucoma (paralectotype, Las Mercedes, Guatemala); 8: A. flavacoma (3 holotype, Ƥ Hitoy Cerere, Costa Rica); 9: A. megaparamera (3 holotype, Ƥ paratype, Cuatro Esquinas, Costa Rica); 10: A. pseudoeucoma (3 holotype, Ƥ paratype, Reserva de San Ramón, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223580/files/figure.png" pageId="5" pageNumber="260">Fig. 7</figureCitation>
), apical tooth long and curved; second tooth at the same level of internal apex of protibia, triangular in shape, acute. Metatibia stout, slightly narrower subapically. Ratio length/width 2.93. First carina somewhat obliterated by elongate and dense punctures, which become rugose below second carina.
</paragraph>
<paragraph id="8BC43697FFF2FFD75AF66404FB3DF9D2" blockId="5.[151,1437,1289,2033]" pageId="5" pageNumber="260">Protarsal claws: external claw strongly curved; ratio length/height: 2.01; internal claw bifurcate, with upper branch slightly shorter and half the width of the lower one. Inferior margin slightly sinuate.</paragraph>
<paragraph id="8BC43697FFF2FFD75AF6645CFE53F942" blockId="5.[151,1437,1289,2033]" pageId="5" pageNumber="260">
Aedeagus (
<figureCitation id="13402A12FFF2FFD75B75645CFE6FF93E" box="[324,455,1649,1673]" captionStart="FIGURES 11 – 25. 11 – 16" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[286,1306,190,1615]" captionTargetId="figure@6.[276,1317,176,1615]" captionTargetPageId="6" captionText="FIGURES 11 – 25. 11 – 16. Aedeagus of A. eucoma (paralectotype, Las Mercedes, Guatemala). 11: Dorsal aspect. 12: Ventral aspect. 13: Lateral aspect; a: dorsal section: part of the dorsal margin that corresponds to the joint between the two parameres; b: dorsal margin: margin of the paramere dorsal with respect to apex; conversely, the ventral part is the ventral margin, which in these species can be divided in c: posterior ventral margin; and d: anterior ventral margin, in relation to the angle formed by the curvature of the margin. Parameres of Costa Rican specimens showing slight variation among populations. 14: San José, Universidad de Costa Rica. 15: Estación Las Pailas, Guanacaste. 16: Estación Sirena, Parque Nacional Corcovado. 17 – 19. Aedeagus of Anomala flavacoma (paratype, Estación Cabro Muco, Costa Rica). 17: Dorsal aspect. 18: Ventral aspect. 19: Lateral aspect. 20 – 22. Aedeagus of Anomala megaparamera (holotype). 20: Dorsal aspect. 21: Ventral aspect. 22: Lateral aspect. 23 – 25. Aedeagus of Anomala pseudoeucoma (paratype, Hitoy Cerere, Costa Rica). 23: Dorsal aspect. 24: Ventral aspect. 25: Lateral aspect. Scale = 1 mm." httpUri="https://zenodo.org/record/223581/files/figure.png" pageId="5" pageNumber="260">Figs 11–13</figureCitation>
): parameres short and wide, with inferior margin slightly sinuate. Endophallus (
<figureCitation id="13402A12FFF2FFD75F72645CFA3BF93E" box="[1347,1427,1649,1673]" captionStart="FIGURES 26 – 32. 26 – 29" captionStartId="7.[151,269,1798,1822]" captionTargetBox="[279,1291,205,1753]" captionTargetId="figure@7.[254,1296,194,1767]" captionTargetPageId="7" captionText="FIGURES 26 – 32. 26 – 29. Endophallus of Anomala eucoma showing variation among populations. 26: Paralectotype, Las Mercedes, Guatemala. 27: San José, Universidad de Costa Rica, Costa Rica. 28: Estación Las Pailas, Guanacaste, Costa Rica. 29: Estación Sirena, Parque Nacional Corcovado, Costa Rica. 30. Endophallus of Anomala flavacoma (holotype). 31. Endophallus of Anomala megaparamera (holotype). 32. Endophallus of Anomala pseudoeucoma (holotype). Scale = 1 mm. The arrows show the position of the gonopore." httpUri="https://zenodo.org/record/223582/files/figure.png" pageId="5" pageNumber="260">Fig. 26</figureCitation>
) with one long sacculus with sclerotized setae in a wide patch on apex; a ring of short but thick setae is present subapically. The rest of surface is covered by microsetation. A smaller diverticle is present at base. Gonopore opening ventrally at basal third.
</paragraph>
<paragraph id="8BC43697FFF2FFD75AF6652CFACDF88A" blockId="5.[151,1437,1289,2033]" pageId="5" pageNumber="260">
Female (
<collectingCountry id="F36C7607FFF2FFD75B19652CFE02F8AE" box="[296,426,1793,1817]" name="Costa Rica" pageId="5" pageNumber="260">Costa Rica</collectingCountry>
): Similar to males. Apical tooth of protibia longer and wider than in male. Inferior margin of internal protarsal claw straight. Metatibia wider than in males. Last abdominal sternite with curved margin.
</paragraph>
<paragraph id="8BC43697FFF2FFD75AF66565FE07F846" blockId="5.[151,1437,1289,2033]" pageId="5" pageNumber="260">
<emphasis id="B90FEA85FFF2FFD75AF66565FEEBF8D6" bold="true" box="[199,323,1864,1889]" pageId="5" pageNumber="260">Variation:</emphasis>
Body length
<quantity id="4C839B72FFF2FFD75BED6564FD27F8D6" box="[476,655,1865,1889]" metricMagnitude="-2" metricUnit="m" metricValue="1.053" metricValueMax="1.262" metricValueMin="0.844" pageId="5" pageNumber="260" unit="mm" value="10.53" valueMax="12.62" valueMin="8.44">8.44–12.62 mm</quantity>
, body width
<quantity id="4C839B72FFF2FFD759106564FC6FF8D6" box="[801,967,1865,1889]" metricMagnitude="-3" metricUnit="m" metricValue="5.9" metricValueMax="6.93" metricValueMin="4.87" pageId="5" pageNumber="260" unit="mm" value="5.9" valueMax="6.93" valueMin="4.87">4.87–6.93 mm</quantity>
. Clypeus w/l: 1.75–2.05. Interocular ratio (interocular width/width of eye): 2.80–3.73. Antenna: ratio funiculus/club 0.66–0.75. Pronotum w/l: 1.54–1.67. Scutellum w/l: 1.21–1.48. Pygidium w/l: 1.53–1.78. Width between mesocoxae:
<quantity id="4C839B72FFF2FFD75E7365BCFB44F81E" box="[1090,1260,1937,1961]" metricMagnitude="-4" metricUnit="m" metricValue="2.1" metricValueMax="2.7" metricValueMin="1.5" pageId="5" pageNumber="260" unit="mm" value="0.21" valueMax="0.27" valueMin="0.15">0.15–0.27 mm</quantity>
. Metatibia w/l: 2.35–3.08. External claw l/h: 1.32–2.01. Colour on head, pronotum, scutellum, and pygidium from greenish to reddish to bronze brown.
</paragraph>
<caption id="DF04661FFFF1FFD45AA6647DFBC6F830" httpUri="https://zenodo.org/record/223581/files/figure.png" pageId="6" pageNumber="261" targetBox="[286,1306,190,1615]" targetPageId="6">
<paragraph id="8BC43697FFF1FFD45AA6647DFBC6F830" blockId="6.[151,1437,1616,1927]" pageId="6" pageNumber="261">
<emphasis id="B90FEA85FFF1FFD45AA6647DFE02F9DF" bold="true" box="[151,426,1616,1640]" pageId="6" pageNumber="261">FIGURES 11–25. 11–16</emphasis>
. Aedeagus of
<taxonomicName id="4C7B4D14FFF1FFD4587F647DFD6AF9D0" box="[590,706,1616,1639]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF1FFD4587F647DFD6AF9D0" box="[590,706,1616,1639]" italics="true" pageId="6" pageNumber="261">A. eucoma</emphasis>
</taxonomicName>
(paralectotype, Las Mercedes, Guatemala). 11: Dorsal aspect. 12: Ventral aspect. 13: Lateral aspect; a: dorsal section: part of the dorsal margin that corresponds to the joint between the two parameres; b: dorsal margin: margin of the paramere dorsal with respect to apex; conversely, the ventral part is the ventral margin, which in these species can be divided in c: posterior ventral margin; and d: anterior ventral margin, in relation to the angle formed by the curvature of the margin. Parameres of Costa Rican specimens showing slight variation among populations. 14: San José, Universidad de Costa Rica. 15: Estación Las Pailas, Guanacaste. 16: Estación Sirena, Parque Nacional Corcovado.
<emphasis id="B90FEA85FFF1FFD4581A653DFDC5F89F" bold="true" box="[555,621,1808,1832]" pageId="6" pageNumber="261">17–19</emphasis>
. Aedeagus of
<taxonomicName id="4C7B4D14FFF1FFD4593F653DFC4EF890" box="[782,998,1808,1831]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="flavacoma">
<emphasis id="B90FEA85FFF1FFD4593F653DFC4EF890" box="[782,998,1808,1831]" italics="true" pageId="6" pageNumber="261">Anomala flavacoma</emphasis>
</taxonomicName>
(paratype, Estación Cabro Muco, Costa Rica). 17: Dorsal aspect. 18: Ventral aspect. 19: Lateral aspect.
<emphasis id="B90FEA85FFF1FFD4590D651DFCD6F8FF" bold="true" box="[828,894,1840,1864]" pageId="6" pageNumber="261">20–22</emphasis>
. Aedeagus of
<taxonomicName id="4C7B4D14FFF1FFD45E26651DFA88F8F0" box="[1047,1312,1840,1863]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="megaparamera">
<emphasis id="B90FEA85FFF1FFD45E26651DFA88F8F0" box="[1047,1312,1840,1863]" italics="true" pageId="6" pageNumber="261">Anomala megaparamera</emphasis>
</taxonomicName>
(holotype). 20: Dorsal aspect. 21: Ventral aspect. 22: Lateral aspect.
<emphasis id="B90FEA85FFF1FFD458CF657DFCE8F8DF" bold="true" box="[766,832,1872,1896]" pageId="6" pageNumber="261">23–25</emphasis>
. Aedeagus of
<taxonomicName id="4C7B4D14FFF1FFD459EF657DFB4CF8D0" box="[990,1252,1872,1895]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="6" pageNumber="261" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFF1FFD459EF657DFB4CF8D0" box="[990,1252,1872,1895]" italics="true" pageId="6" pageNumber="261">Anomala pseudoeucoma</emphasis>
</taxonomicName>
(paratype, Hitoy Cerere, Costa Rica). 23: Dorsal aspect. 24: Ventral aspect. 25: Lateral aspect. Scale = 1 mm.
</paragraph>
</caption>
<caption id="DF04661FFFF0FFD55AA6652BFC0EF82A" httpUri="https://zenodo.org/record/223582/files/figure.png" pageId="7" pageNumber="262" targetBox="[279,1291,205,1753]" targetPageId="7">
<paragraph id="8BC43697FFF0FFD55AA6652BFC0EF82A" blockId="7.[151,1436,1798,1949]" pageId="7" pageNumber="262">
<emphasis id="B90FEA85FFF0FFD55AA6652BFE03F8A9" bold="true" box="[151,427,1798,1822]" pageId="7" pageNumber="262">FIGURES 26–32. 26–29</emphasis>
. Endophallus of
<taxonomicName id="4C7B4D14FFF0FFD5585B652BFC8EF8AA" box="[618,806,1798,1821]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFF0FFD5585B652BFC8EF8AA" box="[618,806,1798,1821]" italics="true" pageId="7" pageNumber="262">Anomala eucoma</emphasis>
</taxonomicName>
showing variation among populations. 26: Paralectotype, Las Mercedes, Guatemala. 27: San José, Universidad de Costa Rica, Costa Rica. 28: Estación Las Pailas, Guanacaste, Costa Rica. 29: Estación Sirena, Parque Nacional Corcovado, Costa Rica.
<emphasis id="B90FEA85FFF0FFD559D3656BFC54F8E9" bold="true" box="[994,1020,1862,1886]" pageId="7" pageNumber="262">30</emphasis>
. Endophallus of
<taxonomicName id="4C7B4D14FFF0FFD55EF0656BFA34F8EA" box="[1217,1436,1862,1885]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="flavacoma">
<emphasis id="B90FEA85FFF0FFD55EF0656BFA34F8EA" box="[1217,1436,1862,1885]" italics="true" pageId="7" pageNumber="262">Anomala flavacoma</emphasis>
</taxonomicName>
(holotype).
<emphasis id="B90FEA85FFF0FFD55B2A654BFE9EF8C9" bold="true" box="[283,310,1894,1918]" pageId="7" pageNumber="262">31</emphasis>
. Endophallus of
<taxonomicName id="4C7B4D14FFF0FFD55832654BFCBDF8CA" box="[515,789,1894,1917]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="megaparamera">
<emphasis id="B90FEA85FFF0FFD55832654BFCBDF8CA" box="[515,789,1894,1917]" italics="true" pageId="7" pageNumber="262">Anomala megaparamera</emphasis>
</taxonomicName>
(holotype).
<emphasis id="B90FEA85FFF0FFD55996654BFC6AF8C9" bold="true" box="[935,962,1894,1918]" pageId="7" pageNumber="262">32</emphasis>
. Endophallus of
<taxonomicName id="4C7B4D14FFF0FFD55EBE654BFA34F8CA" box="[1167,1436,1894,1917]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="7" pageNumber="262" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFF0FFD55EBE654BFA34F8CA" box="[1167,1436,1894,1917]" italics="true" pageId="7" pageNumber="262">Anomala pseudoeucoma</emphasis>
</taxonomicName>
(holotype). Scale = 1 mm. The arrows show the position of the gonopore.
</paragraph>
</caption>
<paragraph id="8BC43697FFFFFFDA5AF662BAFB25FE78" blockId="8.[151,1436,151,788]" pageId="8" pageNumber="263">
Three different populations are present in
<collectingCountry id="F36C7607FFFFFFDA58FD62BAFCF9FF18" box="[716,849,151,175]" name="Costa Rica" pageId="8" pageNumber="263">Costa Rica</collectingCountry>
, with defined geographic ranges (
<figureCitation id="13402A12FFFFFFDA5EC362BAFAE5FF18" box="[1266,1357,151,175]" captionStart="FIGURE 33" captionStartId="8.[151,255,1857,1881]" captionTargetBox="[163,1424,877,1810]" captionTargetId="figure@8.[151,1436,842,1835]" captionTargetPageId="8" captionText="FIGURE 33. Distribution of Anomala eucoma in Costa Rica, with populations differentiated." httpUri="https://zenodo.org/record/223583/files/figure.png" pageId="8" pageNumber="263">Fig. 33</figureCitation>
), each presenting subtle differences with the others and the specimens from Las Mercedes,
<collectingCountry id="F36C7607FFFFFFDA5E986291FA8EFF63" box="[1193,1318,188,212]" name="Guatemala" pageId="8" pageNumber="263">Guatemala</collectingCountry>
. The two
<typeStatus id="54C08835FFFFFFDA5AA662F2FE91FF40" box="[151,313,223,247]" pageId="8" pageNumber="263" type="paralectotype">paralectotypes</typeStatus>
from
<collectingCountry id="F36C7607FFFFFFDA5BB262F2FDA8FF40" box="[387,512,223,247]" name="Costa Rica" pageId="8" pageNumber="263">Costa Rica</collectingCountry>
and the specimens from
<collectingCountry id="F36C7607FFFFFFDA591962F2FC35FF40" box="[808,925,223,247]" name="Saint Pierre and Miquelon" pageId="8" pageNumber="263">San Pedro</collectingCountry>
(Universidad de
<collectingCountry id="F36C7607FFFFFFDA5E5962F2FB40FF40" box="[1128,1256,223,247]" name="Costa Rica" pageId="8" pageNumber="263">Costa Rica</collectingCountry>
, near San José, Central Valley) and Amubri (the Talamanca mountain range) are similar in size (body length
<quantity id="4C839B72FFFFFFDA5EE56329FA30FEAB" box="[1236,1432,260,284]" metricMagnitude="-2" metricUnit="m" metricValue="1.2035" metricValueMax="1.262" metricValueMin="1.145" pageId="8" pageNumber="263" unit="mm" value="12.035" valueMax="12.62" valueMin="11.45">11.45–12.62 mm</quantity>
, width
<quantity id="4C839B72FFFFFFDA5AEC630AFE2BFE88" box="[221,387,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="6.435" metricValueMax="6.93" metricValueMin="5.94" pageId="8" pageNumber="263" unit="mm" value="6.435" valueMax="6.93" valueMin="5.94">5.94–6.93 mm</quantity>
; Guatemalan specimens: body length
<quantity id="4C839B72FFFFFFDA591C630AFC46FE88" box="[813,1006,295,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.2135" metricValueMax="1.253" metricValueMin="1.174" pageId="8" pageNumber="263" unit="mm" value="12.135" valueMax="12.53" valueMin="11.74">11.74–12.53 mm</quantity>
, width
<quantity id="4C839B72FFFFFFDA5E71630AFB4DFE88" box="[1088,1253,295,319]" metricMagnitude="-3" metricUnit="m" metricValue="6.805" metricValueMax="6.9" metricValueMin="6.71" pageId="8" pageNumber="263" unit="mm" value="6.805" valueMax="6.9" valueMin="6.71">6.71–6.90 mm</quantity>
) but are paler in colour; the frons gradually curving towards vertex (
<figureCitation id="13402A12FFFFFFDA58EC6361FC89FED3" box="[733,801,332,356]" captionStart="FIGURE 6" captionStartId="5.[151,255,561,585]" captionTargetBox="[362,1227,193,533]" captionTargetId="figure@5.[340,1248,176,556]" captionTargetPageId="5" captionText="FIGURE 6. Head of a: Anomala eucoma (paralectotype, Las Mercedes, Guatemala) and b: A. eucoma (paralectotype, precise locality unknown, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223579/files/figure.png" pageId="8" pageNumber="263">Fig. 6</figureCitation>
b), clypeus trapezoidal with lateral sides straight. This is in contrast to the Las Mercedes specimens, where they are curved, with basal half perpendicular to the base (see
<figureCitation id="13402A12FFFFFFDA5AA663B9FF74FE1B" box="[151,220,404,428]" captionStart="FIGURE 6" captionStartId="5.[151,255,561,585]" captionTargetBox="[362,1227,193,533]" captionTargetId="figure@5.[340,1248,176,556]" captionTargetPageId="5" captionText="FIGURE 6. Head of a: Anomala eucoma (paralectotype, Las Mercedes, Guatemala) and b: A. eucoma (paralectotype, precise locality unknown, Costa Rica). Scale = 1 mm." httpUri="https://zenodo.org/record/223579/files/figure.png" pageId="8" pageNumber="263">Fig. 6</figureCitation>
). Other differences are: less curved external protarsal claw, metatibia punctate above first carina, which is well defined, slender parameres (
<figureCitation id="13402A12FFFFFFDA583B639AFDCEFE78" box="[522,614,439,463]" captionStart="FIGURES 11 – 25. 11 – 16" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[286,1306,190,1615]" captionTargetId="figure@6.[276,1317,176,1615]" captionTargetPageId="6" captionText="FIGURES 11 – 25. 11 – 16. Aedeagus of A. eucoma (paralectotype, Las Mercedes, Guatemala). 11: Dorsal aspect. 12: Ventral aspect. 13: Lateral aspect; a: dorsal section: part of the dorsal margin that corresponds to the joint between the two parameres; b: dorsal margin: margin of the paramere dorsal with respect to apex; conversely, the ventral part is the ventral margin, which in these species can be divided in c: posterior ventral margin; and d: anterior ventral margin, in relation to the angle formed by the curvature of the margin. Parameres of Costa Rican specimens showing slight variation among populations. 14: San José, Universidad de Costa Rica. 15: Estación Las Pailas, Guanacaste. 16: Estación Sirena, Parque Nacional Corcovado. 17 – 19. Aedeagus of Anomala flavacoma (paratype, Estación Cabro Muco, Costa Rica). 17: Dorsal aspect. 18: Ventral aspect. 19: Lateral aspect. 20 – 22. Aedeagus of Anomala megaparamera (holotype). 20: Dorsal aspect. 21: Ventral aspect. 22: Lateral aspect. 23 – 25. Aedeagus of Anomala pseudoeucoma (paratype, Hitoy Cerere, Costa Rica). 23: Dorsal aspect. 24: Ventral aspect. 25: Lateral aspect. Scale = 1 mm." httpUri="https://zenodo.org/record/223581/files/figure.png" pageId="8" pageNumber="263">Figs. 14</figureCitation>
) and shorter and narrower endophallus (
<figureCitation id="13402A12FFFFFFDA5E1D639AFB28FE78" box="[1068,1152,439,463]" captionStart="FIGURES 26 – 32. 26 – 29" captionStartId="7.[151,269,1798,1822]" captionTargetBox="[279,1291,205,1753]" captionTargetId="figure@7.[254,1296,194,1767]" captionTargetPageId="7" captionText="FIGURES 26 – 32. 26 – 29. Endophallus of Anomala eucoma showing variation among populations. 26: Paralectotype, Las Mercedes, Guatemala. 27: San José, Universidad de Costa Rica, Costa Rica. 28: Estación Las Pailas, Guanacaste, Costa Rica. 29: Estación Sirena, Parque Nacional Corcovado, Costa Rica. 30. Endophallus of Anomala flavacoma (holotype). 31. Endophallus of Anomala megaparamera (holotype). 32. Endophallus of Anomala pseudoeucoma (holotype). Scale = 1 mm. The arrows show the position of the gonopore." httpUri="https://zenodo.org/record/223582/files/figure.png" pageId="8" pageNumber="263">Fig. 27</figureCitation>
).
</paragraph>
<paragraph id="8BC43697FFFFFFDA5AF663F1FED0FD33" blockId="8.[151,1436,151,788]" pageId="8" pageNumber="263">
The specimens from the Guanacaste province are smaller in size (body length
<quantity id="4C839B72FFFFFFDA5E0463F1FB71FE43" box="[1077,1241,476,500]" metricMagnitude="-3" metricUnit="m" metricValue="9.635" metricValueMax="9.96" metricValueMin="9.31" pageId="8" pageNumber="263" unit="mm" value="9.635" valueMax="9.96" valueMin="9.31">9.31–9.96 mm</quantity>
, width
<quantity id="4C839B72FFFFFFDA5F1D63F1FF6FFDA0" metricMagnitude="-3" metricUnit="m" metricValue="5.49" metricValueMax="5.93" metricValueMin="5.05" pageId="8" pageNumber="263" unit="mm" value="5.49" valueMax="5.93" valueMin="5.05">5.05–5.93 mm</quantity>
), with frons gradually curving towards vertex, metatibia punctate above first carina, which is well defined, narrower apical angle of parameres (
<figureCitation id="13402A12FFFFFFDA580B6009FD27FD8B" box="[570,655,548,572]" captionStart="FIGURES 11 – 25. 11 – 16" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[286,1306,190,1615]" captionTargetId="figure@6.[276,1317,176,1615]" captionTargetPageId="6" captionText="FIGURES 11 – 25. 11 – 16. Aedeagus of A. eucoma (paralectotype, Las Mercedes, Guatemala). 11: Dorsal aspect. 12: Ventral aspect. 13: Lateral aspect; a: dorsal section: part of the dorsal margin that corresponds to the joint between the two parameres; b: dorsal margin: margin of the paramere dorsal with respect to apex; conversely, the ventral part is the ventral margin, which in these species can be divided in c: posterior ventral margin; and d: anterior ventral margin, in relation to the angle formed by the curvature of the margin. Parameres of Costa Rican specimens showing slight variation among populations. 14: San José, Universidad de Costa Rica. 15: Estación Las Pailas, Guanacaste. 16: Estación Sirena, Parque Nacional Corcovado. 17 – 19. Aedeagus of Anomala flavacoma (paratype, Estación Cabro Muco, Costa Rica). 17: Dorsal aspect. 18: Ventral aspect. 19: Lateral aspect. 20 – 22. Aedeagus of Anomala megaparamera (holotype). 20: Dorsal aspect. 21: Ventral aspect. 22: Lateral aspect. 23 – 25. Aedeagus of Anomala pseudoeucoma (paratype, Hitoy Cerere, Costa Rica). 23: Dorsal aspect. 24: Ventral aspect. 25: Lateral aspect. Scale = 1 mm." httpUri="https://zenodo.org/record/223581/files/figure.png" pageId="8" pageNumber="263">Fig. 15</figureCitation>
), thinner setation on endophallus (
<figureCitation id="13402A12FFFFFFDA5E2A6009FBD9FD8B" box="[1051,1137,548,572]" captionStart="FIGURES 26 – 32. 26 – 29" captionStartId="7.[151,269,1798,1822]" captionTargetBox="[279,1291,205,1753]" captionTargetId="figure@7.[254,1296,194,1767]" captionTargetPageId="7" captionText="FIGURES 26 – 32. 26 – 29. Endophallus of Anomala eucoma showing variation among populations. 26: Paralectotype, Las Mercedes, Guatemala. 27: San José, Universidad de Costa Rica, Costa Rica. 28: Estación Las Pailas, Guanacaste, Costa Rica. 29: Estación Sirena, Parque Nacional Corcovado, Costa Rica. 30. Endophallus of Anomala flavacoma (holotype). 31. Endophallus of Anomala megaparamera (holotype). 32. Endophallus of Anomala pseudoeucoma (holotype). Scale = 1 mm. The arrows show the position of the gonopore." httpUri="https://zenodo.org/record/223582/files/figure.png" pageId="8" pageNumber="263">Fig. 28</figureCitation>
). The
<typeStatus id="54C08835FFFFFFDA5E8A6009FAF5FD8B" box="[1211,1373,548,572]" pageId="8" pageNumber="263" type="paralectotype">paralectotypes</typeStatus>
from Pantaleon and San Isidro,
<collectingCountry id="F36C7607FFFFFFDA5BF4606AFDE9FDE8" box="[453,577,583,607]" name="Guatemala" pageId="8" pageNumber="263">Guatemala</collectingCountry>
, are more similar to these specimens in size and genitalia than to the ones from Las Mercedes.
</paragraph>
<paragraph id="8BC43697FFFFFFDA5AF660A2FE97FCA3" blockId="8.[151,1436,151,788]" pageId="8" pageNumber="263">
The specimens from the Osa peninsula (P.N. Corcovado) are also smaller in size (body length
<quantity id="4C839B72FFFFFFDA5EC360A2FA3FFD10" box="[1266,1431,655,679]" metricMagnitude="-3" metricUnit="m" metricValue="9.17" metricValueMax="9.9" metricValueMin="8.44" pageId="8" pageNumber="263" unit="mm" value="9.17" valueMax="9.9" valueMin="8.44">8.44–9.90 mm</quantity>
, width
<quantity id="4C839B72FFFFFFDA5AEF6099FE2DFD7B" box="[222,389,692,716]" metricMagnitude="-3" metricUnit="m" metricValue="5.25" metricValueMax="5.63" metricValueMin="4.87" pageId="8" pageNumber="263" unit="mm" value="5.25" valueMax="5.63" valueMin="4.87">4.87–5.63 mm</quantity>
), with frons gradually curving towards vertex, metatibia punctate above first carina, which is well defined, slender parameres with sinuate ventral margin (
<figureCitation id="13402A12FFFFFFDA597E60FAFC0AFD58" box="[847,930,727,751]" captionStart="FIGURES 11 – 25. 11 – 16" captionStartId="6.[151,269,1616,1640]" captionTargetBox="[286,1306,190,1615]" captionTargetId="figure@6.[276,1317,176,1615]" captionTargetPageId="6" captionText="FIGURES 11 – 25. 11 – 16. Aedeagus of A. eucoma (paralectotype, Las Mercedes, Guatemala). 11: Dorsal aspect. 12: Ventral aspect. 13: Lateral aspect; a: dorsal section: part of the dorsal margin that corresponds to the joint between the two parameres; b: dorsal margin: margin of the paramere dorsal with respect to apex; conversely, the ventral part is the ventral margin, which in these species can be divided in c: posterior ventral margin; and d: anterior ventral margin, in relation to the angle formed by the curvature of the margin. Parameres of Costa Rican specimens showing slight variation among populations. 14: San José, Universidad de Costa Rica. 15: Estación Las Pailas, Guanacaste. 16: Estación Sirena, Parque Nacional Corcovado. 17 – 19. Aedeagus of Anomala flavacoma (paratype, Estación Cabro Muco, Costa Rica). 17: Dorsal aspect. 18: Ventral aspect. 19: Lateral aspect. 20 – 22. Aedeagus of Anomala megaparamera (holotype). 20: Dorsal aspect. 21: Ventral aspect. 22: Lateral aspect. 23 – 25. Aedeagus of Anomala pseudoeucoma (paratype, Hitoy Cerere, Costa Rica). 23: Dorsal aspect. 24: Ventral aspect. 25: Lateral aspect. Scale = 1 mm." httpUri="https://zenodo.org/record/223581/files/figure.png" pageId="8" pageNumber="263">Fig. 16</figureCitation>
), endophallus without patches of sclerotized setae (
<figureCitation id="13402A12FFFFFFDA5AEF60D1FE9AFCA3" box="[222,306,764,788]" captionStart="FIGURES 26 – 32. 26 – 29" captionStartId="7.[151,269,1798,1822]" captionTargetBox="[279,1291,205,1753]" captionTargetId="figure@7.[254,1296,194,1767]" captionTargetPageId="7" captionText="FIGURES 26 – 32. 26 – 29. Endophallus of Anomala eucoma showing variation among populations. 26: Paralectotype, Las Mercedes, Guatemala. 27: San José, Universidad de Costa Rica, Costa Rica. 28: Estación Las Pailas, Guanacaste, Costa Rica. 29: Estación Sirena, Parque Nacional Corcovado, Costa Rica. 30. Endophallus of Anomala flavacoma (holotype). 31. Endophallus of Anomala megaparamera (holotype). 32. Endophallus of Anomala pseudoeucoma (holotype). Scale = 1 mm. The arrows show the position of the gonopore." httpUri="https://zenodo.org/record/223582/files/figure.png" pageId="8" pageNumber="263">Fig. 29</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF04661FFFFFFFDA5AA6656CFBD7F8EF" box="[151,1151,1857,1881]" httpUri="https://zenodo.org/record/223583/files/figure.png" pageId="8" pageNumber="263" targetBox="[163,1424,877,1810]" targetPageId="8">
<paragraph id="8BC43697FFFFFFDA5AA6656CFBD7F8EF" blockId="8.[151,1151,1857,1881]" box="[151,1151,1857,1881]" pageId="8" pageNumber="263">
<emphasis id="B90FEA85FFFFFFDA5AA6656CFE88F8EE" bold="true" box="[151,288,1857,1881]" pageId="8" pageNumber="263">FIGURE 33</emphasis>
. Distribution of
<taxonomicName id="4C7B4D14FFFFFFDA5BE0656CFD23F8EF" box="[465,651,1857,1880]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFFFFFDA5BE0656CFD23F8EF" box="[465,651,1857,1880]" italics="true" pageId="8" pageNumber="263">Anomala eucoma</emphasis>
</taxonomicName>
in Costa Rica, with populations differentiated.
</paragraph>
</caption>
<subSubSection id="C361651CFFFFFFDB5AF665A8FF7EFD8B" lastPageId="9" lastPageNumber="264" pageId="8" pageNumber="263" type="diagnosis">
<paragraph id="8BC43697FFFFFFDB5AF665A8FF7EFD8B" blockId="8.[151,1437,1925,2021]" lastBlockId="9.[151,1436,151,644]" lastPageId="9" lastPageNumber="264" pageId="8" pageNumber="263">
<emphasis id="B90FEA85FFFFFFDA5AF665A8FEE9F829" bold="true" box="[199,321,1925,1950]" pageId="8" pageNumber="263">Diagnosis.</emphasis>
<taxonomicName id="4C7B4D14FFFFFFDA5B7965A8FDA3F82A" box="[328,523,1925,1949]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFFFFFDA5B7965A8FDA3F82A" box="[328,523,1925,1949]" italics="true" pageId="8" pageNumber="263">Anomala eucoma</emphasis>
</taxonomicName>
is separated from similar species by the combination of the following characters: two dark bands on elytra, parameres pentagonal in shape, with dorsal and ventral margin not strongly sinuate, endophallus with one long sacculus. It can easily be separated from
<taxonomicName id="4C7B4D14FFFFFFDA59F465E0FBC4F852" box="[965,1132,1997,2021]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="8" pageNumber="263" phylum="Arthropoda" rank="species" species="amphicoma">
<emphasis id="B90FEA85FFFFFFDA59F465E0FBC4F852" box="[965,1132,1997,2021]" italics="true" pageId="8" pageNumber="263">A. amphicoma</emphasis>
</taxonomicName>
, about the same size and dorsally setose, by the latter having the following characteristics: a metallic green pronotum in most specimens, covered by velvety dense setae; the pronotum is largest at 1/3 of height, with base narrower than elytra; elytra uniformly dark in colour, shorter parameres with sinuate ventral margin, endophallus with two long sacculi. It is differentiated from
<taxonomicName id="4C7B4D14FFFEFFDB5B466329FDA5FEAB" box="[375,525,260,284]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="flavacoma">
<emphasis id="B90FEA85FFFEFFDB5B466329FDA5FEAB" box="[375,525,260,284]" italics="true" pageId="9" pageNumber="264">A. flavacoma</emphasis>
</taxonomicName>
by size (
<taxonomicName id="4C7B4D14FFFEFFDB584D6329FCBAFEAB" box="[636,786,260,284]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="flavacoma">
<emphasis id="B90FEA85FFFEFFDB584D6329FCBAFEAB" box="[636,786,260,284]" italics="true" pageId="9" pageNumber="264">A. flavacoma</emphasis>
</taxonomicName>
has an intermediate size between the larger and smaller specimens of
<taxonomicName id="4C7B4D14FFFEFFDB5B046305FE05FEF7" box="[309,429,296,320]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="eucoma">
<emphasis id="B90FEA85FFFEFFDB5B046305FE05FEF7" box="[309,429,296,320]" italics="true" pageId="9" pageNumber="264">A. eucoma</emphasis>
</taxonomicName>
) and its darker colour, proportionally narrower scutellum, endophallus with one narrow and long sacculus and lack of thick setae, in contrast to the two short sacculi of
<taxonomicName id="4C7B4D14FFFEFFDB5E4E6361FAB1FED3" box="[1151,1305,332,356]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="flavacoma">
<emphasis id="B90FEA85FFFEFFDB5E4E6361FAB1FED3" box="[1151,1305,332,356]" italics="true" pageId="9" pageNumber="264">A. flavacoma</emphasis>
</taxonomicName>
. It can be distinguished from
<taxonomicName id="4C7B4D14FFFEFFDB5B44635DFD95FE3F" box="[373,573,368,392]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFFEFFDB5B44635DFD95FE3F" box="[373,573,368,392]" italics="true" pageId="9" pageNumber="264">A. pseudoeucoma</emphasis>
</taxonomicName>
by the longer and more curved apical tooth in male protibia; more convex intermesocoxal space, shorter parameres, endophallus with one long sacculus and lack of thick setae, instead of the two sacculi with patches of thick setae of
<taxonomicName id="4C7B4D14FFFEFFDB58416395FC9DFE67" box="[624,821,440,464]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="pseudoeucoma">
<emphasis id="B90FEA85FFFEFFDB58416395FC9DFE67" box="[624,821,440,464]" italics="true" pageId="9" pageNumber="264">A. pseudoeucoma</emphasis>
</taxonomicName>
.
<taxonomicName id="4C7B4D14FFFEFFDB59756395FBF4FE67" box="[836,1116,440,464]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="megaparamera">
<emphasis id="B90FEA85FFFEFFDB59756395FBF4FE67" box="[836,1116,440,464]" italics="true" pageId="9" pageNumber="264">Anomala megaparamera</emphasis>
</taxonomicName>
is similar in size and colour to the Osa and Guanacaste populations, but
<taxonomicName id="4C7B4D14FFFEFFDB58B263F1FCE6FE43" box="[643,846,476,500]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="264" phylum="Arthropoda" rank="species" species="megaparamera">
<emphasis id="B90FEA85FFFEFFDB58B263F1FCE6FE43" box="[643,846,476,500]" italics="true" pageId="9" pageNumber="264">A. megaparamera</emphasis>
</taxonomicName>
has a proportionally wider pygidium, aedeagus with proportionally larger parameres, endophallus with several inflations and diverticles and copiously covered with setae.
</paragraph>
</subSubSection>
<subSubSection id="C361651CFFFEFFDB5AF6606AFC26FD33" pageId="9" pageNumber="264" type="distribution">
<paragraph id="8BC43697FFFEFFDB5AF6606AFC26FD33" blockId="9.[151,1436,151,644]" pageId="9" pageNumber="264">
<emphasis id="B90FEA85FFFEFFDB5AF6606AFEC8FDD7" bold="true" box="[199,352,583,608]" pageId="9" pageNumber="264">Distribution.</emphasis>
<collectingCountry id="F36C7607FFFEFFDB5B56606AFE4DFDE8" box="[359,485,583,607]" name="Guatemala" pageId="9" pageNumber="264">Guatemala</collectingCountry>
: Las Mercedes;
<collectingCountry id="F36C7607FFFEFFDB58A8606AFCBFFDE8" box="[665,791,583,607]" name="Costa Rica" pageId="9" pageNumber="264">Costa Rica</collectingCountry>
: Guanacaste and the Talamanca mountain ranges; Central Valley; Parque Nacional Corcovado, in the Osa peninsula (
<figureCitation id="13402A12FFFEFFDB591C6041FC28FD33" box="[813,896,620,644]" captionStart="FIGURE 33" captionStartId="8.[151,255,1857,1881]" captionTargetBox="[163,1424,877,1810]" captionTargetId="figure@8.[151,1436,842,1835]" captionTargetPageId="8" captionText="FIGURE 33. Distribution of Anomala eucoma in Costa Rica, with populations differentiated." httpUri="https://zenodo.org/record/223583/files/figure.png" pageId="9" pageNumber="264">Fig. 33</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>
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