325 lines
35 KiB
XML
325 lines
35 KiB
XML
<document ID-DOI="10.11646/zootaxa.5244.6.5" ID-ISSN="1175-5326" ID-Zenodo-Dep="7673187" ID-ZooBank="3ED59A07-2F89-4C9B-B980-E22B5E96557F" approvalRequired="2" approvalRequired_for_taxonomicNames="1" approvalRequired_for_textStreams="1" checkinTime="1677226117531" checkinUser="plazi" docAuthor="Yin, Zi-Wei" docDate="2023" docId="03EB117AE16EFFD0FF53A0776AB7D2F1" docLanguage="en" docName="zootaxa.5244.6.5.pdf" docOrigin="Zootaxa 5244 (6)" docSource="http://dx.doi.org/10.11646/zootaxa.5244.6.5" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pakistatyrus schmidti Yin 2023, sp. nov." docType="treatment" docVersion="2" lastPageNumber="598" masterDocId="FFD26902E16FFFD3FFC4A2006862D634" masterDocTitle="A third species of Pakistatyrus Hlaváč (Coleoptera: Staphylinidae: Pselaphinae)" masterLastPageNumber="599" masterPageNumber="595" pageNumber="596" updateTime="1677226609395" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>A third species of Pakistatyrus Hlaváč (Coleoptera: Staphylinidae: Pselaphinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Yin, Zi-Wei</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2023</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2023-02-23</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>5244</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>6</mods:number>
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||
</mods:detail>
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||
<mods:extent unit="page">
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<mods:start>595</mods:start>
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<mods:end>599</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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||
<mods:location>
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<mods:url>http://dx.doi.org/10.11646/zootaxa.5244.6.5</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.11646/zootaxa.5244.6.5</mods:identifier>
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<mods:identifier type="ISSN">1175-5326</mods:identifier>
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<mods:identifier type="Zenodo-Dep">7673187</mods:identifier>
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<mods:identifier type="ZooBank">3ED59A07-2F89-4C9B-B980-E22B5E96557F</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.7673169" ID-Zenodo-Dep="7673169" LSID="urn:lsid:plazi:treatment:03EB117AE16EFFD0FF53A0776AB7D2F1" httpUri="http://treatment.plazi.org/id/03EB117AE16EFFD0FF53A0776AB7D2F1" lastPageId="3" lastPageNumber="598" pageId="1" pageNumber="596">
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<subSubSection box="[151,512,631,658]" pageId="1" pageNumber="596" type="nomenclature">
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<paragraph blockId="1.[151,512,631,693]" box="[151,512,631,658]" pageId="1" pageNumber="596">
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<heading bold="true" box="[151,512,631,658]" fontSize="11" level="1" pageId="1" pageNumber="596" reason="1">
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<emphasis bold="true" box="[151,512,631,658]" pageId="1" pageNumber="596">
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<taxonomicName authority="Yin, 2023" authorityName="Yin" authorityYear="2023" box="[151,411,631,658]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="schmidti" status="sp. nov.">
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<emphasis bold="true" box="[151,411,631,658]" italics="true" pageId="1" pageNumber="596">Pakistatyrus schmidti</emphasis>
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</taxonomicName>
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<taxonomicNameLabel box="[418,512,631,657]" pageId="1" pageNumber="596" rank="species">sp. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[151,268,667,693]" pageId="1" pageNumber="596" type="description">
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<paragraph blockId="1.[151,512,631,693]" box="[151,268,667,693]" pageId="1" pageNumber="596">
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(
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<figureCitation box="[159,230,667,693]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="1" pageNumber="596">Figs 1</figureCitation>
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,
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<figureCitation box="[244,259,667,693]" captionStart="FIGURE 2" captionStartId="3.[151,250,1990,2014]" captionTargetBox="[151,1436,1245,1967]" captionTargetId="figure-467@3.[151,1436,1245,1967]" captionTargetPageId="3" captionText="FIGURE 2. Distribution of Pakistatyrus species along the Himalaya, with the aedeagi compared." figureDoi="http://doi.org/10.5281/zenodo.7673191" httpUri="https://zenodo.org/record/7673191/files/figure.png" pageId="1" pageNumber="596">2</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection pageId="1" pageNumber="596" type="materials_examined">
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<paragraph blockId="1.[151,1437,739,2025]" pageId="1" pageNumber="596">
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<emphasis bold="true" box="[151,324,739,765]" pageId="1" pageNumber="596">Type material.</emphasis>
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<materialsCitation collectingDate="2005-05-11" collectingDateMax="2015-10-23" collectingDateMin="2005-05-11" collectorName="Manaslu Mts., SW & Schmidt" country="Nepal" county="Nebenfluss des Ngadi Khola" elevation="3250" latitude="28.36139" location="Manaslu Mts." longLatPrecision="20" longitude="84.511665" municipality="Quellgebiet" pageId="1" pageNumber="596" specimenCount="1" typeStatus="holotype">
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<emphasis bold="true" box="[329,605,739,765]" pageId="1" pageNumber="596">
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<typeStatus box="[329,492,739,765]" pageId="1" pageNumber="596">HOLOTYPE</typeStatus>
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:
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<collectingCountry box="[503,600,739,765]" name="Nepal" pageId="1" pageNumber="596">NEPAL</collectingCountry>
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:
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</emphasis>
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J, four labels: ‘NEPAL,
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<collectorName box="[885,1091,739,766]" pageId="1" pageNumber="596">
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<location LSID="urn:lsid:plazi:treatment:03EB117AE16EFFD0FF53A0776AB7D2F1:8E9DF6B7E16EFFD2FCB1A0E36C72D4CA" box="[885,1040,739,766]" country="Nepal" county="Nebenfluss des Ngadi Khola" latitude="28.36139" longLatPrecision="20" longitude="84.511665" municipality="Quellgebiet" name="Manaslu Mts." pageId="1" pageNumber="596">Manaslu Mts.</location>
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, SW
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</collectorName>
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Merne Pokhari,
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<quantity box="[1276,1431,739,766]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="1" pageNumber="596" unit="m" value="3250.0" valueMax="3300.0" valueMin="3200.0">
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<elevation box="[1276,1431,739,766]" metricMagnitude="3" metricUnit="m" metricValue="3.25" metricValueMax="3.3" metricValueMin="3.2" pageId="1" pageNumber="596" unit="m" value="3250.0" valueMax="3300.0" valueMin="3200.0">3200–3300 m</elevation>
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</quantity>
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, leg.
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<collectorName box="[199,295,775,801]" pageId="1" pageNumber="596">Schmidt</collectorName>
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,
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<date box="[305,414,775,802]" pageId="1" pageNumber="596" value="2005-05-11">
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<collectingDate box="[305,414,775,802]" pageId="1" pageNumber="596" value="2005-05-11">11.v.2005</collectingDate>
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</date>
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/
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<geoCoordinate box="[434,568,775,801]" degrees="28" direction="north" minutes="21" orientation="latitude" pageId="1" pageNumber="596" precision="15" seconds="41" value="28.36139">28°21'41"N</geoCoordinate>
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,
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<geoCoordinate box="[579,711,775,801]" degrees="84" direction="east" minutes="30" orientation="longitude" pageId="1" pageNumber="596" precision="15" seconds="42" value="84.511665">84°30'42"E</geoCoordinate>
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,
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<collectingMunicipality box="[721,855,775,801]" pageId="1" pageNumber="596">Quellgebiet</collectingMunicipality>
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, mittl.,
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<collectingCounty box="[938,1249,775,801]" pageId="1" pageNumber="596">Nebenfluss des Ngadi Khola</collectingCounty>
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/ Coll. SENM in MHNG, loan
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<date box="[306,418,811,838]" pageId="1" pageNumber="596" value="2015-10-23">
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<collectingDate box="[306,418,811,838]" pageId="1" pageNumber="596" value="2015-10-23">23.x.2015</collectingDate>
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</date>
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</materialsCitation>
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/
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<materialsCitation collectionCode="SENM" country="Nepal" determinerName="Z. - W. Yin" pageId="1" pageNumber="596" specimenCount="2" typeStatus="holotype">
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<typeStatus box="[440,588,811,837]" pageId="1" pageNumber="596">HOLOTYPE</typeStatus>
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(handwritten) J,
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<taxonomicName authority="Yin, 2023" authorityName="Yin" authorityYear="2023" box="[789,1031,811,837]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="596" phylum="Arthropoda" rank="species" species="schmidti" status="sp. n.">
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<emphasis box="[789,1031,811,837]" italics="true" pageId="1" pageNumber="596">Pakistatyrus schmidti</emphasis>
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</taxonomicName>
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<emphasis bold="true" box="[1038,1102,811,837]" pageId="1" pageNumber="596">
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<taxonomicNameLabel box="[1038,1102,811,837]" pageId="1" pageNumber="596" rank="species">sp. n.</taxonomicNameLabel>
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</emphasis>
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, det.
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<determinerName box="[1163,1274,811,838]" pageId="1" pageNumber="596">Z.-W. Yin</determinerName>
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2023, SENM’ (in
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<collectionCode box="[189,272,847,873]" pageId="1" pageNumber="596">SENM</collectionCode>
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). (Note: right foreleg of
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<typeStatus box="[545,643,847,873]" pageId="1" pageNumber="596">holotype</typeStatus>
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broken at base of femur, and missing)
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</materialsCitation>
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||
</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="1" pageNumber="596" type="diagnosis">
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<paragraph blockId="1.[151,1437,739,2025]" pageId="1" pageNumber="596">
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||
<emphasis bold="true" box="[199,322,883,909]" pageId="1" pageNumber="596">Diagnosis.</emphasis>
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<emphasis box="[326,384,883,909]" italics="true" pageId="1" pageNumber="596">Male</emphasis>
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. Body length approximately
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||
<quantity box="[704,790,883,910]" metricMagnitude="-3" metricUnit="m" metricValue="3.5" pageId="1" pageNumber="596" unit="mm" value="3.5">3.5 mm</quantity>
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, color reddish-brown. Foveae of pronotum and elytra large and setose. Maxillary palpomere 3 2.6 times as long as broad. Antenna approximately
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||
<quantity box="[1115,1200,919,946]" metricMagnitude="-3" metricUnit="m" metricValue="1.7" pageId="1" pageNumber="596" unit="mm" value="1.7">1.7 mm</quantity>
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||
long; antennomere 1 approximately 1.7 times as long as wide, 2–7 each slightly elongate, 8 smallest, as long as wide, 9–11 moderately enlarged to form indistinct club, lacking modifications. Elytra greatly transverse, length/width 0.46. Protrochanter with distinct spine on ventral margin; mesocoxa lacking tubercle; hind leg simple. Aedeagus with broad median lobe narrowing toward apex; parameres asymmetric, much shorter than median lobe, each paramere with four apical setae; lacking obvious endophallus armature.
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<emphasis box="[663,747,1099,1125]" italics="true" pageId="1" pageNumber="596">Female</emphasis>
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. Unknown.
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</paragraph>
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||
</subSubSection>
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<subSubSection lastPageId="3" lastPageNumber="598" pageId="1" pageNumber="596" type="description">
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<paragraph blockId="1.[151,1437,739,2025]" pageId="1" pageNumber="596">
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<emphasis bold="true" box="[199,345,1135,1161]" pageId="1" pageNumber="596">Description.</emphasis>
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<emphasis box="[354,412,1135,1161]" italics="true" pageId="1" pageNumber="596">Male</emphasis>
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. Body (
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<figureCitation box="[506,595,1135,1161]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="1" pageNumber="596">Fig. 1A</figureCitation>
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) length (from anterior margin of clypeus to apex of abdomen)
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||
<quantity box="[1327,1432,1135,1162]" metricMagnitude="-3" metricUnit="m" metricValue="3.49" pageId="1" pageNumber="596" unit="mm" value="3.49">3.49 mm</quantity>
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; color reddish-brown, tarsi and mouthparts lighter. Dorsal surface of body shining, covered with short pubescence.
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</paragraph>
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<paragraph blockId="1.[151,1437,739,2025]" pageId="1" pageNumber="596">
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Head (
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<figureCitation box="[276,363,1207,1233]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="1" pageNumber="596">Fig. 1B</figureCitation>
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) elongate-hexagonal, truncate at apex, slightly longer than wide, length from anterior margin of clypeus to posterior margin (excluding occipital constriction)
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<quantity box="[906,1012,1243,1270]" metricMagnitude="-4" metricUnit="m" metricValue="7.1" pageId="1" pageNumber="596" unit="mm" value="0.71">0.71 mm</quantity>
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, width across eyes
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||
<quantity box="[1244,1351,1243,1270]" metricMagnitude="-4" metricUnit="m" metricValue="6.2" pageId="1" pageNumber="596" unit="mm" value="0.62">0.62 mm</quantity>
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; vertex rough, small asetose vertexal foveae (dorsal tentorial pits) at level posterior to middle of eyes; frons strongly impressed around setose frontal fovea, antennal tubercle weakly convex, forming distinct rostrum, area between tubercles moderately impressed; ocular canthus short, broadly triangular. Gular deeply impressed at middle to form transversely oval depression, where widely separated foveae (posterior tentorial pits) are located. Compound eyes small, each composed of approximately 22 ommatidia. Antenna moderately elongate, length
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<quantity box="[1214,1318,1423,1450]" metricMagnitude="-3" metricUnit="m" metricValue="1.73" pageId="1" pageNumber="596" unit="mm" value="1.73">1.73 mm</quantity>
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, indistinct club formed by three apical moderately enlarged antennomeres, lacking modifications; antennomere 1 shorter than 2 and 3 combined, 1.73 times as long as wide, 2 slightly wider than 3–8, 3–8 each elongate, successively shorter, 9 and 10 subequal in length, 10 slightly wider than 9, 11 suboval, truncate at base, as long as 9 and 10 combined. Maxillary palpus (
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||
<figureCitation box="[360,446,1567,1593]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="1" pageNumber="596">Fig. 1C</figureCitation>
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) with palpomere 1 small, 2 long and strongly curved, 3 and 4 basally pedunculate and apically broadened, 4 densely setose, with short, translucent palpal cone at apex; length/ width of palpomere 2 0.32/
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||
<quantity box="[209,310,1639,1666]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="596" unit="mm" value="0.07">0.07 mm</quantity>
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||
, 3 0.18/
|
||
<quantity box="[400,501,1639,1666]" metricMagnitude="-5" metricUnit="m" metricValue="7" pageId="1" pageNumber="596" unit="mm" value="0.07">0.07 mm</quantity>
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||
, 4 0.21/
|
||
<quantity box="[590,694,1639,1666]" metricMagnitude="-5" metricUnit="m" metricValue="9.0" pageId="1" pageNumber="596" unit="mm" value="0.09">0.09 mm</quantity>
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.
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</paragraph>
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||
<paragraph blockId="1.[151,1437,739,2025]" pageId="1" pageNumber="596">
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Pronotum (
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||
<figureCitation box="[325,409,1675,1701]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="1" pageNumber="596">Fig. 1B</figureCitation>
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) slightly longer than wide, length along middle
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||
<quantity box="[947,1049,1675,1702]" metricMagnitude="-4" metricUnit="m" metricValue="7.7" pageId="1" pageNumber="596" unit="mm" value="0.77">0.77 mm</quantity>
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||
, maximum width
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||
<quantity box="[1250,1352,1675,1702]" metricMagnitude="-4" metricUnit="m" metricValue="7.199999999999999" pageId="1" pageNumber="596" unit="mm" value="0.72">0.72 mm</quantity>
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||
, widest at anterior 1/3; lateral margins rounded at widest point, convergent anteriorly and posteriorly, moderately incised at level of lateral antebasal foveae; disc slightly convex, finely punctate, with shallow median and lateral longitudinal impressions, with large, setose median and lateral antebasal foveae, lacking transverse antebasal impression or mediobasal carina. Prosternum with anterior part as long as coxal part, lateral procoxal foveae in densely setose impression; margin of coxal cavity weakly carinate.
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</paragraph>
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<paragraph blockId="1.[151,1437,739,2025]" pageId="1" pageNumber="596">
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Elytra greatly shortened, subtrapezoidal, truncate at bases, much wider than long, length along suture
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<quantity box="[1330,1432,1891,1918]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="1" pageNumber="596" unit="mm" value="0.59">0.59 mm</quantity>
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, maximum width
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||
<quantity box="[345,448,1927,1954]" metricMagnitude="-3" metricUnit="m" metricValue="1.29" pageId="1" pageNumber="596" unit="mm" value="1.29">1.29 mm</quantity>
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, length/width 0.46, broadest shortly anterior to posterior margin; each elytron with two large, setose basal foveae and broad, deep discal stria; lacking humeral tubercle; subhumeral fovea and marginal stria absent.
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</paragraph>
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||
<caption ID-DOI="http://doi.org/10.5281/zenodo.7673189" ID-Zenodo-Dep="7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="2" pageNumber="597" startId="2.[151,250,1801,1825]" targetBox="[152,1436,180,1776]" targetPageId="2">
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||
<paragraph blockId="2.[151,1437,1801,1898]" pageId="2" pageNumber="597">
|
||
<emphasis bold="true" box="[151,275,1801,1825]" pageId="2" pageNumber="597">FIGURE 1.</emphasis>
|
||
<taxonomicName authority="Yin, 2023" authorityName="Yin" authorityYear="2023" box="[281,499,1802,1826]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="597" phylum="Arthropoda" rank="species" species="schmidti" status="sp. nov.">
|
||
<emphasis box="[281,499,1802,1826]" italics="true" pageId="2" pageNumber="597">Pakistatyrus schmidti</emphasis>
|
||
</taxonomicName>
|
||
<emphasis bold="true" box="[505,585,1802,1826]" pageId="2" pageNumber="597">
|
||
<taxonomicNameLabel box="[505,585,1802,1826]" pageId="2" pageNumber="597" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, male.
|
||
<emphasis bold="true" box="[660,684,1801,1825]" pageId="2" pageNumber="597">A.</emphasis>
|
||
Dorsal Habitus;
|
||
<emphasis bold="true" box="[856,873,1802,1826]" pageId="2" pageNumber="597">B</emphasis>
|
||
. Head and pronotum;
|
||
<emphasis bold="true" box="[1098,1123,1801,1825]" pageId="2" pageNumber="597">C.</emphasis>
|
||
Left maxillary palpus, ventral;
|
||
<emphasis bold="true" box="[151,176,1838,1862]" pageId="2" pageNumber="597">D.</emphasis>
|
||
Protrochanter and profemur;
|
||
<emphasis bold="true" box="[476,499,1838,1862]" pageId="2" pageNumber="597">E.</emphasis>
|
||
Mesotrochanter and mesofemur;
|
||
<emphasis bold="true" box="[840,859,1838,1862]" pageId="2" pageNumber="597">F.</emphasis>
|
||
Sternite IX;
|
||
<emphasis bold="true" box="[990,1048,1837,1862]" pageId="2" pageNumber="597">G, H.</emphasis>
|
||
Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="2.[151,1436,1944,2042]" pageId="2" pageNumber="597">Mesoventrite short, fused with metaventrite, with relatively long and broad mesoventral process. Metaventrite distinctly ridged admesally, broadly impressed at middle, anterior metaventral process short, with truncate anterior margin, posterior process with short slit at middle.</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="598">
|
||
Legs elongate; protibia with two perpendicular rows of spinose setae along mesal margin, protrochanter (
|
||
<figureCitation captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="3" pageNumber="598">Fig. 1D</figureCitation>
|
||
) with blunt ventral spine, two spines on ventral surface of profemur (
|
||
<figureCitation box="[971,1058,187,213]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="3" pageNumber="598">Fig. 1D</figureCitation>
|
||
) supposedly large and long, as in
|
||
<taxonomicName authorityName="Hlavac" authorityYear="2006" box="[151,227,223,249]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="ater">
|
||
<emphasis box="[151,168,223,248]" italics="true" pageId="3" pageNumber="598">P</emphasis>
|
||
.
|
||
<emphasis box="[182,227,223,248]" italics="true" pageId="3" pageNumber="598">ater</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName authorityName="Yin" authorityYear="2022" box="[282,459,223,249]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="inconspicuus">
|
||
<emphasis box="[282,299,223,248]" italics="true" pageId="3" pageNumber="598">P</emphasis>
|
||
.
|
||
<emphasis box="[313,459,223,248]" italics="true" pageId="3" pageNumber="598">inconspicuus</emphasis>
|
||
</taxonomicName>
|
||
(both spines of left profemur broken in
|
||
<typeStatus box="[908,1009,223,249]" pageId="3" pageNumber="598">holotype</typeStatus>
|
||
, as indicated by yellow arrowheads in figure); mesocoxa simple, mesotrochanter (
|
||
<figureCitation box="[643,728,259,285]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="3" pageNumber="598">Fig. 1E</figureCitation>
|
||
) with small ventral spine, mesofemur swollen, mesotibia with short tubercle at apex; hind leg simple, lacking spines or projections.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="598">
|
||
Abdomen much broader than elytra, approximately as long as wide, widest at lateral margins of tergite 1 (IV), length
|
||
<quantity box="[227,328,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.42" pageId="3" pageNumber="598" unit="mm" value="1.42">1.42 mm</quantity>
|
||
, width
|
||
<quantity box="[407,509,367,393]" metricMagnitude="-3" metricUnit="m" metricValue="1.49" pageId="3" pageNumber="598" unit="mm" value="1.49">1.49 mm</quantity>
|
||
. Tergites 1–3 (IV–VI) subequal in length along middle, each with broad paratergites protruding laterally, setose basal sulcus and two basolateral foveae, 4 (VII) slightly longer than 3 (VI) along middle, with triangular paratergites and pair of basolateral foveae in setose lateral impressions, 5 (VIII) transverse, posterior margin broadly truncate and slightly sinuate, with two small basolateral foveae. Sternites 2–4 (IV–VI) subequal in length along middle, 5 (VII) shorter than 4, 2–5 each with setose basal sulcus and two basolateral at lateral margins of sulcus, 6 (VIII) transverse, posterior margin roundly emarginate at middle, 7 (IX) (
|
||
<figureCitation box="[1211,1293,547,573]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="3" pageNumber="598">Fig. 1F</figureCitation>
|
||
) moderately sclerotized, composed of one middle and two lateral plates, middle sclerite with many setae along apical margin.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="598">
|
||
Aedeagus (
|
||
<figureCitation box="[325,446,619,645]" captionStart="FIGURE 1" captionStartId="2.[151,250,1801,1825]" captionTargetBox="[152,1436,180,1776]" captionTargetId="figure-29@2.[151,1436,179,1776]" captionTargetPageId="2" captionText="FIGURE 1. Pakistatyrus schmidti sp. nov., male. A. Dorsal Habitus; B. Head and pronotum; C. Left maxillary palpus, ventral; D. Protrochanter and profemur; E. Mesotrochanter and mesofemur; F. Sternite IX; G, H. Aedeagus, lateral (G) and ventral (H). Scale bars: 1.0 mm in A; 0.5 mm in B, D, E; 0.2 mm in C, F–H." figureDoi="http://doi.org/10.5281/zenodo.7673189" httpUri="https://zenodo.org/record/7673189/files/figure.png" pageId="3" pageNumber="598">Fig. 1G, H</figureCitation>
|
||
)
|
||
<quantity box="[461,561,618,645]" metricMagnitude="-4" metricUnit="m" metricValue="5.8999999999999995" pageId="3" pageNumber="598" unit="mm" value="0.59">0.59 mm</quantity>
|
||
long, dorso-ventrally asymmetric; median lobe broad at base, narrowing from below middle to apex, with bulbous basal capsule and small foramen, dorsal diaphragm distinct, apical area curved to left (position as in figure); parameres greatly asymmetric, left paramere much longer than right one, apical margin of each paramere with four long setae; endophallus armature absent.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,1221]" box="[199,415,763,789]" pageId="3" pageNumber="598">
|
||
<emphasis box="[199,283,763,789]" italics="true" pageId="3" pageNumber="598">Female</emphasis>
|
||
. Unknown.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="598">
|
||
<emphasis bold="true" box="[199,433,798,825]" pageId="3" pageNumber="598">Comparative notes.</emphasis>
|
||
This species, exhibiting the greatest degree of elytral reduction (elytral length/width 0.46 vs.
|
||
<quantity box="[191,270,834,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.3208" pageId="3" pageNumber="598" unit="in" value="0.52">0.52 in</quantity>
|
||
<taxonomicName authorityName="Hlavac" authorityYear="2006" box="[278,355,835,861]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="ater">
|
||
<emphasis box="[278,295,835,860]" italics="true" pageId="3" pageNumber="598">P</emphasis>
|
||
.
|
||
<emphasis box="[310,355,835,860]" italics="true" pageId="3" pageNumber="598">ater</emphasis>
|
||
</taxonomicName>
|
||
, and
|
||
<quantity box="[419,498,835,861]" metricMagnitude="-2" metricUnit="m" metricValue="1.6002" pageId="3" pageNumber="598" unit="in" value="0.63">0.63 in</quantity>
|
||
<taxonomicName authorityName="Yin" authorityYear="2022" box="[506,684,835,861]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="inconspicuus">
|
||
<emphasis box="[506,523,835,860]" italics="true" pageId="3" pageNumber="598">P</emphasis>
|
||
.
|
||
<emphasis box="[538,684,835,860]" italics="true" pageId="3" pageNumber="598">inconspicuus</emphasis>
|
||
</taxonomicName>
|
||
), differs from
|
||
<taxonomicName authorityName="Hlavac" authorityYear="2006" box="[852,929,835,861]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="ater">
|
||
<emphasis box="[852,869,835,860]" italics="true" pageId="3" pageNumber="598">P</emphasis>
|
||
.
|
||
<emphasis box="[884,929,835,860]" italics="true" pageId="3" pageNumber="598">ater</emphasis>
|
||
</taxonomicName>
|
||
by the reddish-brown body and simple male antennal clubs; and from
|
||
<taxonomicName authorityName="Yin" authorityYear="2022" box="[441,619,871,897]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="inconspicuus">
|
||
<emphasis box="[441,458,871,896]" italics="true" pageId="3" pageNumber="598">P</emphasis>
|
||
.
|
||
<emphasis box="[473,619,871,896]" italics="true" pageId="3" pageNumber="598">inconspicuus</emphasis>
|
||
</taxonomicName>
|
||
by the rough vertex, relatively much shorter antennomeres
|
||
<quantity box="[1299,1343,871,897]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="3" pageNumber="598" unit="in" value="1.0">1 in</quantity>
|
||
relation to 2 and 3, and much larger foveae of the pronotum and elytra. The greatly asymmetric aedeagus alone of the new species is characteristic. Geographically,
|
||
<taxonomicName authorityName="Yin" authorityYear="2023" box="[616,743,943,969]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="schmidti">
|
||
<emphasis box="[616,633,943,968]" italics="true" pageId="3" pageNumber="598">P</emphasis>
|
||
.
|
||
<emphasis box="[648,743,943,969]" italics="true" pageId="3" pageNumber="598">schmidti</emphasis>
|
||
</taxonomicName>
|
||
is only
|
||
<quantity box="[835,907,943,969]" metricMagnitude="4" metricUnit="m" metricValue="8.0" pageId="3" pageNumber="598" unit="km" value="80.0">80 km</quantity>
|
||
east to
|
||
<taxonomicName box="[995,1175,943,969]" class="Insecta" family="Staphylinidae" genus="Pakistatyrus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="598" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis box="[995,1135,943,969]" italics="true" pageId="3" pageNumber="598">Pakistatyrus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
distributed in southern
|
||
<collectingRegion box="[151,209,979,1005]" country="China" name="Xizang" pageId="3" pageNumber="598">Tibet</collectingRegion>
|
||
(
|
||
<figureCitation box="[228,300,979,1005]" captionStart="FIGURE 2" captionStartId="3.[151,250,1990,2014]" captionTargetBox="[151,1436,1245,1967]" captionTargetId="figure-467@3.[151,1436,1245,1967]" captionTargetPageId="3" captionText="FIGURE 2. Distribution of Pakistatyrus species along the Himalaya, with the aedeagi compared." figureDoi="http://doi.org/10.5281/zenodo.7673191" httpUri="https://zenodo.org/record/7673191/files/figure.png" pageId="3" pageNumber="598">Fig. 2</figureCitation>
|
||
), but the single female apparently represents another distinct species with smooth vertex, smaller pronotal and elytral foveae, and longer elytra (length/width 0.57). The three known species of the genus can also be quickly separated by using the key provided above.
|
||
</paragraph>
|
||
<paragraph blockId="3.[151,1437,151,1221]" box="[199,456,1086,1113]" pageId="3" pageNumber="598">
|
||
<emphasis bold="true" box="[199,331,1086,1112]" pageId="3" pageNumber="598">Bionomics.</emphasis>
|
||
Unknown.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[199,634,1122,1149]" pageId="3" pageNumber="598" type="distribution">
|
||
<paragraph blockId="3.[151,1437,151,1221]" box="[199,634,1122,1149]" pageId="3" pageNumber="598">
|
||
<emphasis bold="true" box="[199,353,1122,1148]" pageId="3" pageNumber="598">Distribution.</emphasis>
|
||
<collectingCountry box="[359,430,1123,1149]" name="Nepal" pageId="3" pageNumber="598">Nepal</collectingCountry>
|
||
:
|
||
<collectingRegion box="[441,536,1123,1149]" country="Nepal" name="Gandaki" pageId="3" pageNumber="598">Gandaki</collectingRegion>
|
||
(
|
||
<figureCitation box="[551,621,1123,1149]" captionStart="FIGURE 2" captionStartId="3.[151,250,1990,2014]" captionTargetBox="[151,1436,1245,1967]" captionTargetId="figure-467@3.[151,1436,1245,1967]" captionTargetPageId="3" captionText="FIGURE 2. Distribution of Pakistatyrus species along the Himalaya, with the aedeagi compared." figureDoi="http://doi.org/10.5281/zenodo.7673191" httpUri="https://zenodo.org/record/7673191/files/figure.png" pageId="3" pageNumber="598">Fig. 2</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="3" pageNumber="598" type="etymology">
|
||
<paragraph blockId="3.[151,1437,151,1221]" pageId="3" pageNumber="598">
|
||
<emphasis bold="true" box="[199,334,1159,1185]" pageId="3" pageNumber="598">Etymology.</emphasis>
|
||
The new species is named after German carabidologist Joachim Schmidt (University of Rostock, Rostock,
|
||
<collectingCountry box="[257,366,1195,1221]" name="Germany" pageId="3" pageNumber="598">Germany</collectingCountry>
|
||
), who collected the single male.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |