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<document id="2F36ECD1CC16F090DF80898D796EFB95" ID-DOI="10.11646/phytotaxa.630.4.2" ID-ISSN="1179-3163" ID-Zenodo-Dep="10417649" IM.bibliography_approvedBy="karina" IM.illustrations_approvedBy="karina" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="karina" IM.treatments_approvedBy="karina" checkinTime="1703151358317" checkinUser="plazi" docAuthor="Smith, Gideon F." docDate="2023" docId="03DBA95FFFF90B0A2C8F2ECBFE0A08B3" docLanguage="en" docName="phytotaxa.630.4.2.pdf" docOrigin="Phytotaxa 630 (4)" docSource="https://phytotaxa.mapress.com/pt/article/download/phytotaxa.630.4.2/51381" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Kalanchoe dyeri Brown 1904" docType="treatment" docVersion="1" lastPageNumber="272" masterDocId="FFE2D127FFFF0B0C2C072E55FF970C1B" masterDocTitle="A nomenclatural and taxonomic review of the names and new combination published between 1886 and 1931 in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by British botanist Nicholas Edward Brown (1849 - 1934)" masterLastPageNumber="280" masterPageNumber="266" pageNumber="272" updateTime="1703274160119" updateUser="karina" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="CE135F87CD40E79628E9E585800CE2A0">A nomenclatural and taxonomic review of the names and new combination published between 1886 and 1931 in Kalanchoe (Crassulaceae subfam. Kalanchooideae) by British botanist Nicholas Edward Brown (1849 - 1934)</mods:title>
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<mods:date id="3101CC74943AA677B5896CDB8D51D94C">2023</mods:date>
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<treatment id="03DBA95FFFF90B0A2C8F2ECBFE0A08B3" LSID="urn:lsid:plazi:treatment:03DBA95FFFF90B0A2C8F2ECBFE0A08B3" httpUri="http://treatment.plazi.org/id/03DBA95FFFF90B0A2C8F2ECBFE0A08B3" lastPageNumber="272" pageId="6" pageNumber="272">
<subSubSection id="C3684BC2FFF90B0A2C8F2ECBFDC30CA2" box="[136,596,158,185]" pageId="6" pageNumber="272" type="nomenclature">
<paragraph id="8BCD1849FFF90B0A2C8F2ECBFDC30CA2" blockId="6.[136,596,158,185]" box="[136,596,158,185]" pageId="6" pageNumber="272">
<heading id="D085AF25FFF90B0A2C8F2ECBFDC30CA2" box="[136,596,158,185]" fontSize="11" level="2" pageId="6" pageNumber="272" reason="4">
6.
<taxonomicName id="4C7263CAFFF90B0A2CA32ECBFDC70CA2" ID-CoL="3QYJF" authority="Brown (1904 b: 354)" authorityName="Brown" authorityPageNumber="354" authorityYear="1904" box="[164,592,158,185]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2CA32ECBFEF50CA2" bold="true" box="[164,354,158,185]" italics="true" pageId="6" pageNumber="272">Kalanchoe dyeri</emphasis>
<bibRefCitation id="EFE365B8FFF90B0A2D6F2ECAFDC70CA2" author="Brown, N. E." box="[360,592,158,185]" pageId="6" pageNumber="272" refId="ref10692" refString="Brown, N. E. (1904 b [4 June]) New or noteworthy plants. KALANCHOE DYERI, N. E. Brown (n. sp.). The Gardeners' Chronicle [A weekly illustrated journal of horticulture and allied subjects], 3 rd ser., 35 (No. 910): 354. Available from: https: // www. biodiversitylibrary. org / item / 83823 # page / 420 / mode / 1 up (accessed: 20 December 2023)." type="url" year="1904">Brown (1904b: 354)</bibRefCitation>
</taxonomicName>
.
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3684BC2FFF90B0A2C8F2EB2FD910D8A" pageId="6" pageNumber="272" type="materials_examined">
<paragraph id="8BCD1849FFF90B0A2C8F2EB2FDA70D3E" blockId="6.[136,1452,230,1192]" pageId="6" pageNumber="272">
<emphasis id="B906C45BFFF90B0A2C8F2EB2FF2B0D1B" box="[136,188,231,256]" italics="true" pageId="6" pageNumber="272">Type</emphasis>
:—
<collectingCountry id="F36558D9FFF90B0A2CDA2EB3FEC00D1B" box="[221,343,230,256]" name="Malawi" pageId="6" pageNumber="272">MALAWI</collectingCountry>
, “
<emphasis id="B906C45BFFF90B0A2D742EB2FD830D1B" box="[371,532,231,257]" italics="true" pageId="6" pageNumber="272">
<taxonomicName id="4C7263CAFFF90B0A2D742EB2FE7F0D1A" authorityName="Lang" authorityYear="1884" box="[371,488,231,257]" family="Anonymidae" genus="Anonymus" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="272" phylum="Platyhelminthes" rank="genus">Anonymus</taxonomicName>
s.n
</emphasis>
.”, as “
<emphasis id="B906C45BFFF90B0A2E6E2EB3FDEB0D1B" bold="true" box="[617,636,230,256]" pageId="6" pageNumber="272">T</emphasis>
”, without further information, here
<emphasis id="B906C45BFFF90B0A28202EB3FAA50D1B" bold="true" box="[1063,1330,230,256]" pageId="6" pageNumber="272">
corrected to
<typeStatus id="54C9A6EBFFF90B0A28C02EB3FAA50D1B" box="[1223,1330,230,256]" pageId="6" pageNumber="272" type="lectotype">lectotype</typeStatus>
</emphasis>
, Herb. K, designated by
<bibRefCitation id="EFE365B8FFF90B0A2D2D2F5FFDBB0D3E" author="Descoings, B." box="[298,556,266,293]" pageId="6" pageNumber="272" pagination="143 - 181" refId="ref11416" refString="Descoings, B. (2003) Kalanchoe. In: Eggli, U. (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, Heidelberg, New York, pp. 143 - 181; whole work 458 pp. + XLVIII plates." type="book chapter" year="2003">Descoings (2003: 155)</bibRefCitation>
.
</paragraph>
<paragraph id="8BCD1849FFF90B0A2CBA2F7AFD910D8A" blockId="6.[136,1452,230,1192]" pageId="6" pageNumber="272">
<emphasis id="B906C45BFFF90B0A2CBA2F7AFE840D53" box="[189,275,303,328]" italics="true" pageId="6" pageNumber="272">
<typeStatus id="54C9A6EBFFF90B0A2CBA2F7AFE840D53" box="[189,275,303,328]" pageId="6" pageNumber="272" type="epitype">Epitype</typeStatus>
</emphasis>
:—[Nyasaland]
<collectingCountry id="F36558D9FFF90B0A2DC02F7BFDA90D53" box="[455,574,302,328]" name="Malawi" pageId="6" pageNumber="272">MALAWI</collectingCountry>
, received at the Royal Botanic Gardens, Kew, in 1902, specimen prepared on
<date id="FFCC3E89FFF90B0A2C8F2F07FE830D77" box="[136,276,338,365]" pageId="6" pageNumber="272" value="1904-05-02">2 May 1904</date>
,
<emphasis id="B906C45BFFF90B0A2D182F06FE5D0D77" box="[287,458,339,364]" italics="true" pageId="6" pageNumber="272">N.E. Brown s.n</emphasis>
. (
<emphasis id="B906C45BFFF90B0A2DE52F07FDAE0D77" bold="true" box="[482,569,338,364]" pageId="6" pageNumber="272">
<typeStatus id="54C9A6EBFFF90B0A2DE52F07FDAE0D77" box="[482,569,338,364]" pageId="6" pageNumber="272" type="epitype">epitype</typeStatus>
</emphasis>
, K barcode K232910 [image available at http://specimens.kew.org/herbarium/ K000232910]!),
<emphasis id="B906C45BFFF90B0A2D432F23FD970D8B" bold="true" box="[324,512,374,400]" pageId="6" pageNumber="272">here designated</emphasis>
.
</paragraph>
</subSubSection>
<subSubSection id="C3684BC2FFF90B0A2CBA2FCEFE0A08B3" pageId="6" pageNumber="272" type="discussion">
<paragraph id="8BCD1849FFF90B0A2CBA2FCEFB4A0DE7" blockId="6.[136,1452,230,1192]" pageId="6" pageNumber="272">
<emphasis id="B906C45BFFF90B0A2CBA2FCEFEEB0DAF" box="[189,380,411,436]" italics="true" pageId="6" pageNumber="272">Taxonomic notes</emphasis>
:—At present
<taxonomicName id="4C7263CAFFF90B0A2E272FCEFDE90DAF" authorityName="Brown" authorityYear="1904" box="[544,638,410,436]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2E272FCEFDE90DAF" box="[544,638,410,436]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
is treated as an accepted species. The date of publication of the name
<taxonomicName id="4C7263CAFFF90B0A2C8F2FEAFF730DC3" authorityName="Brown" authorityYear="1904" box="[136,228,446,472]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2C8F2FEAFF730DC3" box="[136,228,446,472]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
is sometimes in error given as “1902” (see for example https://www.ipni.org/n/274325-1), likely because
<emphasis id="B906C45BFFF90B0A299F2FEAFEB00DE7" italics="true" pageId="6" pageNumber="272">J. McClounie 35</emphasis>
was collected in
<date id="FFCC3E89FFF90B0A2DED2FB7FD330DE7" box="[490,676,482,508]" pageId="6" pageNumber="272" value="1902-09">September 1902</date>
. However, the species was only described in 1904.
</paragraph>
<paragraph id="8BCD1849FFF90B0A2CBA2C53FCC70ECE" blockId="6.[136,1452,230,1192]" pageId="6" pageNumber="272">
<emphasis id="B906C45BFFF90B0A2CBA2C53FE300E3B" box="[189,423,518,544]" italics="true" pageId="6" pageNumber="272">Nomenclatural notes</emphasis>
:—Herb. K holds
<specimenCount id="9D74D3C0FFF90B0A2E692C52FC840E3B" box="[622,787,518,545]" count="2" pageId="6" pageNumber="272" type="generic">two specimens</specimenCount>
of
<taxonomicName id="4C7263CAFFF90B0A2F302C52FC060E3B" authorityName="Brown" authorityYear="1904" box="[823,913,518,544]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2F302C52FC060E3B" box="[823,913,518,544]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
that predate description of the species on
<date id="FFCC3E89FFF90B0A29632C53FF540E5F" pageId="6" pageNumber="272" value="1904-06-04">4 June 1904</date>
. The first one, which consists of reproductive material only—intact inflorescence branches and flowers—is
<emphasis id="B906C45BFFF90B0A299F2C7EFEB10E73" italics="true" pageId="6" pageNumber="272">J. McClounie 35</emphasis>
, dated
<date id="FFCC3E89FFF90B0A2D712C1BFDBC0E73" box="[374,555,590,616]" pageId="6" pageNumber="272" value="1902-09">September 1902</date>
on the collecting label and 1903 on the herbarium label. Both these dates predate the description of
<taxonomicName id="4C7263CAFFF90B0A2D552C26FE3B0E97" authorityName="Brown" authorityYear="1904" box="[338,428,626,652]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2D552C26FE3B0E97" box="[338,428,626,652]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
in 1904. The material was collected in Northern
<collectingRegion id="49B6D6ABFFF90B0A2FC92C26FBA60E96" box="[974,1073,627,653]" country="Malawi" name="Karonga" pageId="6" pageNumber="272">Karonga</collectingRegion>
, Panda [Mpanda] Peak in
<collectingCountry id="F36558D9FFF90B0A29512C27FA3B0E97" box="[1366,1452,626,652]" name="Malawi" pageId="6" pageNumber="272">Malawi</collectingCountry>
[“British Central Africa”] at an elevation of
<quantity id="4C8AB5ACFFF90B0A2E742CC3FD5D0EAB" box="[627,714,662,688]" metricMagnitude="3" metricUnit="m" metricValue="1.9449288" pageId="6" pageNumber="272" unit="ft" value="6381.0">6381 ft.</quantity>
[~
<quantity id="4C8AB5ACFFF90B0A2EE22CC3FCD50EAA" box="[741,834,662,689]" metricMagnitude="3" metricUnit="m" metricValue="1.944" pageId="6" pageNumber="272" unit="m" value="1944.0">1944 m</quantity>
]. The barcode of this specimen is K000285698 and it is accessible at http://specimens.kew.org/herbarium/K000285698.
</paragraph>
<paragraph id="8BCD1849FFF90B0A2CBA2C8BFBD80FB7" blockId="6.[136,1452,230,1192]" pageId="6" pageNumber="272">
The second specimen of
<taxonomicName id="4C7263CAFFF90B0A2DE62C8AFDA80EE3" authorityName="Brown" authorityYear="1904" box="[481,575,734,760]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2DE62C8AFDA80EE3" box="[481,575,734,760]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
held at Herb. K that predates description of this species on
<date id="FFCC3E89FFF90B0A29062C8BFA190EE3" box="[1281,1422,734,761]" pageId="6" pageNumber="272" value="1904-06-04">4 June 1904</date>
is
<emphasis id="B906C45BFFF90B0A2C8F2D56FEA10F07" box="[136,310,771,796]" italics="true" pageId="6" pageNumber="272">N.E. Brown s.n</emphasis>
. This specimen is dated
<date id="FFCC3E89FFF90B0A2E532D57FD770F07" box="[596,736,770,797]" pageId="6" pageNumber="272" value="1904-05-02">2 May 1904</date>
and was therefore prepared one month before the species was described. The barcode of this material, which consists of both vegetative and reproductive material, is K232910, and it is accessible at http://specimens.kew.org/herbarium/K000232910. A month earlier, on
<date id="FFCC3E89FFF90B0A285A2D1FFB7C0F7F" box="[1117,1259,842,868]" pageId="6" pageNumber="272" value="1904-04-02">2 April 1904</date>
, the material was figured for inclusion in volume 60 [third series] of
<emphasis id="B906C45BFFF90B0A2EC22D3AFB920F93" box="[709,1029,878,905]" italics="true" pageId="6" pageNumber="272">Curtiss Botanical Magazine</emphasis>
, with the painting and accompanying text eventually appearing later in the same year, as plate 7987 (
<bibRefCitation id="EFE365B8FFF90B0A2F4E2DC7FC710FB7" author="Brown, N. E." box="[841,998,914,940]" pageId="6" pageNumber="272" pagination="659" refId="ref10787" refString="Brown, N. E. (1904 c) Kalanchoe dyeri. Native of Nyassaland. [Nat. Ord. Crassulaceae. Genus Kalanchoe, Adans.; (Benth. et Hook. f. Gen. Plant. vol. i. p. 659.)] Curtis's Botanical Magazine 60 [&quot; LX of the third series; CXXX of the whole work &quot;]: t. 7987. Available from: https: // www. biodiversitylibrary. org / item / 14258 # page / 232 / mode / 1 up (accessed: 20 December 2023)." type="journal article" year="1904">Brown 1904c</bibRefCitation>
: t. 7987).
</paragraph>
<paragraph id="8BCD1849FFF90B0A2CBA2DE3FE0A08B3" blockId="6.[136,1452,230,1192]" pageId="6" pageNumber="272">
<bibRefCitation id="EFE365B8FFF90B0A2CBA2DE3FE290FCA" author="Descoings, B." box="[189,446,950,977]" pageId="6" pageNumber="272" pagination="143 - 181" refId="ref11416" refString="Descoings, B. (2003) Kalanchoe. In: Eggli, U. (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, Heidelberg, New York, pp. 143 - 181; whole work 458 pp. + XLVIII plates." type="book chapter" year="2003">Descoings (2003: 155)</bibRefCitation>
was the first to typify the name
<taxonomicName id="4C7263CAFFF90B0A2F352DE2FC190FCB" authorityName="Brown" authorityYear="1904" box="[818,910,950,976]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2F352DE2FC190FCB" box="[818,910,950,976]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
, which he did on a specimen referred to as “
<emphasis id="B906C45BFFF90B0A298B2DE3FA080FCB" bold="true" box="[1420,1439,950,976]" pageId="6" pageNumber="272">T</emphasis>
” (see
<bibRefCitation id="EFE365B8FFF90B0A2CB92D8FFED70FEF" author="Eggli, U." box="[190,320,986,1012]" pageId="6" pageNumber="272" refId="ref11544" refString="Eggli, U. (Ed.) (2003) List of abbreviations and symbols. In: Eggli, U. (Ed.) Illustrated handbook of succulent plants: Crassulaceae. Springer Verlag, Berlin, Heidelberg, New York, pp. IX - X; whole work 458 pp. + XLVIII plates. https: // doi. org / 10.1007 / 978 - 3 - 642 - 55874 - 0" type="book" year="2003">Eggli 2003</bibRefCitation>
: X for a definition of “
<emphasis id="B906C45BFFF90B0A2E492D8FFDF60FEF" bold="true" box="[590,609,986,1012]" pageId="6" pageNumber="272">T</emphasis>
”) “
<collectingCountry id="F36558D9FFF90B0A2E8E2D8FFD760FEF" box="[649,737,986,1012]" name="Malawi" pageId="6" pageNumber="272">Malawi</collectingCountry>
(
<taxonomicName id="4C7263CAFFF90B0A2EF32D8EFCFE0FEE" authorityName="Lang" authorityYear="1884" box="[756,873,987,1013]" family="Anonymidae" genus="Anonymus" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="272" phylum="Platyhelminthes" rank="genus">
<emphasis id="B906C45BFFF90B0A2EF32D8EFCFE0FEE" box="[756,873,987,1013]" italics="true" pageId="6" pageNumber="272">Anonymus</emphasis>
</taxonomicName>
s.n. [K])”. Since
<bibRefCitation id="EFE365B8FFF90B0A28312D8EFAB70FEE" author="Brown, N. E." box="[1078,1312,986,1013]" pageId="6" pageNumber="272" refId="ref10692" refString="Brown, N. E. (1904 b [4 June]) New or noteworthy plants. KALANCHOE DYERI, N. E. Brown (n. sp.). The Gardeners' Chronicle [A weekly illustrated journal of horticulture and allied subjects], 3 rd ser., 35 (No. 910): 354. Available from: https: // www. biodiversitylibrary. org / item / 83823 # page / 420 / mode / 1 up (accessed: 20 December 2023)." type="url" year="1904">Brown (1904b: 354)</bibRefCitation>
did not cite any specimens in the protologue of the name and since, as discussed above, he clearly had more than
<specimenCount id="9D74D3C0FFF90B0A28F02DAAFA070803" box="[1271,1424,1022,1049]" count="1" pageId="6" pageNumber="272" type="generic">one specimen</specimenCount>
at his disposal when publishing the name
<taxonomicName id="4C7263CAFFF90B0A2E432A76FD080827" authorityName="Brown" authorityYear="1904" box="[580,671,1058,1084]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A2E432A76FD080827" box="[580,671,1058,1084]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
, this is interpreted as a lectotypification. The identity of this material [“
<collectingCountry id="F36558D9FFF90B0A2C9E2A13FF64087B" box="[153,243,1094,1120]" name="Malawi" pageId="6" pageNumber="272">Malawi</collectingCountry>
(
<taxonomicName id="4C7263CAFFF90B0A2D022A12FEED087A" authorityName="Lang" authorityYear="1884" box="[261,378,1095,1121]" family="Anonymidae" genus="Anonymus" kingdom="Animalia" order="Polycladida" pageId="6" pageNumber="272" phylum="Platyhelminthes" rank="genus">
<emphasis id="B906C45BFFF90B0A2D022A12FEED087A" box="[261,378,1095,1121]" italics="true" pageId="6" pageNumber="272">Anonymus</emphasis>
</taxonomicName>
s.n. [K])”] could not be determined and, in support of the
<typeStatus id="54C9A6EBFFF90B0A28102A13FB16087B" box="[1047,1153,1094,1120]" pageId="6" pageNumber="272" type="lectotype">lectotype</typeStatus>
, the name
<taxonomicName id="4C7263CAFFF90B0A28FB2A12FACF087B" authorityName="Brown" authorityYear="1904" box="[1276,1368,1094,1120]" class="Magnoliopsida" family="Crassulaceae" genus="Kalanchoe" kingdom="Plantae" order="Saxifragales" pageId="6" pageNumber="272" phylum="Tracheophyta" rank="species" species="dyeri">
<emphasis id="B906C45BFFF90B0A28FB2A12FACF087B" box="[1276,1368,1094,1120]" italics="true" pageId="6" pageNumber="272">K. dyeri</emphasis>
</taxonomicName>
is here epitypified on the specimen
<emphasis id="B906C45BFFF90B0A2DD02A3EFD1E089F" box="[471,649,1131,1156]" italics="true" pageId="6" pageNumber="272">N.E. Brown s.n</emphasis>
., barcode K232910, which is accessible at http://specimens.kew.org/ herbarium/K000232910.
</paragraph>
</subSubSection>
</treatment>
</document>