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<mods:title id="9E9C455AC2809939E962E8FF9E85EFFF">Fossil decapods from the Upper Quaternary in Shinjima Island in Kagoshima Kyushu, Japan, and description of a new species of ghost shrimp (Axiidea Eucalliacidae)</mods:title>
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<mods:affiliation id="F27FB870FC77FF74597F1B65AA5B22DE">Mizunami Fossil Museum, 1 - 47 Yamanouchi, Akeyo-cho, Mizunami City, Gifu 509 - 6132, Japan</mods:affiliation>
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<mods:namePart id="41519F6210158CCE66CCE62931A766D0">Kawano, Shigenori</mods:namePart>
<mods:affiliation id="E3C2290292438B0EECD325F825403614">Tochigi Prefectural Museum, 2 - 2 Mutsumi-cho, Utsunomiya City, Tochigi 320 - 0865, Japan.</mods:affiliation>
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<mods:affiliation id="1168EC7066A71DB30AA4AF0D628D69F9">Fukada Geological Institute, 2 - 13 - 12 Honkomagome, Bunkyo-ku, Tokyo 113 - 0021, Japan.</mods:affiliation>
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<mods:affiliation id="5E75B7A280F896EABCD2B60D3C2DEACF">Graduate School of Science and Technology, Kanazawa University, Kakuma, Kanazawa City, Ishikawa 920 - 1192, Japan. niiyama @ stu. kanazawa-u. ac. jp; https: // orcid. org / 0000 - 0002 - 7141 - 1072</mods:affiliation>
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<mods:namePart id="98FF4FDE74B2AFEF09B9B87ECCCBB97D">Kameyama, Sohiko</mods:namePart>
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<mods:affiliation id="C5BC01DE373BCAABF8D83738958B3200">Graduate School of Environmental Science, Hokkaido University, Kita-ku, Sapporo City, Hokkaido 060 - 0810, Japan. skameyama @ ees. hokudai. ac. jp; https: // orcid. org / 0000 - 0002 - 7104 - 1841</mods:affiliation>
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<mods:affiliation id="836456A0E7B32770072DC9336A29D7E9">Faculty of Science and Engineering, Saga University, 1 Honjo-machi, Saga City, Saga 840 - 8502, Japan. cshmym @ diary. ocn. ne. jp; https: // orcid. org / 0000 - 0002 - 2393 - 0820</mods:affiliation>
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<treatment id="03C987D66A0EB85D83E9FDD95C58FC76" ID-DOI="http://doi.org/10.5281/zenodo.4426214" ID-GBIF-Taxon="180323352" ID-Zenodo-Dep="4426214" LSID="urn:lsid:plazi:treatment:03C987D66A0EB85D83E9FDD95C58FC76" httpUri="http://treatment.plazi.org/id/03C987D66A0EB85D83E9FDD95C58FC76" lastPageNumber="532" pageId="9" pageNumber="532">
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<heading id="D09781AC6A0EB85D83E9FDD95A7CFDEB" bold="true" box="[151,567,583,610]" fontSize="11" level="1" pageId="9" pageNumber="532" reason="1">
<taxonomicName id="4C604D436A0EB85D83E9FDD95A7CFDEB" ID-CoL="7DBPK" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,567,583,610]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="532" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="B914EAD26A0EB85D83E9FDD95A7CFDEB" bold="true" box="[151,567,583,610]" pageId="9" pageNumber="532">
<emphasis id="B914EAD26A0EB85D83E9FDD95915FDEB" bold="true" box="[151,350,583,610]" italics="true" pageId="9" pageNumber="532">Tymolus uncifer</emphasis>
(
<bibRefCitation id="EFF14B316A0EB85D8213FDD95A64FDEB" author="Ortmann, A. E." box="[365,559,583,609]" pageId="9" pageNumber="532" pagination="532 - 588" refId="ref11141" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
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(
<figureCitation id="135B2A456A0EB85D83E1FDF5596AFD0F" box="[159,289,619,646]" captionStart="FIGURE 5" captionStartId="11.[151,250,1643,1667]" captionTargetBox="[151,1435,181,1619]" captionTargetId="figure-17@11.[151,1436,181,1619]" captionTargetPageId="11" captionText="FIGURE 5. Decapods from the Moeshima Silt Bed. A, Munida sp., MFM145609, dorsal view of carapace. B, C, Tymolus un- cifer (Ortmann, 1892); B, MFM145610, dorsal view of carapace; C, MFM145611, lateral view of left cheliped. D, Podocatactes hamifer Ortmann, 1893, MFM145618, dorsal view of carapace. EH, Carcinoplax longimana (de Haan, 1833); E, MFM145620, carapace and chelipeds; E1, dorsal view; E2, ventral view; F, MFM145621, dorsal view of carapace; G, MFM145622, carapace; G1, dorsal view; G2, ventral view; H, MFM145623, lateral view of left cheliped. Scale bars A, DH indicate 5.0 mm, and B and C indicate 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4574060" httpUri="https://zenodo.org/record/4574060/files/figure.png" pageId="9" pageNumber="532">Figs. 5B, C</figureCitation>
)
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<emphasis id="B914EAD26A0EB85D83E9FD2E5924FD42" box="[151,367,688,712]" italics="true" pageId="9" pageNumber="532">Cyclodrippe uncifera</emphasis>
<bibRefCitation id="EFF14B316A0EB85D8208FD2E5A0EFD4D" author="Ortmann, A. E." box="[374,581,687,712]" pageId="9" pageNumber="532" pagination="532 - 588" refId="ref11141" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892: 560</bibRefCitation>
.
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<taxonomicName id="4C604D436A0EB85D83E9FD505970FD6C" ID-CoL="7DBPK" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[151,315,718,742]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="532" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="B914EAD26A0EB85D83E9FD505970FD6C" box="[151,315,718,742]" italics="true" pageId="9" pageNumber="532">Tymolus uncifer</emphasis>
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.—
<treatmentCitation id="0AC110D16A0EB85D8224FD505A49FD6C" author="Balss, H." box="[346,514,718,742]" page="286" pageId="9" pageNumber="532" year="1922">
<bibRefCitation id="EFF14B316A0EB85D8224FD505A49FD6C" author="Balss, H." box="[346,514,718,742]" pageId="9" pageNumber="532" pagination="94 - 166" refId="ref7868" refString="Balss, H. (1922) Ostasiatische decapoden. IV, Die Brachyrhynchen (Cancridae). Archiv fur Naturgeschichte, 88, 94 - 166." type="journal article" year="1922">Balss 1922: 286</bibRefCitation>
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.—Karasawa 1993: 43, pl. 6, fig. 4.—
<treatmentCitation id="0AC110D16A0EB85D80F4FD505C0DFD6C" author="Takeda, M." box="[906,1094,718,742]" page="232" pageId="9" pageNumber="532" year="1997">
<bibRefCitation id="EFF14B316A0EB85D80F4FD505C0DFD6C" author="Takeda, M." box="[906,1094,718,742]" pageId="9" pageNumber="532" pagination="229 - 255" refId="ref12352" refString="Takeda, M. (1997) Deep-sea Decapod Crustacean Fauna of Suruga Bay, Central Japan. National Science Museum Monographs, 12, 229 - 255." type="journal article" year="1997">Takeda 1997: 232</bibRefCitation>
</treatmentCitation>
.—
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<bibRefCitation id="EFF14B316A0EB85D871AFD505D64FD6C" author="Karasawa, H." box="[1124,1327,718,742]" pageId="9" pageNumber="532" pagination="1 - 81" refId="ref9107" refString="Karasawa, H. (1997) A monograph of Cenozoic stomatopod, decapod, isopod and amphipod Crustacea from West Japan. Monograph of the Mizunami Fossil Museum, 8, 1 - 81." type="journal article" year="1997">Karasawa 1997: 41</bibRefCitation>
</treatmentCitation>
, pl. 8, fig. 10.—
<treatmentCitation id="0AC110D16A0EB85D8383FD7359D7FC8F" author="Ikeda, H." box="[253,412,749,773]" page="21" pageId="9" pageNumber="532" year="1998">
<bibRefCitation id="EFF14B316A0EB85D8383FD7359D7FC8F" author="Ikeda, H." box="[253,412,749,773]" pageId="9" pageNumber="532" refId="ref8726" refString="Ikeda, H. (1998) The deep-sea crabs of Sagami Bay. Hayama Shiosai Museum, Kanagawa, 180 pp." type="book" year="1998">Ikeda 1998: 21</bibRefCitation>
</treatmentCitation>
, pl. 5, figs. 13.
<treatmentCitation id="0AC110D16A0EB85D811AFD735B51FC8F" author="Takeda, M." box="[612,794,749,773]" page="223" pageId="9" pageNumber="532" year="2001">
<bibRefCitation id="EFF14B316A0EB85D811AFD735B51FC8F" author="Takeda, M." box="[612,794,749,773]" pageId="9" pageNumber="532" pagination="217 - 262" refId="ref12381" refString="Takeda, M. (2001) Annotated List of Crabs from Tosa Bay, Southwest Japan, Collected by the R / V / Kotaka Maru during the Years 1997 - 2000. National Science Museum Monographs, 20, 217 - 262." type="journal article" year="2001">Takeda 2001: 223</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0AC110D16A0EB85D8046FD735CC3FC8F" author="Spiridonov, V. A. &amp; Turkay, M." box="[824,1160,749,773]" page="6" pageId="9" pageNumber="532" year="2007">
<bibRefCitation id="EFF14B316A0EB85D8046FD735CC3FC8F" author="Spiridonov, V. A. &amp; Turkay, M." box="[824,1160,749,773]" pageId="9" pageNumber="532" pagination="1 - 9" refId="ref12085" refString="Spiridonov, V. A. &amp; Turkay, M. (2007) Podotreme crabs from the western India Ocean: description of a new species of Tymolus Stimpson, 1858 with a review of regional distribution of the Cyclodorippidae (Crustacea: Decapoda: Brachyura: Podotremata). Arthropoda Selecta, 16, 1 - 9." type="journal article" year="2007">Spiridonov &amp; Türkay 2007: 67</bibRefCitation>
</treatmentCitation>
, figs. 4DE.—
<treatmentCitation id="0AC110D16A0EB85D8661FD735916FCA9" author="Kitamura, K. &amp; Shiba, M." page="15" pageId="9" pageNumber="532" year="2008">
<bibRefCitation id="EFF14B316A0EB85D8661FD735916FCA9" author="Kitamura, K. &amp; Shiba, M." pageId="9" pageNumber="532" pagination="11 - 18" refId="ref9683" refString="Kitamura, K. &amp; Shiba, M. (2008) Fossil Decapod Crustaceans from the Hijikata Formation of the Kakegawa Group, distributed in Nagaya, Kakegawa City, Shizuoka Prefecture, Central Japan. Science Reports of The Museum, Tokai University, 9, 11 - 18." type="journal article" year="2008">Kitamura &amp; Shiba 2008: 15</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0AC110D16A0EB85D8202FC955AE4FCA9" author="Komatsu, H. &amp; Komai, T." box="[380,687,779,804]" page="598" pageId="9" pageNumber="532" year="2009">
<bibRefCitation id="EFF14B316A0EB85D8202FC955AE4FCA9" author="Komatsu, H. &amp; Komai, T." box="[380,687,779,804]" pageId="9" pageNumber="532" pagination="581 - 613" refId="ref9920" refString="Komatsu, H. &amp; Komai, T. (2009) Thalassinidea, Anomura and Brachyura (Crustacea: Decapoda) from Northeastern Japan Collected during the &quot; Research on Deep-sea Fauna and Pollutants off Pacific Coast of Northern Japan &quot; Project. National Museum of Nature and Science Monographs, 39, 581 - 613." type="journal article" year="2009">Komatsu &amp; Komai 2009: 598</bibRefCitation>
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, fig.
<specimenCount id="9D66FD496A0EB85D819DFC925B48FCAE" box="[739,771,780,804]" pageId="9" pageNumber="532" type="generic">3F.</specimenCount>
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</paragraph>
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<emphasis id="B914EAD26A0EB85D83E9FCCF59CAFCE1" bold="true" box="[151,385,849,875]" pageId="9" pageNumber="532">Material examined.</emphasis>
<collectionCode id="ED71AE056A0EB85D82F9FCCC5980FCE7" box="[391,459,850,877]" pageId="9" pageNumber="532">MFM</collectionCode>
145610,
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145611 and five other referred specimens.
</paragraph>
</subSubSection>
<subSubSection id="C37A654B6A0EB85D83B9FCE85C58FC76" pageId="9" pageNumber="532" type="discussion">
<paragraph id="8BDF36C06A0EB85D83B9FCE85C58FC76" blockId="9.[151,1437,849,1020]" pageId="9" pageNumber="532">
<emphasis id="B914EAD26A0EB85D83B9FCE85970FC1A" bold="true" box="[199,315,886,912]" pageId="9" pageNumber="532">Remarks.</emphasis>
Although MFM145610 lacks the rostrum and six other specimens are chelipeds, their features match those of
<taxonomicName id="4C604D436A0EB85D8387FC045A3EFC3E" authority="(Ortmann, 1892)" baseAuthorityName="Ortmann" baseAuthorityYear="1892" box="[249,629,922,948]" class="Malacostraca" family="Cyclodorippidae" genus="Tymolus" kingdom="Animalia" order="Decapoda" pageId="9" pageNumber="532" phylum="Arthropoda" rank="species" species="uncifer">
<emphasis id="B914EAD26A0EB85D8387FC0459E6FC3E" box="[249,429,922,948]" italics="true" pageId="9" pageNumber="532">Tymolus uncifer</emphasis>
(
<bibRefCitation id="EFF14B316A0EB85D82C3FC045A26FC3E" author="Ortmann, A. E." box="[445,621,922,948]" pageId="9" pageNumber="532" pagination="532 - 588" refId="ref11141" refString="Ortmann, A. E. (1892) Die Decapoden-Krebse des Strassburger Museums, mit besonderer Berucksichtigung der von Herrn Dr. Doderlein bei Japan und bei den Liu-Kiu-Inseln gesammelten und zur Zeit im Strassburger Museum aufbewahrten Formen. V Theil. Die Abtheilungen Hippidea, Dromiidea und Oxystomata. Zoologische Jahrbucher. Abteilung fur Systematik, Geographie und Biologie der Thiere, 6, 532 - 588. https: // doi. org / 10.5962 / bhl. part. 26456" type="journal article" year="1892">Ortmann, 1892</bibRefCitation>
)
</taxonomicName>
. The fossil specimens are known from the Lower Pleistocene Soga and Hijikata formations of Kakegawa Group (Karasawa 1993, 1997;
<bibRefCitation id="EFF14B316A0EB85D80FFFC205CD2FC52" author="Kitamura, K. &amp; Shiba, M." box="[897,1177,958,984]" pageId="9" pageNumber="532" pagination="11 - 18" refId="ref9683" refString="Kitamura, K. &amp; Shiba, M. (2008) Fossil Decapod Crustaceans from the Hijikata Formation of the Kakegawa Group, distributed in Nagaya, Kakegawa City, Shizuoka Prefecture, Central Japan. Science Reports of The Museum, Tokai University, 9, 11 - 18." type="journal article" year="2008">Kitamura &amp; Shiba 2008</bibRefCitation>
). The extant species is known around Japanese coasts (e.g.,
<bibRefCitation id="EFF14B316A0EB85D814BFC7C5A8CFC76" author="Takeda, M." box="[565,711,994,1020]" pageId="9" pageNumber="532" pagination="217 - 262" refId="ref12381" refString="Takeda, M. (2001) Annotated List of Crabs from Tosa Bay, Southwest Japan, Collected by the R / V / Kotaka Maru during the Years 1997 - 2000. National Science Museum Monographs, 20, 217 - 262." type="journal article" year="2001">Takeda 2001</bibRefCitation>
;
<bibRefCitation id="EFF14B316A0EB85D81ADFC7C5C43FC76" author="Spiridonov, V. A. &amp; Turkay, M." box="[723,1032,994,1020]" pageId="9" pageNumber="532" pagination="1 - 9" refId="ref12085" refString="Spiridonov, V. A. &amp; Turkay, M. (2007) Podotreme crabs from the western India Ocean: description of a new species of Tymolus Stimpson, 1858 with a review of regional distribution of the Cyclodorippidae (Crustacea: Decapoda: Brachyura: Podotremata). Arthropoda Selecta, 16, 1 - 9." type="journal article" year="2007">Spiridonov &amp; Türkay 2007</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>