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<mods:title id="6BD47D32506694E0D9A901640B008A49">Fossil decapods from the Upper Quaternary in Shinjima Island in Kagoshima Kyushu, Japan, and description of a new species of ghost shrimp (Axiidea Eucalliacidae)</mods:title>
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<mods:affiliation id="A9F77A46661A1CE9B4C3DDA496184CA3">Graduate School of Environmental Science, Hokkaido University, Kita-ku, Sapporo City, Hokkaido 060 - 0810, Japan. skameyama @ ees. hokudai. ac. jp; https: // orcid. org / 0000 - 0002 - 7104 - 1841</mods:affiliation>
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<treatment id="03C987D66A04B85083E9F9C45C8EFDF3" ID-DOI="http://doi.org/10.5281/zenodo.4574166" ID-GBIF-Taxon="180323374" ID-Zenodo-Dep="4574166" LSID="urn:lsid:plazi:treatment:03C987D66A04B85083E9F9C45C8EFDF3" httpUri="http://treatment.plazi.org/id/03C987D66A04B85083E9F9C45C8EFDF3" lastPageId="4" lastPageNumber="527" pageId="3" pageNumber="526">
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<taxonomicName id="4C604D436A04B85783E9F9C45AB1F9FE" authority="(Karasawa &amp; Goda, 1996)" baseAuthorityName="Karasawa &amp; Goda" baseAuthorityYear="1996" box="[151,762,1626,1653]" class="Malacostraca" family="Callianassidae" genus="Laticallichirus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B914EAD26A04B85783E9F9C45AB1F9FE" bold="true" box="[151,762,1626,1653]" pageId="3" pageNumber="526">
<emphasis id="B914EAD26A04B85783E9F9C459E0F9FF" bold="true" box="[151,427,1626,1653]" italics="true" pageId="3" pageNumber="526">Laticallichirus grandis</emphasis>
(
<bibRefCitation id="EFF14B316A04B85782C5F9C45AB9F9FE" author="Karasawa, H. &amp; Goda, T." box="[443,754,1626,1652]" pageId="3" pageNumber="526" pagination="1 - 4" refId="ref9143" refString="Karasawa, H. &amp; Goda, T. (1996) Two species of decapod crustaceans from the Middle Pleistocene Atsumi Group, Japan. Scientific Report of the Toyohashi Museum of Natural History, 6, 1 - 4." type="journal article" year="1996">Karasawa &amp; Goda, 1996</bibRefCitation>
)
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<paragraph id="8BDF36C06A04B85783E9F9E15968F910" blockId="3.[151,762,1626,1690]" box="[151,291,1663,1690]" pageId="3" pageNumber="526">
(
<figureCitation id="135B2A456A04B85783E1F9E158A7F910" box="[159,236,1663,1690]" captionStart="FIGURE 6" captionStartId="13.[151,250,1499,1523]" captionTargetBox="[151,1436,344,1476]" captionTargetId="figure-85@13.[151,1436,344,1476]" captionTargetPageId="13" captionText="FIGURE 6. Decapods from the Moeshima Shell Bed. AJ, Laticallichirus grandis (Karasawa &amp; Goda, 1996); A, 145597, lateral view of left major cheliped; B, MFM145598, lateral view of right major cheliped; C, MFM145599, lateral view of right carpus; D, MFM145600, lateral view of left minor cheliped; E, MFM145601, lateral view of right dactylus; F, MFM145602, lateral view of right dactylus; G, MFM145603, lateral view of right dactylus; H, MFM145604, lateral view of left dactylus; I, MFM145605, lateral view of right merus; J, MFM145606, lateral view of left merus. K, Paguridae gen. et sp. indet., MFM145607, lateral view of fragment of the fixed finger. L, Pagurus cf. megalops (Stimpson, 1858b), MFM 145608, lateral view of left cheliped. M, Ebalia tuberculosa (A. Milne-Edwards, 1873), MFM145612, dorsal view of carapace. N, Myra sp., MFM145613, lateral view of fragment of the merus. O, Philyra cf. syndactyla Ortmann, 1892, MFM145614, lateral view of right cheliped. P, Majidae gen. et sp. indet., MFM145615, dorsal view of fragment of the carapace. Q, Leptomithrax sp., MFM 145616, lateral view of left cheliped. R, Romaleon sp., MFM 145617, lateral view of fragment of the dactylus. S, Charibdis sp. MFM145619, lateral view of fragment of the fixed finger. Scale bars of AH, K, M, O, Q and S indicate 5.0 mm, and I, J, L, N, P and R indicate 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.4425396" httpUri="https://zenodo.org/record/4425396/files/figure.png" pageId="3" pageNumber="526">Figs. 6</figureCitation>
AJ)
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<taxonomicName id="4C604D436A04B85783E9F95D595BF951" box="[151,272,1731,1755]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="sp.">
<emphasis id="B914EAD26A04B85783E9F95D595BF951" box="[151,272,1731,1755]" italics="true" pageId="3" pageNumber="526">Callianassa</emphasis>
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sp.
<typeStatus id="54DB88626A04B8578244F95A592DF956" box="[314,358,1732,1756]" pageId="3" pageNumber="526">type</typeStatus>
B.—Tokai Fossil Society (Ed.) 1977: 60, pl. 27, fig. 6.
<taxonomicName id="4C604D436A04B85783E9F97C595BF970" authority="Kato &amp; Koizumi 1992" box="[151,272,1762,1786]" class="Malacostraca" family="Callianassidae" genus="Callianassa" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="sp.">
<emphasis id="B914EAD26A04B85783E9F97C595BF970" box="[151,272,1762,1786]" italics="true" pageId="3" pageNumber="526">Callianassa</emphasis>
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sp.—
<treatmentCitation id="0AC110D16A04B8578235F97C5A12F970" author="Kato, H. &amp; Koizumi, A." box="[331,601,1762,1786]" page="49" pageId="3" pageNumber="526" year="1992">
<bibRefCitation id="EFF14B316A04B8578235F97C5A12F970" author="Kato, H. &amp; Koizumi, A." box="[331,601,1762,1786]" pageId="3" pageNumber="526" pagination="45 - 53" refId="ref9639" refString="Kato, H. &amp; Koizumi, A. (1992) Decapod fossils from the Pleistocene Shimosueyoshi Formation in the northern part of Yokoyama City. Bulletin of the Kanagawa Prefectural Museum (Natural Science), 21, 45 - 53." type="journal article" year="1992">Kato &amp; Koizumi 1992: 49</bibRefCitation>
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, fig. 3-3.
<taxonomicName id="4C604D436A04B85783E9F89E58A9F892" authorityName="de Saint Laurent" authorityYear="1973" box="[151,226,1792,1816]" class="Malacostraca" family="Callianassidae" genus="Calliax" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="526" phylum="Arthropoda" rank="genus">
<emphasis id="B914EAD26A04B85783E9F89E58A9F892" box="[151,226,1792,1816]" italics="true" pageId="3" pageNumber="526">Calliax</emphasis>
</taxonomicName>
sp.—
<treatmentCitation id="0AC110D16A04B8578262F89F5A07F893" author="Karasawa, H. &amp; Tanaka, T." box="[284,588,1792,1817]" page="12" pageId="3" pageNumber="526" year="1994">
<bibRefCitation id="EFF14B316A04B8578262F89F5A07F893" author="Karasawa, H. &amp; Tanaka, T." box="[284,588,1792,1817]" pageId="3" pageNumber="526" pagination="11 - 19" refId="ref9377" refString="Karasawa, H. &amp; Tanaka, T. (1994) Decapod Crustacea from the Atsumi Group (Middle Pleistocene) of Aichi Prefecture, Central Japan. Scientific Report of the Toyohashi Museum of Natural History, 4, 11 - 19." type="journal article" year="1994">Karasawa &amp; Tanaka 1994: 12</bibRefCitation>
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, figs. 2-112.
<taxonomicName id="4C604D436A04B85783E9F8815967F8BD" authorityName="K.Sakai" authorityYear="1988" box="[151,300,1823,1847]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="sp.">
<emphasis id="B914EAD26A04B85783E9F8815967F8BD" box="[151,300,1823,1847]" italics="true" pageId="3" pageNumber="526">Neocallichirus</emphasis>
</taxonomicName>
sp.—
<treatmentCitation id="0AC110D16A04B8578218F8BE5A3FF8BD" author="Karasawa, H. &amp; Nohara, T. &amp; Shimoji, K." box="[358,628,1823,1847]" page="128" pageId="3" pageNumber="526" year="1995">
<bibRefCitation id="EFF14B316A04B8578218F8BE5A3FF8BD" author="Karasawa, H. &amp; Nohara, T. &amp; Shimoji, K." box="[358,628,1823,1847]" pageId="3" pageNumber="526" pagination="127 - 132" refId="ref9334" refString="Karasawa, H. Nohara, T. &amp; Shimoji, K. (1995) Decapod Crustacea from the Ryukyu Group (Pleistocene) of Okinawa-jima, Japan. Bulletin of the Mizunami Fossil Museum, 22, 127 - 132." type="journal article" year="1995">
Karasawa
<emphasis id="B914EAD26A04B85782ADF8BE5A4BF8BD" box="[467,512,1823,1847]" italics="true" pageId="3" pageNumber="526">et al</emphasis>
. 1995: 128
</bibRefCitation>
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, fig. 2. “
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<emphasis id="B914EAD26A04B85783DCF8A0597CF8DC" box="[162,311,1854,1878]" italics="true" pageId="3" pageNumber="526">Neocallichirus</emphasis>
<treatmentCitation id="0AC110D16A04B8578237F8A05AF9F8DC" author="Karasawa, H. &amp; Goda, T." box="[329,690,1854,1878]" page="1" pageId="3" pageNumber="526" year="1996">
<emphasis id="B914EAD26A04B8578237F8A059DDF8DC" box="[329,406,1854,1878]" italics="true" pageId="3" pageNumber="526">grandis</emphasis>
<bibRefCitation id="EFF14B316A04B85782E3F8A05AF9F8DC" author="Karasawa, H. &amp; Goda, T." box="[413,690,1854,1878]" pageId="3" pageNumber="526" pagination="1 - 4" refId="ref9143" refString="Karasawa, H. &amp; Goda, T. (1996) Two species of decapod crustaceans from the Middle Pleistocene Atsumi Group, Japan. Scientific Report of the Toyohashi Museum of Natural History, 6, 1 - 4." type="journal article" year="1996">Karasawa &amp; Goda, 1996: 1</bibRefCitation>
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</taxonomicName>
, fig. 1.—
<treatmentCitation id="0AC110D16A04B8578073F8A05B93F8DC" author="Karasawa, H." box="[781,984,1854,1878]" page="33" pageId="3" pageNumber="526" year="1997">
<bibRefCitation id="EFF14B316A04B8578073F8A05B93F8DC" author="Karasawa, H." box="[781,984,1854,1878]" pageId="3" pageNumber="526" pagination="1 - 81" refId="ref9107" refString="Karasawa, H. (1997) A monograph of Cenozoic stomatopod, decapod, isopod and amphipod Crustacea from West Japan. Monograph of the Mizunami Fossil Museum, 8, 1 - 81." type="journal article" year="1997">Karasawa 1997: 33</bibRefCitation>
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, pl. 5, figs. 14.
<taxonomicName id="4C604D436A04B85783E9F8C259CBF8FE" authorityName="Karasawa &amp; Goda" authorityYear="1996" box="[151,384,1884,1908]" class="Malacostraca" family="Callianassidae" genus="Neocallichirus" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="526" phylum="Arthropoda" rank="species" species="grandis">
<emphasis id="B914EAD26A04B85783E9F8C259CBF8FE" box="[151,384,1884,1908]" italics="true" pageId="3" pageNumber="526">Neocallichirus grandis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC110D16A04B85782E0F8C35AE1F8FE" author="Kato, H. &amp; Karasawa, H." box="[414,682,1884,1909]" page="5" pageId="3" pageNumber="526" year="1998">
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</treatmentCitation>
, pl. 2, figs. 716. —
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<bibRefCitation id="EFF14B316A04B8578005F8C35C47F8FE" author="Kato, H." box="[891,1036,1884,1909]" pageId="3" pageNumber="526" pagination="37 - 48" refId="ref9491" refString="Kato, H. (2001) Fossil decapod assemblages from the Pleistocene Kiyokawa and Kioroshi Formations, Shimosa Group, central Japan. Journal of the Natural History Museum and Institute, Chiba, Special Issue 4, 37 - 48." type="journal article" year="2001">Kato 2001: 40</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0AC110D16A04B8578757F8C25D08F8FE" author="Schweitzer, C. E. &amp; Iturralde-Vinent, M. &amp; Hetler, J. L. &amp; Velez-Juarbe, J." box="[1065,1347,1884,1908]" page="115" pageId="3" pageNumber="526" year="2006">
<bibRefCitation id="EFF14B316A04B8578757F8C25D08F8FE" author="Schweitzer, C. E. &amp; Iturralde-Vinent, M. &amp; Hetler, J. L. &amp; Velez-Juarbe, J." box="[1065,1347,1884,1908]" pageId="3" pageNumber="526" pagination="111 - 136" refId="ref11967" refString="Schweitzer, C. E., Iturralde-Vinent, M., Hetler, J. L. &amp; Velez-Juarbe, J. (2006) Oligocene and Miocene decapods (Thalassinidea &amp; Brachyura) from the Caribbean. Annals of Carnegie Museum, 75, 111 - 136. https: // doi. org / 10.2992 / 0097 - 4463 (2006) 75 [111: OAMDTA] 2.0. CO; 2" type="journal article" year="2006">
Schweitzer
<emphasis id="B914EAD26A04B85787DFF8C35C85F8FE" box="[1185,1230,1884,1908]" italics="true" pageId="3" pageNumber="526">et al</emphasis>
. 2006: 115
</bibRefCitation>
</treatmentCitation>
.
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<emphasis id="B914EAD26A04B85783E9F8E25927F819" box="[151,364,1915,1939]" italics="true" pageId="3" pageNumber="526">Grynaminna grandis</emphasis>
</taxonomicName>
.—
<treatmentCitation id="0AC110D16A04B85782F4F8E55AE9F819" author="Obata, K. &amp; Hayashi, Y." box="[394,674,1915,1939]" page="46" pageId="3" pageNumber="526" year="2001">
<bibRefCitation id="EFF14B316A04B85782F4F8E55AE9F819" author="Obata, K. &amp; Hayashi, Y." box="[394,674,1915,1939]" pageId="3" pageNumber="526" pagination="45 - 52" refId="ref11032" refString="Obata, K. &amp; Hayashi, Y. (2001) Decapod Crustacea from the upper Pleistocene Kioroshi Formation in Showa-machi, Saitama Prefecture, central Japan. Bulletin of the Saitama Museum of Natural History, 19, 45 - 52." type="journal article" year="2001">Obata &amp; Hayashi 2001: 46</bibRefCitation>
</treatmentCitation>
, figs. 3.112.—
<treatmentCitation id="0AC110D16A04B857803AF8E55BA2F819" author="Kato, H." box="[836,1001,1915,1939]" page="159" pageId="3" pageNumber="526" year="2003">
<bibRefCitation id="EFF14B316A04B857803AF8E55BA2F819" author="Kato, H." box="[836,1001,1915,1939]" pageId="3" pageNumber="526" pagination="157 - 160" refId="ref9534" refString="Kato, H. (2003) Cenozoic decapod crustacean assemblages from northeast Japan. Contributions to Zoology, 72, 157 - 160. https: // doi. org / 10.1163 / 18759866 - 0720203018" type="journal article" year="2003">Kato 2003: 159</bibRefCitation>
</treatmentCitation>
. —
<treatmentCitation id="0AC110D16A04B8578770F8E55D70F819" author="Hyzny, M. &amp; Karasawa, H." box="[1038,1339,1915,1939]" page="65" pageId="3" pageNumber="526" year="2012">
<bibRefCitation id="EFF14B316A04B8578770F8E55D70F819" author="Hyzny, M. &amp; Karasawa, H." box="[1038,1339,1915,1939]" pageId="3" pageNumber="526" pagination="59 - 68" refId="ref8633" refString="Hyzny, M. &amp; Karasawa, H. (2012) How to distinguish Neocallichirus, Sergio, Podocallichirus and Grynaminna (Decapoda: Callianassidae: Callichirinae) from each other in the fossil record? Bulletin of the Mizunami Fossil Museum, 38, 59 - 68." type="journal article" year="2012">Hyžný &amp; Karasawa 2012: 65</bibRefCitation>
</treatmentCitation>
, fig. 1A
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDF36C06A04B85783B9F8045DD6F838" blockId="3.[151,1437,1731,2001]" box="[199,1437,1946,1970]" pageId="3" pageNumber="526">
H.—
<bibRefCitation id="EFF14B316A04B8578389F80459B2F838" author="Karasawa, H. &amp; Kobayashi, N. &amp; Goda, T. &amp; Ohira, N. &amp; Ando, Y." box="[247,505,1946,1970]" pageId="3" pageNumber="526" pagination="55 - 73" refId="ref9185" refString="Karasawa, H., Kobayashi, N., Goda, T., Ohira, N. &amp; Ando, Y. (2014) A diversity for crabs (Decapoda) from the middle Pleistocene Atsumi Group, Japan. Bulletin of the Mizunami Fossil Museum, 40, 55 - 73." type="journal article" year="2014">
Karasawa
<emphasis id="B914EAD26A04B857821DF80559DBF838" box="[355,400,1946,1970]" italics="true" pageId="3" pageNumber="526">et al</emphasis>
. 2014: 59
</bibRefCitation>
, fig. 5.15.3.—
<bibRefCitation id="EFF14B316A04B85781E8F8045BF4F838" author="Hyzny, M. &amp; Klompmaker, A." box="[662,959,1946,1970]" pageId="3" pageNumber="526" pagination="401 - 437" refId="ref8683" refString="Hyzny, M. &amp; Klompmaker, A. (2015) Systematics, phylogeny, and taphonomy of ghost shrimp (Decapoda): a perspective from the fossil record. Arthropod systematics &amp; Phylogeny, 73, 401 - 437." type="journal article" year="2015">Hyžný &amp; Klompmaker 2015</bibRefCitation>
: fig. 7A, 412 (Table 1).—
<bibRefCitation id="EFF14B316A04B85787C1F8045DD3F838" author="Ando, Y. &amp; Kawano, S. &amp; Komatsu, T. &amp; Niitani, M." box="[1215,1432,1946,1970]" pageId="3" pageNumber="526" pagination="16 - 25" refId="ref7819" refString="Ando, Y., Kawano, S., Komatsu, T. &amp; Niitani, M. (2016) Decapod crustaceans from the Pleistocene Oe Formation in Minamishimabara City, Nagasaki Prefecture, Japan. Journal of Fossil Research, 48, 16 - 25." type="journal article" year="2016">
Ando
<emphasis id="B914EAD26A04B857867DF8055D7BF838" box="[1283,1328,1946,1970]" italics="true" pageId="3" pageNumber="526">et al</emphasis>
. 2016: 18
</bibRefCitation>
,
</paragraph>
</subSubSection>
<subSubSection id="C37A654B6A04B85783B9F8265B4AF85B" box="[199,769,1976,2001]" pageId="3" pageNumber="526" type="description">
<paragraph id="8BDF36C06A04B85783B9F8265B4AF85B" blockId="3.[151,1437,1731,2001]" box="[199,769,1976,2001]" pageId="3" pageNumber="526">
figs. 3.310.—
<bibRefCitation id="EFF14B316A04B8578223F8265A25F85A" author="Ando, Y. &amp; Kawano, S." box="[349,622,1976,2001]" pageId="3" pageNumber="526" pagination="83 - 92" refId="ref7779" refString="Ando, Y. &amp; Kawano, S. (2017) Decapods from the lower Pleistocene Masuda Formation in Minamitane-cho, Kagoshima Prefecture, Japan. Bulletin of the Mizunami Fossil Museum, 43, 83 - 92." type="journal article" year="2017">Ando &amp; Kawano 2017: 85</bibRefCitation>
, fig. 2.92.20.
</paragraph>
</subSubSection>
<subSubSection id="C37A654B6A03B85083E9FF095C70FF66" pageId="4" pageNumber="527" type="reference_group">
<paragraph id="8BDF36C06A03B85083E9FF0958ACFF47" blockId="4.[151,1437,151,236]" pageId="4" pageNumber="527">
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<emphasis id="B914EAD26A03B85083E9FF0959C6FF25" box="[151,397,151,175]" italics="true" pageId="4" pageNumber="527">Podocallichirus grandis</emphasis>
</taxonomicName>
.—
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<bibRefCitation id="EFF14B316A03B85082D5FF095AF7FF25" author="Karasawa, H. &amp; Tanaka, T. &amp; Kobayashi, N. &amp; Goda, T. &amp; Ohira, N. &amp; Shinya, J." box="[427,700,151,175]" pageId="4" pageNumber="527" pagination="127 - 133" refId="ref9422" refString="Karasawa, H., Tanaka, T., Kobayashi, N., Goda, T., Ohira, N. &amp; Shinya, J. (2006) Podocallichirus grandis (Crustacea: Decapoda: Thalassinidae) preserved within burrows from the middle Pleistocene Atsumi Group of Aichi Prefecture, Japan. Bulletin of the Mizunami Fossil Museum, 33, 127 - 133." type="journal article" year="2006">
Karasawa
<emphasis id="B914EAD26A03B8508166FF095A0DFF25" box="[536,582,151,175]" italics="true" pageId="4" pageNumber="527">et al</emphasis>
. 2006: 127
</bibRefCitation>
</treatmentCitation>
, pl. 2, figs. 16.—
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Kobayashi
<emphasis id="B914EAD26A03B850808EFF095C55FF25" box="[1008,1054,151,175]" italics="true" pageId="4" pageNumber="527">et al</emphasis>
. 2008: 111
</bibRefCitation>
</treatmentCitation>
.—
<treatmentCitation id="0AC110D16A03B85087D2FF0958A8FF47" author="Schweitzer, C. E. &amp; Feldmann, R. M. &amp; Garassino, A. &amp; Karasawa, H. &amp; Schweigert, G." page="39" pageId="4" pageNumber="527" year="2010">
<bibRefCitation id="EFF14B316A03B85087D2FF0958A8FF47" author="Schweitzer, C. E. &amp; Feldmann, R. M. &amp; Garassino, A. &amp; Karasawa, H. &amp; Schweigert, G." pageId="4" pageNumber="527" refId="ref11894" refString="Schweitzer, C. E., Feldmann, R. M., Garassino, A., Karasawa, H. &amp; Schweigert, G. (2010) Systematic list of fossil decapod crustacean species. In: Crustaceana Monographs. Vol. 10. Brill, Leiden, pp. 1 - 22. https: // doi. org / 10.1163 / ej. 9789004178915. i- 222" type="book" year="2010">
Schweitzer
<emphasis id="B914EAD26A03B8508656FF095D1EFF25" box="[1320,1365,151,175]" italics="true" pageId="4" pageNumber="527">et al</emphasis>
. 2010: 39
</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BDF36C06A03B85083E9FF4A5C70FF66" blockId="4.[151,1437,151,236]" box="[151,1083,212,236]" pageId="4" pageNumber="527">
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<emphasis id="B914EAD26A03B85083E9FF4A59CBFF66" box="[151,384,212,236]" italics="true" pageId="4" pageNumber="527">Laticallichirus grandis</emphasis>
</taxonomicName>
.—
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<bibRefCitation id="EFF14B316A03B85082E0FF4A5A8BFF66" author="Komai, T. &amp; Yokooka, H. &amp; Henmi, Y. &amp; Itani, G." box="[414,704,212,236]" pageId="4" pageNumber="527" pagination="461 - 481" refId="ref9843" refString="Komai, T., Yokooka, H., Henmi, Y. &amp; Itani, G. (2019) A new genus for &quot; Neocallichirus &quot; grandis Karasawa &amp; Goda, 1996, a ghost shrimp species (Decapoda: Axiidea: Callianassidae) heretofore known only by fossil materials. Zootaxa, 4604 (3), 461 - 481. https: // doi. org / 10.11646 / zootaxa. 4604.3.4" type="journal article" year="2019">
Komai
<emphasis id="B914EAD26A03B8508295FF4B5A53FF66" box="[491,536,212,236]" italics="true" pageId="4" pageNumber="527">et al</emphasis>
. 2019: 466477
</bibRefCitation>
</treatmentCitation>
, figs. 16.—
<treatmentCitation id="0AC110D16A03B8508042FF4A5BBEFF66" author="Mimoto, K." box="[828,1013,212,236]" page="51" pageId="4" pageNumber="527" year="2020">
<bibRefCitation id="EFF14B316A03B8508042FF4A5BBEFF66" author="Mimoto, K." box="[828,1013,212,236]" pageId="4" pageNumber="527" pagination="50 - 52" refId="ref10676" refString="Mimoto, K. (2020) Laticallichirus grandis (Karasawa &amp; Goda, 1996) (Decapoda: Axiidea: Callianassidae) from the Pleistocene Ashizuri Formation of Kochi Prefecture, Shikoku, Japan. Kaseki-no-tomo (Publication of the Tokai Fossil Society), 65, 50 - 52." type="journal article" year="2020">Mimoto 2020: 51</bibRefCitation>
</treatmentCitation>
, fig. 3.
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</paragraph>
</subSubSection>
<subSubSection id="C37A654B6A03B85083E9FE845C36FEBE" box="[151,1149,282,309]" pageId="4" pageNumber="527" type="materials_examined">
<paragraph id="8BDF36C06A03B85083E9FE845C36FEBE" blockId="4.[151,1437,282,633]" box="[151,1149,282,309]" pageId="4" pageNumber="527">
<emphasis id="B914EAD26A03B85083E9FE8459CAFEBE" bold="true" box="[151,385,282,308]" pageId="4" pageNumber="527">Material examined.</emphasis>
<collectionCode id="ED71AE056A03B85082F9FE845981FEBF" box="[391,458,282,309]" pageId="4" pageNumber="527">MFM</collectionCode>
145597
<collectionCode id="ED71AE056A03B8508155FE845A25FEBF" box="[555,622,282,309]" pageId="4" pageNumber="527">MFM</collectionCode>
145606 and over 600 other referred specimens.
</paragraph>
</subSubSection>
<subSubSection id="C37A654B6A03B85083B9FEA05C8EFDF3" pageId="4" pageNumber="527" type="discussion">
<paragraph id="8BDF36C06A03B85083B9FEA05C8EFDF3" blockId="4.[151,1437,282,633]" pageId="4" pageNumber="527">
<emphasis id="B914EAD26A03B85083B9FEA05970FED2" bold="true" box="[199,315,318,344]" pageId="4" pageNumber="527">Remarks.</emphasis>
The present species has been described from the Lower Pleistocene to Recent in the Kyushu to Kanto district,
<collectingCountry id="F37776506A03B850838DFEFD5978FEF7" box="[243,307,355,381]" name="Japan" pageId="4" pageNumber="527">Japan</collectingCountry>
(e.g.,
<bibRefCitation id="EFF14B316A03B8508205FEFD5AC6FEF6" author="Karasawa, H. &amp; Goda, T." box="[379,653,354,381]" pageId="4" pageNumber="527" pagination="1 - 4" refId="ref9143" refString="Karasawa, H. &amp; Goda, T. (1996) Two species of decapod crustaceans from the Middle Pleistocene Atsumi Group, Japan. Scientific Report of the Toyohashi Museum of Natural History, 6, 1 - 4." type="journal article" year="1996">Karasawa &amp; Goda 1996</bibRefCitation>
;
<bibRefCitation id="EFF14B316A03B85081E4FEFD5BEDFEF6" author="Kato, H. &amp; Karasawa, H." box="[666,934,354,381]" pageId="4" pageNumber="527" pagination="1 - 31" refId="ref9571" refString="Kato, H. &amp; Karasawa, H. (1998) Pleistocene fossil decapod Crustacea from the Boso Peninsula, Japan. In: Yonebayashi, C., Yagi, R. &amp; Harada, H. (Eds.), The Fossil Record in the Boso Peninsula, Japan. Natural History Research, Special Number 5, pp. 1 - 31." type="book chapter" year="1998">Kato &amp; Karasawa 1998</bibRefCitation>
;
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;
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Komai
<emphasis id="B914EAD26A03B850866AFEFD5D0DFEF6" box="[1300,1350,354,380]" italics="true" pageId="4" pageNumber="527">et al</emphasis>
. 2019
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).
<bibRefCitation id="EFF14B316A03B85083E9FE185937FE2A" author="Komai, T. &amp; Yokooka, H. &amp; Henmi, Y. &amp; Itani, G." box="[151,380,390,417]" pageId="4" pageNumber="527" pagination="461 - 481" refId="ref9843" refString="Komai, T., Yokooka, H., Henmi, Y. &amp; Itani, G. (2019) A new genus for &quot; Neocallichirus &quot; grandis Karasawa &amp; Goda, 1996, a ghost shrimp species (Decapoda: Axiidea: Callianassidae) heretofore known only by fossil materials. Zootaxa, 4604 (3), 461 - 481. https: // doi. org / 10.11646 / zootaxa. 4604.3.4" type="journal article" year="2019">
Komai
<emphasis id="B914EAD26A03B8508393FE19596AFE2A" box="[237,289,390,416]" italics="true" pageId="4" pageNumber="527">et al</emphasis>
. (2019)
</bibRefCitation>
collected the specimens of extant
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<emphasis id="B914EAD26A03B8508073FE185C59FE2A" box="[781,1042,390,416]" italics="true" pageId="4" pageNumber="527">Laticallichirus grandis</emphasis>
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from the muddy-sand mixed with gravel bottom of subtidal zone, and
<bibRefCitation id="EFF14B316A03B8508151FE345B08FE4E" author="Ando, Y. &amp; Kawano, S." box="[559,835,426,453]" pageId="4" pageNumber="527" pagination="83 - 92" refId="ref7779" refString="Ando, Y. &amp; Kawano, S. (2017) Decapods from the lower Pleistocene Masuda Formation in Minamitane-cho, Kagoshima Prefecture, Japan. Bulletin of the Mizunami Fossil Museum, 43, 83 - 92." type="journal article" year="2017">Ando &amp; Kawano (2017)</bibRefCitation>
showed that the species was living under the coarsegrained sandy and gravelly bottoms mixed with shells of the upper sublittoral zone. The preservation of the present specimens is similar to other described specimens that seemed to be living near the sedimentary area (
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Ando
<emphasis id="B914EAD26A03B850861DFE6D5DDEFD86" box="[1379,1429,498,524]" italics="true" pageId="4" pageNumber="527">et al</emphasis>
. 2016
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;
<bibRefCitation id="EFF14B316A03B85083A3FD8859A9FDBA" author="Ando, Y. &amp; Kawano, S." box="[221,482,534,561]" pageId="4" pageNumber="527" pagination="83 - 92" refId="ref7779" refString="Ando, Y. &amp; Kawano, S. (2017) Decapods from the lower Pleistocene Masuda Formation in Minamitane-cho, Kagoshima Prefecture, Japan. Bulletin of the Mizunami Fossil Museum, 43, 83 - 92." type="journal article" year="2017">Ando &amp; Kawano 2017</bibRefCitation>
). Therefore, the present specimens were not, in all likelihood, transported from other localities and shallower environment, and the present record suggests that the species has been living under coarsegrained sandy and gravelly bottoms mixed with shells of the lower sublittoral to subtidal zones.
</paragraph>
</subSubSection>
</treatment>
</document>