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95 lines
13 KiB
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<document id="803D353C8B7F6E1711BABECC8D30A48C" ID-DOI="10.11646/zootaxa.3973.2.10" ID-GBIF-Dataset="0b0db3fd-d54b-432b-a661-408f1ac32abc" ID-ISSN="1175-5326" ID-Zenodo-Dep="245230" ID-ZooBank="4E6E002C-309C-47EB-964C-BF5A06FEC62E" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461209209336" checkinUser="plazi" docAuthor="Landim, Maria Isabel, Moreira, Cristiano R. & Figueiredo, Carlos A." docDate="2015" docId="03C987AF3D70FF9E0DBF1BE3FA4BF80A" docLanguage="en" docName="zt03973p380.pdf" docOrigin="Zootaxa 3973 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Retroculus Eigenmann & Bray 1894" docType="treatment" docVersion="7" lastPageNumber="370" masterDocId="FFF0FFD73D71FF9F0D281E29FFC9FFD0" masterDocTitle="Retroculus acherontos, a new species of cichlid fish (Teleostei) from the Rio Tocantins basin" masterLastPageNumber="380" masterPageNumber="369" pageNumber="370" updateTime="1698601664754" updateUser="plazi">
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<mods:mods id="F7FBF043A4304EE83C583D75FC4B6EE5" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="53DA03A60FDC940F4CBCD6C69C6D0C9D">
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<mods:title id="B3262E6BCC54AFA2CACC68D21EE13531">Retroculus acherontos, a new species of cichlid fish (Teleostei) from the Rio Tocantins basin</mods:title>
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<mods:name id="4689049EAB25AA6FD3A6D3212C2E1F9E" type="personal">
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<mods:roleTerm id="4E67FD18DA19F10E347EF9285F55860E">Author</mods:roleTerm>
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<mods:namePart id="9031AABAAB0B91B528A18BA59934E5DF">Landim, Maria Isabel</mods:namePart>
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<mods:namePart id="00AB80BA227751F09E763D0F884DDA6A">Moreira, Cristiano R.</mods:namePart>
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<mods:name id="7E4AC61A9345D7D06B9A6AA8D0162C39" type="personal">
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<mods:namePart id="6608C4EBFE13F16BCE6DB3E0F63E95A7">Figueiredo, Carlos A.</mods:namePart>
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<mods:typeOfResource id="022C01D40519C9DA4AB958C0286E7427">text</mods:typeOfResource>
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<mods:title id="17AE0EB4F9BEB90F6EB25D3A1D8875AE">Zootaxa</mods:title>
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<mods:part id="AD2334CA3385C91C849ECEF448A6AFB2">
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<mods:date id="F1CD19C62EA4ED4F6FC19F6AE3D311C1">2015</mods:date>
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<mods:detail id="E5575F9AE95B712EFA1D1F34AED69FE3" type="volume">
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<mods:number id="1F66ED872D39988D228941CED232CFED">3973</mods:number>
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<mods:detail id="A992389BA5925DDC1BB09E6B5D7C596E" type="issue">
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<mods:classification id="48E26B39973E9E897ABBC036FD7C43ED">journal article</mods:classification>
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<mods:identifier id="B6089635EADA7FBC8328134001146E2F" type="DOI">10.11646/zootaxa.3973.2.10</mods:identifier>
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<mods:identifier id="297C4E174DD8005DA4A247CE3D195CDC" type="GBIF-Dataset">0b0db3fd-d54b-432b-a661-408f1ac32abc</mods:identifier>
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<treatment id="03C987AF3D70FF9E0DBF1BE3FA4BF80A" ID-DOI="http://doi.org/10.5281/zenodo.6112967" ID-GBIF-Taxon="119630152" ID-Zenodo-Dep="6112967" LSID="urn:lsid:plazi:treatment:03C987AF3D70FF9E0DBF1BE3FA4BF80A" httpUri="http://treatment.plazi.org/id/03C987AF3D70FF9E0DBF1BE3FA4BF80A" lastPageNumber="370" pageId="1" pageNumber="370">
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<subSubSection id="C37A65323D70FF9E0DBF1BE3FD97FA35" box="[151,606,1482,1509]" pageId="1" pageNumber="370" type="nomenclature">
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<paragraph id="8BDF36B93D70FF9E0DBF1BE3FD97FA35" blockId="1.[151,606,1482,1509]" box="[151,606,1482,1509]" pageId="1" pageNumber="370">
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<heading id="D09781D53D70FF9E0DBF1BE3FD97FA35" bold="true" box="[151,606,1482,1509]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
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<taxonomicName id="4C604D3A3D70FF9E0DBF1BE3FD97FA35" ID-CoL="77X2" ID-ENA="81350" authority="Eigenmann & Bray, 1894" authorityName="Eigenmann & Bray" authorityYear="1894" box="[151,606,1482,1509]" class="Actinopterygii" family="Cichlidae" genus="Retroculus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="370" phylum="Chordata" rank="genus">
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<emphasis id="B914EAAB3D70FF9E0DBF1BE3FD97FA35" bold="true" box="[151,606,1482,1509]" pageId="1" pageNumber="370">
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<emphasis id="B914EAAB3D70FF9E0DBF1BE3FED2FA34" bold="true" box="[151,283,1482,1508]" italics="true" pageId="1" pageNumber="370">Retroculus</emphasis>
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<bibRefCitation id="EFF14B483D70FF9E0C0A1BE2FD97FA35" author="Eigenmann" box="[290,606,1483,1509]" pageId="1" pageNumber="370" refString="Eigenmann, C. H. & Bray, W. L. (1894) A revision of the American Cichlidae. Annals of the New York Academy of Science, 7, 607 - 624." type="journal article" year="1894">Eigenmann & Bray, 1894</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C37A65323D70FF9E0DBF1838FA4BF80A" pageId="1" pageNumber="370" type="diagnosis">
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<paragraph id="8BDF36B93D70FF9E0DBF1838FA4BF80A" blockId="1.[151,1437,1553,2010]" pageId="1" pageNumber="370">
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<emphasis id="B914EAAB3D70FF9E0DBF1838FED8F9FA" bold="true" box="[151,273,1553,1578]" pageId="1" pageNumber="370">Diagnosis.</emphasis>
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<taxonomicName id="4C604D3A3D70FF9E0C321838FE5BF9FA" box="[282,402,1553,1578]" class="Actinopterygii" family="Cichlidae" genus="Retroculus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="370" phylum="Chordata" rank="genus">
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<emphasis id="B914EAAB3D70FF9E0C321838FE5BF9FA" box="[282,402,1553,1578]" italics="true" pageId="1" pageNumber="370">Retroculus</emphasis>
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</taxonomicName>
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is distinguished from all other cichlid genera by the presence of two exclusive characters: epibranchial lobe supported by a thickening of the first epibranchial (vs. epibranchial lobe supported by the second epibranchial, or by a laminar expansion of the first epibranchial;
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<figureCitation id="135B2A3C3D70FF9E0EAF1870FC00F9A2" box="[903,969,1625,1650]" captionStart="FIGURE 1" captionStartId="2.[151,249,1559,1581]" captionTargetBox="[372,1201,210,1511]" captionTargetId="figure@2.[365,1229,195,1536]" captionTargetPageId="2" captionText="FIGURE 1. First epibranchial of (A) Retroculus acherontos, MZUSP 85838, (B) R. s eptentrionalis, MZUSP 38947, paratype, and (C) Satanoperca lilith, MZUSP 8503. Ventral view, anterior to top. Cartilage black. Scale bar = 1 mm." httpUri="https://zenodo.org/record/245231/files/figure.png" pageId="1" pageNumber="370">Fig. 1</figureCitation>
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); and the presence of a vomero-palatine apparatus on roof of the mouth (vs. roof of mouth not distinctively specialized;
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<figureCitation id="135B2A3C3D70FF9E09111854FB49F946" box="[1081,1152,1661,1686]" captionStart="FIGURE 2" captionStartId="3.[151,249,1412,1434]" captionTargetBox="[151,1436,1021,1390]" captionTargetId="figure@3.[151,1436,1021,1391]" captionTargetPageId="3" captionText="FIGURE 2. Vomero-Palatine apparatus of (A) Retroculus acherontos, MZUSP 85826, 161.3 mm SL, (B) R. xinguensis, MZUSP 40682, 78.2 mm SL, and (C) R. septentrionalis, paratype, MZUSP 38947, 106.2 mm SL. Ventral view, anterior to top. Arrows indicate the lateral depressions. V indicate the ventral projection of vomer." httpUri="https://zenodo.org/record/245232/files/figure.png" pageId="1" pageNumber="370">Fig. 2</figureCitation>
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).
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<taxonomicName id="4C604D3A3D70FF9E09BF1854FAC6F946" box="[1175,1295,1661,1686]" class="Actinopterygii" family="Cichlidae" genus="Retroculus" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="370" phylum="Chordata" rank="genus">
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<emphasis id="B914EAAB3D70FF9E09BF1854FAC6F946" box="[1175,1295,1661,1686]" italics="true" pageId="1" pageNumber="370">Retroculus</emphasis>
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</taxonomicName>
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can also be distinguished by a combination of the following diagnostic characters: the presence of two lachrymal plates plus five tubular infraorbitals (
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<figureCitation id="135B2A3C3D70FF9E0C9118ECFE34F90E" box="[441,509,1733,1758]" captionStart="FIGURE 3" captionStartId="3.[151,249,1945,1967]" captionTargetBox="[306,1281,1545,1923]" captionTargetId="figure@3.[306,1281,1545,1924]" captionTargetPageId="3" captionText="FIGURE 3. Lachrymals and tubular infraorbitals of Retroculus acherontos, MZUSP 85838. Lateral view, anterior to right. Scale bar = 3 mm." httpUri="https://zenodo.org/record/245233/files/figure.png" pageId="1" pageNumber="370">Fig. 3</figureCitation>
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); a first tubular infraorbital only posteriorly associated with the orbit; the lip folds meeting without overlap at the corner of the mouth (African lips of Kullander, 1986; or type-I lip of
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<bibRefCitation id="EFF14B483D70FF9E081218C0FF1AF8F6" author="Stiassny" pageId="1" pageNumber="370" refString="Stiassny, M. (1987) Cichlid familial intrarelationships and the placement of neotropical genus Cichla (Perciformes, Labroidei). Journal of Natural History, 21, 1311 - 1331." type="journal article" year="1987">Stiassny, 1987</bibRefCitation>
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); presence of two supraneurals; the seven
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<taxonomicName id="4C604D3A3D70FF9E0F921924FCDAF8F6" box="[698,787,1805,1830]" class="Actinopterygii" family="Cichlidae" genus="Tylochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="370" phylum="Chordata" rank="species" species="lateralis">lateralis</taxonomicName>
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canal foramina in the preopercle; the five
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<taxonomicName id="4C604D3A3D70FF9E09D71924FA91F8F6" box="[1279,1368,1805,1830]" class="Actinopterygii" family="Cichlidae" genus="Tylochromis" kingdom="Animalia" order="Perciformes" pageId="1" pageNumber="370" phylum="Chordata" rank="species" species="lateralis">lateralis</taxonomicName>
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canal foramina on the dentary; a black blotch on the anterior soft dorsal-fin rays; the orobranchial mucosa laterally covered by papillae; by the presence of skin flaps and a lack of microbranchiospines along the external margin of the first ceratobranchial; attachment of the ligament hypohyal-hypobranchial 1 along the antero-medial spiny process of the first hypobranchial (
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<figureCitation id="135B2A3C3D70FF9E0F0619B4FDBEF866" box="[558,631,1949,1974]" captionStart="FIGURE 4" captionStartId="4.[151,249,663,685]" captionTargetBox="[294,1292,193,640]" captionTargetId="figure@4.[294,1292,193,641]" captionTargetPageId="4" captionText="FIGURE 4. First hypobranchial of (A) Retroculus acherontos, MZUSP 85838, and (B) R. septentrionalis, MZUSP 38947, paratype. Dorsal view, anterior to top. Cartilage black. Scale bar = 1 mm." httpUri="https://zenodo.org/record/245234/files/figure.png" pageId="1" pageNumber="370">Fig. 4</figureCitation>
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); supracleithrum laterosensory canal posteroventrally angled (
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<figureCitation id="135B2A3C3D70FF9E086D19B4FA44F866" box="[1349,1421,1949,1974]" captionStart="FIGURE 5" captionStartId="4.[151,249,1899,1921]" captionTargetBox="[606,969,785,1851]" captionTargetId="figure@4.[606,969,785,1851]" captionTargetPageId="4" captionText="FIGURE 5. Supracleithrum of Retroculus acherontos, MZUSP 85838. Lateral view, anterior to left. Scale bar = 1 mm." httpUri="https://zenodo.org/record/245235/files/figure.png" pageId="1" pageNumber="370">Fig. 5</figureCitation>
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); posterior portion of the dentigerous arm of the dentary slender; and rakers at base of epibranchial lobe (
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<figureCitation id="135B2A3C3D70FF9E083519E8FAA9F80A" box="[1309,1376,1985,2010]" captionStart="FIGURE 6" captionStartId="5.[151,249,1470,1492]" captionTargetBox="[340,1247,193,1448]" captionTargetId="figure@5.[340,1248,193,1449]" captionTargetPageId="5" captionText="FIGURE 6. Epibranchial lobe on the first branchial arch of (A) Geophagus altifrons, MZUSP 8547, 148 mm SL, and (B) Retroculus acherontos, MZUSP 85826, 161.3 mm SL. Lateral view, anterior to left." httpUri="https://zenodo.org/record/245236/files/figure.png" pageId="1" pageNumber="370">Fig. 6</figureCitation>
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B).
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |