treatments-xml/data/03/C9/87/03C987A6FF9BFFE53AB6FBB71C53D8BD.xml
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<document id="092C86272944377271ADB6205670D39D" ID-CLB-Dataset="22568" ID-DOI="10.11646/zootaxa.4504.1.4" ID-GBIF-Dataset="2ffe9030-0b6d-48d9-88a0-819dc0c8a941" ID-ISSN="1175-5326" ID-Zenodo-Dep="3770961" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588019052635" checkinUser="plazi" docAuthor="Jordal, Bjarte H." docDate="2018" docId="03C987A6FF9BFFE53AB6FBB71C53D8BD" docLanguage="en" docName="zootaxa.4504.1.4.pdf" docOrigin="Zootaxa 4504 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Scolytodes curvicostatus Jordal 2018, sp. nov." docType="treatment" docUuid="BC1B067C-6418-466E-BE89-9A3DB3534D94" docUuidSource="ZooBank" docVersion="8" lastPageNumber="88" masterDocId="FFF0FFDEFF92FFE93A21F8631D77D868" masterDocTitle="Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae)" masterLastPageNumber="104" masterPageNumber="76" pageNumber="85" updateTime="1698837411584" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="06842D1F045C06E46CED8F1CF49ADEEA">Hidden gems in museum cabinets: new species and new distributional records of Scolytodes (Coleoptera: Scolytinae)</mods:title>
</mods:titleInfo>
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<mods:namePart id="046D5F1A6B55C30447F4C47D51E810BD">Jordal, Bjarte H.</mods:namePart>
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<mods:title id="3FD1E45A2A9F4055F3038B6950A50A2C">Zootaxa</mods:title>
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<mods:date id="4CF1E9342AC35B5FE18DAC0B8AE17345">2018</mods:date>
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<mods:number id="C3F8CC7A76F670C0A62723AFC5D05165">2018-10-23</mods:number>
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<mods:number id="5D9EEF7B6A6E899CBBF459ABD4F3F95C">4504</mods:number>
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<mods:identifier id="65C4E0EE05BF1B079A6D45EE9B51B7FA" type="DOI">10.11646/zootaxa.4504.1.4</mods:identifier>
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<treatment id="03C987A6FF9BFFE53AB6FBB71C53D8BD" ID-DOI="http://doi.org/10.5281/zenodo.3798792" ID-GBIF-Taxon="163813169" ID-Zenodo-Dep="3798792" LSID="urn:lsid:zoobank.org:act:BC1B067C-6418-466E-BE89-9A3DB3534D94" httpUri="http://treatment.plazi.org/id/03C987A6FF9BFFE53AB6FBB71C53D8BD" lastPageId="12" lastPageNumber="88" pageId="9" pageNumber="85">
<subSubSection id="C37A653BFF9BFFE03AB6FBB71EBEDC67" pageId="9" pageNumber="85" type="nomenclature">
<paragraph id="8BDF36B0FF9BFFE03AB6FBB71FF6DB86" blockId="9.[151,969,980,1075]" box="[151,641,980,1006]" pageId="9" pageNumber="85">
<heading id="D09781DCFF9BFFE03AB6FBB71FF6DB86" bold="true" box="[151,641,980,1006]" fontSize="11" level="1" pageId="9" pageNumber="85" reason="1">
<emphasis id="B914EAA2FF9BFFE03AB6FBB71FF6DB86" bold="true" box="[151,641,980,1006]" pageId="9" pageNumber="85">
<taxonomicName id="4C604D33FF9BFFE03AB6FBB71F60DB86" authority="Jordal" authorityName="Jordal" authorityYear="2018" box="[151,535,980,1006]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="85" phylum="Arthropoda" rank="species" species="curvicostatus" status="sp. nov.">
<emphasis id="B914EAA2FF9BFFE03AB6FBB71CCBDB86" bold="true" box="[151,444,980,1006]" italics="true" pageId="9" pageNumber="85">Scolytodes curvicostatus</emphasis>
Jordal
</taxonomicName>
,
<taxonomicNameLabel id="A22757D9FF9BFFE03805FBB71FF6DB86" box="[548,641,980,1006]" pageId="9" pageNumber="85" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BDF36B0FF9BFFE03AB6FB951EBEDC67" blockId="9.[151,969,980,1075]" box="[151,969,1014,1039]" pageId="9" pageNumber="85">
<uri id="FFF13AB2FF9BFFE03AB6FB951EBEDC67" box="[151,969,1014,1039]" pageId="9" pageNumber="85">
urn:lsid:zoobank.org:act:
<uuid id="FFC60C65FF9BFFE03B8BFB951EBEDC67" box="[426,969,1014,1039]" pageId="9" pageNumber="85">BC1B067C-6418-466E-BE89-9A3DB3534D94</uuid>
</uri>
</paragraph>
</subSubSection>
<subSubSection id="C37A653BFF9BFFE03AB6FC791C26DC5B" box="[151,337,1050,1075]" pageId="9" pageNumber="85" type="description">
<paragraph id="8BDF36B0FF9BFFE03AB6FC791C26DC5B" blockId="9.[151,969,980,1075]" box="[151,337,1050,1075]" pageId="9" pageNumber="85">
(
<figureCitation id="135B2A35FF9BFFE03ABEFC791C3FDC5B" box="[159,328,1050,1075]" captionStart="FIGURES 2836" captionStartId="11.[151,264,1962,1984]" captionTargetBox="[154,1429,249,1896]" captionTargetId="figure@11.[151,1435,241,1897]" captionTargetPageId="11" captionText="FIGURES 2836. Dorsal, lateral and front view of the holotype of female Scolytodes curvicostatus (28, 31, 34), female S. vellus (29, 32, 35), and S. granulatus (30, 33, 36)." figureDoi="http://doi.org/10.5281/zenodo.3770969" httpUri="https://zenodo.org/record/3770969/files/figure.png" pageId="9" pageNumber="85">Figs 28, 31, 34</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C37A653BFF9BFFE03AB6FC01182BDCF7" pageId="9" pageNumber="85" type="materials_examined">
<paragraph id="8BDF36B0FF9BFFE03AB6FC01182BDCF7" blockId="9.[151,1437,1122,2010]" pageId="9" pageNumber="85">
<emphasis id="B914EAA2FF9BFFE03AB6FC011C33DC13" bold="true" box="[151,324,1122,1147]" pageId="9" pageNumber="85">Type material.</emphasis>
<materialsCitation id="3B083CEDFF9BFFE03B6AFC01182BDCF7" ID-GBIF-Occurrence="2609009876" collectingDate="1997-06-07" collectionCode="FSCA" collectorName="de Junio &amp; R. Anderson" country="Costa Rica" elevation="2600" latitude="9.658333" location="Tres de Junio" longLatPrecision="21" longitude="-83.85833" pageId="9" pageNumber="85" specimenCount="1" stateProvince="San Jose" typeStatus="holotype">
<emphasis id="B914EAA2FF9BFFE03B6AFC011F6ADC13" bold="true" box="[331,541,1122,1147]" pageId="9" pageNumber="85">
<typeStatus id="54DB8812FF9BFFE03B6AFC011CCEDC13" box="[331,441,1122,1147]" pageId="9" pageNumber="85" type="holotype">Holotype</typeStatus>
, female:
</emphasis>
<collectingCountry id="F3777620FF9BFFE03805FC001FD7DC13" box="[548,672,1122,1147]" name="Costa Rica" pageId="9" pageNumber="85">Costa Rica</collectingCountry>
,
<collectingRegion id="49A4F852FF9BFFE0388AFC001E7ADC13" box="[683,781,1123,1147]" country="Costa Rica" name="San Jose" pageId="9" pageNumber="85">San Jose</collectingRegion>
/
<collectingRegion id="49A4F852FF9BFFE03934FC001E18DC13" box="[789,879,1123,1147]" country="Costa Rica" name="Cartago" pageId="9" pageNumber="85">Cartago</collectingRegion>
, km 69, Int. Amer. Hwy. nr.
<location id="8EBF606BFF9BFFE03E8DFC001833DC13" LSID="urn:lsid:plazi:treatment:03C987A6FF9BFFE53AB6FBB71C53D8BD:8EBF606BFF9BFFE03E8DFC001833DC13" box="[1196,1348,1122,1147]" country="Costa Rica" latitude="9.658333" longLatPrecision="21" longitude="-83.85833" name="Tres de Junio" pageId="9" pageNumber="85" stateProvince="San Jose">
Tres
<collectorName id="26955366FF9BFFE03EC2FC011833DC13" box="[1251,1348,1122,1147]" pageId="9" pageNumber="85">de Junio</collectorName>
</location>
,
<quantity id="4C989B55FF9BFFE03F6FFC0118EBDC13" box="[1358,1436,1122,1147]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="85" unit="m" value="2600.0">
<elevation id="004DD183FF9BFFE03F6FFC0118EBDC13" box="[1358,1436,1122,1147]" metricMagnitude="3" metricUnit="m" metricValue="2.6" pageId="9" pageNumber="85" unit="m" value="2600.0">2600m</elevation>
</quantity>
<geoCoordinate id="EE545077FF9BFFE03AB6FCE41C50DCF7" box="[151,295,1159,1183]" degrees="09" direction="north" minutes="39" orientation="latitude" pageId="9" pageNumber="85" precision="15" seconds="30" value="9.658333">09°3930N</geoCoordinate>
,
<geoCoordinate id="EE545077FF9BFFE03B13FCE51CB2DCF7" box="[306,453,1158,1183]" degrees="83" direction="west" minutes="51" orientation="longitude" pageId="9" pageNumber="85" precision="15" seconds="30" value="-83.85833">83°5130W</geoCoordinate>
,
<date id="FFDE1070FF9BFFE03BF0FCE51F4ADCF7" box="[465,573,1158,1183]" pageId="9" pageNumber="85" value="1997-06-07">
<collectingDate id="EF9AE998FF9BFFE03BF0FCE51F4ADCF7" box="[465,573,1158,1183]" pageId="9" pageNumber="85" value="1997-06-07">7.vi.1997</collectingDate>
</date>
,
<collectorName id="26955366FF9BFFE03868FCE41FADDCF7" box="[585,730,1158,1183]" pageId="9" pageNumber="85">R. Anderson</collectorName>
, wet cloud forest litter, 97-004A. HT deposited in
<collectionCode id="ED71AE75FF9BFFE03F2EFCE4182EDCF7" box="[1295,1369,1159,1183]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="9" pageNumber="85" type="Museum">FSCA</collectionCode>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="C37A653BFF9BFFE03AE6FCC91C70DD3B" pageId="9" pageNumber="85" type="diagnosis">
<paragraph id="8BDF36B0FF9BFFE03AE6FCC91C70DD3B" blockId="9.[151,1437,1122,2010]" pageId="9" pageNumber="85">
<emphasis id="B914EAA2FF9BFFE03AE6FCC91C36DCAB" bold="true" box="[199,321,1194,1219]" pageId="9" pageNumber="85">Diagnosis.</emphasis>
Interstriae 10 sharply elevated to level of ventrite 1; protibiae with a distinct additional mesal tooth near tarsal insertion. Lateral carina on pronotum strongly curved dorso-ventrally; pronotum from above appearing constricted on posterior fourth. Distinguished from a group of distantly related species near
<taxonomicName id="4C604D33FF9BFFE03EB6FC9018EBDD63" authority="Wood" authorityName="Wood" authorityYear="1978" box="[1175,1436,1266,1291]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="85" phylum="Arthropoda" rank="species" species="semipunctatus">
<emphasis id="B914EAA2FF9BFFE03EB6FC901824DD62" box="[1175,1363,1267,1290]" italics="true" pageId="9" pageNumber="85">S. semipunctatus</emphasis>
Wood
</taxonomicName>
and
<taxonomicName id="4C604D33FF9BFFE03AE7FD741CC1DD47" authority="Wood" authorityName="Wood" authorityYear="1967" box="[198,438,1302,1327]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="85" phylum="Arthropoda" rank="species" species="clusiacolens">
<emphasis id="B914EAA2FF9BFFE03AE7FD741C1ADD46" box="[198,365,1302,1326]" italics="true" pageId="9" pageNumber="85">S. clusiacolens</emphasis>
Wood
</taxonomicName>
by erect interstrial setae in rows combined with the presence of very short strial setae on the elytra.
</paragraph>
</subSubSection>
<subSubSection id="C37A653BFF9BFFE03AE6FD3D1C30DF23" pageId="9" pageNumber="85" type="description">
<paragraph id="8BDF36B0FF9BFFE03AE6FD3D1FA8DF6B" blockId="9.[151,1437,1122,2010]" pageId="9" pageNumber="85">
<emphasis id="B914EAA2FF9BFFE03AE6FD3D1CC5DD1F" bold="true" box="[199,434,1374,1399]" pageId="9" pageNumber="85">Description female.</emphasis>
Length
<quantity id="4C989B55FF9BFFE03834FD3C1F07DD1E" box="[533,624,1375,1399]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="9" pageNumber="85" unit="mm" value="2.7">2.7 mm</quantity>
, 2.45 × longer than wide; colour reddish brown to black.
<emphasis id="B914EAA2FF9BFFE03F35FD3D1823DD1F" bold="true" box="[1300,1364,1374,1399]" pageId="9" pageNumber="85">Head</emphasis>
. Eyes entire, separated above by 2.8 × their width. Frons convex on upper two-thirds, weakly impressed below; surface reticulate with few scattered punctures; vestiture consisting of a brush of short golden setae in impressed area, denser on epistoma. Antennal club with two obliquely procurved sutures marked by fine setae, first two segments corneous. Funiculus 6-segmented.
<emphasis id="B914EAA2FF9BFFE0383EFD8D1FE0DE6F" bold="true" box="[543,663,1518,1543]" pageId="9" pageNumber="85">Pronotum</emphasis>
strongly reticulate, small punctures spaced by 3¯5 × their diameter. Vestiture consisting of 6 erect setae (420), and scant, short, fine, interspersed setae.
<emphasis id="B914EAA2FF9BFFE03E79FE7119D3DE43" bold="true" box="[1112,1188,1554,1579]" pageId="9" pageNumber="85">Elytra</emphasis>
generally smooth and shiny, striae weakly impressed, punctures distinct, in regular rows, separated by 1¯2 × their diameter; interstriae 3 × wider than striae, punctures minute. Vestiture consisting of erect interstrial setae of variable length, and much shorter recumbent strial setae.
<emphasis id="B914EAA2FF9BFFE03BD3FE1D1F46DEFF" bold="true" box="[498,561,1662,1687]" pageId="9" pageNumber="85">Legs.</emphasis>
Procoxae separated by 0.8 × and mesocoxae 1.2 × the width of one procoxa. Protibiae narrow, distal teeth 1 and 2 of equal length, with 4¯5 tiny additional granules along the edge towards base; protibial mucro obtuse. Meso- and metatibiae with 6 lateral, socketed teeth on distal third.
<emphasis id="B914EAA2FF9BFFE03EECFEA518E2DEB7" bold="true" box="[1229,1429,1734,1759]" pageId="9" pageNumber="85">Ventral vestiture</emphasis>
. Setae on metasternum and metanepisternum simple.
</paragraph>
<paragraph id="8BDF36B0FF9BFFE03AE6FF6D1C30DF23" blockId="9.[151,1437,1122,2010]" pageId="9" pageNumber="85">
<emphasis id="B914EAA2FF9BFFE03AE6FF6D1C7CDF4F" bold="true" box="[199,267,1806,1831]" pageId="9" pageNumber="85">Male.</emphasis>
Unknown. The sex of the
<typeStatus id="54DB8812FF9BFFE03819FF6C1F1FDF4F" box="[568,616,1807,1831]" pageId="9" pageNumber="85">type</typeStatus>
(female) was determined based on the single sclerotized terminal tergite (seven in total).
</paragraph>
</subSubSection>
<subSubSection id="C37A653BFF9BFFE03AE6FF351943DF07" box="[199,1076,1878,1903]" pageId="9" pageNumber="85" type="key">
<paragraph id="8BDF36B0FF9BFFE03AE6FF351943DF07" blockId="9.[151,1437,1122,2010]" box="[199,1076,1878,1903]" pageId="9" pageNumber="85">
<emphasis id="B914EAA2FF9BFFE03AE6FF351CEADF07" bold="true" box="[199,413,1878,1903]" pageId="9" pageNumber="85">Key (Wood 1982).</emphasis>
Keys to couplet 25,
<taxonomicName id="4C604D33FF9BFFE038A5FF341E5CDF06" ID-CoL="4VT3L" authorityName="Wood" authorityYear="1967" box="[644,811,1878,1902]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="9" pageNumber="85" phylum="Arthropoda" rank="species" species="clusiacolens">
<emphasis id="B914EAA2FF9BFFE038A5FF341E5CDF06" box="[644,811,1878,1902]" italics="true" pageId="9" pageNumber="85">S. clusiacolens</emphasis>
</taxonomicName>
, with no further match.
</paragraph>
</subSubSection>
<subSubSection id="C37A653BFF9BFFE03AE6FF191C7CDFB2" pageId="9" pageNumber="85" type="etymology">
<paragraph id="8BDF36B0FF9BFFE03AE6FF191C7CDFB2" blockId="9.[151,1437,1122,2010]" pageId="9" pageNumber="85">
<emphasis id="B914EAA2FF9BFFE03AE6FF191C39DFFB" bold="true" box="[199,334,1914,1939]" pageId="9" pageNumber="85">Etymology.</emphasis>
The name is composed by the Latin masculine genitive adjective
<emphasis id="B914EAA2FF9BFFE03E78FF1819E5DFFA" box="[1113,1170,1915,1938]" italics="true" pageId="9" pageNumber="85">curvi</emphasis>
(from
<emphasis id="B914EAA2FF9BFFE03EC9FF181844DFFA" box="[1256,1331,1915,1938]" italics="true" pageId="9" pageNumber="85">curvus</emphasis>
) and the masculine nominative adjective
<emphasis id="B914EAA2FF9BFFE03835FFFC1F07DFDE" box="[532,624,1951,1974]" italics="true" pageId="9" pageNumber="85">costatus</emphasis>
, referring to the strongly dorsoventrally curved lateral costa of the pronotum.
</paragraph>
</subSubSection>
<caption id="DF1F6638FF98FFE33AB6FFCB1E76DFB4" ID-DOI="http://doi.org/10.5281/zenodo.3770967" ID-Zenodo-Dep="3770967" httpUri="https://zenodo.org/record/3770967/files/figure.png" pageId="10" pageNumber="86" startId="10.[151,264,1960,1982]" targetBox="[164,1422,285,1891]" targetPageId="10">
<paragraph id="8BDF36B0FF98FFE33AB6FFCB1E76DFB4" blockId="10.[151,1436,1960,2012]" pageId="10" pageNumber="86">
<emphasis id="B914EAA2FF98FFE33AB6FFCB1C2EDFD6" bold="true" box="[151,345,1960,1982]" pageId="10" pageNumber="86">FIGURES 1927.</emphasis>
Dorsal, lateral and front view of the holotype of male
<taxonomicName id="4C604D33FF98FFE3399DFFCB19EEDFD6" authorityName="Jordal" authorityYear="2018" box="[956,1177,1960,1982]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="86" phylum="Arthropoda" rank="species" species="monticola">
<emphasis id="B914EAA2FF98FFE3399DFFCB19EEDFD6" box="[956,1177,1960,1982]" italics="true" pageId="10" pageNumber="86">Scolytodes monticola</emphasis>
</taxonomicName>
(19, 22, 25), female
<taxonomicName id="4C604D33FF98FFE33FA8FFCA1D88DFB4" authorityName="Jordal" authorityYear="2018" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="86" phylum="Arthropoda" rank="species" species="aureifrons">
<emphasis id="B914EAA2FF98FFE33FA8FFCA1D88DFB4" italics="true" pageId="10" pageNumber="86">S. aureifrons</emphasis>
</taxonomicName>
(20, 23, 26), and female
<taxonomicName id="4C604D33FF98FFE33BDCFFA41F09DFB4" authorityName="Jordal" authorityYear="2018" box="[509,638,1990,2012]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="10" pageNumber="86" phylum="Arthropoda" rank="species" species="piliscapus">
<emphasis id="B914EAA2FF98FFE33BDCFFA41F09DFB4" box="[509,638,1990,2012]" italics="true" pageId="10" pageNumber="86">S. piliscapus</emphasis>
</taxonomicName>
(21, 24, 27).
</paragraph>
</caption>
<caption id="DF1F6638FF99FFE23AB6FFC91FE7DF8B" ID-DOI="http://doi.org/10.5281/zenodo.3770969" ID-Zenodo-Dep="3770969" httpUri="https://zenodo.org/record/3770969/files/figure.png" pageId="11" pageNumber="87" startId="11.[151,264,1962,1984]" targetBox="[154,1429,249,1896]" targetPageId="11">
<paragraph id="8BDF36B0FF99FFE23AB6FFC91FE7DF8B" blockId="11.[151,1436,1962,2019]" pageId="11" pageNumber="87">
<emphasis id="B914EAA2FF99FFE23AB6FFC91C21DFA8" bold="true" box="[151,342,1962,1984]" pageId="11" pageNumber="87">FIGURES 2836.</emphasis>
Dorsal, lateral and front view of the holotype of female
<taxonomicName id="4C604D33FF99FFE2398DFFC919DEDFA8" authorityName="Jordal" authorityYear="2018" box="[940,1193,1962,1984]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="species" species="curvicostatus">
<emphasis id="B914EAA2FF99FFE2398DFFC919DEDFA8" box="[940,1193,1962,1984]" italics="true" pageId="11" pageNumber="87">Scolytodes curvicostatus</emphasis>
</taxonomicName>
(28, 31, 34), female
<taxonomicName id="4C604D33FF99FFE23FA8FFC81DA5DF8B" authorityName="Jordal" authorityYear="2018" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="species" species="vellus">
<emphasis id="B914EAA2FF99FFE23FA8FFC81DA5DF8B" italics="true" pageId="11" pageNumber="87">S. vellus</emphasis>
</taxonomicName>
(29, 32, 35), and
<taxonomicName id="4C604D33FF99FFE23BA7FFAD1F7ADF8B" authorityName="Jordal" authorityYear="2018" box="[390,525,1997,2019]" class="Insecta" family="Curculionidae" genus="Scolytodes" kingdom="Animalia" order="Coleoptera" pageId="11" pageNumber="87" phylum="Arthropoda" rank="species" species="granulatus">
<emphasis id="B914EAA2FF99FFE23BA7FFAD1F7ADF8B" box="[390,525,1997,2019]" italics="true" pageId="11" pageNumber="87">S. granulatus</emphasis>
</taxonomicName>
(30, 33, 36).
</paragraph>
</caption>
<subSubSection id="C37A653BFF9EFFE53AE6F8F41C53D8BD" pageId="12" pageNumber="88" type="biology_ecology">
<paragraph id="8BDF36B0FF9EFFE53AE6F8F41C53D8BD" blockId="12.[151,1436,151,213]" pageId="12" pageNumber="88">
<emphasis id="B914EAA2FF9EFFE53AE6F8F41C82D8D8" bold="true" box="[199,501,151,176]" pageId="12" pageNumber="88">Biology and distribution.</emphasis>
This species is only known from the
<typeStatus id="54DB8812FF9EFFE5398DF8FB1EABD8D8" box="[940,988,152,176]" pageId="12" pageNumber="88">type</typeStatus>
locality in
<collectingCountry id="F3777620FF9EFFE53E40F8FB1997D8D8" box="[1121,1248,151,176]" name="Costa Rica" pageId="12" pageNumber="88">Costa Rica</collectingCountry>
, in high altitude cloud forest.
</paragraph>
</subSubSection>
</treatment>
</document>