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<document id="CE7D6816495AA5DD27965BE7ED50ADA3" ID-CLB-Dataset="53324" ID-DOI="10.11646/phytotaxa.576.2.1" ID-GBIF-Dataset="1a9bd59f-7293-432f-b3c1-6df0bc659474" ID-ISSN="1179-3163" ID-Zenodo-Dep="7461280" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" checkinTime="1671521402645" checkinUser="plazi" docAuthor="Hassan, Walaa A., Al-Shaye, Najla A., Alghamdi, Salma, Korany, Shereen M. &amp; Iamonico, Duilio" docDate="2022" docId="03C987964B41FFA706B902B854E166A2" docLanguage="en" docName="phytotaxa.576.2.1.pdf" docOrigin="Phytotaxa 576 (2)" docSource="http://dx.doi.org/10.11646/phytotaxa.576.2.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.7:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="7" docTitle="Amaranthus viridis L." docType="treatment" docVersion="7" lastPageNumber="141" masterDocId="FFF0FFEE4B47FFA106310226570A6226" masterDocTitle="Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia" masterLastPageNumber="157" masterPageNumber="135" pageNumber="141" updateTime="1699413697922" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="53C7823E17102FF5CE8854C65C738319">Taxonomic revision of the genus Amaranthus (Amaranthaceae) in Saudi Arabia</mods:title>
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<mods:namePart id="C27EDD12AB7735E5AC2514A9C1ECB7F7">Hassan, Walaa A.</mods:namePart>
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<mods:namePart id="0CDF4A3476266918F270B4405F5EE429">Al-Shaye, Najla A.</mods:namePart>
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<mods:affiliation id="4ED44A41849AA458599DAA2111A6157E">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; naaalshaye @ pnu. edu. sa; https: // orcid. org / 0000 - 0002 - 0447 - 8613</mods:affiliation>
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<mods:namePart id="EF2C9EACB8660E42738C5AB168A9D6A9">Alghamdi, Salma</mods:namePart>
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<mods:affiliation id="7715514E657B307858E6891D8B10F723">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; saatalghamdi @ pnu. edu. sa; https: // orcid. org / 0000 - 0001 - 7716 - 2233</mods:affiliation>
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<mods:namePart id="45B1B8619FE6D6290042AD9EDC6DFEF3">Korany, Shereen M.</mods:namePart>
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<mods:affiliation id="20D5CEA745D86E2F0621CC11504949A4">Department of Biology, College of Science, Princess Nourah bint Abdulrahman University, P. O. Box 84428, Riyadh 11671, Saudi Arabia &amp; smkorany @ pnu. edu. as; https: // orcid. org / 0000 - 0002 - 9355 - 3952</mods:affiliation>
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<mods:namePart id="AEFE2EA6F244688951304C798BB00145">Iamonico, Duilio</mods:namePart>
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<mods:affiliation id="D3626A686BD4D48BCDC1112CBB1D3D17">Ce. R. S. I. Te. S., University of Rome Sapienza, viale XXIV Maggio 7, 04100 - Latina, Italy &amp; duilio. iamonico @ uniroma 1. it; https: // orcid. org / 0000 - 0001 - 5491 - 7568 &amp; Authors that contributed equally to the work</mods:affiliation>
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<treatment id="03C987964B41FFA706B902B854E166A2" ID-DOI="http://doi.org/10.5281/zenodo.7465652" ID-GBIF-Taxon="206458943" ID-Zenodo-Dep="7465652" LSID="urn:lsid:plazi:treatment:03C987964B41FFA706B902B854E166A2" httpUri="http://treatment.plazi.org/id/03C987964B41FFA706B902B854E166A2" lastPageNumber="141" pageId="6" pageNumber="141">
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<emphasis id="B914EA924B41FFA706B902B85797629E" bold="true" box="[136,157,158,184]" pageId="6" pageNumber="141">4.</emphasis>
<taxonomicName id="4C604D034B41FFA7069502B856AE629F" ID-CoL="CH7N" authority="L." authorityName="L." box="[164,420,158,185]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="141" phylum="Tracheophyta" rank="species" species="viridis">
<emphasis id="B914EA924B41FFA7069502B8568E629F" bold="true" box="[164,388,158,185]" italics="true" pageId="6" pageNumber="141">Amaranthus viridis</emphasis>
L.
</taxonomicName>
, Sp. Pl., ed. 2. 2: 1405. 1763
</heading>
</paragraph>
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<paragraph id="8BDF36804B41FFA706B902E054FE62D9" blockId="6.[136,1452,158,256]" pageId="6" pageNumber="141">
<materialsCitation id="3B083CDD4B41FFA706B902E054FB62D9" ID-GBIF-Occurrence="3999584315" collectorName="Herb. Linn. No." location="Habitat in Europa" pageId="6" pageNumber="141" specimenCount="2" typeStatus="lectotype">
Type (
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designated by
<bibRefCitation id="EFF14B714B41FFA7078B02E055D662FD" author="Fawcett, W. &amp; Rendle, A. B." box="[442,732,198,220]" pageId="6" pageNumber="141" refId="ref15699" refString="Fawcett, W. &amp; Rendle, A. B. (1914) Flora of Jamaica containing descriptions of the flowering plants known from the island, vol. 3. Order of the Trustees of the British Museum, London, 208 pp." type="book" year="1914">Fawcett &amp; Rendle 1914: 131</bibRefCitation>
):—UNKNOWN ORIGIN.
<emphasis id="B914EA924B41FFA705DD02E053F762FD" box="[1004,1277,198,219]" italics="true" pageId="6" pageNumber="141">
<location id="8EBF605B4B41FFA705DD02E053AE62FD" LSID="urn:lsid:plazi:treatment:03C987964B41FFA706B902B854E166A2:8EBF605B4B41FFA705DD02E053AE62FD" box="[1004,1188,198,219]" name="Habitat in Europa" pageId="6" pageNumber="141">Habitat in Europa</location>
, Brasilia
</emphasis>
,
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1117.15 (LINN!, image of the lectotype available at http://linnean-online.org/11641/)
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.
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<emphasis id="B914EA924B41FFA706B903085632636E" bold="true" box="[136,312,302,328]" pageId="6" pageNumber="141">Description:—</emphasis>
Herbs 17(8) dm tall, monoecious, annual (therophyte). Stems erect, glabrous, green to brownish, branched. Leaves black-green, ovate, rhomboidal [(2.04.5) × (1.5)2.07.0 cm], with entire (rarely undulate) margins, apex obtuse or rounded (rarely slightly emarginate) and sometimens mucronate, base usually cuneate, usually glabrous, petioled [petiole 1.55.0 cm long]. Synflorescences terminal, spike- or panicle-like (sometimes axillary glomerules also occur), the main florescence
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long, green to brown, usually thin (
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in diameter). Floral bracts yellowish or greenish, ovate to lanceolate [0.51.0 ×
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], shorter (up to ⅓) than the perianth, acuminate, margin entire, glabrous. Staminate flowers with 3 tepals, ovate; stamens 3. Pistillate flowers with 3 tepals, ovatelanceolate or obovate-spathulate (1.21.5 ×
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), with rounded apex (sometimes acute), mucronate or not; stigmas (2)3. Fruit brownish, subglobose [(1.2)1.41.7(1.9) × 1.41.6(1.8) mm] as long as or slighly longer (up to ¼) than the perianth, clearly rugose, indehiscent. Seed lenticular (
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in diameter), black or brownish-black.
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<emphasis id="B914EA924B41FFA7068C00B056716096" bold="true" box="[189,379,662,688]" pageId="6" pageNumber="141">Iconography:—</emphasis>
<bibRefCitation id="EFF14B714B41FFA7074A00B055706096" author="Chaudhary, S. A." box="[379,634,662,688]" pageId="6" pageNumber="141" refId="ref15255" refString="Chaudhary, S. A. (1998) Flora of the Kingdom of the Saudi Arabia, vol. 1. Ministry of Agriculture &amp; Water, Riyadh, 692 pp." type="book" year="1998">Chaudhary (1998: 239</bibRefCitation>
, Plate. 129b),
<bibRefCitation id="EFF14B714B41FFA7052B00B054ED6096" author="Bayon, N. D." box="[794,999,662,689]" pageId="6" pageNumber="141" pagination="261 - 383" refId="ref15004" refString="Bayon, N. D. (2015) Revision taxonomica de las especies monoicas de Amaranthus (Amaranthaceae): Amaranthus subg. Amaranthus and Amaranthus subg. Albersia. Ann. Missouri Bot. Garden 101 (2): 261 - 383. https: // doi. org / 10.3417 / 2010080" type="journal article" year="2015">Bayón (2015: 371</bibRefCitation>
, Figura 64).
</paragraph>
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<emphasis id="B914EA924B41FFA7068C009C565560F2" bold="true" box="[189,351,698,724]" pageId="6" pageNumber="141">Phenology:—</emphasis>
Flowering time february.
</paragraph>
<paragraph id="8BDF36804B41FFA7068C00F8549360DE" blockId="6.[136,1452,302,1157]" box="[189,921,734,761]" pageId="6" pageNumber="141">
<emphasis id="B914EA924B41FFA7068C00F856EF60DE" bold="true" box="[189,485,734,760]" pageId="6" pageNumber="141">Habitat and elevation:—</emphasis>
Human-made habitat,
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a.s.l.
</paragraph>
<paragraph id="8BDF36804B41FFA7068C0124554F613A" blockId="6.[136,1452,302,1157]" box="[189,581,770,797]" pageId="6" pageNumber="141">
<emphasis id="B914EA924B41FFA7068C012456E2613A" bold="true" box="[189,488,770,796]" pageId="6" pageNumber="141">Chromosome number:—</emphasis>
2n = 34.
</paragraph>
<paragraph id="8BDF36804B41FFA7068C010056B16142" blockId="6.[136,1452,302,1157]" pageId="6" pageNumber="141">
<emphasis id="B914EA924B41FFA7068C0100567B6166" bold="true" box="[189,369,806,832]" pageId="6" pageNumber="141">Alien status:—</emphasis>
Neophyte species native to South America, it can be considered as naturalized in
<collectingCountry id="F37776104B41FFA70329010052A16166" box="[1304,1451,806,832]" name="Saudi Arabia" pageId="6" pageNumber="141">Saudi Arabia</collectingCountry>
(see also
<bibRefCitation id="EFF14B714B41FFA706C1016C56A46142" author="Chaudhary, S. A." box="[240,430,842,868]" pageId="6" pageNumber="141" refId="ref15255" refString="Chaudhary, S. A. (1998) Flora of the Kingdom of the Saudi Arabia, vol. 1. Ministry of Agriculture &amp; Water, Riyadh, 692 pp." type="book" year="1998">Chaudhary 1998</bibRefCitation>
).
</paragraph>
<paragraph id="8BDF36804B41FFA7068C01485413618A" blockId="6.[136,1452,302,1157]" pageId="6" pageNumber="141">
<emphasis id="B914EA924B41FFA7068C0148559261AE" bold="true" box="[189,664,878,904]" pageId="6" pageNumber="141">
Occurrence in
<collectingCountry id="F37776104B41FFA707430148551861AE" box="[370,530,878,904]" name="Saudi Arabia" pageId="6" pageNumber="141">Saudi Arabia</collectingCountry>
(
<figureCitation id="135B2A054B41FFA704120148556161AE" box="[547,619,878,904]" captionStart="FIGURE 3" captionStartId="6.[136,229,1837,1859]" captionTargetId="figure-393@6.[331,1256,1181,1813]" captionTargetPageId="6" captionText="FIGURE 3. Map of Amaranthus viridis." figureDoi="http://doi.org/10.5281/zenodo.7465622" httpUri="https://zenodo.org/record/7465622/files/figure.png" pageId="6" pageNumber="141">Fig. 3</figureCitation>
):—
</emphasis>
<collectingRegion id="49A4F8624B41FFA704A9014854B561AE" box="[664,959,878,904]" country="Saudi Arabia" name="Al Hudud ash Shamaliyah" pageId="6" pageNumber="141">Al hudud ash Shamaliyah</collectingRegion>
(
<bibRefCitation id="EFF14B714B41FFA705E10148525C61AE" author="Osman, A. K. E. &amp; El-Ameid Abedin, M. A." box="[976,1366,878,904]" pageId="6" pageNumber="141" pagination="772 - 789" refId="ref17597" refString="Osman, A. K. E. &amp; El-Ameid Abedin, M. A. (2019) Floristic diversity of Wadi Ar'ar, Saudi Arabia. Journal of Taibah University for Science 13 (1): 772 - 789. https: // doi. org / 10.1080 / 16583655.2019.1634177" type="journal article" year="2019">Osman &amp; El-Ameid Abedin 2019</bibRefCitation>
), Bisha (
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Abbas
<emphasis id="B914EA924B41FFA706D101B55612618A" box="[224,280,914,940]" italics="true" pageId="6" pageNumber="141">et al.</emphasis>
2020
</bibRefCitation>
),
<collectingRegion id="49A4F8624B41FFA7075F01B456A0618A" box="[366,426,914,940]" country="Saudi Arabia" name="Jizan" pageId="6" pageNumber="141">Jizan</collectingRegion>
, and Taif (
<bibRefCitation id="EFF14B714B41FFA7041101B45407618A" author="Abdullah, M. A. &amp; Al &amp; Yasee, H. &amp; Al-Sudani, Y. &amp; Moneruzaman, M." box="[544,781,914,940]" pageId="6" pageNumber="141" pagination="762 - 769" refId="ref14472" refString="Abdullah, M. A., Al. Yasee, H., Al-Sudani, Y. &amp; Moneruzaman, M. (2017) Developing weeds of agricultural crops at different levels of heights, in taif area of Saudia Arabia. Bulgarian Journal of Agricultural Science 23 (5): 762 - 769." type="journal article" year="2017">
Abdullah
<emphasis id="B914EA924B41FFA704A001B555C3618A" box="[657,713,914,940]" italics="true" pageId="6" pageNumber="141">et al.</emphasis>
2017
</bibRefCitation>
).
</paragraph>
<paragraph id="8BDF36804B41FFA7068C019054FE661B" blockId="6.[136,1452,302,1157]" pageId="6" pageNumber="141">
<emphasis id="B914EA924B41FFA7068C019056E561F6" bold="true" box="[189,495,950,976]" pageId="6" pageNumber="141">Taxonomic annotation:—</emphasis>
The name
<taxonomicName id="4C604D034B41FFA704560190549B61F6" authority="Desf." authorityName="Desf." box="[615,913,950,976]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="141" phylum="Tracheophyta" rank="species" species="gracilis">
<emphasis id="B914EA924B41FFA704560190544761F6" box="[615,845,950,976]" italics="true" pageId="6" pageNumber="141">Amaranthus gracilis</emphasis>
Desf.
</taxonomicName>
was cited by
<bibRefCitation id="EFF14B714B41FFA7021F019053F561F6" author="Mandaville, J. P." box="[1070,1279,950,976]" pageId="6" pageNumber="141" refId="ref17351" refString="Mandaville, J. P. (2011) Flora of eastern saudi Arabia. Routledge, London, 276 pp." type="book" year="2011">Mandaville (2011)</bibRefCitation>
as synonym of
<taxonomicName id="4C604D034B41FFA706B901FD57FA61D2" box="[136,240,986,1013]" class="Magnoliopsida" family="Amaranthaceae" genus="Amaranthus" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="141" phylum="Tracheophyta" rank="species" species="viridis">
<emphasis id="B914EA924B41FFA706B901FD57FA61D2" box="[136,240,986,1013]" italics="true" pageId="6" pageNumber="141">A. viridis</emphasis>
</taxonomicName>
. Desfontaines name was widely discussed by
<bibRefCitation id="EFF14B714B41FFA7053701FC54DA61D2" author="Iamonico, D." box="[774,976,986,1012]" pageId="6" pageNumber="141" pagination="61 - 68" refId="ref16729" refString="Iamonico, D. (2016 b) Nomenclature survey of the genus Amaranthus (Amaranthaceae). 4. Detailed questions arising around the name Amaranthus gracilis. Botanica Serbica 40 (1): 61 - 68." type="journal article" year="2016">Iamonico (2016b)</bibRefCitation>
who reached to the conclusion that it is a
<emphasis id="B914EA924B41FFA706B901D95658663E" box="[136,338,1022,1048]" italics="true" pageId="6" pageNumber="141">nomen ambiguum</emphasis>
published by
<bibRefCitation id="EFF14B714B41FFA707C601D855D7663E" author="Desfontaines, R. L." box="[503,733,1022,1048]" pageId="6" pageNumber="141" refId="ref15473" refString="Desfontaines, R. L. (1804) Tableau de l'ecole de Botanique du Museum d'Histoire Naturelle. J. A. Brosson, Paris, 238 pp. https: // doi. org / 10.5962 / bhl. title. 13828" type="book" year="1804">Desfontaines (1804)</bibRefCitation>
as
<emphasis id="B914EA924B41FFA7053601D954DE663E" box="[775,980,1023,1048]" italics="true" pageId="6" pageNumber="141">nomen novum pro</emphasis>
<taxonomicName id="4C604D034B41FFA705EF01D8523F663F" authority="Jacq." authorityName="Jacq." box="[990,1333,1022,1049]" class="Magnoliopsida" family="Amaranthaceae" genus="Chenopodium" kingdom="Plantae" order="Caryophyllales" pageId="6" pageNumber="141" phylum="Tracheophyta" rank="species" species="caudatum">
<emphasis id="B914EA924B41FFA705EF01D853F8663E" box="[990,1266,1022,1048]" italics="true" pageId="6" pageNumber="141">Chenopodium caudatum</emphasis>
Jacq.
</taxonomicName>
The latter Jaquins name was proposed as
<emphasis id="B914EA924B41FFA707DC060555C2661A" box="[493,712,1058,1084]" italics="true" pageId="6" pageNumber="141">nomen rejectendum</emphasis>
by
<bibRefCitation id="EFF14B714B41FFA704C3060454FA661B" author="Iamonico, D. &amp; Sukhorukov, A. P. &amp; Reveal, J. L." box="[754,1008,1058,1085]" pageId="6" pageNumber="141" pagination="638 - 639" refId="ref17220" refString="Iamonico, D., Sukhorukov, A. P. &amp; Reveal, J. L. (2015) (2360) Proposal to reject the name Chenopodium caudatum (Amaranthaceae / Chenopodiaceae). Taxon 64 (3): 638 - 639. https: // dx. doi. org / 10.12705 / 643.18" type="journal article" year="2015">
Iamonico
<emphasis id="B914EA924B41FFA7055206055491661A" box="[867,923,1058,1084]" italics="true" pageId="6" pageNumber="141">et al.</emphasis>
(2015)
</bibRefCitation>
.
</paragraph>
<paragraph id="8BDF36804B41FFA7068C066054E166A2" blockId="6.[136,1452,302,1157]" pageId="6" pageNumber="141">
<emphasis id="B914EA924B41FFA7068C066056ED6646" bold="true" box="[189,487,1094,1120]" pageId="6" pageNumber="141">Specimina visa selecta:—</emphasis>
<materialsCitation id="3B083CDD4B41FFA707D6066054ED66A2" ID-GBIF-Occurrence="3999584321" collectingDate="2021-02-17" collectionCode="RO" collectorName="Masrhai &amp; Al-shaye" country="Saudi Arabia" determinerName="Masrhai" elevation="10" location="Jizan" pageId="6" pageNumber="141" specimenCount="1" stateProvince="Jizan">
<collectingCountry id="F37776104B41FFA707D6066055A46646" box="[487,686,1094,1120]" name="Saudi Arabia" pageId="6" pageNumber="141">SAUDI ARABIA</collectingCountry>
,
<collectingRegion id="49A4F8624B41FFA70486066055F96646" box="[695,755,1094,1120]" country="Saudi Arabia" name="Jizan" pageId="6" pageNumber="141">Jizan</collectingRegion>
, human-made habitat, (coastal plain)
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a.s.l.
</elevation>
,
<date id="FFDE10404B41FFA70314066057C966A2" pageId="6" pageNumber="141" value="2021-02-17">
<collectingDate id="EF9AE9A84B41FFA70314066057C966A2" pageId="6" pageNumber="141" value="2021-02-17">17 February 2021</collectingDate>
</date>
, leg.
<emphasis id="B914EA924B41FFA706CF064C554C66A3" box="[254,582,1130,1157]" italics="true" pageId="6" pageNumber="141">
<collectorName id="269553564B41FFA706CF064C565666A2" box="[254,348,1130,1156]" pageId="6" pageNumber="141">Masrhai</collectorName>
et
<collectorName id="269553564B41FFA7074F064C56EA66A2" box="[382,480,1130,1156]" pageId="6" pageNumber="141">Al-shaye</collectorName>
(PNUH)
</emphasis>
, det.
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<determinerName id="7E03DA444B41FFA704B5064C55E866A2" box="[644,738,1130,1156]" pageId="6" pageNumber="141">Masrhai</determinerName>
</emphasis>
, conf.
<emphasis id="B914EA924B41FFA70500064D549166A2" box="[817,923,1131,1156]" italics="true" pageId="6" pageNumber="141">
<collectorName id="269553564B41FFA70500064D549166A2" box="[817,923,1131,1156]" pageId="6" pageNumber="141">Iamonico</collectorName>
</emphasis>
(
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!)
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>