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<mods:title id="98A58DD5CDD7EE63C76367D426BA1E63">Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific</mods:title>
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<paragraph id="8BDFDE46FFC1FFAA75DFE4EFFD2AC3CE" blockId="82.[151,749,151,214]" box="[151,749,151,178]" pageId="82" pageNumber="83">
<taxonomicName id="4C60A5C5FFC1FFAA75DFE4EFFDBCC3CD" authority="(O'Connor, 1997)" baseAuthorityName="O'Connor" baseAuthorityYear="1997" box="[151,635,151,178]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" order="Nassellaria" pageId="82" pageNumber="83" phylum="Sarcomastigophora" rank="species" species="tekopua" status="comb. nov.">
<emphasis id="B9140254FFC1FFAA75DFE4EFFE5DC3CE" box="[151,410,152,178]" italics="true" pageId="82" pageNumber="83">Pelagomanes tekopua</emphasis>
<emphasis id="B9140254FFC1FFAA74E9E4E0FDBCC3CD" bold="true" box="[417,635,151,177]" pageId="82" pageNumber="83">
(
<bibRefCitation id="EFF1A3B7FFC1FFAA74E1E4E0FDB4C3CD" author="O'Connor, B." box="[425,627,151,177]" pageId="82" pageNumber="83" pagination="101 - 128" refId="ref92709" refString="O'Connor, B. (1997) Lower Miocene Radiolaria from the Kopua Point, Kaipara Harbour, New Zealand. Micropaleontology, 43 (2), 101 - 128. https: // doi. org / 10.2307 / 1485777" type="journal article" year="1997">OConnor, 1997</bibRefCitation>
)
</emphasis>
</taxonomicName>
<emphasis id="B9140254FFC1FFAA77CBE4EFFD2AC3CE" bold="true" box="[643,749,152,178]" pageId="82" pageNumber="83">
<taxonomicNameLabel id="A227BF2FFFC1FFAA77CBE4EFFD2AC3CE" box="[643,749,152,178]" pageId="82" pageNumber="83" rank="species">n. comb.</taxonomicNameLabel>
</emphasis>
</paragraph>
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<subSubSection id="C37A8DCDFFC1FFAA75DFE4CBFE91C3AA" box="[151,342,188,214]" pageId="82" pageNumber="83" type="description">
<paragraph id="8BDFDE46FFC1FFAA75DFE4CBFE91C3AA" blockId="82.[151,749,151,214]" box="[151,342,188,214]" pageId="82" pageNumber="83">
Plate 40,
<figureCitation id="135BC2C3FFC1FFAA7448E4CBFE94C3AA" box="[256,339,188,214]" captionStart="FIGURE 13" captionStartId="56.[151,250,1336,1360]" captionTargetBox="[169,1418,495,1309]" captionTargetId="figure-127@56.[162,1426,489,1312]" captionTargetPageId="56" captionText="FIGURE 13. Chart illustrating the differences between Lophophaena gozui n. sp. (orange diamonds) and L. ushionii n. sp. (green squares), in terms of cephalis height and the ratio between thorax and cephalis width. L. gozui has a shorter average cephalis height (40 microns) compared to L. ushionii (average cephalis height = 54 microns). In addition, the thorax is proportionally wider compared to the cephalis in L. gozui (average ratio of thorax:cephalis width = 2.5). L. ushionii has a proportionally narrower thorax (average ratio of thorax: cephalis width = 1.5)." figureDoi="http://doi.org/10.5281/zenodo.10544082" httpUri="https://zenodo.org/record/10544082/files/figure.png" pageId="82" pageNumber="83">Fig. 13</figureCitation>
.
</paragraph>
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<subSubSection id="C37A8DCDFFC1FFAA75DFE577FC73C22A" pageId="82" pageNumber="83" type="reference_group">
<paragraph id="8BDFDE46FFC1FFAA75DFE577FCD3C264" blockId="82.[151,977,256,342]" box="[151,788,256,280]" pageId="82" pageNumber="83">
<taxonomicName id="4C60A5C5FFC1FFAA75DFE577FD90C264" box="[151,599,256,280]" class="Polycystina" family="Plagiacanthidae" genus="Pseudodictyophimus" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="undefined-b">
<emphasis id="B9140254FFC1FFAA75DFE577FEAEC264" box="[151,361,256,280]" italics="true" pageId="82" pageNumber="83">Pseudodictyophimus</emphasis>
sp.
<bibRefCitation id="EFF1A3B7FFC1FFAA74DAE576FD90C264" author="O'Connor, B." box="[402,599,256,280]" pageId="82" pageNumber="83" refId="ref92673" refString="O'Connor, B. (1993) Radiolaria from the Mahurangi Limestone, Northland, New Zealand. M. Sc. thesis, Geology Department, University of Auckland, Auckland, 135 pp." type="book" year="1993">B, OConnor, 1993</bibRefCitation>
</taxonomicName>
, pl. 6, figs. 1415.
</paragraph>
<paragraph id="8BDFDE46FFC1FFAA75DFE568FC16C24B" blockId="82.[151,977,256,342]" box="[151,977,287,311]" pageId="82" pageNumber="83">
<taxonomicName id="4C60A5C5FFC1FFAA75DFE568FEBEC24B" baseAuthorityName="O'Connor" baseAuthorityYear="1997" box="[151,377,287,311]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="tekopua" status="sp. nov.">
<emphasis id="B9140254FFC1FFAA75DFE568FEBEC24B" box="[151,377,287,311]" italics="true" pageId="82" pageNumber="83">Lophophaena tekopua</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A227BF2FFFC1FFAA7437E568FE71C24A" box="[383,438,287,310]" pageId="82" pageNumber="83" rank="species">n. sp.</taxonomicNameLabel>
,
<bibRefCitation id="EFF1A3B7FFC1FFAA748AE568FDADC24B" author="O'Connor, B." box="[450,618,287,311]" pageId="82" pageNumber="83" pagination="101 - 128" refId="ref92709" refString="O'Connor, B. (1997) Lower Miocene Radiolaria from the Kopua Point, Kaipara Harbour, New Zealand. Micropaleontology, 43 (2), 101 - 128. https: // doi. org / 10.2307 / 1485777" type="journal article" year="1997">OConnor, 1997</bibRefCitation>
, pl. 2, figs. 1114, pl. 7, figs. 710.
</paragraph>
<paragraph id="8BDFDE46FFC1FFAA75DFE54AFC73C22A" blockId="82.[151,977,256,342]" box="[151,948,317,342]" pageId="82" pageNumber="83">
<taxonomicName id="4C60A5C5FFC1FFAA75DFE54AFD52C22A" authority="O'Connor, Kamikuri, 2019" authorityName="O'Connor, Kamikuri" authorityYear="2019" box="[151,661,317,342]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="tekopua">
<emphasis id="B9140254FFC1FFAA75DFE54AFEBEC229" box="[151,377,317,341]" italics="true" pageId="82" pageNumber="83">Lophophaena tekopua</emphasis>
OConnor,
<bibRefCitation id="EFF1A3B7FFC1FFAA74BAE54AFD52C22A" author="Kamikuri, S." box="[498,661,317,342]" pageId="82" pageNumber="83" pagination="163 - 194" refId="ref90498" refString="Kamikuri, S. (2019) Middle to late Miocene radiolarians from ODP Site 1021 in the eastern North Pacific. Bulletin of the Geological Survey of Japan, 70, 163 - 194." type="journal article" year="2019">Kamikuri, 2019</bibRefCitation>
</taxonomicName>
, pl. 14, fig. 14 (non fig. 13).
</paragraph>
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<subSubSection id="C37A8DCDFFC1FFAA75DFE5F3FEEAC032" pageId="82" pageNumber="83" type="discussion">
<paragraph id="8BDFDE46FFC1FFAA75DFE5F3FEEAC032" blockId="82.[151,1437,388,882]" pageId="82" pageNumber="83">
<emphasis id="B9140254FFC1FFAA75DFE5F3FECCC2E2" bold="true" box="[151,267,388,414]" pageId="82" pageNumber="83">Remarks.</emphasis>
This species is very similar to
<taxonomicName id="4C60A5C5FFC1FFAA77C5E5F3FC06C2E2" baseAuthorityName="Sugiyama and Furutani" baseAuthorityYear="1992" box="[653,961,388,414]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" order="Nassellaria" pageId="82" pageNumber="83" phylum="Sarcomastigophora" rank="species" species="cantharoides">
<emphasis id="B9140254FFC1FFAA77C5E5F3FC06C2E2" box="[653,961,388,414]" italics="true" pageId="82" pageNumber="83">Pelagomanes cantharoides</emphasis>
</taxonomicName>
(Pl. 40,
<figureCitation id="135BC2C3FFC1FFAA7179E5F3FB66C2E2" box="[1073,1185,388,414]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="6.[151,250,1340,1364]" captionStartId-1="7.[151,250,1538,1562]" captionStartId-2="9.[151,250,1885,1909]" captionTargetBox-0="[151,1430,743,1316]" captionTargetBox-1="[157,1428,745,1510]" captionTargetBox-2="[189,1398,1065,1862]" captionTargetId-0="figure-284@6.[151,1436,741,1317]" captionTargetId-1="figure-251@7.[151,1436,741,1514]" captionTargetId-2="figure-391@9.[189,1398,1065,1862]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionTargetPageId-2="9" captionText-0="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." captionText-1="FIGURE 2. Relative abundance of specimens and taxa belonging to Plagiacanthoidea and Lophophaenidae per sample, from IODP Site U1337 in the EEP over the last ~10 Ma. The relative percentages of total taxa are represented by the blue lines and the red lines represent the percentages of total specimens (triangles = Plagiacanthoidea; circles = Lophophaenidae). The Plagiacanthoidea was among the most abundant and speciose of radiolarian groups encountered in Trubovitz et al. (2020), with at least 10% of the specimens and 20% of the taxonomic richness in every sample. Within the Plagiacanthoidea, lophophaenids consistently made up the majority of specimens (~67%) and taxa (~61%). Data are from Trubovitz et al. (2020)." captionText-2="FIGURE 3. Locality map of IODP Site U1337, where all samples for this study were collected. Map from Google EarthTM." figureDoi-0="http://doi.org/10.5281/zenodo.10544060" figureDoi-1="http://doi.org/10.5281/zenodo.10544062" figureDoi-2="http://doi.org/10.5281/zenodo.10544064" httpUri-0="https://zenodo.org/record/10544060/files/figure.png" httpUri-1="https://zenodo.org/record/10544062/files/figure.png" httpUri-2="https://zenodo.org/record/10544064/files/figure.png" pageId="82" pageNumber="83">Figs. 13</figureCitation>
,
<figureCitation id="135BC2C3FFC1FFAA71FBE5F3FB18C2E2" box="[1203,1247,388,414]" captionStart-0="FIGURE 7" captionStart-1="FIGURE 8" captionStartId-0="18.[151,250,1317,1341]" captionStartId-1="24.[151,250,1090,1114]" captionTargetBox-0="[164,1422,562,1294]" captionTargetBox-1="[178,1411,181,1066]" captionTargetId-0="figure-182@18.[164,1422,561,1294]" captionTargetId-1="figure-22@24.[177,1411,181,1066]" captionTargetPageId-0="18" captionTargetPageId-1="24" captionText-0="FIGURE 7. Internal skeletal structure of Arachnocorys umbellifera (Haeckel, 1862) Petrushevskaya, 1971." captionText-1="FIGURE 8. Internal skeletal structure of Botryopera amabie n. sp." figureDoi-0="http://doi.org/10.5281/zenodo.10544072" figureDoi-1="http://doi.org/10.5281/zenodo.10544074" httpUri-0="https://zenodo.org/record/10544072/files/figure.png" httpUri-1="https://zenodo.org/record/10544074/files/figure.png" pageId="82" pageNumber="83">78</figureCitation>
,
<figureCitation id="135BC2C3FFC1FFAA71B8E5F3FAC9C2E2" box="[1264,1294,388,414]" captionStart="FIGURE 10" captionStartId="40.[151,250,1263,1287]" captionTargetBox="[164,1423,418,1240]" captionTargetId="figure-130@40.[164,1423,417,1240]" captionTargetPageId="40" captionText="FIGURE 10. Internal skeletal structure of Lithomelissa alkonost n. sp." figureDoi="http://doi.org/10.5281/zenodo.10544078" httpUri="https://zenodo.org/record/10544078/files/figure.png" pageId="82" pageNumber="83">10</figureCitation>
,
<figureCitation id="135BC2C3FFC1FFAA7057E5F3FAFBC2E2" box="[1311,1340,388,414]" captionStart="FIGURE 14" captionStartId="64.[151,250,1182,1206]" captionTargetBox="[169,1418,386,1151]" captionTargetId="figure-94@64.[163,1423,381,1159]" captionTargetPageId="64" captionText="FIGURE 14. Chart illustrating the different cephalis sizes in Lophophaena shishigae n. sp. and Arachnocorys jorogumoae n. sp. The yellow circles indicate L. shishigae, with a small cephalis, and the purple triangles represent A. jorogumoae, with a significantly larger and proportionally higher cephalis." figureDoi="http://doi.org/10.5281/zenodo.10544084" httpUri="https://zenodo.org/record/10544084/files/figure.png" pageId="82" pageNumber="83">14</figureCitation>
)
<emphasis id="B9140254FFC1FFAA7019E5F3FEFEC2BD" italics="true" pageId="82" pageNumber="83">
and
<taxonomicName id="4C60A5C5FFC1FFAA70C0E5F3FEFEC2BD" authorityName="Funakawa, Renaudie" authorityYear="2014" class="Polycystina" family="Plagiacanthidae" genus="Ceratocyrtis" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="morawanensis">P. morawanensis</taxonomicName>
</emphasis>
(Pl. 40,
<figureCitation id="135BC2C3FFC1FFAA74DFE5DFFE3FC2BD" box="[407,504,423,450]" captionStart="FIGURE 4" captionStartId="10.[151,250,1579,1603]" captionTargetBox="[164,1423,849,1556]" captionTargetId="figure-347@10.[164,1423,849,1556]" captionTargetPageId="10" captionText="FIGURE 4. Examples of specimen measurements discussed in the Systematic Paleontology section of this study." figureDoi="http://doi.org/10.5281/zenodo.10544066" httpUri="https://zenodo.org/record/10544066/files/figure.png" pageId="82" pageNumber="83">Figs. 4A</figureCitation>
<figureCitation id="135BC2C3FFC1FFAA7740E5D0FDD2C2BD" box="[520,533,423,449]" captionStart="FIGURE 6" captionStartId="15.[151,250,1463,1487]" captionTargetBox="[153,1436,737,1439]" captionTargetId="figure-139@15.[151,1436,736,1440]" captionTargetPageId="15" captionText="FIGURE 6. Internal skeletal structure of Arachnocorallium calvata (Haeckel, 1887) Petrushevskaya, 1971." figureDoi="http://doi.org/10.5281/zenodo.10544070" httpUri="https://zenodo.org/record/10544070/files/figure.png" pageId="82" pageNumber="83">6</figureCitation>
,
<figureCitation id="135BC2C3FFC1FFAA7769E5DFFDF6C2BE" box="[545,561,424,450]" captionStart="FIGURE 9" captionStartId="30.[151,250,1556,1580]" captionTargetBox="[152,1436,742,1532]" captionTargetId="figure-252@30.[151,1436,741,1532]" captionTargetPageId="30" captionText="FIGURE 9. Internal skeletal structure of Ceratocyrtis sp. C." figureDoi="http://doi.org/10.5281/zenodo.10544076" httpUri="https://zenodo.org/record/10544076/files/figure.png" pageId="82" pageNumber="83">9</figureCitation>
, 1112), and
<bibRefCitation id="EFF1A3B7FFC1FFAA778FE5DFFC4DC2BE" author="O'Connor, B." box="[711,906,424,450]" pageId="82" pageNumber="83" pagination="101 - 128" refId="ref92709" refString="O'Connor, B. (1997) Lower Miocene Radiolaria from the Kopua Point, Kaipara Harbour, New Zealand. Micropaleontology, 43 (2), 101 - 128. https: // doi. org / 10.2307 / 1485777" type="journal article" year="1997">OConnor (1997)</bibRefCitation>
provided no differential diagnosis between any of these species. The
<typeStatus id="54DB60E4FFC1FFAA74C4E5BBFE7BC29A" box="[396,444,460,486]" pageId="82" pageNumber="83">type</typeStatus>
material of this species appears to indicate a relatively larger cephalis:thorax ratio than either
<taxonomicName id="4C60A5C5FFC1FFAA7596E587FDCAC176" baseAuthorityName="Sugiyama and Furutani" baseAuthorityYear="1992" box="[222,525,496,522]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" order="Nassellaria" pageId="82" pageNumber="83" phylum="Sarcomastigophora" rank="species" species="cantharoides">
<emphasis id="B9140254FFC1FFAA7596E587FDCAC176" box="[222,525,496,522]" italics="true" pageId="82" pageNumber="83">Pelagomanes cantharoides</emphasis>
</taxonomicName>
or
<taxonomicName id="4C60A5C5FFC1FFAA777AE587FD28C175" authorityName="Funakawa, Renaudie" authorityYear="2014" box="[562,751,496,521]" class="Polycystina" family="Plagiacanthidae" genus="Ceratocyrtis" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="morawanensis">
<emphasis id="B9140254FFC1FFAA777AE587FD28C175" box="[562,751,496,521]" italics="true" pageId="82" pageNumber="83">P. morawanensis</emphasis>
</taxonomicName>
. However, the range of variability is not clear and we are not certain that this is indeed a separate species.
<bibRefCitation id="EFF1A3B7FFC1FFAA77EBE663FC88C152" author="Renaudie, J." box="[675,847,532,558]" pageId="82" pageNumber="83" refId="ref94207" refString="Renaudie, J. (2014) A synthesis of Antarctic Neogene radiolarians: taxonomy, macroevolution and biostratigraphy. PhD Thesis, Humboldt-Universitat zu Berlin, Berlin, 300 pp. https: // doi. org / 10.18452 / 16985" type="book" year="2014">Renaudie 2014</bibRefCitation>
considered
<taxonomicName id="4C60A5C5FFC1FFAA7694E663FA93C152" authority="O'Connor" authorityName="O'Connor" authorityYear="1997" box="[988,1364,532,558]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="tekopua">
<emphasis id="B9140254FFC1FFAA7694E663FB1EC152" box="[988,1241,532,558]" italics="true" pageId="82" pageNumber="83">Lophophaena tekopua</emphasis>
OConnor
</taxonomicName>
along with
<taxonomicName id="4C60A5C5FFC1FFAA7599E64FFE3AC12E" box="[209,509,568,594]" class="Polycystina" family="Plagiacanthidae" genus="Pseudodictyophimus" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="undefined-b">
<emphasis id="B9140254FFC1FFAA7599E64FFE7EC12E" box="[209,441,568,594]" italics="true" pageId="82" pageNumber="83">Pseudodictyophimus</emphasis>
sp. B
</taxonomicName>
OConnor to be conspecific with
<taxonomicName id="4C60A5C5FFC1FFAA76CAE64FFB15C12E" authority="(Funakawa)" baseAuthorityName="Funakawa" box="[898,1234,568,594]" class="Polycystina" family="Plagiacanthidae" genus="Ceratocyrtis" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="morawanensis">
<emphasis id="B9140254FFC1FFAA76CAE64FFB86C12D" box="[898,1089,568,593]" italics="true" pageId="82" pageNumber="83">P. morawanensis</emphasis>
(Funakawa)
</taxonomicName>
.
<bibRefCitation id="EFF1A3B7FFC1FFAA7196E64FFA5BC12E" author="Kamikuri, S." box="[1246,1436,568,594]" pageId="82" pageNumber="83" pagination="163 - 194" refId="ref90498" refString="Kamikuri, S. (2019) Middle to late Miocene radiolarians from ODP Site 1021 in the eastern North Pacific. Bulletin of the Geological Survey of Japan, 70, 163 - 194." type="journal article" year="2019">Kamikuri (2019)</bibRefCitation>
identified
<specimenCount id="9D6615CFFFC1FFAA7442E62BFE76C10A" box="[266,433,604,630]" pageId="82" pageNumber="83" type="generic">two specimens</specimenCount>
as
<taxonomicName id="4C60A5C5FFC1FFAA7490E62BFC88C10A" authority="O'Connor" authorityName="O'Connor" authorityYear="1997" box="[472,847,604,630]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="tekopua">
<emphasis id="B9140254FFC1FFAA7490E62BFD15C10A" box="[472,722,604,630]" italics="true" pageId="82" pageNumber="83">Lophophaena tekopua</emphasis>
OConnor
</taxonomicName>
; one of these is almost identical to the
<typeStatus id="54DB60E4FFC1FFAA7040E62BFAFFC10A" box="[1288,1336,604,630]" pageId="82" pageNumber="83">type</typeStatus>
material illustrated by OConnor, and the other appears to more closely match the original illustrations of
<taxonomicName id="4C60A5C5FFC1FFAA7190E6F7FA52C1E5" box="[1240,1429,640,665]" class="Polycystina" family="Plagiacanthidae" genus="Ceratocyrtis" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="morawanensis">
<emphasis id="B9140254FFC1FFAA7190E6F7FA52C1E5" box="[1240,1429,640,665]" italics="true" pageId="82" pageNumber="83">P. morawanensis</emphasis>
</taxonomicName>
. It is not evident to us whether these are two endpoints on a morphological continuum, or two distinct morphotypes that should be considered separate species. The group of taxa including
<taxonomicName id="4C60A5C5FFC1FFAA7683E6BFFBBEC19E" baseAuthorityName="Sugiyama and Furutani" baseAuthorityYear="1992" box="[971,1145,712,738]" family="Lophophaenidae" genus="Pelagomanes" higherTaxonomySource="GBIF" kingdom="Protozoa" pageId="82" pageNumber="83" phylum="Sarcomastigophora" rank="species" species="cantharoides">
<emphasis id="B9140254FFC1FFAA7683E6BFFBBEC19E" box="[971,1145,712,738]" italics="true" pageId="82" pageNumber="83">P. cantharoides</emphasis>
</taxonomicName>
,
<emphasis id="B9140254FFC1FFAA71C1E6BFFA89C19D" box="[1161,1358,712,737]" italics="true" pageId="82" pageNumber="83">
<taxonomicName id="4C60A5C5FFC1FFAA71C1E6BFFA8CC19D" box="[1161,1355,712,737]" class="Polycystina" family="Plagiacanthidae" genus="Ceratocyrtis" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="morawanensis">P. morawanensis</taxonomicName>
,
</emphasis>
and
<taxonomicName id="4C60A5C5FFC1FFAA70C0E6BFFF37C07A" authorityName="O'Connor" authorityYear="1997" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="82" pageNumber="83" phylum="Radiozoa" rank="species" species="tekopua">
<emphasis id="B9140254FFC1FFAA70C0E6BFFF37C07A" italics="true" pageId="82" pageNumber="83">P. tekopua</emphasis>
</taxonomicName>
will require further work to determine the consistent differences between species, if there are any. For the purposes of this manuscript, we are tentatively retaining the three separate species concepts until more material can be examined.
</paragraph>
</subSubSection>
<subSubSection id="C37A8DCDFFC1FFAA758FE72FFC7BC00E" box="[199,956,856,882]" pageId="82" pageNumber="83" type="biology_ecology">
<paragraph id="8BDFDE46FFC1FFAA758FE72FFC7BC00E" blockId="82.[151,1437,388,882]" box="[199,956,856,882]" pageId="82" pageNumber="83">
<emphasis id="B9140254FFC1FFAA758FE72FFEDDC00E" bold="true" box="[199,282,856,882]" pageId="82" pageNumber="83">Range.</emphasis>
Middle Miocene, EEP (
<tableCitation id="C6E2EBFDFFC1FFAA7760E72FFDBBC00E" box="[552,636,856,882]" captionStart="TABLE 1" captionStartId="84.[151,248,834,860]" captionTargetPageId="84" captionText="TABLE 1. Occurrence chart of Late NeogeneRecent lophophaenids in the EEP." httpUri="http://table.plazi.org/id/DF1F8ECEFFC7FFAC75DFE735FB46C021" pageId="82" pageNumber="83" tableUuid="DF1F8ECEFFC7FFAC75DFE735FB46C021">Table 1</tableCitation>
). Range poorly constrained.
</paragraph>
</subSubSection>
</treatment>
</document>