194 lines
20 KiB
XML
194 lines
20 KiB
XML
<document id="DC4C320CA42416F2FD074B5E18B9A396" ID-DOI="10.11646/zootaxa.5160.1.1" ID-ISSN="11755334" ID-Zenodo-Dep="10544058" ID-ZooBank="A9179C79-EE43-44E4-8723-919505500049" IM.bibliography_approvedBy="juliana" IM.illustrations_approvedBy="juliana" IM.materialsCitations_approvedBy="juliana" IM.metadata_approvedBy="juliana" IM.tables_approvedBy="juliana" IM.taxonomicNames_approvedBy="juliana" IM.treatmentCitations_approvedBy="juliana" IM.treatments_approvedBy="juliana" checkinTime="1656928390227" checkinUser="plazi" docAuthor="Trubovitz, Sarah, Renaudie, Johan, Lazarus, David & Noble, Paula" docDate="2022" docId="03C96F50FFA4FFC075DFE127FBD8C2AD" docLanguage="en" docName="zootaxa.5160.1.1.pdf" docOrigin="Zootaxa 5160 (1)" docSource="http://dx.doi.org/10.11646/zootaxa.5160.1.1" docStyle="DocumentStyle:8B0D3ECF822058C8413568C103B59429.6:Zootaxa.2001-2006.monograph" docStyleId="8B0D3ECF822058C8413568C103B59429" docStyleName="Zootaxa.2001-2006.monograph" docStyleVersion="6" docTitle="Lophophaena domovoi, n. sp." docType="treatment" docUuid="2C507BDD-D308-40B4-A308-4A8A9E987A94" docUuidSource="ZooBank" docVersion="4" lastPageNumber="57" masterDocId="FFF01728FF93FFF87548E477FFC7C37C" masterDocTitle="Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific" masterLastPageNumber="158" masterPageNumber="1" pageNumber="56" updateTime="1705931330204" updateUser="juliana" zenodo-license-document="CLOSED">
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<mods:titleInfo id="93D1CCCBD2EE5F0B6D1A65E4ECA37693">
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<mods:title id="0AA01C2A4478E09DCC56D23779132575">Late Neogene Lophophaenidae (Nassellaria, Radiolaria) from the eastern equatorial Pacific</mods:title>
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<mods:name id="4480B3AEC4A4C9A0C8924845E0F29520" type="personal">
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<mods:namePart id="ABFB71BEA88D4E7A2CC275F3ABF7D9DA">Trubovitz, Sarah</mods:namePart>
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<mods:namePart id="A104974C0B784E6AE5676AD56771E62E">Renaudie, Johan</mods:namePart>
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<mods:name id="2D8B4376B4DFB412A53C4A837FC1F5B1" type="personal">
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<mods:namePart id="5D5700875C63EBA57CB0312D22279332">Lazarus, David</mods:namePart>
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<mods:name id="F4121EB008F12B79653F575D6810DE72" type="personal">
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<mods:roleTerm id="607D5564FAAFEC4DB770CEC17801F661">Author</mods:roleTerm>
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<mods:namePart id="CA07DBFAD403C0E6C15CAD6AB326EFDC">Noble, Paula</mods:namePart>
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<mods:typeOfResource id="F32D9915FE94D765F53E9F146BB4FA9E">text</mods:typeOfResource>
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<mods:title id="ECEDFCF3C9430D6254ED86A68EA2FBFB">Zootaxa</mods:title>
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<mods:part id="F1B9B4C6A7D449797D9BACA779E69D45">
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<mods:date id="885BC8DA0859340BC9DA78F8D8233838">2022</mods:date>
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<mods:number id="A0FE026A7EDFCEC4571D922C8BB927C9">2022-07-04</mods:number>
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<mods:number id="2C2A36DBB93C1639638F5F0A4F468545">5160</mods:number>
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<mods:url id="41C0C1D5412E7E3BB86915F2568A7BFE">http://dx.doi.org/10.11646/zootaxa.5160.1.1</mods:url>
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<mods:classification id="8A8E445183393A3898A8F88B772BACAC">journal article</mods:classification>
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<mods:identifier id="E7AC3F1A96F5EC2234147A8269BEA3E9" type="DOI">10.11646/zootaxa.5160.1.1</mods:identifier>
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<mods:identifier id="3E94BB702AC1BB51BFF7B81EF92EE430" type="ZooBank">A9179C79-EE43-44E4-8723-919505500049</mods:identifier>
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</mods:mods>
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<treatment id="03C96F50FFA4FFC075DFE127FBD8C2AD" LSID="urn:lsid:zoobank.org:act:2C507BDD-D308-40B4-A308-4A8A9E987A94" httpUri="http://treatment.plazi.org/id/03C96F50FFA4FFC075DFE127FBD8C2AD" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56">
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<subSubSection id="C37A8DCDFFA4FFCF75DFE127FE28C616" box="[151,495,1360,1386]" pageId="55" pageNumber="56" type="nomenclature">
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<paragraph id="8BDFDE46FFA4FFCF75DFE127FE28C616" blockId="55.[151,495,1360,1422]" box="[151,495,1360,1386]" pageId="55" pageNumber="56">
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<taxonomicName id="4C60A5C5FFA4FFCF75DFE127FE62C616" authority="Trubovitz, Renaudie, Lazarus & Noble, 2022" box="[151,421,1360,1386]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="55" pageNumber="56" phylum="Radiozoa" rank="species" species="domovoi" status="sp. nov.">
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<emphasis id="B9140254FFA4FFCF75DFE127FE62C616" box="[151,421,1360,1386]" italics="true" pageId="55" pageNumber="56">Lophophaena domovoi</emphasis>
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</taxonomicName>
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<emphasis id="B9140254FFA4FFCF74E4E127FE28C616" bold="true" box="[428,495,1360,1386]" pageId="55" pageNumber="56">
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<taxonomicNameLabel id="A227BF2FFFA4FFCF74E4E127FE28C616" box="[428,495,1360,1386]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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||
</paragraph>
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</subSubSection>
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<subSubSection id="C37A8DCDFFA4FFCF75DFE103FC7AC692" pageId="55" pageNumber="56" type="description">
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<paragraph id="8BDFDE46FFA4FFCF75DFE103FE50C6F2" blockId="55.[151,495,1360,1422]" box="[151,407,1396,1422]" pageId="55" pageNumber="56">
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Plate 30,
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<figureCitation id="135BC2C3FFA4FFCF7448E103FE98C6F2" box="[256,351,1396,1422]" captionStart="FIGURE 1" captionStartId="6.[151,250,1340,1364]" captionTargetBox="[151,1430,743,1316]" captionTargetId="figure-284@6.[151,1436,741,1317]" captionTargetPageId="6" captionText="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." figureDoi="http://doi.org/10.5281/zenodo.10544060" httpUri="https://zenodo.org/record/10544060/files/figure.png" pageId="55" pageNumber="56">Figs. 1A</figureCitation>
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–
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<figureCitation id="135BC2C3FFA4FFCF7438E103FE54C6F2" box="[368,403,1396,1422]" captionStart="FIGURE 7" captionStartId="18.[151,250,1317,1341]" captionTargetBox="[164,1422,562,1294]" captionTargetId="figure-182@18.[164,1422,561,1294]" captionTargetPageId="18" captionText="FIGURE 7. Internal skeletal structure of Arachnocorys umbellifera (Haeckel, 1862) Petrushevskaya, 1971." figureDoi="http://doi.org/10.5281/zenodo.10544072" httpUri="https://zenodo.org/record/10544072/files/figure.png" pageId="55" pageNumber="56">7B</figureCitation>
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.
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</paragraph>
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<paragraph id="8BDFDE46FFA4FFCF75DFE1CFFC44C6AC" blockId="55.[151,957,1464,1519]" box="[151,899,1464,1488]" pageId="55" pageNumber="56">
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<uri id="FFF1D244FFA4FFCF75DFE1CFFC44C6AC" box="[151,899,1464,1488]" pageId="55" pageNumber="56">
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urn:lsid:zoobank.org:act:
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<uuid id="FFC6E493FFA4FFCF74C7E1CFFC44C6AC" box="[399,899,1464,1488]" pageId="55" pageNumber="56">2C507BDD-D308-40B4-A308-4A8A9E987A94</uuid>
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</uri>
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</paragraph>
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<paragraph id="8BDFDE46FFA4FFCF75DFE1A0FC7AC692" blockId="55.[151,957,1464,1519]" box="[151,957,1495,1519]" pageId="55" pageNumber="56">
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unknown plagonid group C sp 20,
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<bibRefCitation id="EFF1A3B7FFA4FFCF74BEE1A0FD13C693" author="Trubovitz, S. & Lazarus, D. & Renaudie, J. & Noble, P. J." box="[502,724,1495,1519]" pageId="55" pageNumber="56" pagination="1 - 10" refId="ref96304" refString="Trubovitz, S., Lazarus, D., Renaudie, J. & Noble, P. J. (2020) Marine plankton show threshold extinction response to Neogene climate change. Nature Communications, 11 (5069), 1 - 10. https: // doi. org / 10.1038 / s 41467 - 020 - 18879 - 7" type="journal article" year="2020">
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Trubovitz
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<emphasis id="B9140254FFA4FFCF7717E1A0FD5FC693" box="[607,664,1495,1519]" italics="true" pageId="55" pageNumber="56">et al.,</emphasis>
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2020
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</bibRefCitation>
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, supplementary data 7.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37A8DCDFFA4FFCF75DFE26AFA48C54B" box="[151,1423,1565,1591]" pageId="55" pageNumber="56" type="diagnosis">
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<paragraph id="8BDFDE46FFA4FFCF75DFE26AFA48C54B" blockId="55.[151,1437,1565,2023]" box="[151,1423,1565,1591]" pageId="55" pageNumber="56">
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||
<emphasis id="B9140254FFA4FFCF75DFE26AFED5C54B" bold="true" box="[151,274,1565,1591]" pageId="55" pageNumber="56">Diagnosis.</emphasis>
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<taxonomicName id="4C60A5C5FFA4FFCF7450E26AFE75C54B" authorityName="Ehrenberg, 1847 emend. Petrushevskaya" authorityYear="1971" box="[280,434,1565,1591]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="55" pageNumber="56" phylum="Radiozoa" rank="genus">
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<emphasis id="B9140254FFA4FFCF7450E26AFE75C54B" box="[280,434,1565,1591]" italics="true" pageId="55" pageNumber="56">Lophophaena</emphasis>
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</taxonomicName>
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with a curved, hook-like apical spine and a thin, conical horn on the top of the cephalis.
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C37A8DCDFFA4FFCF758FE236FE0AC44F" pageId="55" pageNumber="56" type="description">
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||
<paragraph id="8BDFDE46FFA4FFCF758FE236FE0AC44F" blockId="55.[151,1437,1565,2023]" pageId="55" pageNumber="56">
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<emphasis id="B9140254FFA4FFCF758FE236FE9EC527" bold="true" box="[199,345,1601,1627]" pageId="55" pageNumber="56">Description.</emphasis>
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The cephalis of this species has two spines. One originates from the apical spine and is curved upward, resembling a hook or thorn, and extends outward from the dorsal side of the cephalis, well below the apex. The second spine is conical with a pointed tip, extends from the apex of the cephalis, and is not related to any of the primary skeletal spines. Pores on the cephalis are small and round, and decrease in size from the base to the top of the cephalis. The thorax is wider than the cephalis to varying degrees, but not dramatically so. There are strong indentations on the thorax where the lateral and dorsal spines run along the shell wall before extending outside the thorax as short appendages.
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</paragraph>
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</subSubSection>
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<subSubSection id="C37A8DCDFFA4FFCF758FE34AFC75C4E3" pageId="55" pageNumber="56" type="discussion">
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<paragraph id="8BDFDE46FFA4FFCF758FE34AFC75C4E3" blockId="55.[151,1437,1565,2023]" pageId="55" pageNumber="56">
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<emphasis id="B9140254FFA4FFCF758FE34AFEFCC42B" bold="true" box="[199,315,1853,1879]" pageId="55" pageNumber="56">Remarks.</emphasis>
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Some specimens of this species can have few, or very small pores, such as the specimen in
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<figureCitation id="135BC2C3FFA4FFCF7076E34AFA5BC424" box="[1342,1436,1853,1880]" captionStart="FIGURE 5" captionStartId="12.[151,250,1242,1266]" captionTargetBox="[151,1436,520,1219]" captionTargetId="figure-153@12.[151,1436,520,1219]" captionTargetPageId="12" captionText="FIGURE 5. Internal skeletal structure of Amphiplecta acrostoma Haeckel, 1887." figureDoi="http://doi.org/10.5281/zenodo.10544068" httpUri="https://zenodo.org/record/10544068/files/figure.png" pageId="55" pageNumber="56">Figure 5</figureCitation>
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(seen from dorsal side). The size of this species is significantly smaller than
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<taxonomicName id="4C60A5C5FFA4FFCF76A3E316FB03C407" box="[1003,1220,1889,1915]" class="Polycystina" family="Plagiacanthidae" genus="Lophophaena" kingdom="Chromista" order="Nassellaria" pageId="55" pageNumber="56" phylum="Radiozoa" rank="species" species="ikota" status="sp. nov.">
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<emphasis id="B9140254FFA4FFCF76A3E316FB03C407" box="[1003,1220,1889,1915]" italics="true" pageId="55" pageNumber="56">Lophophaena ikota</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A227BF2FFFA4FFCF7182E316FAC1C407" box="[1226,1286,1889,1915]" pageId="55" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
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(Pl. 33,
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<figureCitation id="135BC2C3FFA4FFCF702DE316FF7FC4E3" captionStart="FIGURE 1" captionStartId="6.[151,250,1340,1364]" captionTargetBox="[151,1430,743,1316]" captionTargetId="figure-284@6.[151,1436,741,1317]" captionTargetPageId="6" captionText="FIGURE 1. Diagram illustrating the general skeletal morphology of Lophophaenidae, and terminology used in this manuscript. Dashed lines indicate spines that are not visible from the angle shown, as they extend into the skeleton. An example specimen of Lophophaena casperi n. sp. is included for comparison to the diagrams." figureDoi="http://doi.org/10.5281/zenodo.10544060" httpUri="https://zenodo.org/record/10544060/files/figure.png" pageId="55" pageNumber="56">Figs. 1A</figureCitation>
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–
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<figureCitation id="135BC2C3FFA4FFCF7581E3F2FF2CC4E3" box="[201,235,1925,1951]" captionStart="FIGURE 7" captionStartId="18.[151,250,1317,1341]" captionTargetBox="[164,1422,562,1294]" captionTargetId="figure-182@18.[164,1422,561,1294]" captionTargetPageId="18" captionText="FIGURE 7. Internal skeletal structure of Arachnocorys umbellifera (Haeckel, 1862) Petrushevskaya, 1971." figureDoi="http://doi.org/10.5281/zenodo.10544072" httpUri="https://zenodo.org/record/10544072/files/figure.png" pageId="55" pageNumber="56">7B</figureCitation>
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), and it also differs in that the apical spine is more pronounced.
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</paragraph>
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</subSubSection>
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||
<subSubSection id="C37A8DCDFFA4FFC0758FE3DEFC4EC23D" lastPageId="56" lastPageNumber="57" pageId="55" pageNumber="56" type="materials_examined">
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||
<paragraph id="8BDFDE46FFA4FFCF758FE3DEFD71C49B" blockId="55.[151,1437,1565,2023]" pageId="55" pageNumber="56">
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||
<emphasis id="B9140254FFA4FFCF758FE3DEFE77C4BF" bold="true" box="[199,432,1961,1987]" pageId="55" pageNumber="56">Material examined.</emphasis>
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18 specimens observed from samples 321-1337A-31X-6W, 4–6cm (Middle Miocene) and 321-1337D-23H-6, 134–137cm (Late Miocene).
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</paragraph>
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<paragraph id="8BDFDE46FFABFFC07586E4E0FB74C3CD" blockId="56.[151,1437,151,465]" box="[206,1203,151,177]" pageId="56" pageNumber="57">
|
||
<materialsCitation id="3B08D41BFFABFFC07586E4E0FB68C3CD" box="[206,1199,151,177]" pageId="56" pageNumber="57" specimenCode="321-1337D-23H-6" specimenCount="1" typeStatus="holotype">
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||
<emphasis id="B9140254FFABFFC07586E4E0FE85C3CD" bold="true" box="[206,322,151,177]" pageId="56" pageNumber="57">
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||
<typeStatus id="54DB60E4FFABFFC07586E4E0FEFAC3CD" box="[206,317,151,177]" pageId="56" pageNumber="57" type="holotype">Holotype</typeStatus>
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||
.
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</emphasis>
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Pl. 30, figs. 1A–C; sample
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<specimenCode id="DBC6763DFFABFFC07730E4E0FC8BC3CD" box="[632,844,151,177]" pageId="56" pageNumber="57">
|
||
321-1337D-23H-
|
||
<quantity id="4C9873A3FFABFFC07676E4E0FC18C3CD" box="[830,991,151,177]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="56" pageNumber="57" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">
|
||
6, 134–
|
||
<quantity id="4C9873A3FFABFFC076D8E4E0FC18C3CD" box="[912,991,151,177]" metricMagnitude="0" metricUnit="m" metricValue="1.3699999999999999" pageId="56" pageNumber="57" unit="cm" value="137.0">137cm</quantity>
|
||
</quantity>
|
||
</specimenCode>
|
||
; ECO-147; K30-4
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||
</materialsCitation>
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||
.
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||
</paragraph>
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||
<paragraph id="8BDFDE46FFABFFC0758FE4CCFC4EC23D" blockId="56.[151,1437,151,465]" pageId="56" pageNumber="57">
|
||
<emphasis id="B9140254FFABFFC0758FE4CCFE81C3A9" bold="true" box="[199,326,187,213]" pageId="56" pageNumber="57">
|
||
<typeStatus id="54DB60E4FFABFFC0758FE4CCFE86C3A9" box="[199,321,187,213]" pageId="56" pageNumber="57" type="paratype">Paratypes</typeStatus>
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||
.
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||
</emphasis>
|
||
(1) Pl. 30, fig. 2; sample 321-1337D-23H-
|
||
<quantity id="4C9873A3FFABFFC07675E4CCFC26C3A9" box="[829,993,187,213]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="56" pageNumber="57" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134–137cm</quantity>
|
||
; ECO-147; F8-1. (2) Pl. 30, figs. 3AB; sample 321-1337D-23H-
|
||
<quantity id="4C9873A3FFABFFC07492E4A8FDB9C385" box="[474,638,223,249]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="56" pageNumber="57" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134–137cm</quantity>
|
||
; ECO-146;
|
||
<collectionCode id="ED714683FFABFFC07640E4A9FCDAC385" box="[776,797,222,249]" name="Tavera, Department of Geology and Geophysics" pageId="56" pageNumber="57">T</collectionCode>
|
||
35-1. (3) Pl. 30, figs. 7A–B; sample 321-1337D-23H-
|
||
<quantity id="4C9873A3FFABFFC070C2E4A8FED8C261" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="56" pageNumber="57" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134–137cm</quantity>
|
||
; ECO-145; K25-2. (4) Pl. 30, fig. 5; sample 321-1337D-23H-
|
||
<quantity id="4C9873A3FFABFFC0769EE574FBB0C261" box="[982,1143,259,285]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="56" pageNumber="57" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134–137cm</quantity>
|
||
; ECO-146; M9-1. (5) [not figured] sample 321-1337D-23H-
|
||
<quantity id="4C9873A3FFABFFC0775CE550FD72C23D" box="[532,693,295,321]" metricMagnitude="-1" metricUnit="m" metricValue="7.1567" metricValueMax="13.7" metricValueMin="0.6134" pageId="56" pageNumber="57" unit="cm" value="71.567" valueMax="137.0" valueMin="6.134">6, 134–137cm</quantity>
|
||
; ECO-146; N37-3.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A8DCDFFABFFC0758FE53CFCEFC2F5" pageId="56" pageNumber="57" type="description">
|
||
<paragraph id="8BDFDE46FFABFFC0758FE53CFCEFC2F5" blockId="56.[151,1437,151,465]" pageId="56" pageNumber="57">
|
||
<emphasis id="B9140254FFABFFC0758FE53CFEB9C219" bold="true" box="[199,382,331,357]" pageId="56" pageNumber="57">Measurements.</emphasis>
|
||
Cephalis height 40–49 (45)μm; cephalis width 29–38 (35)μm; neck width 25–31 (28)μm; width at shoulders 40–57 (46)μm. Based on
|
||
<specimenCount id="9D6615CFFFABFFC077C0E519FCE4C2F5" box="[648,803,366,393]" pageId="56" pageNumber="57" type="generic">15 specimens</specimenCount>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A8DCDFFABFFC0758FE5E4FC36C2D1" box="[199,1009,403,429]" pageId="56" pageNumber="57" type="etymology">
|
||
<paragraph id="8BDFDE46FFABFFC0758FE5E4FC36C2D1" blockId="56.[151,1437,151,465]" box="[199,1009,403,429]" pageId="56" pageNumber="57">
|
||
<emphasis id="B9140254FFABFFC0758FE5E4FE89C2D1" bold="true" box="[199,334,403,429]" pageId="56" pageNumber="57">Etymology.</emphasis>
|
||
Named for the house spirit in Slavic folklore, the Domovoi.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C37A8DCDFFABFFC0758FE5C0FBD8C2AD" box="[199,1055,439,465]" pageId="56" pageNumber="57" type="biology_ecology">
|
||
<paragraph id="8BDFDE46FFABFFC0758FE5C0FBD8C2AD" blockId="56.[151,1437,151,465]" box="[199,1055,439,465]" pageId="56" pageNumber="57">
|
||
<emphasis id="B9140254FFABFFC0758FE5C0FEDDC2AD" bold="true" box="[199,282,439,465]" pageId="56" pageNumber="57">Range.</emphasis>
|
||
Middle—Late Miocene, EEP (
|
||
<tableCitation id="C6E2EBFDFFABFFC0773EE5C0FD0DC2AD" box="[630,714,439,465]" captionStart="TABLE 1" captionStartId="84.[151,248,834,860]" captionTargetPageId="84" captionText="TABLE 1. Occurrence chart of Late Neogene–Recent lophophaenids in the EEP." httpUri="http://table.plazi.org/id/DF1F8ECEFFC7FFAC75DFE735FB46C021" pageId="56" pageNumber="57" tableUuid="DF1F8ECEFFC7FFAC75DFE735FB46C021">Table 1</tableCitation>
|
||
). Lower limit not determined.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |