481 lines
71 KiB
XML
481 lines
71 KiB
XML
<document id="63F7548A51BBD392307CADFA6F54E71D" ID-CLB-Dataset="28029" ID-DOI="10.11646/zootaxa.4508.4.5" ID-GBIF-Dataset="15e2857b-23b4-4974-9813-8930dcd65a75" ID-ISSN="1175-5326" ID-Zenodo-Dep="2607544" ID-ZooBank="0CCD6063-4B3E-48E0-85D4-9A8015BEB9AC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553586415817" checkinUser="plazi" docAuthor="Ng, Peter K. L. & Grinang, Jongkar" docDate="2018" docId="03987A17FFDEFF96789EF1FA9E27F859" docLanguage="en" docName="zootaxa.4508.4.5.pdf" docOrigin="Zootaxa 4508 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Geosesarma larsi Ng & Grinang 2018, n. sp." docType="treatment" docVersion="8" lastPageNumber="573" masterDocId="FFA1026FFFDFFF927809F16D9A13FFF0" masterDocTitle="A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak" masterLastPageNumber="575" masterPageNumber="569" pageNumber="570" updateTime="1698713135093" updateUser="ExternalLinkService">
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<mods:mods id="BB900634ADC0E6280C86C246168150B1" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="4E0A5097EBE71701721DE5A4F7A5A98A">
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<mods:title id="479173C3B38EDE8744688C9C09E19E95">A new species of highland vampire crab (Crustacea: Brachyura: Sesarmidae: Geosesarma) from Serian, Sarawak</mods:title>
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<mods:name id="BD3E644995C71FBD68BDB48AFCCCEE6F" type="personal">
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<mods:roleTerm id="D2E5BF75843152C9C9B43C97A3156136">Author</mods:roleTerm>
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<mods:namePart id="4730C01F76FDEA51E1A6308036765CEE">Ng, Peter K. L.</mods:namePart>
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<mods:name id="53E35DCDE1A81A5BCB7446E5B5F554C4" type="personal">
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<mods:roleTerm id="87501921954901FA07331D1BDE0ED624">Author</mods:roleTerm>
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<mods:namePart id="862A1FA8797DF2141480B0939C7CBA85">Grinang, Jongkar</mods:namePart>
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<mods:typeOfResource id="49171286E0D15E2759C8C3FC8B74FEF4">text</mods:typeOfResource>
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<mods:title id="4D5F52590798F60068D7C016FEFB789A">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="12A5C7574926F986309C0D6CAFF5A13E">
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<mods:date id="A8E4D6782711A0AE99803449CEFF4C71">2018</mods:date>
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<mods:number id="123CDD01B7C999ECB6372AEF673C6A2E">2018-11-02</mods:number>
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<mods:number id="65F9884DA76A61F1914BB7880CE63332">4508</mods:number>
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<mods:number id="E96397AB16395F85D324A82C9227DF52">4</mods:number>
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<mods:identifier id="D634E277C704912478539DE08D676754" type="CLB-Dataset">28029</mods:identifier>
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<mods:identifier id="01C648317B66CD37E1B06E7980E8D54E" type="DOI">10.11646/zootaxa.4508.4.5</mods:identifier>
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<treatment id="03987A17FFDEFF96789EF1FA9E27F859" ID-DOI="http://doi.org/10.5281/zenodo.6494648" ID-GBIF-Taxon="154496068" ID-Zenodo-Dep="6494648" LSID="urn:lsid:plazi:treatment:03987A17FFDEFF96789EF1FA9E27F859" httpUri="http://treatment.plazi.org/id/03987A17FFDEFF96789EF1FA9E27F859" lastPageId="4" lastPageNumber="573" pageId="1" pageNumber="570">
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<subSubSection id="C32B988AFFDEFF93789EF1FA9BA1FF42" box="[151,434,151,178]" pageId="1" pageNumber="570" type="nomenclature">
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<paragraph id="8B8ECB01FFDEFF93789EF1FA9BA1FF42" blockId="1.[151,434,151,211]" box="[151,434,151,178]" pageId="1" pageNumber="570">
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<heading id="D0C67C6DFFDEFF93789EF1FA9BA1FF42" bold="true" box="[151,434,151,178]" fontSize="11" level="1" pageId="1" pageNumber="570" reason="1">
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<emphasis id="B9451713FFDEFF93789EF1FA9BA1FF42" bold="true" box="[151,434,151,178]" pageId="1" pageNumber="570">
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<taxonomicName id="4C31B082FFDEFF93789EF1FA9B74FF41" ID-CoL="97ec6452-b6f7-4951-b4f8-f18236a0b515" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[151,359,151,177]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="570" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
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<emphasis id="B9451713FFDEFF93789EF1FA9B74FF41" bold="true" box="[151,359,151,177]" italics="true" pageId="1" pageNumber="570">Geosesarma larsi</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A276AA68FFDEFF937966F1F59BA1FF42" box="[367,434,152,178]" pageId="1" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C32B988AFFDEFF93789EF1D79B01FF23" box="[151,274,186,211]" pageId="1" pageNumber="570" type="description">
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<paragraph id="8B8ECB01FFDEFF93789EF1D79B01FF23" blockId="1.[151,434,151,211]" box="[151,274,186,211]" pageId="1" pageNumber="570">
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(
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<figureCitation id="130AD784FFDEFF937896F1D79B1AFF23" box="[159,265,186,211]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStart-3="FIGURE 4" captionStartId-0="1.[151,250,1959,1981]" captionStartId-1="2.[151,250,1379,1401]" captionStartId-2="3.[151,250,1132,1154]" captionStartId-3="4.[151,250,1700,1722]" captionTargetBox-0="[339,1247,1039,1925]" captionTargetBox-1="[339,1247,193,1358]" captionTargetBox-2="[151,1435,193,1110]" captionTargetBox-3="[151,1435,661,1678]" captionTargetId-0="figure@1.[339,1247,1039,1925]" captionTargetId-1="figure@2.[339,1247,193,1358]" captionTargetId-2="figure@3.[151,1435,193,1110]" captionTargetId-3="figure@4.[151,1435,661,1678]" captionTargetPageId-0="1" captionTargetPageId-1="2" captionTargetPageId-2="3" captionTargetPageId-3="4" captionText-0="FIGURE 1. Geosesarma larsi n. sp., colour in life. A–D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); E, F, female (12.6 × 12.5 mm) (ZRC 2017.1297). Specimens from Gunung Ampungan. Photographs by P.Y.C. Ng." captionText-1="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." captionText-2="FIGURE 3. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, outer view of left chela; B, inner view of left chela; C, dorsal view of chela showing tubercles on dactylus; D, left fourth ambulatory leg; E, right fourth ambulatory leg; F, right third ambulatory leg." captionText-3="FIGURE 4. Geosesarma larsi n. sp. A–C, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); D, E, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A–C, male thoracic sternum and pleon; D, female pleon; E, female sternopleonal cavity showing vulvae." httpUri-0="https://zenodo.org/record/2607546/files/figure.png" httpUri-1="https://zenodo.org/record/2607548/files/figure.png" httpUri-2="https://zenodo.org/record/2607550/files/figure.png" httpUri-3="https://zenodo.org/record/2607552/files/figure.png" pageId="1" pageNumber="570">Figs. 1–4</figureCitation>
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)
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</paragraph>
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||
</subSubSection>
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<subSubSection id="C32B988AFFDEFF93789EF06F9828FE3F" pageId="1" pageNumber="570" type="materials_examined">
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<paragraph id="8B8ECB01FFDEFF93789EF06F9828FE3F" blockId="1.[151,1436,258,967]" pageId="1" pageNumber="570">
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<emphasis id="B9451713FFDEFF93789EF06F9B69FEEB" bold="true" box="[151,378,258,283]" pageId="1" pageNumber="570">Material examined</emphasis>
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.
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<materialsCitation id="3B59C15CFFDEFF937980F06F99DDFECF" ID-GBIF-Occurrence="2028139807" collectingDate="2017-06-21" collectionCode="ZRC" collectorName="J. Grinang & L. Fehlandt" country="Malaysia" elevation="600" location="Sarawak" pageId="1" pageNumber="570" specimenCode="ZRC 2017.1296" specimenCount="1" typeStatus="holotype">
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<typeStatus id="548A75A3FFDEFF937980F06F9BE5FEEB" box="[393,502,258,283]" pageId="1" pageNumber="570" type="holotype">Holotype</typeStatus>
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: male (13.6 ×
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<quantity id="4CC966E4FFDEFF937A92F06E9911FEEB" box="[667,770,259,283]" metricMagnitude="-2" metricUnit="m" metricValue="1.34" pageId="1" pageNumber="570" unit="mm" value="13.4">13.4 mm</quantity>
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) (
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||
<specimenCode id="DB97637AFFDEFF937B11F06E99C6FEEB" box="[792,981,258,283]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="570">ZRC 2017.1296</specimenCode>
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), Gunung Ampungan,
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||
<elevation id="001C2C32FFDEFF937CDFF06F9F5BFEEB" box="[1238,1352,258,283]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="570" unit="m" value="600.0">
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<quantity id="4CC966E4FFDEFF937CDFF06F9F0DFEEA" box="[1238,1310,258,283]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="570" unit="m" value="600.0">600 m</quantity>
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asl
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||
</elevation>
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||
, Serian Division,
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||
<collectingCountry id="F3268B91FFDEFF93790CF04B9B7BFECF" box="[261,360,294,319]" name="Malaysia" pageId="1" pageNumber="570">Sarawak</collectingCountry>
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||
, at night, coll.
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||
<collectorName id="26C4AED7FFDEFF937A07F04A9890FECF" box="[526,643,294,319]" pageId="1" pageNumber="570">J. Grinang</collectorName>
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||
&
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||
<collectorName id="26C4AED7FFDEFF937AAFF04A9939FECF" box="[678,810,294,319]" pageId="1" pageNumber="570">L. Fehlandt</collectorName>
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||
,
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||
<date id="FF8FEDC1FFDEFF937B3CF04A99DDFECF" box="[821,974,295,319]" pageId="1" pageNumber="570" value="2017-06-21">
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||
<collectingDate id="EFCB1429FFDEFF937B3CF04A99DDFECF" box="[821,974,295,319]" pageId="1" pageNumber="570" value="2017-06-21">21 June 2017</collectingDate>
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||
</date>
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||
</materialsCitation>
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||
.
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<materialsCitation id="3B59C15CFFDEFF937BD1F04A9908FE93" ID-GBIF-Occurrence="2028139806" collectionCode="ZRC" country="Malaysia" pageId="1" pageNumber="570" specimenCode="ZRC 2017.1297" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
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||
<typeStatus id="548A75A3FFDEFF937BD1F04A9E59FECF" box="[984,1098,295,319]" pageId="1" pageNumber="570" type="paratype">Paratypes</typeStatus>
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||
:
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||
<specimenCount id="9D370088FFDEFF937C5DF04A9EA6FECF" box="[1108,1205,294,319]" pageId="1" pageNumber="570" type="female">1 female</specimenCount>
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||
(14.5 ×
|
||
<quantity id="4CC966E4FFDEFF937D1DF04B9F6FFECF" box="[1300,1404,294,319]" metricMagnitude="-2" metricUnit="m" metricValue="1.35" pageId="1" pageNumber="570" unit="mm" value="13.5">13.5 mm</quantity>
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||
), 1 ex-ovigerous female (12.6 ×
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||
<quantity id="4CC966E4FFDEFF9379D4F0279851FE93" box="[477,578,330,355]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="570" unit="mm" value="12.5">12.5 mm</quantity>
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) (
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||
<specimenCode id="DB97637AFFDEFF937A50F0269900FE93" box="[601,787,331,355]" collectionCode="ZRC" country="Singapore" httpUri="http://grbio.org/cool/16z4-1tbx" name="Zoological Reference Collection, National University of Singapore" pageId="1" pageNumber="570">ZRC 2017.1297</specimenCode>
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)
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||
</materialsCitation>
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||
,
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||
<specimenCount id="9D370088FFDEFF937B2FF026996EFE93" box="[806,893,330,355]" pageId="1" pageNumber="570" type="male">3 males</specimenCount>
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||
(largest 13.4 × 13.0 mm) (SBC),
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||
<specimenCount id="9D370088FFDEFF937CF0F0269F49FE93" box="[1273,1370,330,355]" pageId="1" pageNumber="570" type="female">1 female</specimenCount>
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||
(14.1 ×
|
||
<quantity id="4CC966E4FFDEFF9378A4F0029B01FE77" box="[173,274,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="1.28" pageId="1" pageNumber="570" unit="mm" value="12.8">12.8 mm</quantity>
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) (SBC),
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||
<materialsCitation id="3B59C15CFFDEFF93797FF0029828FE3F" ID-GBIF-Occurrence="2028139805" collectingDate="2016-12-22" collectionCode="ZRC" collectorName="L. Fehlandt" country="Malaysia" elevation="600" location="Gunung Ampungan" municipality="Serian Division" pageId="1" pageNumber="570" specimenCount="4" specimenCount-female="4" typeStatus="paratype">
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<specimenCount id="9D370088FFDEFF93797FF0029BF1FE77" box="[374,482,366,391]" pageId="1" pageNumber="570" type="female">3 females</specimenCount>
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||
with young crabs (largest 12.7 ×
|
||
<quantity id="4CC966E4FFDEFF937B51F00299D3FE77" box="[856,960,367,391]" metricMagnitude="-2" metricUnit="m" metricValue="1.21" pageId="1" pageNumber="570" unit="mm" value="12.1">12.1 mm</quantity>
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||
), same data as holotype, male (13.5 ×
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||
<quantity id="4CC966E4FFDEFF937D62F0039AD0FE5A" metricMagnitude="-2" metricUnit="m" metricValue="1.25" pageId="1" pageNumber="570" unit="mm" value="12.5">12.5 mm</quantity>
|
||
(SBC),
|
||
<specimenCount id="9D370088FFDEFF93792AF0FE9B95FE5B" box="[291,390,402,427]" pageId="1" pageNumber="570" type="female">1 female</specimenCount>
|
||
(15.0 × 15.0 mm) (SBC),
|
||
<location id="8EEE9DDAFFDEFF937AB3F0FE998FFE5B" LSID="urn:lsid:plazi:treatment:03987A17FFDEFF96789EF1FA9E27F859:8EEE9DDAFFDEFF937AB3F0FE998FFE5B" box="[698,924,403,427]" country="Malaysia" municipality="Serian Division" name="Gunung Ampungan" pageId="1" pageNumber="570">Gunung Ampungan</location>
|
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,
|
||
<elevation id="001C2C32FFDEFF937BA3F0FF9E0EFE5B" box="[938,1053,402,427]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="570" unit="m" value="600.0">
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<quantity id="4CC966E4FFDEFF937BA3F0FF99E0FE5A" box="[938,1011,402,427]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="1" pageNumber="570" unit="m" value="600.0">600 m</quantity>
|
||
asl
|
||
</elevation>
|
||
,
|
||
<collectingMunicipality id="6BEA517BFFDEFF937C23F0FF9ECDFE5B" box="[1066,1246,402,427]" pageId="1" pageNumber="570">Serian Division</collectingMunicipality>
|
||
,
|
||
<collectingCountry id="F3268B91FFDEFF937CE2F0FF9F5DFE5B" box="[1259,1358,402,427]" name="Malaysia" pageId="1" pageNumber="570">Sarawak</collectingCountry>
|
||
, night, coll.
|
||
<collectorName id="26C4AED7FFDEFF9378C6F0DA9B40FE3F" box="[207,339,438,463]" pageId="1" pageNumber="570">L. Fehlandt</collectorName>
|
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,
|
||
<date id="FF8FEDC1FFDEFF937957F0DA9824FE3F" box="[350,567,438,463]" pageId="1" pageNumber="570" value="2016-12-22">
|
||
<collectingDate id="EFCB1429FFDEFF937957F0DA9824FE3F" box="[350,567,438,463]" pageId="1" pageNumber="570" value="2016-12-22">22 December 2016</collectingDate>
|
||
</date>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B988AFFDEFF9078CEF0B79B5AF992" lastPageId="2" lastPageNumber="571" pageId="1" pageNumber="570" type="diagnosis">
|
||
<paragraph id="8B8ECB01FFDEFF9378CEF0B79AAFFC37" blockId="1.[151,1436,258,967]" pageId="1" pageNumber="570">
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||
<emphasis id="B9451713FFDEFF9378CEF0B79B8EFE03" bold="true" box="[199,413,474,499]" pageId="1" pageNumber="570">Diagnosis of male</emphasis>
|
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. Carapace almost square, width to length ratio 1.01–1.07, lateral margins gently sinuous, subparallel (
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<figureCitation id="130AD784FFDEFF93792BF0939B8DFDE7" box="[290,414,510,535]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="1" pageNumber="570">Fig. 2A, C</figureCitation>
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); dorsal surface with well-defined regions, anterior regions covered with small rounded or flattened granules (
|
||
<figureCitation id="130AD784FFDEFF937974F34F9BCDFDCB" box="[381,478,546,571]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="1" pageNumber="570">Fig. 2C</figureCitation>
|
||
); front deflexed, with 2 broad subtruncate lobes, separated by shallow median concavity; postfrontal, protogastric lobe prominent, very sharp, strong (
|
||
<figureCitation id="130AD784FFDEFF937BF0F32B9E97FDAF" box="[1017,1156,582,607]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="1" pageNumber="570">Fig. 2C, D</figureCitation>
|
||
); external orbital tooth triangular, outer margins curving anteriorly, tooth directed obliquely laterally, clearly extending beyond lateral margin (
|
||
<figureCitation id="130AD784FFDEFF9378FEF3E39B28FD57" box="[247,315,654,679]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="1" pageNumber="570">Fig. 2</figureCitation>
|
||
A–C). Merus of third maxilliped subovate, subequal in length to ischium; exopod slender, without trace of flagellum (
|
||
<figureCitation id="130AD784FFDEFF937970F3DF9BCBFD3B" box="[377,472,690,715]" captionStart="FIGURE 5" captionStartId="5.[151,250,1298,1320]" captionTargetBox="[354,1237,222,1261]" captionTargetId="figure@5.[340,1248,193,1277]" captionTargetPageId="5" captionText="FIGURE 5. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, left third maxilliped (setae denuded except for exopod); B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (ventro-mesial view); F, distal part of left G1 (dorsal view); G, distal part of left G1 (dorso-mesial view); H, left G2. Structures denuded except for exopod on third maxilliped and G2. Scales: A = 1.0 mm, C, C, H = 0.5 mm; D–G = 0.25 mm." httpUri="https://zenodo.org/record/2607554/files/figure.png" pageId="1" pageNumber="570">Fig. 5A</figureCitation>
|
||
). Outer surface of palm of adult male covered with small rounded granules; inner surface with low median ridge; dorsal margin of dactylus with 11 or 12 tubercles (distal one smallest), each with corneous tip (
|
||
<figureCitation id="130AD784FFDEFF93793EF3979B68FCE3" box="[311,379,762,787]" captionStart="FIGURE 3" captionStartId="3.[151,250,1132,1154]" captionTargetBox="[151,1435,193,1110]" captionTargetId="figure@3.[151,1435,193,1110]" captionTargetPageId="3" captionText="FIGURE 3. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, outer view of left chela; B, inner view of left chela; C, dorsal view of chela showing tubercles on dactylus; D, left fourth ambulatory leg; E, right fourth ambulatory leg; F, right third ambulatory leg." httpUri="https://zenodo.org/record/2607550/files/figure.png" pageId="1" pageNumber="570">Fig. 3</figureCitation>
|
||
A–C). Ambulatory legs with relatively slender merus, with sharp subdistal spine on dorsal margin, surfaces slightly rugose (
|
||
<figureCitation id="130AD784FFDEFF937A05F2739882FCC6" box="[524,657,798,823]" captionStart="FIGURE 1" captionStartId="1.[151,250,1959,1981]" captionTargetBox="[339,1247,1039,1925]" captionTargetId="figure@1.[339,1247,1039,1925]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma larsi n. sp., colour in life. A–D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); E, F, female (12.6 × 12.5 mm) (ZRC 2017.1297). Specimens from Gunung Ampungan. Photographs by P.Y.C. Ng." httpUri="https://zenodo.org/record/2607546/files/figure.png" pageId="1" pageNumber="570">Figs. 1A, B</figureCitation>
|
||
,
|
||
<figureCitation id="130AD784FFDEFF937A94F27298BAFCC7" box="[669,681,799,823]" captionStart="FIGURE 3" captionStartId="3.[151,250,1132,1154]" captionTargetBox="[151,1435,193,1110]" captionTargetId="figure@3.[151,1435,193,1110]" captionTargetPageId="3" captionText="FIGURE 3. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, outer view of left chela; B, inner view of left chela; C, dorsal view of chela showing tubercles on dactylus; D, left fourth ambulatory leg; E, right fourth ambulatory leg; F, right third ambulatory leg." httpUri="https://zenodo.org/record/2607550/files/figure.png" pageId="1" pageNumber="570">3</figureCitation>
|
||
D–F). Male pleon triangular; somite 6 relatively broad, with gently convex lateral margins; telson triangular with convex lateral margins (
|
||
<figureCitation id="130AD784FFDEFF937BA3F22F99FFFCAB" box="[938,1004,834,859]" captionStart="FIGURE 4" captionStartId="4.[151,250,1700,1722]" captionTargetBox="[151,1435,661,1678]" captionTargetId="figure@4.[151,1435,661,1678]" captionTargetPageId="4" captionText="FIGURE 4. Geosesarma larsi n. sp. A–C, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); D, E, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A–C, male thoracic sternum and pleon; D, female pleon; E, female sternopleonal cavity showing vulvae." httpUri="https://zenodo.org/record/2607552/files/figure.png" pageId="1" pageNumber="570">Fig. 4</figureCitation>
|
||
A–C). G1 stout, subdistal part of outer margin (ventral view) with distinct sloping hump-like structure; corneous distal part elongate from lateral view, bent at angle of ca. 45° from longitudinal axis, spatuliform from marginal view, with small median cleft (
|
||
<figureCitation id="130AD784FFDEFF937D30F2E79F6AFC53" box="[1337,1401,906,931]" captionStart="FIGURE 5" captionStartId="5.[151,250,1298,1320]" captionTargetBox="[354,1237,222,1261]" captionTargetId="figure@5.[340,1248,193,1277]" captionTargetPageId="5" captionText="FIGURE 5. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, left third maxilliped (setae denuded except for exopod); B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (ventro-mesial view); F, distal part of left G1 (dorsal view); G, distal part of left G1 (dorso-mesial view); H, left G2. Structures denuded except for exopod on third maxilliped and G2. Scales: A = 1.0 mm, C, C, H = 0.5 mm; D–G = 0.25 mm." httpUri="https://zenodo.org/record/2607554/files/figure.png" pageId="1" pageNumber="570">Fig. 5</figureCitation>
|
||
B– G).
|
||
</paragraph>
|
||
<caption id="DF4E9B89FFDEFF93789EF6CA9E8CF82C" httpUri="https://zenodo.org/record/2607546/files/figure.png" pageId="1" pageNumber="570" startId="1.[151,250,1959,1981]" targetBox="[339,1247,1039,1925]" targetPageId="1">
|
||
<paragraph id="8B8ECB01FFDEFF93789EF6CA9E8CF82C" blockId="1.[151,1436,1959,2012]" pageId="1" pageNumber="570">
|
||
<emphasis id="B9451713FFDEFF93789EF6CA9B1CF84D" bold="true" box="[151,271,1959,1981]" pageId="1" pageNumber="570">FIGURE 1</emphasis>
|
||
.
|
||
<taxonomicName id="4C31B082FFDEFF937914F6C59BC1F84D" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[285,466,1959,1981]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="570" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDEFF937914F6C59BC1F84D" box="[285,466,1959,1981]" italics="true" pageId="1" pageNumber="570">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDEFF9379D2F6CA9804F84D" bold="true" box="[475,535,1959,1981]" pageId="1" pageNumber="570">
|
||
<taxonomicNameLabel id="A276AA68FFDEFF9379D2F6CA9804F84D" box="[475,535,1959,1981]" pageId="1" pageNumber="570" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, colour in life. A–D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); E, F, female (12.6 × 12.5 mm) (ZRC 2017.1297). Specimens from Gunung Ampungan. Photographs by P.Y.C. Ng.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF4E9B89FFDDFF90789EF40E9B93FA46" httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="2" pageNumber="571" startId="2.[151,250,1379,1401]" targetBox="[339,1247,193,1358]" targetPageId="2">
|
||
<paragraph id="8B8ECB01FFDDFF90789EF40E9B93FA46" blockId="2.[151,1436,1379,1462]" pageId="2" pageNumber="571">
|
||
<emphasis id="B9451713FFDDFF90789EF40E9B1FFA89" bold="true" box="[151,268,1379,1401]" pageId="2" pageNumber="571">FIGURE 2</emphasis>
|
||
.
|
||
<taxonomicName id="4C31B082FFDDFF907913F4099BDDFA89" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[282,462,1379,1401]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDDFF907913F4099BDDFA89" box="[282,462,1379,1401]" italics="true" pageId="2" pageNumber="571">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDDFF9079DCF40E981CFA89" bold="true" box="[469,527,1379,1401]" pageId="2" pageNumber="571">
|
||
<taxonomicNameLabel id="A276AA68FFDDFF9079DCF40E981CFA89" box="[469,527,1379,1401]" pageId="2" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8ECB01FFDDFF9078CEF4B39B5AF992" blockId="2.[151,1437,1502,2031]" pageId="2" pageNumber="571">
|
||
<emphasis id="B9451713FFDDFF9078CEF4B39B3AFA07" bold="true" box="[199,297,1502,1527]" pageId="2" pageNumber="571">Females</emphasis>
|
||
. In larger females, the carapace shape is relatively more trapezoidal, with the lateral margins more divergent towards posterior carapace margin (
|
||
<figureCitation id="130AD784FFDDFF907ABDF76F9901F9EA" box="[692,786,1538,1563]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="2" pageNumber="571">Fig. 2B</figureCitation>
|
||
). The female pleon is almost round, covering most of thoracic sternum (
|
||
<figureCitation id="130AD784FFDDFF90796CF74B9BD2F9CF" box="[357,449,1574,1599]" captionStart="FIGURE 4" captionStartId="4.[151,250,1700,1722]" captionTargetBox="[151,1435,661,1678]" captionTargetId="figure@4.[151,1435,661,1678]" captionTargetPageId="4" captionText="FIGURE 4. Geosesarma larsi n. sp. A–C, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); D, E, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A–C, male thoracic sternum and pleon; D, female pleon; E, female sternopleonal cavity showing vulvae." httpUri="https://zenodo.org/record/2607552/files/figure.png" pageId="2" pageNumber="571">Fig. 4D</figureCitation>
|
||
), and the vulva is ovate, positioned on the anterior half of sternite 6 with an opercular cover (
|
||
<figureCitation id="130AD784FFDDFF9078EDF7279B2FF992" box="[228,316,1610,1635]" captionStart="FIGURE 4" captionStartId="4.[151,250,1700,1722]" captionTargetBox="[151,1435,661,1678]" captionTargetId="figure@4.[151,1435,661,1678]" captionTargetPageId="4" captionText="FIGURE 4. Geosesarma larsi n. sp. A–C, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); D, E, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A–C, male thoracic sternum and pleon; D, female pleon; E, female sternopleonal cavity showing vulvae." httpUri="https://zenodo.org/record/2607552/files/figure.png" pageId="2" pageNumber="571">Fig. 4E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B988AFFDDFF9078CEF7039B12F8E7" pageId="2" pageNumber="571" type="description">
|
||
<paragraph id="8B8ECB01FFDDFF9078CEF7039B12F8E7" blockId="2.[151,1437,1502,2031]" pageId="2" pageNumber="571">
|
||
<emphasis id="B9451713FFDDFF9078CEF7039B9FF977" bold="true" box="[199,396,1646,1671]" pageId="2" pageNumber="571">Live colouration</emphasis>
|
||
. Adult
|
||
<taxonomicName id="4C31B082FFDDFF9079D6F702983DF976" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[479,558,1646,1670]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDDFF9079D6F702983DF976" box="[479,558,1646,1670]" italics="true" pageId="2" pageNumber="571">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDDFF907A31F7039864F977" bold="true" box="[568,631,1646,1671]" pageId="2" pageNumber="571">
|
||
<taxonomicNameLabel id="A276AA68FFDDFF907A31F7039864F977" box="[568,631,1646,1671]" pageId="2" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130AD784FFDDFF907A8FF70398D5F977" box="[646,710,1646,1671]" captionStart="FIGURE 1" captionStartId="1.[151,250,1959,1981]" captionTargetBox="[339,1247,1039,1925]" captionTargetId="figure@1.[339,1247,1039,1925]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma larsi n. sp., colour in life. A–D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); E, F, female (12.6 × 12.5 mm) (ZRC 2017.1297). Specimens from Gunung Ampungan. Photographs by P.Y.C. Ng." httpUri="https://zenodo.org/record/2607546/files/figure.png" pageId="2" pageNumber="571">Fig. 1</figureCitation>
|
||
) have the anterior half of the carapace, ambulatory meri, and the merus, carpus and palm of the cheliped purple to purplish-red; with the posterior carapace surface pink to reddishwhite, the ambulatory prodi and dactyli orange to red, the fingers of the chela orange. The ventral surfaces are a pale purplish-red to pink (
|
||
<figureCitation id="130AD784FFDDFF9079C9F7B79808F902" box="[448,539,1754,1779]" captionStart="FIGURE 1" captionStartId="1.[151,250,1959,1981]" captionTargetBox="[339,1247,1039,1925]" captionTargetId="figure@1.[339,1247,1039,1925]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma larsi n. sp., colour in life. A–D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); E, F, female (12.6 × 12.5 mm) (ZRC 2017.1297). Specimens from Gunung Ampungan. Photographs by P.Y.C. Ng." httpUri="https://zenodo.org/record/2607546/files/figure.png" pageId="2" pageNumber="571">Fig. 1D</figureCitation>
|
||
). The eyes are yellow to reddish-yellow (Fig. A). The eggs are reddish-orange (
|
||
<figureCitation id="130AD784FFDDFF907896F7939AE0F8E7" box="[159,243,1790,1815]" captionStart="FIGURE 1" captionStartId="1.[151,250,1959,1981]" captionTargetBox="[339,1247,1039,1925]" captionTargetId="figure@1.[339,1247,1039,1925]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma larsi n. sp., colour in life. A–D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); E, F, female (12.6 × 12.5 mm) (ZRC 2017.1297). Specimens from Gunung Ampungan. Photographs by P.Y.C. Ng." httpUri="https://zenodo.org/record/2607546/files/figure.png" pageId="2" pageNumber="571">Fig. 1F</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B988AFFDDFF9078CEF64F9B2EF8AF" pageId="2" pageNumber="571" type="etymology">
|
||
<paragraph id="8B8ECB01FFDDFF9078CEF64F9B2EF8AF" blockId="2.[151,1437,1502,2031]" pageId="2" pageNumber="571">
|
||
<emphasis id="B9451713FFDDFF9078CEF64F9B55F8CB" bold="true" box="[199,326,1826,1851]" pageId="2" pageNumber="571">Etymology</emphasis>
|
||
. The species is named after Lars Fehlandt who first observed this species and helped the authors get specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B988AFFDDFF9678CEF6079F03FEB0" lastPageId="4" lastPageNumber="573" pageId="2" pageNumber="571" type="discussion">
|
||
<paragraph id="8B8ECB01FFDDFF9078CEF6079AF3F81F" blockId="2.[151,1437,1502,2031]" pageId="2" pageNumber="571">
|
||
<emphasis id="B9451713FFDDFF9078CEF6079B27F873" bold="true" box="[199,308,1898,1923]" pageId="2" pageNumber="571">Remarks</emphasis>
|
||
. The purplish-red colour of
|
||
<taxonomicName id="4C31B082FFDDFF907A7CF607992FF872" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[629,828,1898,1922]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDDFF907A7CF607992FF872" box="[629,828,1898,1922]" italics="true" pageId="2" pageNumber="571">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDDFF907B4DF6079997F873" bold="true" box="[836,900,1898,1923]" pageId="2" pageNumber="571">
|
||
<taxonomicNameLabel id="A276AA68FFDDFF907B4DF6079997F873" box="[836,900,1898,1923]" pageId="2" pageNumber="571" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is not known in any other congener in Borneo. Morphologically,
|
||
<taxonomicName id="4C31B082FFDDFF907968F6E29BA6F856" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[353,437,1934,1958]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="571" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDDFF907968F6E29BA6F856" box="[353,437,1934,1958]" italics="true" pageId="2" pageNumber="571">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A276AA68FFDDFF9079B4F6E39BEDF857" box="[445,510,1934,1959]" pageId="2" pageNumber="571" rank="species">
|
||
<emphasis id="B9451713FFDDFF9079B4F6E39BEDF857" bold="true" box="[445,510,1934,1959]" pageId="2" pageNumber="571">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
belongs to a group in which the exopod of the third maxilliped has no trace of a flagellum, the G1is relatively stout and short, and whose members often occur at higher altitudes (see Ng 2015, 2017).
|
||
</paragraph>
|
||
<caption id="DF4E9B89FFDCFF91789EF50198CFFB4F" httpUri="https://zenodo.org/record/2607550/files/figure.png" pageId="3" pageNumber="572" startId="3.[151,250,1132,1154]" targetBox="[151,1435,193,1110]" targetPageId="3">
|
||
<paragraph id="8B8ECB01FFDCFF91789EF50198CFFB4F" blockId="3.[151,1436,1132,1215]" pageId="3" pageNumber="572">
|
||
<emphasis id="B9451713FFDCFF91789EF5019B1FFB72" bold="true" box="[151,268,1132,1154]" pageId="3" pageNumber="572">FIGURE 3</emphasis>
|
||
.
|
||
<taxonomicName id="4C31B082FFDCFF917911F5019BD9FB72" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[280,458,1132,1154]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDCFF917911F5019BD9FB72" box="[280,458,1132,1154]" italics="true" pageId="3" pageNumber="572">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDCFF9179D8F5019819FB72" bold="true" box="[465,522,1132,1154]" pageId="3" pageNumber="572">
|
||
<taxonomicNameLabel id="A276AA68FFDCFF9179D8F5019819FB72" box="[465,522,1132,1154]" pageId="3" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, outer view of left chela; B, inner view of left chela; C, dorsal view of chela showing tubercles on dactylus; D, left fourth ambulatory leg; E, right fourth ambulatory leg; F, right third ambulatory leg.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8ECB01FFDCFF9178CEF5869EFCF9B8" blockId="3.[151,1436,1259,2004]" pageId="3" pageNumber="572">
|
||
The western part of
|
||
<collectingCountry id="F3268B91FFDCFF9179A3F5869819FAF4" box="[426,522,1259,1284]" name="Malaysia" pageId="3" pageNumber="572">Sarawak</collectingCountry>
|
||
has two other species,
|
||
<taxonomicName id="4C31B082FFDCFF917B06F5819E78FAF4" authority="(Serene, 1968)" authorityName="Serene" authorityYear="1968" baseAuthorityName="Serene" baseAuthorityYear="1968" box="[783,1131,1259,1284]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="B9451713FFDCFF917B06F58199ACFAF4" box="[783,959,1260,1284]" italics="true" pageId="3" pageNumber="572">G. sarawakense</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA0B6F0FFDCFF917BC7F5819E70FAF4" author="Serene, R." box="[974,1123,1259,1284]" pageId="3" pageNumber="572" pagination="1084 - 1095" refId="ref3140" refString="Serene, R. (1968) Note preliminaire sur de nouvelles especes de Sesarma (Decapoda Brachyura). Bulletin du Museum naturelle d'Histoire national, Paris, Series 2, 39 (5), 1084 - 1095, pls. 1, 2." type="book chapter" year="1968">Serène, 1968</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C31B082FFDCFF917CABF5819AC1FAD8" authority="Ng & Grinang, 2004" authorityName="Ng & Grinang" authorityYear="2004" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="B9451713FFDCFF917CABF5819EF8FAF4" box="[1186,1259,1260,1284]" italics="true" pageId="3" pageNumber="572">G. bau</emphasis>
|
||
<bibRefCitation id="EFA0B6F0FFDCFF917CFAF5819AC1FAD8" author="Ng, P. K. L. & Grinang, J." pageId="3" pageNumber="572" pagination="299 - 325" refId="ref2864" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
, both from lowlands.
|
||
<taxonomicName id="4C31B082FFDCFF9179C0F47D9883FAD8" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[457,656,1296,1320]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDCFF9179C0F47D9883FAD8" box="[457,656,1296,1320]" italics="true" pageId="3" pageNumber="572">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDCFF917A91F46298CBFAD8" bold="true" box="[664,728,1295,1320]" pageId="3" pageNumber="572">
|
||
<taxonomicNameLabel id="A276AA68FFDCFF917A91F46298CBFAD8" box="[664,728,1295,1320]" pageId="3" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is immediately separated from
|
||
<taxonomicName id="4C31B082FFDCFF917C34F47D9EFDFAD8" baseAuthorityName="Serene" baseAuthorityYear="1968" box="[1085,1262,1296,1320]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="sarawakense">
|
||
<emphasis id="B9451713FFDCFF917C34F47D9EFDFAD8" box="[1085,1262,1296,1320]" italics="true" pageId="3" pageNumber="572">G. sarawakense</emphasis>
|
||
</taxonomicName>
|
||
by the absence of a flagellum on the third maxilliped exopod.
|
||
<taxonomicName id="4C31B082FFDCFF917AA5F4599960FABC" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[684,883,1332,1356]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDCFF917AA5F4599960FABC" box="[684,883,1332,1356]" italics="true" pageId="3" pageNumber="572">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDCFF917B72F45E99A8FABC" bold="true" box="[891,955,1331,1356]" pageId="3" pageNumber="572">
|
||
<taxonomicNameLabel id="A276AA68FFDCFF917B72F45E99A8FABC" box="[891,955,1331,1356]" pageId="3" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
more closely resembles
|
||
<taxonomicName id="4C31B082FFDCFF917CDCF4599F08FABC" authorityName="Ng & Grinang" authorityYear="2004" box="[1237,1307,1332,1356]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="bau">
|
||
<emphasis id="B9451713FFDCFF917CDCF4599F08FABC" box="[1237,1307,1332,1356]" italics="true" pageId="3" pageNumber="572">G. bau</emphasis>
|
||
</taxonomicName>
|
||
(from Bau and Kuching areas) in the squarish carapace shape, obliquely directed external orbital teeth and absence of a flagellum on the exopod of the third maxilliped, but can be distinguished by its more granular lateral carapace surfaces (
|
||
<figureCitation id="130AD784FFDCFF917905F4F29B44FA48" box="[268,343,1439,1464]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="3" pageNumber="572">Fig. 2</figureCitation>
|
||
A–C) (versus smoother lateral carapace surfaces;
|
||
<bibRefCitation id="EFA0B6F0FFDCFF917BB6F4CD9ED1FA48" author="Ng, P. K. L. & Grinang, J." box="[959,1218,1439,1464]" pageId="3" pageNumber="572" pagination="299 - 325" refId="ref2864" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: figs. 8, 9A, B); proportionately stouter and shorter ambulatory meri (
|
||
<figureCitation id="130AD784FFDCFF917AE4F4AE9961FA2B" box="[749,882,1475,1500]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="3" pageNumber="572">Figs. 2A, B</figureCitation>
|
||
,
|
||
<figureCitation id="130AD784FFDCFF917B77F4A99999FA2C" box="[894,906,1476,1500]" captionStart="FIGURE 3" captionStartId="3.[151,250,1132,1154]" captionTargetBox="[151,1435,193,1110]" captionTargetId="figure@3.[151,1435,193,1110]" captionTargetPageId="3" captionText="FIGURE 3. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, outer view of left chela; B, inner view of left chela; C, dorsal view of chela showing tubercles on dactylus; D, left fourth ambulatory leg; E, right fourth ambulatory leg; F, right third ambulatory leg." httpUri="https://zenodo.org/record/2607550/files/figure.png" pageId="3" pageNumber="572">3</figureCitation>
|
||
D–F) (versus proportionately more slender and longer ambulatory meri;
|
||
<bibRefCitation id="EFA0B6F0FFDCFF9179B9F4859884F9F0" author="Ng, P. K. L. & Grinang, J." box="[432,663,1511,1536]" pageId="3" pageNumber="572" pagination="299 - 325" refId="ref2864" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: fig. 8); and taxonomically significant, G1 being relatively short and stout with the distal corneous process bent at angle of 45° from the longitudinal axis (
|
||
<figureCitation id="130AD784FFDCFF917C53F7669F01F9D4" box="[1114,1298,1547,1572]" captionStart="FIGURE 5" captionStartId="5.[151,250,1298,1320]" captionTargetBox="[354,1237,222,1261]" captionTargetId="figure@5.[340,1248,193,1277]" captionTargetPageId="5" captionText="FIGURE 5. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, left third maxilliped (setae denuded except for exopod); B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (ventro-mesial view); F, distal part of left G1 (dorsal view); G, distal part of left G1 (dorso-mesial view); H, left G2. Structures denuded except for exopod on third maxilliped and G2. Scales: A = 1.0 mm, C, C, H = 0.5 mm; D–G = 0.25 mm." httpUri="https://zenodo.org/record/2607554/files/figure.png" pageId="3" pageNumber="572">Fig. 5B, C, D, F</figureCitation>
|
||
) (versus G1 slender and long, with corneous distal part bent at an angle of 30°;
|
||
<bibRefCitation id="EFA0B6F0FFDCFF917B8BF75D9E74F9B8" author="Ng, P. K. L. & Grinang, J." box="[898,1127,1583,1608]" pageId="3" pageNumber="572" pagination="299 - 325" refId="ref2864" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang 2004</bibRefCitation>
|
||
: fig. 9D, F).
|
||
</paragraph>
|
||
<paragraph id="8B8ECB01FFDCFF9678CEF7399882FF24" blockId="3.[151,1436,1259,2004]" lastBlockId="4.[151,1437,151,608]" lastPageId="4" lastPageNumber="573" pageId="3" pageNumber="572">
|
||
In its squarish male carapace, obliquely directed external orbital teeth, absence of a flagellum on the exopod of the third maxilliped and stout G1 with the corneous part directed at about 45°,
|
||
<taxonomicName id="4C31B082FFDCFF917BF7F7159ED7F960" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[1022,1220,1656,1680]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDCFF917BF7F7159ED7F960" box="[1022,1220,1656,1680]" italics="true" pageId="3" pageNumber="572">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDCFF917CC2F71A9F1AF960" bold="true" box="[1227,1289,1655,1680]" pageId="3" pageNumber="572">
|
||
<taxonomicNameLabel id="A276AA68FFDCFF917CC2F71A9F1AF960" box="[1227,1289,1655,1680]" pageId="3" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
most closely resembles
|
||
<taxonomicName id="4C31B082FFDCFF917904F7F1980BF944" authority="Ng, 2015" authorityName="Ng" authorityYear="2015" box="[269,536,1691,1716]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B9451713FFDCFF917904F7F19BB4F944" box="[269,423,1692,1716]" italics="true" pageId="3" pageNumber="572">G. ambawang</emphasis>
|
||
Ng, 2015
|
||
</taxonomicName>
|
||
, from Pontianak in West Kalimantan in Indonesian Borneo.
|
||
<taxonomicName id="4C31B082FFDCFF917CB0F7F19F6CF944" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[1209,1407,1692,1716]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDCFF917CB0F7F19F6CF944" box="[1209,1407,1692,1716]" italics="true" pageId="3" pageNumber="572">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDCFF917D8CF7F69AABF928" bold="true" pageId="3" pageNumber="572">
|
||
<taxonomicNameLabel id="A276AA68FFDCFF917D8CF7F69AABF928" pageId="3" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, however, can be distinguished from
|
||
<taxonomicName id="4C31B082FFDCFF917A6CF7AD9913F928" authorityName="Ng" authorityYear="2015" box="[613,768,1728,1752]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B9451713FFDCFF917A6CF7AD9913F928" box="[613,768,1728,1752]" italics="true" pageId="3" pageNumber="572">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
by the carapace surfaces with proportionately much fewer granules (
|
||
<figureCitation id="130AD784FFDCFF917903F78E9B76F90C" box="[266,357,1763,1788]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="3" pageNumber="572">Fig. 2C</figureCitation>
|
||
) (versus anterior part of the carapace being strongly granulated on the frontal, gastric and branchial surfaces, with prominent rounded granules in
|
||
<taxonomicName id="4C31B082FFDCFF917B0DF665998EF8D0" authorityName="Ng" authorityYear="2015" box="[772,925,1800,1824]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B9451713FFDCFF917B0DF665998EF8D0" box="[772,925,1800,1824]" italics="true" pageId="3" pageNumber="572">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
; Ng 2015: fig. 8B); the male pleonal somite 6 is proportionately broader (
|
||
<figureCitation id="130AD784FFDCFF9179C4F6469854F8B4" box="[461,583,1835,1860]" captionStart="FIGURE 4" captionStartId="4.[151,250,1700,1722]" captionTargetBox="[151,1435,661,1678]" captionTargetId="figure@4.[151,1435,661,1678]" captionTargetPageId="4" captionText="FIGURE 4. Geosesarma larsi n. sp. A–C, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); D, E, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A–C, male thoracic sternum and pleon; D, female pleon; E, female sternopleonal cavity showing vulvae." httpUri="https://zenodo.org/record/2607552/files/figure.png" pageId="3" pageNumber="572">Fig. 4A, B</figureCitation>
|
||
) (versus proportionately more narrow in
|
||
<taxonomicName id="4C31B082FFDCFF917C14F6419EABF8B4" authorityName="Ng" authorityYear="2015" box="[1053,1208,1836,1860]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B9451713FFDCFF917C14F6419EABF8B4" box="[1053,1208,1836,1860]" italics="true" pageId="3" pageNumber="572">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
; Ng 2015: fig. 8D), and the G1 is relatively shorter and stouter (
|
||
<figureCitation id="130AD784FFDCFF917A8CF62298EFF898" box="[645,764,1871,1896]" captionStart="FIGURE 5" captionStartId="5.[151,250,1298,1320]" captionTargetBox="[354,1237,222,1261]" captionTargetId="figure@5.[340,1248,193,1277]" captionTargetPageId="5" captionText="FIGURE 5. Geosesarma larsi n. sp., holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); Gunung Ampungan. A, left third maxilliped (setae denuded except for exopod); B, left G1 (ventral view); C, left G1 (dorsal view); D, distal part of left G1 (ventral view); E, distal part of left G1 (ventro-mesial view); F, distal part of left G1 (dorsal view); G, distal part of left G1 (dorso-mesial view); H, left G2. Structures denuded except for exopod on third maxilliped and G2. Scales: A = 1.0 mm, C, C, H = 0.5 mm; D–G = 0.25 mm." httpUri="https://zenodo.org/record/2607554/files/figure.png" pageId="3" pageNumber="572">Fig. 5B, C</figureCitation>
|
||
) (versus relatively more slender in
|
||
<taxonomicName id="4C31B082FFDCFF917C83F63D9F37F898" authorityName="Ng" authorityYear="2015" box="[1162,1316,1872,1896]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B9451713FFDCFF917C83F63D9F37F898" box="[1162,1316,1872,1896]" italics="true" pageId="3" pageNumber="572">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
; Ng 2015: fig. 9D, E). The colour of
|
||
<taxonomicName id="4C31B082FFDCFF9179CAF6199805F87C" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[451,534,1908,1932]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDCFF9179CAF6199805F87C" box="[451,534,1908,1932]" italics="true" pageId="3" pageNumber="572">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDCFF917A16F61E984CF87C" bold="true" box="[543,607,1907,1932]" pageId="3" pageNumber="572">
|
||
<taxonomicNameLabel id="A276AA68FFDCFF917A16F61E984CF87C" box="[543,607,1907,1932]" pageId="3" pageNumber="572" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is purplish-red on the anterior half of the carapace, ambulatory meri and chelipeds (
|
||
<figureCitation id="130AD784FFDCFF917918F6FA9B47F840" box="[273,340,1943,1968]" captionStart="FIGURE 1" captionStartId="1.[151,250,1959,1981]" captionTargetBox="[339,1247,1039,1925]" captionTargetId="figure@1.[339,1247,1039,1925]" captionTargetPageId="1" captionText="FIGURE 1. Geosesarma larsi n. sp., colour in life. A–D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); E, F, female (12.6 × 12.5 mm) (ZRC 2017.1297). Specimens from Gunung Ampungan. Photographs by P.Y.C. Ng." httpUri="https://zenodo.org/record/2607546/files/figure.png" pageId="3" pageNumber="572">Fig. 1</figureCitation>
|
||
), whereas
|
||
<taxonomicName id="4C31B082FFDCFF9179D8F6F5987FF840" authorityName="Ng" authorityYear="2015" box="[465,620,1944,1968]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="3" pageNumber="572" phylum="Arthropoda" rank="species" species="ambawang">
|
||
<emphasis id="B9451713FFDCFF9179D8F6F5987FF840" box="[465,620,1944,1968]" italics="true" pageId="3" pageNumber="572">G. ambawang</emphasis>
|
||
</taxonomicName>
|
||
is brown to reddish-brown on the anterior half of the carapace with the chelipeds and ambulatory legs reddish-brown to red (Ng 2015: fig. 10). The morphology of the carapace in larger females of
|
||
<taxonomicName id="4C31B082FFDBFF96791BF1F59B70FF40" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[274,355,152,176]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDBFF96791BF1F59B70FF40" box="[274,355,152,176]" italics="true" pageId="4" pageNumber="573">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDBFF967963F1FA9BBBFF40" bold="true" box="[362,424,151,176]" pageId="4" pageNumber="573">
|
||
<taxonomicNameLabel id="A276AA68FFDBFF967963F1FA9BBBFF40" box="[362,424,151,176]" pageId="4" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
(
|
||
<figureCitation id="130AD784FFDBFF9679BEF1FA981FFF40" box="[439,524,151,176]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="4" pageNumber="573">Fig. 2B</figureCitation>
|
||
) has also been reported for montane species like
|
||
<taxonomicName id="4C31B082FFDBFF967C27F1F59F0AFF40" authority="(Kemp, 1918)" authorityName="Kemp" authorityYear="1918" baseAuthorityName="Kemp" baseAuthorityYear="1918" box="[1070,1305,152,176]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="foxi">
|
||
<emphasis id="B9451713FFDBFF967C27F1F59E61FF40" box="[1070,1138,152,176]" italics="true" pageId="4" pageNumber="573">G. foxi</emphasis>
|
||
(
|
||
<bibRefCitation id="EFA0B6F0FFDBFF967C8DF1F59F02FF40" author="Kemp, S." box="[1156,1297,152,176]" pageId="4" pageNumber="573" pagination="217 - 297" refId="ref2453" refString="Kemp, S. (1918) Zoological results of a tour in the Far East. Crustacea Decapoda and Stomatopoda. Memoirs of the Asiatic Society of Bengal, 6, 217 - 297." type="journal article" year="1918">Kemp, 1918</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from Pulau Langkawi in Peninsular
|
||
<collectingCountry id="F3268B91FFDBFF9679A1F1D1981CFF25" box="[424,527,188,213]" name="Malaysia" pageId="4" pageNumber="573">Malaysia</collectingCountry>
|
||
(
|
||
<bibRefCitation id="EFA0B6F0FFDBFF967A14F1D19896FF24" author="Ng, P. K. L." box="[541,645,188,212]" pageId="4" pageNumber="573" pagination="226 - 242" refId="ref2796" refString="Ng, P. K. L. (2017) On the identities of the highland vampire crabs, Geosesarma foxi (Kemp, 1918) and G. serenei Ng, 1986, with description of a new phytotelmic species from Penang, Peninsular Malaysia (Crustacea: Decapoda: Brachyura: Sesarmidae). Raffles Bulletin of Zoology, 65, 226 - 242." type="journal article" year="2017">Ng 2017</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8B8ECB01FFDBFF9678CEF18D9F03FEB0" blockId="4.[151,1437,151,608]" pageId="4" pageNumber="573">
|
||
From
|
||
<taxonomicName id="4C31B082FFDBFF967902F18D987CFF08" authority="(De Man, 1902)" authorityName="De Man" authorityYear="1902" baseAuthorityName="De Man" baseAuthorityYear="1902" box="[267,623,224,248]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="gracillimum">
|
||
<emphasis id="B9451713FFDBFF967902F18D9BA0FF08" box="[267,435,224,248]" italics="true" pageId="4" pageNumber="573">G. gracillimum</emphasis>
|
||
(De Man, 1902)
|
||
</taxonomicName>
|
||
which occurs in lowlands in eastern
|
||
<collectingCountry id="F3268B91FFDBFF967C19F1B29E63FF08" box="[1040,1136,223,248]" name="Malaysia" pageId="4" pageNumber="573">Sarawak</collectingCountry>
|
||
and
|
||
<collectingCountry id="F3268B91FFDBFF967CA1F1B29EE4FF08" box="[1192,1271,223,248]" name="Brunei" pageId="4" pageNumber="573">Brunei</collectingCountry>
|
||
,
|
||
<taxonomicName id="4C31B082FFDBFF967D0AF18D9F46FF08" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[1283,1365,224,248]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDBFF967D0AF18D9F46FF08" box="[1283,1365,224,248]" italics="true" pageId="4" pageNumber="573">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDBFF967D55F1B29F8FFF08" bold="true" box="[1372,1436,223,248]" pageId="4" pageNumber="573">
|
||
<taxonomicNameLabel id="A276AA68FFDBFF967D55F1B29F8FFF08" box="[1372,1436,223,248]" pageId="4" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
can be easily be distinguished by its less quadrate carapace, proportionately stouter and shorter ambulatory legs, and relatively stouter G1 (
|
||
<figureCitation id="130AD784FFDBFF9679B5F04A981AFEB0" box="[444,521,295,320]" captionStart="FIGURE 2" captionStartId="2.[151,250,1379,1401]" captionTargetBox="[339,1247,193,1358]" captionTargetId="figure@2.[339,1247,193,1358]" captionTargetPageId="2" captionText="FIGURE 2. Geosesarma larsi n. sp. A, C, D, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); B, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A, B, overall habitus; C, dorsal view of carapace; D, frontal view of cephalothorax." httpUri="https://zenodo.org/record/2607548/files/figure.png" pageId="4" pageNumber="573">Figs. 2</figureCitation>
|
||
A–C, 3D–F, 5B–G) (cf.
|
||
<bibRefCitation id="EFA0B6F0FFDBFF967B11F045996EFEB0" author="Ng, P. K. L." box="[792,893,295,320]" pageId="4" pageNumber="573" pagination="57 - 72" refId="ref2677" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng 1995</bibRefCitation>
|
||
: fig. 13B–E; Ng 2015: fig. 14A, B).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C32B988AFFDBFF9678CEF0219E27F859" pageId="4" pageNumber="573" type="description">
|
||
<paragraph id="8B8ECB01FFDBFF9678CEF0219E93FE5D" blockId="4.[151,1437,151,608]" pageId="4" pageNumber="573">
|
||
<emphasis id="B9451713FFDBFF9678CEF0219B94FE95" bold="true" box="[199,391,332,357]" pageId="4" pageNumber="573">Ecological notes</emphasis>
|
||
.
|
||
<taxonomicName id="4C31B082FFDBFF96799CF021984EFE94" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[405,605,332,356]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDBFF96799CF021984EFE94" box="[405,605,332,356]" italics="true" pageId="4" pageNumber="573">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDBFF967A6DF02198B0FE95" bold="true" box="[612,675,332,357]" pageId="4" pageNumber="573">
|
||
<taxonomicNameLabel id="A276AA68FFDBFF967A6DF02198B0FE95" box="[612,675,332,357]" pageId="4" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
lives in shallow soil burrows on a steep slope under shaded canopy at relatively higher altitudes. Individuals were observed foraging at night on the ground among the bases of ferns. They were also found inside the phytotelms formed by the water-filled bases of the ferns.
|
||
</paragraph>
|
||
<paragraph id="8B8ECB01FFDBFF9678CEF0D59B17FD90" blockId="4.[151,1437,151,608]" pageId="4" pageNumber="573">
|
||
From habitats just lower down Gunung Ampungan,
|
||
<bibRefCitation id="EFA0B6F0FFDBFF967B49F0DA9E5BFE20" author="Grinang, J. & Ng, P. K. L." box="[832,1096,439,464]" pageId="4" pageNumber="573" pagination="564 - 582" refId="ref2391" refString="Grinang, J. & Ng, P. K. L. (2015) Taxonomy of the semiterrestrial crab Lepidothelphusa cognettii (Nobili, 1903) (Crustacea: Decapoda: Brachyura: Gecarcinucidae), with descriptions of five new species from Sarawak, Malaysia, Borneo. Raffles Bulletin of Zoology, 63, 564 - 582." type="journal article" year="2015">Grinang & Ng (2015)</bibRefCitation>
|
||
described a new species of semiterrestrial gecarcinucid crab,
|
||
<taxonomicName id="4C31B082FFDBFF967A2DF0B1991AFE04" authorityName="Grinang & Ng" authorityYear="2015" box="[548,777,476,500]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="loi">
|
||
<emphasis id="B9451713FFDBFF967A2DF0B1991AFE04" box="[548,777,476,500]" italics="true" pageId="4" pageNumber="573">Lepidothelphusa loi</emphasis>
|
||
</taxonomicName>
|
||
, from around
|
||
<quantity id="4CC966E4FFDBFF967BB2F0B19E1BFE04" box="[955,1032,476,501]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="4" pageNumber="573" unit="m" value="500.0">500 m</quantity>
|
||
asl. The aquatic potamid
|
||
<taxonomicName id="4C31B082FFDBFF967D4FF0B19BE4FDE8" authority="Ng & Grinang, 2004" authorityName="Ng & Grinang" authorityYear="2004" class="Malacostraca" family="Potamidae" genus="Ibanum" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="pilimanus">
|
||
<emphasis id="B9451713FFDBFF967D4FF0B19B16FDE8" italics="true" pageId="4" pageNumber="573">Ibanum pilimanus</emphasis>
|
||
<bibRefCitation id="EFA0B6F0FFDBFF967905F36D9BE4FDE8" author="Ng, P. K. L. & Grinang, J." box="[268,503,511,536]" pageId="4" pageNumber="573" pagination="299 - 325" refId="ref2864" refString="Ng, P. K. L. & Grinang, J. (2004) Decapod crustaceans with descriptions of three new species. In: Yong, H. S., Ng, F. S. P. & Yen, E. E. L. (Eds.), Sarawak Bau Limestone Biodiversity. Sarawak Museum Journal, New Series, 59 (80), Special Issue No. 6, pp. 299 - 325." type="journal article" year="2004">Ng & Grinang, 2004</bibRefCitation>
|
||
</taxonomicName>
|
||
, is also present, as is a potentially new species of gecarcinucid (genus
|
||
<taxonomicName id="4C31B082FFDBFF967D02F36D9AC1FDCD" authority="Ng, 1995" authorityName="Ng" authorityYear="1995" class="Malacostraca" family="Gecarcinucidae" genus="Bakousa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B9451713FFDBFF967D02F36D9F7FFDE8" box="[1291,1388,512,536]" italics="true" pageId="4" pageNumber="573">Bakousa</emphasis>
|
||
<bibRefCitation id="EFA0B6F0FFDBFF967D7BF36D9AC1FDCD" author="Ng, P. K. L." pageId="4" pageNumber="573" pagination="57 - 72" refId="ref2677" refString="Ng, P. K. L. (1995) On one new genus and three new species of freshwater crabs (Crustacea: Decapoda: Brachyura: Potamidae and Grapsidae) from Lanjak-Entimau, Sarawak, East Malaysia, Borneo. Zoologische Mededelingen, 69 (5), 57 - 72." type="journal article" year="1995">Ng, 1995</bibRefCitation>
|
||
</taxonomicName>
|
||
), which was also collected with
|
||
<taxonomicName id="4C31B082FFDBFF967A35F348986BFDCC" authorityName="Grinang & Ng" authorityYear="2015" box="[572,632,548,572]" class="Malacostraca" family="Gecarcinucidae" genus="Lepidothelphusa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="loi">
|
||
<emphasis id="B9451713FFDBFF967A35F348986BFDCC" box="[572,632,548,572]" italics="true" pageId="4" pageNumber="573">L. loi</emphasis>
|
||
</taxonomicName>
|
||
. At this site, however, no other species of
|
||
<emphasis id="B9451713FFDBFF967C5AF3499ECCFDCB" box="[1107,1247,548,571]" italics="true" pageId="4" pageNumber="573">Geosesarma</emphasis>
|
||
was collected or observed.
|
||
</paragraph>
|
||
<caption id="DF4E9B89FFDBFF96789EF7C9984FF907" httpUri="https://zenodo.org/record/2607552/files/figure.png" pageId="4" pageNumber="573" startId="4.[151,250,1700,1722]" targetBox="[151,1435,661,1678]" targetPageId="4">
|
||
<paragraph id="8B8ECB01FFDBFF96789EF7C9984FF907" blockId="4.[151,1435,1700,1783]" pageId="4" pageNumber="573">
|
||
<emphasis id="B9451713FFDBFF96789EF7C99B1EF94A" bold="true" box="[151,269,1700,1722]" pageId="4" pageNumber="573">FIGURE 4</emphasis>
|
||
.
|
||
<taxonomicName id="4C31B082FFDBFF967912F7C89BDDF94A" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[283,462,1700,1722]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDBFF967912F7C89BDDF94A" box="[283,462,1700,1722]" italics="true" pageId="4" pageNumber="573">Geosesarma larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDBFF9679DFF7C99803F94A" bold="true" box="[470,528,1700,1722]" pageId="4" pageNumber="573">
|
||
<taxonomicNameLabel id="A276AA68FFDBFF9679DFF7C99803F94A" box="[470,528,1700,1722]" pageId="4" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
A–C, holotype male (13.6 × 13.4 mm) (ZRC 2017.1296); D, E, paratype female (14.5 × 13.5 mm) (ZRC 2017.1297); specimens from Gunung Ampungan. A–C, male thoracic sternum and pleon; D, female pleon; E, female sternopleonal cavity showing vulvae.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B8ECB01FFDBFF9678CEF6499E27F859" blockId="4.[151,1436,1828,1961]" pageId="4" pageNumber="573">
|
||
<emphasis id="B9451713FFDBFF9678CEF6499BA3F8CD" bold="true" box="[199,432,1828,1853]" pageId="4" pageNumber="573">Conservation notes</emphasis>
|
||
. From the conservation perspective,
|
||
<taxonomicName id="4C31B082FFDBFF967B6AF64999ABF8CC" authority="Ng & Grinang, 2018" authorityName="Ng & Grinang" authorityYear="2018" box="[867,952,1828,1852]" class="Malacostraca" family="Sesarmidae" genus="Geosesarma" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="4" pageNumber="573" phylum="Arthropoda" rank="species" species="larsi" status="sp. nov.">
|
||
<emphasis id="B9451713FFDBFF967B6AF64999ABF8CC" box="[867,952,1828,1852]" italics="true" pageId="4" pageNumber="573">G. larsi</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B9451713FFDBFF967BCDF6499E15F8CD" bold="true" box="[964,1030,1828,1853]" pageId="4" pageNumber="573">
|
||
<taxonomicNameLabel id="A276AA68FFDBFF967BCDF6499E15F8CD" box="[964,1030,1828,1853]" pageId="4" pageNumber="573" rank="species">n. sp.</taxonomicNameLabel>
|
||
</emphasis>
|
||
is a species of concern because it appears to be confined to a relatively small highland area. The habitat is not protected and has a high chance of being degraded if adjacent agricultural activities such as oil palm plantations and orchards are expanded. The area will need to be more thoroughly surveyed to see if the species is more widespread.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |