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<document id="BAA0E474C34A292A6AB9523F9A0ABA1F" ID-DOI="10.3853/j.0067-1975.63.2011.1581" ID-ISSN="2201-4349" ID-Zenodo-Dep="5239547" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1629575327011" checkinUser="felipe" docAuthor="Köhler, Frank" docDate="2011" docId="0384503AFFB73C50FC65FE42FDA46935" docLanguage="en" docName="RecAustMus.63.2.167-202.pdf" docOrigin="Records of the Australian Museum 63 (2)" docSource="http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581" docStyle="DocumentStyle:7863F7301B7BEE922F541F6F9B5AEEC6.2:RecAustMus.2008-2018.journal_article.0cover" docStyleId="7863F7301B7BEE922F541F6F9B5AEEC6" docStyleName="RecAustMus.2008-2018.journal_article.0cover" docStyleVersion="2" docTitle="Amplirhagada vialae Köhler 2011, n.sp." docType="treatment" docUuid="3B6C8C66-39D7-4059-ACA5-0197BDDE7D7C" docUuidSource="ZooBank" docVersion="3" lastPageNumber="180" masterDocId="FFBD2842FFBB3C5EFFB8FFD2FFED6E4A" masterDocTitle="Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)" masterLastPageNumber="202" masterPageNumber="167" pageNumber="179" updateTime="1699078749944" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="6E048DAB43E566655A3C8645D0426E05">Descriptions of New Species of the Diverse and Endemic Land Snail Amplirhagada Iredale, 1933 from Rainforest Patches across the Kimberley, Western Australia (Pulmonata: Camaenidae)</mods:title>
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<mods:name id="AC4C3F3F3B1EA5AC3C1CBCB3A995F5F1" type="personal">
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<mods:namePart id="49E7CC7DCD3E8045F6136992A3605D1A">Köhler, Frank</mods:namePart>
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<mods:title id="6F6CD8E12D439E4E262CDE20C884A07A">Records of the Australian Museum</mods:title>
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<mods:date id="3FB6D075B2E4B4A70C4D6BBAD01B0E14">2011</mods:date>
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<mods:number id="81E88C4E176814B9ECE2D730122E3395">2011-11-30</mods:number>
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<mods:url id="875F5D10F9FA8B2E62F3C04C9CD86F1B">http://dx.doi.org/10.3853/j.0067-1975.63.2011.1581</mods:url>
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<mods:classification id="4F22BA5667ECFFC2BD256ABE5B17C71F">journal article</mods:classification>
<mods:identifier id="94968B88D5829A7938360EFE5A272006" type="DOI">10.3853/j.0067-1975.63.2011.1581</mods:identifier>
<mods:identifier id="220612BAA6D9284FA8D917E6AA8E2B62" type="ISSN">2201-4349</mods:identifier>
<mods:identifier id="3A12EC89865309EEB50D5DAF3A1028FC" type="Zenodo-Dep">5239547</mods:identifier>
<mods:identifier id="91EE04DC8CC844C9B68A4E089C445827" type="ZooBank">1BCD4085-D2B9-400D-B504-8C85C30303D6</mods:identifier>
</mods:mods>
<treatment id="0384503AFFB73C50FC65FE42FDA46935" LSID="urn:lsid:zoobank.org:act:3B6C8C66-39D7-4059-ACA5-0197BDDE7D7C" httpUri="http://treatment.plazi.org/id/0384503AFFB73C50FC65FE42FDA46935" lastPageId="14" lastPageNumber="180" pageId="12" pageNumber="179">
<subSubSection id="C337B2A7FFB73C52FC65FE42FAC96FE0" box="[989,1316,400,426]" pageId="12" pageNumber="179" type="nomenclature">
<paragraph id="8B92E12CFFB73C52FC65FE42FAC96FE0" blockId="12.[989,1316,400,426]" box="[989,1316,400,426]" pageId="12" pageNumber="179">
<heading id="D0DA5640FFB73C52FC65FE42FAC96FE0" box="[989,1316,400,426]" centered="true" fontSize="11" level="2" pageId="12" pageNumber="179" reason="2">
<emphasis id="B9593D3EFFB73C52FC65FE42FAC96FE0" bold="true" box="[989,1316,400,426]" pageId="12" pageNumber="179">
<taxonomicName id="4C2D9AAFFFB73C52FC65FE42FB0D6FE0" ID-CoL="66GSP" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[989,1248,400,426]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="179" phylum="Mollusca" rank="species" species="vialae" status="sp. nov.">
<emphasis id="B9593D3EFFB73C52FC65FE42FB0D6FE0" bold="true" box="[989,1248,400,426]" italics="true" pageId="12" pageNumber="179">Amplirhagada vialae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFB73C52FB50FE42FAC96FE0" box="[1256,1316,400,426]" pageId="12" pageNumber="179" rank="species">n.sp.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB73C52FCFFFE15FBF86D4E" pageId="12" pageNumber="179" type="materials_examined">
<paragraph id="8B92E12CFFB73C52FCFFFE15FB7C6C7D" blockId="12.[839,1467,455,567]" pageId="12" pageNumber="179">
<materialsCitation id="3B45EB71FFB73C52FCFFFE15FB616C7D" collectingDate="1987-06-05" collectionCode="V" collectorName="NW Kimberley" latitude="-14.620833" location="Savage Hill" longLatPrecision="21" longitude="125.254166" pageId="12" pageNumber="179" specimenCount="1" stateProvince="Western Australia" typeStatus="holotype">
<emphasis id="B9593D3EFFB73C52FCFFFE15FC386F95" bold="true" box="[839,981,455,479]" pageId="12" pageNumber="179">
<typeStatus id="54965F8EFFB73C52FCFFFE15FC926F95" box="[839,895,455,479]" pageId="12" pageNumber="179">Type</typeStatus>
locality
</emphasis>
.
<collectingRegion id="49E92FCEFFB73C52FC58FE1AFB716F95" box="[992,1180,456,479]" country="Australia" name="Western Australia" pageId="12" pageNumber="179">Western Australia</collectingRegion>
,
<collectorName id="26D884FAFFB73C52FB1CFE1AFAAB6F95" box="[1188,1350,456,479]" pageId="12" pageNumber="179">NW Kimberley</collectorName>
, Bonaparte Archipelago, Montague Sound, Scott Strait,
<quantity id="4CD54CC9FFB73C52FA8AFE37FA6D6FB6" box="[1330,1408,485,508]" metricMagnitude="3" metricUnit="m" metricValue="8.3" pageId="12" pageNumber="179" unit="km" value="8.3">8.3 km</quantity>
E of
<location id="8EF2B7F7FFB73C52FCFFFDD0FC256C53" LSID="urn:lsid:plazi:treatment:0384503AFFB73C50FC65FE42FDA46935:8EF2B7F7FFB73C52FCFFFDD0FC256C53" box="[839,968,514,537]" latitude="-14.620833" longLatPrecision="21" longitude="125.254166" name="Savage Hill" pageId="12" pageNumber="179" stateProvince="Western Australia">Savage Hill</location>
;
<geoCoordinate id="EE1987EBFFB73C52FC6DFDD0FBA36C53" box="[981,1102,514,537]" degrees="14" direction="south" minutes="37" orientation="latitude" pageId="12" pageNumber="179" precision="15" seconds="15" value="-14.620833">14°37'15&quot;S</geoCoordinate>
<geoCoordinate id="EE1987EBFFB73C52FBEFFDD0FB326C53" box="[1111,1247,514,537]" degrees="125" direction="east" minutes="15" orientation="longitude" pageId="12" pageNumber="179" precision="15" seconds="15" value="125.254166">125°15'15&quot;E</geoCoordinate>
(RFS-052; coll.
<collectionCode id="ED3C79E9FFB73C52FA1CFDD0FA5A6C53" box="[1444,1463,514,537]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="12" pageNumber="179" type="Museum">V</collectionCode>
. Kessner,
<date id="FF93C7ECFFB73C52FC11FDF2FBC06C7D" box="[937,1069,544,567]" pageId="12" pageNumber="179" value="1987-06-05">
<collectingDate id="EFD73E04FFB73C52FC11FDF2FBC06C7D" box="[937,1069,544,567]" pageId="12" pageNumber="179" value="1987-06-05">05 Jun 1987</collectingDate>
</date>
) (
<figureCitation id="1316FDA9FFB73C52FBFAFDF2FB6C6C7D" box="[1090,1153,544,567]" captionStart="Figure 1" captionStartId="1.[121,184,1877,1898]" captionTargetBox="[149,1389,975,1848]" captionTargetId="figure-423@1.[136,1403,961,1856]" captionTargetPageId="1" captionText="Figure 1. Distribution of Amplirhagada species described in the present work. (A) A. atlantis n.sp. (B) A. carsoniana n.sp. (C) A. alicunda n.sp. (D) A. davidsoniana n.sp. (E) A. vialae n.sp. and A. discoidea n.sp. (F) A. forrestiana n.sp. (G) A. inusitata n.sp. and A. epiphallica n.sp. (H) A. lindsayae n.sp. (I) A. angustocauda n.sp. (K) A. moraniana n.sp. (L) A. gardneriana n.sp. (M) A. bendraytoni n.sp. (N) A. coffea n.sp. (P) A. carinata Solem, 1981. When species occur at more than one locality, the type locality of newly described taxa is marked with an asterisk." figureDoi="http://doi.org/10.5281/zenodo.5239549" httpUri="https://zenodo.org/record/5239549/files/figure.png" pageId="12" pageNumber="179">Fig. 1</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B92E12CFFB73C52FCFFFD88FBF86D4E" blockId="12.[839,1467,602,772]" pageId="12" pageNumber="179">
<emphasis id="B9593D3EFFB73C52FCFFFD88FC166C38" bold="true" box="[839,1019,602,626]" pageId="12" pageNumber="179">Type material</emphasis>
.
<materialsCitation id="3B45EB71FFB73C52FBA8FD88FB7D6CC5" accessionNumber="S34719" collectionCode="WAM" pageId="12" pageNumber="179" specimenCount="1" typeStatus="holotype">
<typeStatus id="54965F8EFFB73C52FBA8FD88FB936C3B" box="[1040,1150,602,625]" pageId="12" pageNumber="179" type="holotype">Holotype</typeStatus>
<collectionCode id="ED3C79E9FFB73C52FB30FD88FB206C3B" box="[1160,1229,602,625]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="179" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB73C52FB61FD88FADE6C3B" box="[1241,1331,602,625]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34719" pageId="12" pageNumber="179" type="EnaNcbi">S34719</accessionNumber>
(preserved specimen) (Pl. 1.6;
<tableCitation id="C6AFD497FFB73C52FB96FDAAFB696CC5" box="[1070,1156,632,655]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="12" pageNumber="179">Table 1</tableCitation>
)
</materialsCitation>
.
<materialsCitation id="3B45EB71FFB73C52FB18FDAAFC2C6C83" accessionNumber="S34720" collectionCode="AM, WAM" pageId="12" pageNumber="179" specimenCount="1" typeStatus="paratype">
<typeStatus id="54965F8EFFB73C52FB18FDAAFAE06CC5" box="[1184,1293,632,655]" pageId="12" pageNumber="179" type="paratype">Paratypes</typeStatus>
<collectionCode id="ED3C79E9FFB73C52FAAFFDAAFAAE6CC5" box="[1303,1347,632,655]" country="Australia" name="Australian Museum" pageId="12" pageNumber="179" type="Museum">AM</collectionCode>
C.472925 (2 preserved specimens),
<collectionCode id="ED3C79E9FFB73C52FBC0FD47FB516CE6" box="[1144,1212,661,684]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="179" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB73C52FB7EFD47FAF06CE6" box="[1222,1309,661,684]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34720" pageId="12" pageNumber="179" type="EnaNcbi">S34720</accessionNumber>
(5 preserved specimens)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB73C52FC76FD60FA696C83" accessionNumber="S34720" box="[974,1412,690,713]" collectionCode="FMNH" isTailEnumeration="true" pageId="12" pageNumber="179" specimenCode="FMNH 220066" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8B4957FFB73C52FC76FD60FB9E6C83" box="[974,1139,690,713]" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="179" type="Museum">FMNH 220066</specimenCode>
(9 preserved specimens)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB73C52FA37FD60FA886CAD" accessionNumber="S34721" collectionCode="AM, WAM" pageId="12" pageNumber="179" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED3C79E9FFB73C52FA37FD60FA576C83" box="[1423,1466,690,713]" country="Australia" name="Australian Museum" pageId="12" pageNumber="179" type="Museum">AM</collectionCode>
C.472926 (4 shells),
<collectionCode id="ED3C79E9FFB73C52FBA6FD02FBB26CAD" box="[1054,1119,720,743]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34981" lsid="urn:lsid:biocol.org:col:34981" name="Western Australian Museum" pageId="12" pageNumber="179" type="Museum">WAM</collectionCode>
<accessionNumber id="947E7CCFFFB73C52FBDCFD02FB586CAD" box="[1124,1205,720,743]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/S34721" pageId="12" pageNumber="179" type="EnaNcbi">S34721</accessionNumber>
(10 dried shells)
</materialsCitation>
,
<materialsCitation id="3B45EB71FFB73C52FAD6FD02FBFC6D4E" accessionNumber="S34721" collectionCode="FMNH" isTailEnumeration="true" pageId="12" pageNumber="179" specimenCode="FMNH 220065" specimenCount="1" typeStatus="paratype">
<specimenCode id="DB8B4957FFB73C52FAD6FD02FC7A6D4E" collectionCode="FMNH" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34795" lsid="urn:lsid:biocol.org:col:34795" name="Field Museum of Natural History" pageId="12" pageNumber="179" type="Museum">FMNH 220065</specimenCode>
(12 shells)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB73C52FCFFFCF5FB0B6D16" pageId="12" pageNumber="179" type="etymology">
<paragraph id="8B92E12CFFB73C52FCFFFCF5FB0B6D16" blockId="12.[839,1466,807,860]" pageId="12" pageNumber="179">
<emphasis id="B9593D3EFFB73C52FCFFFCF5FC2D6D75" bold="true" box="[839,960,807,831]" pageId="12" pageNumber="179">Etymology</emphasis>
. Named in honour of Marlene Vial in recognition for her help with SEM and other work.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB73C53FB8FFC54FE1E6935" lastPageId="13" lastPageNumber="180" pageId="12" pageNumber="179" type="description">
<paragraph id="8B92E12CFFB73C52FB8FFC54FB266DEA" blockId="12.[1079,1227,902,928]" box="[1079,1227,902,928]" pageId="12" pageNumber="179">
<emphasis id="B9593D3EFFB73C52FB8FFC54FB266DEA" bold="true" box="[1079,1227,902,928]" pageId="12" pageNumber="179">Description</emphasis>
</paragraph>
<paragraph id="8B92E12CFFB73C52FCFFFC6FFC33684B" blockId="12.[839,1468,957,2007]" pageId="12" pageNumber="179">
<emphasis id="B9593D3EFFB73C52FCFFFC6FFC936D9E" box="[839,894,957,980]" italics="true" pageId="12" pageNumber="179">Shell</emphasis>
(Pl. 1.6;
<figureCitation id="1316FDA9FFB73C52FC56FC6FFB6E6D9E" box="[1006,1155,957,980]" captionStart="Figure 17" captionStartId="13.[121,184,1560,1581]" captionTargetBox="[121,1421,151,1546]" captionTargetId="figure-50@13.[121,1421,151,1546]" captionTargetPageId="13" captionText="Figure 17. Amplirhagada vialae n.sp. SEM of shell (AM C.472926), radula and jaw (AM C.472925). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Lateral view of shell. (D) Lateral view of last whorl. (E) Jaw. (F) Central and inner lateral teeth viewed from above. (G) Marginal teeth viewed from above. Scale bars: A, 200 µm, BD, 1 mm; E, 100 µm, F, 20 µm; G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239580" httpUri="https://zenodo.org/record/5239580/files/figure.png" pageId="12" pageNumber="179">Fig. 17 AD</figureCitation>
). Semi-globose with highly elevated spire. Solid, not translucent. Periphery well rounded to slightly compressed, occasionally slightly angulate; upper sector of whorl flattened to slightly shouldered underneath suture, basal sectors rounded. Umbilicus completely concealed by columellar reflection. Background colour pale yellowish brown to horn; peripheral band conspicuous, well developed, moderately broad, clearly visible on last whorls; sub-sutural band broad, diffuse to well developed, brown, clearly visible on last whorls; both bands may blend into each other covering shell with a dark brown tone; outer and inner lip colour white, conspicuously contrasting shell colour. Protoconch
<quantity id="4CD54CC9FFB73C52FBA3FACFFB806B7E" box="[1051,1133,1309,1332]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" pageId="12" pageNumber="179" unit="mm" value="3.4">3.4 mm</quantity>
in diameter, comprising about 1.5 whorls, with weak radially elongated pustulations. Teleoconch with faint, regular axial growth lines, evenly distributed across shell. Angle of aperture about 30 degrees; outer lip moderately thick, well rounded, slightly expanded, not reflected; basal node absent or weak. Parietal wall of inner lip inconspicuous. Average shell size 17.1±1.8 × 19.8±1.0 mm (
<tableCitation id="C6AFD497FFB73C52FCC6FA38FC23684B" box="[894,974,1514,1537]" captionStart="Table 1" captionStartId="33.[121,184,1071,1095]" captionText="Table 1. Shell measurements of the type series. Given are maximumminimum (mean±standard deviation) of N measured adult shells. Abbreviations used: HT = holotype." pageId="12" pageNumber="179">Table 1</tableCitation>
).
</paragraph>
<paragraph id="8B92E12CFFB73C52FCDDF9DAFBA06813" blockId="12.[839,1468,957,2007]" pageId="12" pageNumber="179">
<emphasis id="B9593D3EFFB73C52FCDDF9DAFB546855" box="[869,1209,1544,1567]" italics="true" pageId="12" pageNumber="179">Radular and jaw morphology</emphasis>
(
<figureCitation id="1316FDA9FFB73C52FB76F9DAFA886855" box="[1230,1381,1544,1567]" captionStart="Figure 17" captionStartId="13.[121,184,1560,1581]" captionTargetBox="[121,1421,151,1546]" captionTargetId="figure-50@13.[121,1421,151,1546]" captionTargetPageId="13" captionText="Figure 17. Amplirhagada vialae n.sp. SEM of shell (AM C.472926), radula and jaw (AM C.472925). (A) Protoconch and first teleoconch whorls viewed from above. (B) Detail of first teleoconch whorls viewed from above. (C) Lateral view of shell. (D) Lateral view of last whorl. (E) Jaw. (F) Central and inner lateral teeth viewed from above. (G) Marginal teeth viewed from above. Scale bars: A, 200 µm, BD, 1 mm; E, 100 µm, F, 20 µm; G, 10 µm." figureDoi="http://doi.org/10.5281/zenodo.5239580" httpUri="https://zenodo.org/record/5239580/files/figure.png" pageId="12" pageNumber="179">Fig. 17 EG</figureCitation>
). Tooth formula C + 1416 + 4 + 2022, with 160 rows of teeth (n = 1). Jaw with 10 plates.
</paragraph>
<paragraph id="8B92E12CFFB73C53FCDDF9B2FE5E6928" blockId="12.[839,1468,957,2007]" lastBlockId="13.[121,749,1749,1919]" lastPageId="13" lastPageNumber="180" pageId="12" pageNumber="179">
<emphasis id="B9593D3EFFB73C52FCDDF9B2FBB4683D" box="[869,1113,1632,1655]" italics="true" pageId="12" pageNumber="179">Genital morphology</emphasis>
(
<figureCitation id="1316FDA9FFB73C52FBD6F9B2FB13683D" box="[1134,1278,1632,1655]" captionStart-0="Figure 18" captionStart-1="Figure 19" captionStartId-0="14.[166,228,565,586]" captionStartId-1="14.[166,230,1610,1631]" captionTargetBox-0="[167,780,153,545]" captionTargetBox-1="[187,762,673,1589]" captionTargetId-0="figure-9@14.[166,793,138,553]" captionTargetId-1="figure-32@14.[166,793,661,1597]" captionTargetPageId-0="14" captionTargetPageId-1="14" captionText-0="Figure 18. Genitalia of Amplirhagada vialae n.sp. (AM C.472925). For labelling of structures see Fig. 3. Scale bar = 10 mm." captionText-1="Figure 19. Penial anatomy of Amplirhagada vialae n.sp. (AM C.472925).For labelling of structures see Fig.4. Scale bar = 3 mm." figureDoi-0="http://doi.org/10.5281/zenodo.5239582" figureDoi-1="http://doi.org/10.5281/zenodo.5239584" httpUri-0="https://zenodo.org/record/5239582/files/figure.png" httpUri-1="https://zenodo.org/record/5239584/files/figure.png" pageId="12" pageNumber="179">Figs 1819</figureCitation>
). Penis curved, medially inflated, about as long as anterior part of oviduct. Penial sheath very delicate. Length of penial retractor muscle equivalent to about 1/3 of length of penial complex. Penial verge elongately conical, with pointed tip, short, extending about 1/10 of length of penial chamber. Inner penial wall entirely covered by well-developed, elongate pustules densely arranged in longitudinal rows. Main stimulatory pilaster huge, cone-shaped comprising 3/4 of length of inner penial wall; with regular lateral corrugations that support dense and regular rows of hooks. Vas deferens thick, not undulating, entering penial sheath close to penial apex. Vagina long, inflated; inner vaginal wall and wall of bursa copulatrix with smooth longitudinal pilasters. Bursa copulatrix reaching base of spermoviduct. Free oviduct rather long, tubular, comprising about half of length of anterior part of oviduct. Spermoviduct not much longer than anterior part of oviduct.
</paragraph>
<caption id="DF52B1A4FFB73C52FF1EF858FF17699F" ID-DOI="http://doi.org/10.5281/zenodo.5239578" ID-Zenodo-Dep="5239578" httpUri="https://zenodo.org/record/5239578/files/figure.png" pageId="12" pageNumber="179" startId="12.[166,230,1930,1951]" targetBox="[186,766,931,1906]" targetPageId="12">
<paragraph id="8B92E12CFFB73C52FF1EF858FF17699F" blockId="12.[166,794,1930,2005]" pageId="12" pageNumber="179">
Figure 16. Penial anatomy of
<taxonomicName id="4C2D9AAFFFB73C52FE75F858FD0869D5" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[461,741,1930,1951]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="12" pageNumber="179" phylum="Mollusca" rank="species" species="davidsoniana" status="sp. nov.">
<emphasis id="B9593D3EFFB73C52FE75F858FD0869D5" box="[461,741,1930,1951]" italics="true" pageId="12" pageNumber="179">Amplirhagada davidsoniana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFB73C52FD54F858FCF769D5" box="[748,794,1930,1951]" pageId="12" pageNumber="179" rank="species">n.sp.</taxonomicNameLabel>
(AM C.472923). For labelling of structures see Fig. 4. Scale bar = 3 mm.
</paragraph>
</caption>
<caption id="DF52B1A4FFB63C53FFC1F9CAFDC76837" ID-DOI="http://doi.org/10.5281/zenodo.5239580" ID-Zenodo-Dep="5239580" httpUri="https://zenodo.org/record/5239580/files/figure.png" pageId="13" pageNumber="180" startId="13.[121,184,1560,1581]" targetBox="[121,1421,151,1546]" targetPageId="13">
<paragraph id="8B92E12CFFB63C53FFC1F9CAFDC76837" blockId="13.[121,1421,1560,1661]" pageId="13" pageNumber="180">
Figure 17.
<taxonomicName id="4C2D9AAFFFB63C53FF58F9CAFE466867" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[224,427,1560,1581]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="180" phylum="Mollusca" rank="species" species="vialae" status="sp. nov.">
<emphasis id="B9593D3EFFB63C53FF58F9CAFE466867" box="[224,427,1560,1581]" italics="true" pageId="13" pageNumber="180">Amplirhagada vialae</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFB63C53FE17F9CAFE306867" box="[431,477,1560,1581]" pageId="13" pageNumber="180" rank="species">n.sp.</taxonomicNameLabel>
SEM of shell (AM C.472926), radula and jaw (AM C.472925).
<emphasis id="B9593D3EFFB63C53FB85F9CAFBB66867" box="[1085,1115,1560,1581]" italics="true" pageId="13" pageNumber="180">(A)</emphasis>
Protoconch and first teleoconch whorls viewed from above.
<emphasis id="B9593D3EFFB63C53FE3FF9E1FE486802" box="[391,421,1587,1608]" italics="true" pageId="13" pageNumber="180">(B)</emphasis>
Detail of first teleoconch whorls viewed from above.
<emphasis id="B9593D3EFFB63C53FC08F9E1FC3D6802" box="[944,976,1587,1608]" italics="true" pageId="13" pageNumber="180">(C)</emphasis>
Lateral view of shell.
<emphasis id="B9593D3EFFB63C53FB12F9E1FB266802" box="[1194,1227,1587,1608]" italics="true" pageId="13" pageNumber="180">(D)</emphasis>
Lateral view of last whorl.
<emphasis id="B9593D3EFFB63C53FF07F99FFF306828" box="[191,221,1613,1634]" italics="true" pageId="13" pageNumber="180">(E)</emphasis>
Jaw.
<emphasis id="B9593D3EFFB63C53FEADF99FFED96828" box="[277,308,1613,1634]" italics="true" pageId="13" pageNumber="180">(F)</emphasis>
Central and inner lateral teeth viewed from above.
<emphasis id="B9593D3EFFB63C53FC94F99FFCA06828" box="[812,845,1613,1634]" italics="true" pageId="13" pageNumber="180">(G)</emphasis>
Marginal teeth viewed from above. Scale bars:
<emphasis id="B9593D3EFFB63C53FA99F99FFADD6828" box="[1313,1328,1613,1634]" italics="true" pageId="13" pageNumber="180">A</emphasis>
, 200 µm,
<emphasis id="B9593D3EFFB63C53FFC1F9BAFF486837" box="[121,165,1640,1661]" italics="true" pageId="13" pageNumber="180">BD</emphasis>
, 1 mm;
<emphasis id="B9593D3EFFB63C53FF4DF9BAFEE96837" box="[245,260,1640,1661]" italics="true" pageId="13" pageNumber="180">E</emphasis>
, 100 µm,
<emphasis id="B9593D3EFFB63C53FEDDF9BAFE996837" box="[357,372,1640,1661]" italics="true" pageId="13" pageNumber="180">F</emphasis>
, 20 µm;
<emphasis id="B9593D3EFFB63C53FE71F9BAFE376837" box="[457,474,1640,1661]" italics="true" pageId="13" pageNumber="180">G</emphasis>
, 10 µm.
</paragraph>
</caption>
<paragraph id="8B92E12CFFB63C53FF2FF8BAFE1E6935" blockId="13.[121,749,1749,1919]" box="[151,499,1896,1919]" pageId="13" pageNumber="180">
<emphasis id="B9593D3EFFB63C53FF2FF8BAFE876935" box="[151,362,1896,1919]" italics="true" pageId="13" pageNumber="180">Aestivation strategy</emphasis>
: Free sealer.
</paragraph>
</subSubSection>
<subSubSection id="C337B2A7FFB63C50FCA2F907FDA46935" lastPageId="14" lastPageNumber="181" pageId="13" pageNumber="180" type="discussion">
<paragraph id="8B92E12CFFB63C50FCA2F907FDA46935" blockId="13.[794,1422,1749,1919]" lastBlockId="14.[166,795,1749,1919]" lastPageId="14" lastPageNumber="181" pageId="13" pageNumber="180">
<emphasis id="B9593D3EFFB63C53FCA2F907FC6E68A7" bold="true" box="[794,899,1749,1773]" pageId="13" pageNumber="180">Remarks</emphasis>
. Anatomical description based on dissections of
<specimenCount id="9D2B2AA5FFB63C53FCA2F921FC506940" box="[794,957,1779,1802]" pageId="13" pageNumber="180" type="generic">two specimens</specimenCount>
. Listed by Solem (1991) as “
<taxonomicName id="4C2D9AAFFFB63C53FB48F921FCA2696D" authority="NSP" authorityName="NSP" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="180" phylum="Mollusca" rank="genus">
<emphasis id="B9593D3EFFB63C53FB48F921FA606940" box="[1264,1421,1779,1802]" italics="true" pageId="13" pageNumber="180">Amplirhagada</emphasis>
NSP
</taxonomicName>
22”. The “bee-hive” shaped shell is similar to that of
<taxonomicName id="4C2D9AAFFFB63C53FACEF8C2FC14690E" authority="Kohler, 2010" authorityName="Kohler" authorityYear="2010" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="180" phylum="Mollusca" rank="species" species="decora">
<emphasis id="B9593D3EFFB63C53FACEF8C2FC88690F" italics="true" pageId="13" pageNumber="180">A. decora</emphasis>
Köhler, 2010
</taxonomicName>
(Bigge Island) and
<taxonomicName id="4C2D9AAFFFB63C53FB69F8FCFC496928" authority="Kohler, 2010" authorityName="Kohler" authorityYear="2010" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="13" pageNumber="180" phylum="Mollusca" rank="species" species="boongareensis">
<emphasis id="B9593D3EFFB63C53FB69F8FCFA60690F" box="[1233,1421,1838,1861]" italics="true" pageId="13" pageNumber="180">A. boongareensis</emphasis>
Köhler, 2010
</taxonomicName>
(Boongaree Island), but these species differ in shell colour. All three species are characterized by a huge, cone-shaped main pilaster and by their inner penial wall being densely covered with large pustules.
<taxonomicName id="4C2D9AAFFFB53C50FDC5F921FEAD696D" authorityName="Kohler" authorityYear="2010" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="181" phylum="Mollusca" rank="species" species="boongareensis">
<emphasis id="B9593D3EFFB53C50FDC5F921FEAD696D" italics="true" pageId="14" pageNumber="181">Amplirhagada boongareensis</emphasis>
</taxonomicName>
differs by possessing two additional pilasters and
<taxonomicName id="4C2D9AAFFFB53C50FF6CF8FCFE22690E" authority="Kohler, 2010" authorityName="Kohler" authorityYear="2010" box="[212,463,1837,1861]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="181" phylum="Mollusca" rank="species" species="decora">
<emphasis id="B9593D3EFFB53C50FF6CF8FCFED1690F" box="[212,316,1838,1861]" italics="true" pageId="14" pageNumber="181">A. decora</emphasis>
Köhler, 2010
</taxonomicName>
has a shorter bursa copulatrix. The present species occurs in sympatry with
<taxonomicName id="4C2D9AAFFFB53C50FD2BF899FCF46928" authority="Köhler, 2011" authorityName="Köhler" authorityYear="2011" box="[659,793,1867,1890]" class="Gastropoda" family="Camaenidae" genus="Amplirhagada" kingdom="Animalia" order="Stylommatophora" pageId="14" pageNumber="181" phylum="Mollusca" rank="species" species="discoidea" status="sp. nov.">
<emphasis id="B9593D3EFFB53C50FD2BF899FCF46928" box="[659,793,1867,1890]" italics="true" pageId="14" pageNumber="181">A. discoidea</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A26A8045FFB53C50FF1EF8BAFF376935" box="[166,218,1896,1919]" pageId="14" pageNumber="181" rank="species">n.sp.</taxonomicNameLabel>
being described in the following.
</paragraph>
</subSubSection>
</treatment>
</document>