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<mods:title id="502CA95AB8E61EF629662F75CEBE622F">The Staphylinidae of Armenia and Nagorno-Karabakh (Coleoptera)</mods:title>
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<mods:namePart id="05D7A9A7B650CD8F5EB78CF8C90CA638">Assing, Volker</mods:namePart>
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<heading id="23265061721E7103FF2B6532FB58F9E0" box="[249,589,1546,1574]" centered="true" fontSize="12" level="2" pageId="72" pageNumber="163" reason="2">
<taxonomicName id="BFD19C8E721E7103FF2B6532FB58F9E0" authority="REITTER, 1909" authorityName="REITTER" authorityYear="1909" box="[249,589,1546,1574]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="rufopacus">
<emphasis id="4AA53B1F721E7103FF2B6532F8ACF9E0" box="[249,441,1546,1574]" italics="true" pageId="72" pageNumber="163">Astenus rufopacus</emphasis>
<bibRefCitation id="1C409AFC721E7103FE126532FB58F9E0" author="REITTER, E." box="[448,589,1546,1574]" pageId="72" pageNumber="163" pagination="1 - 392" refId="ref81359" refString="REITTER, E. 1909: Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. 2. - K. G. Lutz, Stuttgart: 1 - 392." type="book chapter" year="1909">REITTER, 1909</bibRefCitation>
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<treatmentCitationGroup id="58C1C023721E7103FF06657AFB50F991" box="[212,581,1601,1623]" pageId="72" pageNumber="163">
<treatmentCitation id="F970C11C721E7103FF06657AFB54F991" author="REITTER, E." box="[212,577,1601,1623]" page="150" pageId="72" pageNumber="163" year="1909">
<taxonomicName id="BFD19C8E721E7103FF06657AFB54F991" authority="REITTER, 1909: 150" authorityName="REITTER" authorityPageNumber="150" authorityYear="1909" box="[212,577,1601,1623]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="rufopacus">
<emphasis id="4AA53B1F721E7103FF06657AF86AF991" box="[212,383,1602,1623]" italics="true" pageId="72" pageNumber="163">Astenus rufopacus</emphasis>
<bibRefCitation id="1C409AFC721E7103FE56657AFB54F991" author="REITTER, E." box="[388,577,1601,1623]" pageId="72" pageNumber="163" pagination="1 - 392" refId="ref81359" refString="REITTER, E. 1909: Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. 2. - K. G. Lutz, Stuttgart: 1 - 392." type="book chapter" year="1909">REITTER, 1909: 150</bibRefCitation>
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.
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</paragraph>
<paragraph id="786EE70D721E7103FF06655AFB6BF9B1" blockId="72.[212,638,1601,1656]" box="[212,638,1633,1656]" pageId="72" pageNumber="163">
<treatmentCitationGroup id="58C1C023721E7103FF06655AFB6BF9B1" box="[212,638,1633,1656]" pageId="72" pageNumber="163">
<treatmentCitation id="F970C11C721E7103FF06655AFB1EF9B1" author="COIFFAIT, H." box="[212,523,1633,1655]" page="94" pageId="72" pageNumber="163" year="1960">
<taxonomicName id="BFD19C8E721E7103FF06655AFB1EF9B1" authority="COIFFAIT, 1960: 94" authorityName="COIFFAIT" authorityPageNumber="94" authorityYear="1960" box="[212,523,1633,1655]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="baali">
<emphasis id="4AA53B1F721E7103FF06655AF847F9B1" box="[212,338,1634,1655]" italics="true" pageId="72" pageNumber="163">Astenus baali</emphasis>
<bibRefCitation id="1C409AFC721E7103FE85655AFB1EF9B1" author="COIFFAIT, H." box="[343,523,1633,1655]" pageId="72" pageNumber="163" pagination="49 - 99" refId="ref76603" refString="COIFFAIT, H. 1960: Les Astenus d'Europe et de la mediterrraneenne (Coleopteres, Staphylinidae). - Bulletin de la Societe d'Histoire Naturelle de Toulouse 95: 49 - 99." type="journal article" year="1960">
COIFFAIT, 1960:
<specimenCount id="6ED72C84721E7103FE26655AFB1EF9B1" box="[500,523,1634,1655]" pageId="72" pageNumber="163" type="generic">94</specimenCount>
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f.;
<emphasis id="4AA53B1F721E7103FDFA655AFB6BF9B1" bold="true" box="[552,638,1634,1655]" pageId="72" pageNumber="163">syn. nov.</emphasis>
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</paragraph>
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The original description of
<taxonomicName id="BFD19C8E721E7103FDC56598FB8AF97E" authorityName="REITTER" authorityYear="1909" box="[535,671,1696,1720]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="rufopacus">
<emphasis id="4AA53B1F721E7103FDC56598FB8AF97E" box="[535,671,1696,1720]" italics="true" pageId="72" pageNumber="163">A. rufopacus</emphasis>
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is based on an unspecified number of
<typeStatus id="A76A59AF721E7103FDD765F8FB76F91E" box="[517,611,1728,1752]" pageId="72" pageNumber="163" type="syntype">syntypes</typeStatus>
from “Araxesthal, Lenkoran” (
<bibRefCitation id="1C409AFC721E7103FE8665D8F8E2F93E" author="REITTER, E." box="[340,503,1760,1784]" pageId="72" pageNumber="163" pagination="1 - 392" refId="ref81359" refString="REITTER, E. 1909: Fauna Germanica. Die Kafer des Deutschen Reiches. Nach der analytischen Methode bearbeitet. 2. - K. G. Lutz, Stuttgart: 1 - 392." type="book chapter" year="1909">REITTER 1909</bibRefCitation>
), that of
<taxonomicName id="BFD19C8E721E7103FD8D65D8FBA7F93E" authorityName="COIFFAIT" authorityYear="1960" box="[607,690,1760,1784]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="baali">
<emphasis id="4AA53B1F721E7103FD8D65D8FBA7F93E" box="[607,690,1760,1784]" italics="true" pageId="72" pageNumber="163">A. baali</emphasis>
</taxonomicName>
on a
<typeStatus id="A76A59AF721E7103FD2465D8F816F8DE" pageId="72" pageNumber="163" type="holotype">holotype</typeStatus>
and several male
<typeStatus id="A76A59AF721E7103FE156438FB25F8DE" box="[455,560,1792,1816]" pageId="72" pageNumber="163" type="paratype">paratypes</typeStatus>
from
<collectingCountry id="00C6A79D721E7103FDA66438FBC3F8DE" box="[628,726,1792,1816]" name="Lebanon" pageId="72" pageNumber="163">Lebanon</collectingCountry>
,
<collectingCountry id="00C6A79D721E7103FD326438FA27F8DE" box="[736,818,1792,1816]" name="Turkey" pageId="72" pageNumber="163">Turkey</collectingCountry>
, and
<collectingCountry id="00C6A79D721E7103FEDA6418F87CF8FE" box="[264,361,1824,1848]" name="Armenia" pageId="72" pageNumber="163">Armenia</collectingCountry>
(“Delizhan” = Dilijan) (
<bibRefCitation id="1C409AFC721E7103FDA46418FA33F8FE" author="COIFFAIT, H." box="[630,806,1824,1848]" pageId="72" pageNumber="163" pagination="49 - 99" refId="ref76603" refString="COIFFAIT, H. 1960: Les Astenus d'Europe et de la mediterrraneenne (Coleopteres, Staphylinidae). - Bulletin de la Societe d'Histoire Naturelle de Toulouse 95: 49 - 99." type="journal article" year="1960">COIFFAIT 1960</bibRefCitation>
).
<bibRefCitation id="1C409AFC721E7103FF066478F87EF89E" author="ANLAS, S." box="[212,363,1856,1880]" pageId="72" pageNumber="163" pagination="405 - 413" refId="ref72453" refString="ANLAS, S. 2017: Notes on the genus Astenus DEJEAN, 1833 from the Palearctic region (Coleoptera: Staphylinidae: Paederinae). - Turkiye Entomoloji Dergisi 41 (4): 405 - 413. - http: // dergipark. gov. tr / download / articlefile / 451101." type="journal article" year="2017">ANLAŞ (2017)</bibRefCitation>
studied the type material of
<taxonomicName id="BFD19C8E721E7103FD716478FA3EF89E" authorityName="REITTER" authorityYear="1909" box="[675,811,1856,1880]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="rufopacus">
<emphasis id="4AA53B1F721E7103FD716478FA3EF89E" box="[675,811,1856,1880]" italics="true" pageId="72" pageNumber="163">A. rufopacus</emphasis>
</taxonomicName>
, designated a
<typeStatus id="A76A59AF721E7103FEBB6458F8A8F8BE" box="[361,445,1888,1912]" pageId="72" pageNumber="163" type="syntype">syntype</typeStatus>
from Lenkoran as the
<typeStatus id="A76A59AF721E7103FD156458FA3FF8BE" box="[711,810,1888,1912]" pageId="72" pageNumber="163" type="lectotype">lectotype</typeStatus>
, and illustrated the habitus and the male sexual characters. The type material of
<taxonomicName id="BFD19C8E721E7103FDD86498FB4BF87E" authorityName="COIFFAIT" authorityYear="1960" box="[522,606,1952,1976]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="baali">
<emphasis id="4AA53B1F721E7103FDD86498FB4BF87E" box="[522,606,1952,1976]" italics="true" pageId="72" pageNumber="163">A. baali</emphasis>
</taxonomicName>
is deposited in the Coiffait collection in the Natural History Museum in Paris and consequently inaccessible for scientific study (see
<bibRefCitation id="1C409AFC721E7103FED76B38F88DF7DE" author="ASSING, V." box="[261,408,2048,2072]" pageId="72" pageNumber="163" pagination="129 - 182" refId="ref75112" refString="ASSING, V. 2018 f: On the taxonomy and zoogeography of some West Palaearctic Quedius species, with a focus on the East Mediterranean and the species allied to Quedius umbrinus and Q. nivicola (Coleoptera: Staphylinidae: Staphylininae). - Linzer Biologische Beitrage 50 (1): 129 - 182." type="journal article" year="2018">ASSING 2018f</bibRefCitation>
). However, based on the details indicated in the original description of
<taxonomicName id="BFD19C8E721E7103FB286238FC5BFEDE" authorityName="COIFFAIT" authorityYear="1960" box="[1274,1358,256,280]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="baali">
<emphasis id="4AA53B1F721E7103FB286238FC5BFEDE" box="[1274,1358,256,280]" italics="true" pageId="72" pageNumber="163">A. baali</emphasis>
</taxonomicName>
and the illustrations of the aedeagus (
<bibRefCitation id="1C409AFC721E7103FB7D6218FC4DFEFE" author="COIFFAIT, H." box="[1199,1368,288,312]" pageId="72" pageNumber="163" pagination="49 - 99" refId="ref76603" refString="COIFFAIT, H. 1960: Les Astenus d'Europe et de la mediterrraneenne (Coleopteres, Staphylinidae). - Bulletin de la Societe d'Histoire Naturelle de Toulouse 95: 49 - 99." type="journal article" year="1960">COIFFAIT 1960</bibRefCitation>
: figures 6162), there is little doubt that the type material of
<taxonomicName id="BFD19C8E721E7103FC846258FABFFEBE" authorityName="COIFFAIT" authorityYear="1960" box="[854,938,352,376]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="baali">
<emphasis id="4AA53B1F721E7103FC846258FABFFEBE" box="[854,938,352,376]" italics="true" pageId="72" pageNumber="163">A. baali</emphasis>
</taxonomicName>
is conspecific with that of
<taxonomicName id="BFD19C8E721E7103FB1F6258FC40FEBE" authorityName="REITTER" authorityYear="1909" box="[1229,1365,352,376]" class="Insecta" family="Staphylinidae" genus="Astenus" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="species" species="rufopacus">
<emphasis id="4AA53B1F721E7103FB1F6258FC40FEBE" box="[1229,1365,352,376]" italics="true" pageId="72" pageNumber="163">A. rufopacus</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
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<paragraph id="786EE70D721E7103FCAE62F0FC2FFE21" blockId="72.[892,1338,456,487]" box="[892,1338,456,487]" pageId="72" pageNumber="163">
3.6.9
<taxonomicName id="BFD19C8E721E7103FC6962F0FD4DFE21" authorityName="Latreille" authorityYear="1802" box="[955,1112,456,487]" class="Insecta" family="Staphylinidae" kingdom="Animalia" order="Coleoptera" pageId="72" pageNumber="163" phylum="Arthropoda" rank="subFamily" subFamily="Staphylininae">Staphylininae</taxonomicName>
(by VOLKER ASSING)
</paragraph>
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</treatment>
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