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<document ID-CLB-Dataset="9577" ID-DOI="http://dx.doi.org/10.3897/zookeys.993.49434" ID-GBIF-Dataset="9e79a4e8-6a72-449d-a986-d3061121ab29" ID-PMC="PMC7683492" ID-Pensoft-Pub="1313-2970-993-35" ID-Pensoft-UUID="EF942C1D7CBF5E34BE934BDAA776776D" ID-PubMed="33262674" ID-ZooBank="F50BE60A8CFA4A659A990C9E908F526F" ModsDocID="1313-2970-993-35" checkinTime="1605571769446" checkinUser="pensoft" docAuthor="Delgado, Leonardo &amp; Toledo-Hernandez, Victor Hugo" docDate="2020" docId="898C96D11C00556EB17502B347ECF393" docLanguage="en" docName="ZooKeys 993: 35-46" docOrigin="ZooKeys 993" docSource="http://dx.doi.org/10.3897/zookeys.993.49434" docTitle="Diplotaxis balam Delgado &amp; Toledo-Hernández 2020, sp. nov." docType="treatment" docUuid="75CDF203-6811-4A1E-8D85-40547CEC4FBD" docUuidSource="ZooBank" docVersion="7" id="EF942C1D7CBF5E34BE934BDAA776776D" lastPageNumber="35" masterDocId="EF942C1D7CBF5E34BE934BDAA776776D" masterDocTitle="Three new species of Diplotaxis Kirby from Guatemala and Mexico (Coleoptera, Scarabaeidae, Melolonthinae), with a key to the species of the trapezifera group" masterLastPageNumber="46" masterPageNumber="35" pageNumber="35" updateTime="1736217479064" updateUser="felipe">
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<mods:title id="4DD4EE6EE7C8FA85D6EA3F2781EDAF3B">Three new species of Diplotaxis Kirby from Guatemala and Mexico (Coleoptera, Scarabaeidae, Melolonthinae), with a key to the species of the trapezifera group</mods:title>
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<mods:name id="BE3B7516E04210F7A201D167231301B8" type="personal">
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<mods:namePart id="C5A269FD2FF2ED3DDDFDE1414F3A1632">Delgado, Leonardo</mods:namePart>
<mods:affiliation id="7E81BCBBC34A17E366464C88A1B03BFD">Centro de Investigacion en Biodiversidad y Conservacion, Universidad Autonoma del Estado de Morelos, Avenida Universidad No. 1001, Col. Chamilpa, 62209 Cuernavaca, Morelos, Mexico &amp; Instituto de Ecologia, A. C. Carretera Antigua a Coatepec 351, 91070 Xalapa, Veracruz, Mexico</mods:affiliation>
<mods:nameIdentifier id="CD440DD7D7320EAE8226FED7D841AA8C" type="email">leonardo.delgado@inecol.mx</mods:nameIdentifier>
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<mods:namePart id="70A4C5E604D9C1A887572D792DCDB568">Toledo-Hernandez, Victor Hugo</mods:namePart>
<mods:affiliation id="D9E4FA5BEAC936361BF547CEF807B0BD">Centro de Investigacion en Biodiversidad y Conservacion, Universidad Autonoma del Estado de Morelos, Avenida Universidad No. 1001, Col. Chamilpa, 62209 Cuernavaca, Morelos, Mexico</mods:affiliation>
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<mods:date id="ED0490F0CA99F20EEBAF1CAB8370A3E1">2020</mods:date>
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<mods:number id="595C5C151BDBD85CF98D9CE00EDBDCDC">993</mods:number>
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<mods:classification id="FC1AE36D0A90C8CC70C41211B8DF24AF">journal article</mods:classification>
<mods:identifier id="1569FDA5DA1448C3DE65A4056FE3DE94" type="DOI">http://dx.doi.org/10.3897/zookeys.993.49434</mods:identifier>
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<treatment id="898C96D11C00556EB17502B347ECF393" ID-GBIF-Taxon="169091797" LSID="urn:lsid:zoobank.org:act:75CDF203-6811-4A1E-8D85-40547CEC4FBD" httpUri="http://treatment.plazi.org/id/898C96D11C00556EB17502B347ECF393" lastPageNumber="35" pageId="0" pageNumber="35" scope_family="Scarabaeidae" scope_order="Coleoptera" scope_subFamily="Melolonthinae">
<subSubSection id="BEFFCC74696D065F668465BEFEE0A164" pageId="0" pageNumber="35" type="nomenclature">
<paragraph id="9E57BDE6F1A6096E89B80D474A464635" pageId="0" pageNumber="35">
<taxonomicName id="9E501458681B4CE6975631234250D25D" ID-CoL="7VRV4" LSID="898C96D1-1C00-556E-B175-02B347ECF393" authority="Delgado &amp; Toledo-Hernández, 2020" authorityName="Delgado &amp; Toledo-Hernández" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplotaxis balam" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="balam" status="sp. nov." subFamily="Melolonthinae">Diplotaxis balam</taxonomicName>
<taxonomicNameLabel id="51D65BC1C52C1AC058824B8C19B3278C" pageId="0" pageNumber="35">sp. nov.</taxonomicNameLabel>
<figureCitation id="8B34C6324AF60D4389213139E2316B77" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">Figs 1-5</figureCitation>
</paragraph>
</subSubSection>
<subSubSection id="2D24681E677C5D82195409CB9FDBD087" pageId="0" pageNumber="35" type="materials_examined">
<paragraph id="451E48CC262517786785C4DADFCAD774" pageId="0" pageNumber="35">Material examined.</paragraph>
<paragraph id="C35E6ACC9560C113830D4B165C0A60AD" pageId="0" pageNumber="35">
<emphasis id="0AF4406EA08DDB292F9BECA86B425DD0" bold="true" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
male,&quot;Guatemala: Zacapa, arriba de La
<normalizedToken id="B9F6835B5C6FF00ADC9E50E5F3675CA0" originalValue="Unión">Union</normalizedToken>
, 16-III-1996, Alt. 1,550 m, bosque nuboso, J. C. Schuster col.&quot; (UVGC).
<emphasis id="727CD24F9CB67685248BB6CC6441AFE2" bold="true" italics="true" pageId="0" pageNumber="35">Paratype</emphasis>
female, same data as holotype (LLDC).
</paragraph>
</subSubSection>
<subSubSection id="BA23C30F2627F0CD7F2F90DE4265E4B9" pageId="0" pageNumber="35" type="diagnosis">
<paragraph id="875F655BBAFAF999C6CE406EFA73392E" pageId="0" pageNumber="35">Diagnosis.</paragraph>
<paragraph id="3C5B71923127D3D284DB5FDE4857258B" pageId="0" pageNumber="35">
This species is easily recognized by the color of the elytra (Figs
<figureCitation id="8CB6A11CF8E65A25F4C06027B5AC0D5C" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">1-2</figureCitation>
,
<figureCitation id="A9128737073F6CEE228FCEC6EBE784C9" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">5</figureCitation>
): ground color yellow with black, irregular foveae distributed throughout entire surface. No other described species of this genus shows this color pattern.
</paragraph>
<caption id="D6555A5F19817A70ED81A0ADBC2E14F6" doi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35" start="Figures 15" startId="F1">
<paragraph id="345555F5F2061C4B1F583BDF824A7EA7" pageId="0" pageNumber="35">
<emphasis id="11B70F60702ED05F25164576FF6BDE90" bold="true" pageId="0" pageNumber="35">Figures 1-5.</emphasis>
<taxonomicName id="1D7634EF6124B591315994C4E01B3DBD" authorityName="Delgado &amp; Toledo-Hernández" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplotaxis balam" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="balam" subFamily="Melolonthinae">
<emphasis id="3912404331944A7AAA1913804EAF9427" italics="true" pageId="0" pageNumber="35">Diplotaxis balam</emphasis>
</taxonomicName>
sp. nov.
<emphasis id="C2B731E01A3F753B143AA620786DAFD5" bold="true" pageId="0" pageNumber="35">1</emphasis>
male habitus, dorsal view
<emphasis id="8BE01A66DFFA06FB5AAE6A950CC2BD31" bold="true" pageId="0" pageNumber="35">2</emphasis>
male habitus, lateral view
<emphasis id="72C55DA0551BA98AA28FCC7C873EAFDE" bold="true" pageId="0" pageNumber="35">3</emphasis>
aedeagus, dorsal view
<emphasis id="C747F30F3F0B908D720F08ABBFE2CB1E" bold="true" pageId="0" pageNumber="35">4</emphasis>
aedeagus, lateral view
<emphasis id="851B5B2B73C07B7F2ECA2A6062D19DF9" bold="true" pageId="0" pageNumber="35">5</emphasis>
female habitus. Scale bars: 1 mm (
<emphasis id="FD6CEAF4CDC38492F1E1BCED5D8BF984" bold="true" pageId="0" pageNumber="35">1, 2, 5</emphasis>
); 0.5 mm (
<emphasis id="BF7E2C526521384F37437EE39161B17B" bold="true" pageId="0" pageNumber="35">3, 4</emphasis>
).
</paragraph>
</caption>
</subSubSection>
<subSubSection id="012B320330ED470FB2F3050355D0211B" pageId="0" pageNumber="35" type="description">
<paragraph id="5E1F36C655155CC65176086239BA1D29" pageId="0" pageNumber="35">Description.</paragraph>
<paragraph id="E336410648A50EC22AD1507E9EA65380" pageId="0" pageNumber="35">
<emphasis id="015DCD3FB0C35A4BF9ADC7483D8017AC" bold="true" pageId="0" pageNumber="35">
<emphasis id="31D0B2CE4500E2497A884A456089435C" italics="true" pageId="0" pageNumber="35">Holotype</emphasis>
. Male
</emphasis>
(Figs
<figureCitation id="CBAB8FF651C7F566F11C3962A26A20B2" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">1-4</figureCitation>
). Length 8.55 mm; width 4.21 mm. Body elongate and dorsoventrally flattened. Clypeus red with a metallic green tinge, frons and pronotum metallic green, elytra yellow and black with irregular foveae throughout entire surface, venter metallic green, legs tan with a weak metallic green tinge.
</paragraph>
<paragraph id="C91A0B3C75FD693FF44D99134E9C4BC3" pageId="0" pageNumber="35">
<emphasis id="00972174092882B08B57DACF139017C7" bold="true" pageId="0" pageNumber="35">
<emphasis id="A51389172FAF76419907041393594BC9" italics="true" pageId="0" pageNumber="35">Head</emphasis>
.
</emphasis>
Clypeus trapezoidal in shape, 0.4 length of frons, apex broadly sinuated with anterior angles rounded, sides indented in front of eyes; clypeal surface concave, rugose, with distinct setae; frontoclypeal suture absent; frons flat, gradually declivous to clypeus, with coarse, dense punctures; transverse eye diameter 0.33 interocular width; antennae 10-segmented; labrum flat, flush with, and 0.20 times longer than reflexed underside face of clypeus, surface with dense punctures; mandibles slender in frontal view; mentum slightly convex, with weak anterior declivity marked by a suture; last article of maxillary palps not impressed dorsally.
</paragraph>
<paragraph id="B5F4ECB80301873A8C30F97983B4F9E3" pageId="0" pageNumber="35">
<emphasis id="66365FBC5BB66EF64EBD28847ABACB46" bold="true" pageId="0" pageNumber="35">
<emphasis id="682EF9A8BC00816A76F7A83139935AA8" italics="true" pageId="0" pageNumber="35">Pronotum</emphasis>
.
</emphasis>
Hexagonal in shape; with anterior angles acute, lateral margins angled and situated behind middle, posterior angles obtuse; pronotal surface almost evenly convex, with three lateral foveae on each side; pronotal punctuation regular with dense, medium-sized punctures on disc, confluent near sides; basal margin with bead not cariniform, with a continuous row of punctures; most punctures bearing a minute seta slightly longer than one puncture diameter.
</paragraph>
<paragraph id="79B7F59C90F8F703D0ED2D7B42EE1DD0" pageId="0" pageNumber="35">
<emphasis id="82BA0CE1819DDF4A701EDE50CF34EE58" bold="true" pageId="0" pageNumber="35">
<emphasis id="166E0493CA9982F39BBF22B1BC8C78EA" italics="true" pageId="0" pageNumber="35">Scutellum</emphasis>
.
</emphasis>
With medium-sized punctures at sides.
<emphasis id="2DB38FD3B6176749124610F6ED47B97E" bold="true" italics="true" pageId="0" pageNumber="35">Elytra</emphasis>
. 1.7 times longer than width, elytral surface with irregular, shallow, black foveae, most of them on the intervals; elytral striae with separated, ocellate punctures, intervals with small, moderately dense punctures; elytral punctures with setae minute but slightly longer than those of pronotum.
</paragraph>
<paragraph id="F18EFE34513A9875D7A9D2DCCB7BBB45" pageId="0" pageNumber="35">
<emphasis id="3CF02D09D99EBE4C66C9367FD8479C02" bold="true" pageId="0" pageNumber="35">
<emphasis id="0205D5C956E4C726F8EB47B1BEDF2791" italics="true" pageId="0" pageNumber="35">Abdomen</emphasis>
.
</emphasis>
Without lateral carina; propygidium without groove above of pygidium; ventrites 2-5 subequal in length, surface with setae medially and with whitish scales laterally; pygidium 1.7 times wider than long, surface with coarse, deep, setigerous punctures; setae dense, longer on apical half.
</paragraph>
<paragraph id="3DA5DBE3732BE8D7F414F441E6B73120" pageId="0" pageNumber="35">
<emphasis id="4110457E3C4DFB1CE25078185F4685BD" bold="true" pageId="0" pageNumber="35">
<emphasis id="0FB0EBFE72405B0826D09BCD7C3E0BD9" italics="true" pageId="0" pageNumber="35">Legs</emphasis>
.
</emphasis>
Protibiae tridentate, basal tooth weak and situated in distal half; claws long, slightly curved, cleft subapically, inner rami of claws shorter than apex; tarsi longer than respective tibiae; mesotarsomere 1 slightly shorter than 2; metacoxal plate rounded and margined laterally; metafemora straight and slender; metatibial spurs slender, long, acute; metatarsomere 1 shorter than the 2 and slightly longer than longest spur.
</paragraph>
<paragraph id="4B1618D5751CD412A056D3340CAE4912" pageId="0" pageNumber="35">
<emphasis id="A3FB6B3B9C5F8DF499E48820BDAFD624" bold="true" pageId="0" pageNumber="35">
<emphasis id="1D349CADA132F9C9E4B16AC2DDACB7FC" italics="true" pageId="0" pageNumber="35">Genitalia</emphasis>
.
</emphasis>
Basal piece shorter than parameres, which are joined on inner margin at basal fifth, moderately widened at middle, apices blunt (Figs
<figureCitation id="4BEEDB456DCCDC1078E49672A158AC41" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">3-4</figureCitation>
).
</paragraph>
<paragraph id="26AD7A521445DC7E84A9B6D33FDB36DF" pageId="0" pageNumber="35">
<emphasis id="E2CCAAB3E76477CA637E1E8BB7DAA54F" bold="true" pageId="0" pageNumber="35">Female</emphasis>
. One female paratype (Fig.
<figureCitation id="D682C5EDB03DCADE91E5EE10D3524BB5" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">5</figureCitation>
). Length 6.97 mm; width 3.53 mm. The female differs from the male in the following respects: clypeus slightly shorter; frons and vertex more convex; transverse eye diameter 0.31 interocular width; pronotum with anterior angles obtuse and lateral angles rounded; elytra 1.3 times longer than width; abdomen nearly flat; pygidium 1.6 times wider than long; tibiae broader and robust; metafemora slightly broader; inner metatibial spur wider and longer than metatarsomere 1.
</paragraph>
</subSubSection>
<subSubSection id="53BA77FCC3B8721253D6FE380C7AF71C" pageId="0" pageNumber="35" type="etymology">
<paragraph id="F626BD434B4AA2656E923F974A17EF50" pageId="0" pageNumber="35">Etymology.</paragraph>
<paragraph id="ED6C3B406C71B5D408BFA4D105FCEA2A" pageId="0" pageNumber="35">
The specific epithet
<taxonomicName id="346FB3E548F01D6DB4D94BD2710012E3" authorityName="Delgado &amp; Toledo-Hernández" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="balam" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="balam" subFamily="Melolonthinae">
<emphasis id="625AF3C31ACF9F3A7DEB7B6C549F0159" italics="true" pageId="0" pageNumber="35">balam</emphasis>
</taxonomicName>
, meaning jaguar in the Mayan language, refers to the color pattern of the elytra, similar to the skin of this feline.
</paragraph>
</subSubSection>
<subSubSection id="2521B7083F3C0711F396C9881CE48501" pageId="0" pageNumber="35" type="distribution">
<paragraph id="A4344623B368F5E67902B7304CE8045F" pageId="0" pageNumber="35">Distribution.</paragraph>
<paragraph id="163E3BC3802A0D1F4521AAD6B42AF631" pageId="0" pageNumber="35">
This species is only known from the type locality, situated in the Sierra de Las Minas, Guatemala, near the border with Honduras (
<geoCoordinate id="F68AC9174CD64905A6DAF49FFF40794C" degrees="14" direction="north" minutes="56" orientation="latitude" precision="1" seconds="45.6" value="14.946">14°56'45.6&quot;N</geoCoordinate>
,
<geoCoordinate id="188EA6C99638AA476FC045DA90ADF0FC" degrees="89" direction="west" minutes="16" orientation="longitude" precision="1" seconds="40.1" value="-89.27781">89°16'40.1&quot;W</geoCoordinate>
) (Fig.
<figureCitation id="AD244242F7D1C77392298107B96104E4" captionStart="Figure 17" captionStartId="F4" captionText="Figure 17. Distribution of Diplotaxis species. Diplotaxis balam sp. nov. (black circle), D. chiapasensis sp. nov. (black triangle), D. complanatis sp. nov. (white circle), D. xalapensis Delgado and Capistran (white triangle)." figureDoi="10.3897/zookeys.993.49434.figure17" httpUri="https://binary.pensoft.net/fig/475355" pageId="0" pageNumber="35">17</figureCitation>
). The locality is at 1550 m altitude, covered by a cloud forest.
</paragraph>
</subSubSection>
<subSubSection id="7FE3DC8C7EB40CBF440F4A5E4EAA7C95" pageId="0" pageNumber="35" type="taxonomic remarks">
<paragraph id="94A9F2B4FB3ABAA7788C90BCD1931852" pageId="0" pageNumber="35">Taxonomic remarks.</paragraph>
<paragraph id="521C0F9DCD4E255318AEEEA04F786697" pageId="0" pageNumber="35">
The features of
<taxonomicName id="34D8FD23B5A3EE1E1DFD215864B48C61" lsidName="D. balam" pageId="0" pageNumber="35" rank="species" species="balam">
<emphasis id="E7A191080C5EDA4761F24EE4C5358EB9" italics="true" pageId="0" pageNumber="35">D. balam</emphasis>
</taxonomicName>
sp. nov. agree in part with those of the
<emphasis id="9BBCE2347C0A119D33E5D4669D02113D" italics="true" pageId="0" pageNumber="35">trapezifera</emphasis>
species group [see key to species groups by
<bibRefCitation id="8261D0CB38789B84AA238FA0645F3C12" author="Vaurie, P" journalOrPublisher="Bulletin of the American Museum of Natural History" pageId="0" pageNumber="35" pagination="161 - 434" refId="B9" refString="Vaurie, P, 1960. A revision of the genus Diplotaxis (Coleoptera, Scarabaeidae, Melolonthinae), Part II. Bulletin of the American Museum of Natural History 120 (2): 161 - 434" title="A revision of the genus Diplotaxis (Coleoptera, Scarabaeidae, Melolonthinae), Part II." volume="120" year="1960">Vaurie (1960)</bibRefCitation>
]. The group is mainly characterized by the presence of setae on the clypeal surface, the rest of the dorsum being glabrous or with minute setae only. However,
<taxonomicName id="43B10BC460CE65E19AA0A58FA6815BC3" lsidName="D. balam" pageId="0" pageNumber="35" rank="species" species="balam">
<emphasis id="8805E1178E9B7C5E943D653DBAEC77CD" italics="true" pageId="0" pageNumber="35">D. balam</emphasis>
</taxonomicName>
sp. nov. (as well as
<taxonomicName id="353CD1D5FD73BD27DD41ED9E4588B33F" lsidName="D. xalapensis" pageId="0" pageNumber="35" rank="species" species="xalapensis">
<emphasis id="4F2F2AAD6FFF49660B2944EAC3CA11F5" italics="true" pageId="0" pageNumber="35">D. xalapensis</emphasis>
</taxonomicName>
and the two new species described below) has a dorsoventrally flattened body, unlike species of the
<emphasis id="6767314DE103A8CDCEC55E9050A7E09E" italics="true" pageId="0" pageNumber="35">trapezifera</emphasis>
group which have a convex body.
<taxonomicName id="DDB7D9E3CAF9EC91971BF95E9ED34521" authorityName="Delgado &amp; Toledo-Hernández" authorityYear="2020" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplotaxis balam" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="species" species="balam" subFamily="Melolonthinae">
<emphasis id="6637AD9DCAFFFAAA7234B729D68DCF58" italics="true" pageId="0" pageNumber="35">Diplotaxis balam</emphasis>
</taxonomicName>
sp. nov. is distinguished from all other
<taxonomicName id="4D2C641964A20D76E94F8F3D06C3B871" class="Insecta" family="Scarabaeidae" genus="Diplotaxis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Diplotaxis" order="Coleoptera" pageId="0" pageNumber="35" phylum="Arthropoda" rank="genus" subFamily="Melolonthinae">
<emphasis id="33F5311ADFF77AA01270D11CB47CCE98" italics="true" pageId="0" pageNumber="35">Diplotaxis</emphasis>
</taxonomicName>
by the unique color pattern of the elytra (Figs
<figureCitation id="C96799AF3EA3310AA9D8B65BE068B760" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">1-2</figureCitation>
,
<figureCitation id="BE303B5D9E647F411D720C261B0D951F" captionStart="Figures 15" captionStartId="F1" captionText="Figures 1 - 5. Diplotaxis balam sp. nov. 1 male habitus, dorsal view 2 male habitus, lateral view 3 aedeagus, dorsal view 4 aedeagus, lateral view 5 female habitus. Scale bars: 1 mm (1, 2, 5); 0.5 mm (3, 4)." figureDoi="10.3897/zookeys.993.49434.figures1-5" httpUri="https://binary.pensoft.net/fig/475352" pageId="0" pageNumber="35">5</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>